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<document id="5D63A028EDEF5B4F5B349349ADDA25AD" ID-CLB-Dataset="8143" ID-DOI="10.2478/aemnp-2019-0020" ID-GBIF-Dataset="a03ecce2-b526-4f94-aa65-620016956099" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488918" ID-ZooBank="9F309FCC-A2ED-47B9-BC37-D0C4A3B482E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612219281920" checkinUser="felipe" docAuthor="Fikáček, Martin &amp; Liu, Hsing-Che" docDate="2019" docId="03F387C57A21FFAFFC0D05D6350EB41C" docLanguage="en" docName="ActaEntMusNatPra.59.1.229-252.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 59 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Thysanarthria madurensis Hebauer 2001" docType="treatment" docVersion="10" lastPageNumber="246" masterDocId="FFCAFFBD7A2EFFBEFF8901283431B709" masterDocTitle="A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)" masterLastPageNumber="252" masterPageNumber="229" pageNumber="244" updateTime="1698926077238" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="3F68F7F38878EBFB2EB43CE480A9A728">A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)</mods:title>
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<mods:namePart id="94624653DF81DA5097E30CC45CF61E72">Fikáček, Martin</mods:namePart>
<mods:affiliation id="C74079528F57CC31E27E2AAD5490ECF2">Department of Entomology, National Museum, Cirkusová 1740, CZ- 19100 Praha 9 Horní Počernice, Czech Republic &amp; Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844 Praha 2, Czech Republic</mods:affiliation>
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<mods:namePart id="0C707831F369F47197DBBE9065A9BB7F">Liu, Hsing-Che</mods:namePart>
<mods:affiliation id="3A301AB9577E9230DD78D86436A3A1C3">Department of Environmental Engineering and Management, Chaoyang University of Technology, No. 168, Jifeng E. Rd., Taichung City 413, Taiwan</mods:affiliation>
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<mods:date id="6538F6F5E2910C34209D1475B9510F3B">2019</mods:date>
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<treatment id="03F387C57A21FFAFFC0D05D6350EB41C" ID-DOI="http://doi.org/10.5281/zenodo.4488717" ID-GBIF-Taxon="177423426" ID-Zenodo-Dep="4488717" LSID="urn:lsid:plazi:treatment:03F387C57A21FFAFFC0D05D6350EB41C" httpUri="http://treatment.plazi.org/id/03F387C57A21FFAFFC0D05D6350EB41C" lastPageId="17" lastPageNumber="246" pageId="15" pageNumber="244">
<subSubSection id="C34065587A21FFB1FC0D05D63162B21C" box="[900,1363,1278,1301]" pageId="15" pageNumber="244" type="nomenclature">
<paragraph id="8BE536D37A21FFB1FC0D05D63162B21C" blockId="15.[900,1363,1278,1301]" box="[900,1363,1278,1301]" pageId="15" pageNumber="244">
<heading id="D0AD81BF7A21FFB1FC0D05D63162B21C" box="[900,1363,1278,1301]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="244" reason="3">
<taxonomicName id="4C5A4D507A21FFB1FC0D05D63162B21C" ID-CoL="56TH6" authority="Hebauer, 2001" authorityName="Hebauer" authorityYear="2001" box="[900,1363,1278,1301]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="madurensis">
<emphasis id="B92EEAC17A21FFB1FC0D05D63162B21C" bold="true" box="[900,1363,1278,1301]" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FC0D05D63098B21C" bold="true" box="[900,1193,1278,1301]" italics="true" pageId="15" pageNumber="244">Thysanarthria madurensis</emphasis>
<bibRefCitation id="EFCB4B227A21FFB1FB3905D63162B21C" author="Hebauer" box="[1200,1363,1278,1301]" firstAuthor="Hebauer" pageId="15" pageNumber="244" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">Hebauer, 2001</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C34065587A21FFB1FBAF04343083B226" box="[1062,1202,1308,1327]" pageId="15" pageNumber="244" type="description">
<paragraph id="8BE536D37A21FFB1FBAF04343083B226" blockId="15.[828,1437,1308,1617]" box="[1062,1202,1308,1327]" pageId="15" pageNumber="244">
(
<figureCitation id="13612A567A21FFB1FBA504343051B226" box="[1068,1120,1308,1327]" captionStart="Fig" captionStartId="16.[151,182,1976,1995]" captionTargetBox="[204,1420,202,1944]" captionTargetId="figure-108@16.[162,1439,189,1956]" captionTargetPageId="16" captionText="Fig. 8. Male genitalia of Thysanarthria species, holotypes.AE T. chui sp. nov.(AC whole aedeagus in dorsal, lateral and ventral view; DE detail of median lobe and parameres in dorsal and ventral view). FJ T. hongsonensis Hebauer, 2001 (FH whole aedeagus in dorsal, lateral and ventral view; IJ details of parameres and median lobe in dorsal and ventral view). KO T. madurensis Hebauer, 2001 (KM whole aedeagus in dorsal, lateral and ventral view; NO details of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488940" httpUri="https://zenodo.org/record/4488940/files/figure.png" pageId="15" pageNumber="244">Figs 8</figureCitation>
KO,
<figureCitation id="13612A567A21FFB1FB1F0434309AB226" box="[1174,1195,1308,1327]" captionStart="Fig" captionStartId="20.[151,182,1949,1968]" captionTargetBox="[151,1436,401,1931]" captionTargetId="figure-13@20.[151,1437,401,1931]" captionTargetPageId="20" captionText="Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (AB) and Near East (C), and examples of habitats of Thysanarthria (DF). Localities: D India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; FG Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, FG by H.-C. Liu." figureDoi="http://doi.org/10.5281/zenodo.4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="15" pageNumber="244">11</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BE536D37A21FFB1FCB5041630E0B258" blockId="15.[828,1437,1308,1617]" box="[828,1233,1342,1361]" pageId="15" pageNumber="244">
<treatmentCitationGroup id="AB4A11FD7A21FFB1FCB5041630E0B258" box="[828,1233,1342,1361]" pageId="15" pageNumber="244">
<treatmentCitation id="0AFB10C27A21FFB1FCB5041630FCB258" author="HEBAUER F." box="[828,1229,1342,1361]" page="398" pageId="15" pageNumber="244" year="2001">
<taxonomicName id="4C5A4D507A21FFB1FCB5041630FCB258" ID-CoL="56TH6" authority="Hebauer, 2001: 398" authorityName="Hebauer" authorityPageNumber="398" authorityYear="2001" box="[828,1229,1342,1361]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="madurensis">
<emphasis id="B92EEAC17A21FFB1FCB50416302CB258" box="[828,1053,1342,1361]" italics="true" pageId="15" pageNumber="244">Thysanarthria madurensis</emphasis>
<bibRefCitation id="EFCB4B227A21FFB1FBAB041630FCB258" author="Hebauer" box="[1058,1229,1342,1361]" firstAuthor="Hebauer" page="398" pageId="15" pageNumber="244" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">Hebauer, 2001: 398</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34065587A21FFB1FCB50440302CB158" pageId="15" pageNumber="244" type="materials_examined">
<paragraph id="8BE536D37A21FFB1FCB5044037C0B2EF" blockId="15.[828,1437,1308,1617]" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FCB504403024B272" bold="true" box="[828,1045,1384,1403]" pageId="15" pageNumber="244">Type material examined.</emphasis>
<materialsCitation id="3B323C8E7A21FFB1FB91044037C0B2EF" collectingDate="1962-03-13" collectionCode="MZLU" country="Sri Lanka" location="near river" municipality="Madura Oya" pageId="15" pageNumber="244" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54E188717A21FFB1FB9104403058B270" box="[1048,1129,1384,1403]" pageId="15" pageNumber="244" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D5CFD5A7A21FFB1FBFB044A30B3B272" box="[1138,1154,1378,1403]" pageId="15" pageNumber="244" type="male"></specimenCount>
(
<collectionCode id="ED4BAE167A21FFB1FB02044030FAB272" box="[1163,1227,1384,1403]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="15" pageNumber="244">MZLU</collectionCode>
),
<emphasis id="B92EEAC17A21FFB1FB5E04403160B272" bold="true" box="[1239,1361,1384,1403]" pageId="15" pageNumber="244">
<collectingCountry id="F34D76437A21FFB1FB5E0440317CB272" box="[1239,1357,1384,1403]" name="Sri Lanka" pageId="15" pageNumber="244">SRI LANKA</collectingCountry>
:
</emphasis>
ʻCeylon, E. Prov. /
<collectingMunicipality id="6B81ACA97A21FFB1FC1904AB37CAB29F" box="[912,1019,1411,1430]" pageId="15" pageNumber="244">Madura Oya</collectingMunicipality>
/
<locationDeviation id="9A572FF17A21FFB1FB8204AB3084B29F" box="[1035,1205,1411,1430]" pageId="15" pageNumber="244">15 mls NNW Bibile</locationDeviation>
/ 13.III.62 Loc. 138 //
<location id="8E8560087A21FFB1FAFE04AB3755B2B8" LSID="urn:lsid:plazi:treatment:03F387C57A21FFAFFC0D05D6350EB41C:8E8560087A21FFB1FAFE04AB3755B2B8" country="Sri Lanka" municipality="Madura Oya" name="near river" pageId="15" pageNumber="244">near river</location>
// Lund University / Ceylon Expedition 1962 / Brinck-Andersson- / Cederholm // MZLU / Type no. / 3121:1 // Photo 2017 / by MZLU // MZLU / 2017 / 466.
</materialsCitation>
</paragraph>
<paragraph id="8BE536D37A21FFB1FCB504C6302CB158" blockId="15.[828,1437,1308,1617]" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FCB504C63060B108" bold="true" box="[828,1105,1518,1537]" pageId="15" pageNumber="244">Additional material examined.</emphasis>
<materialsCitation id="3B323C8E7A21FFB1FBDC04C6313FB112" ID-GBIF-Occurrence="3756360324" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FBDC04C630DDB108" bold="true" box="[1109,1260,1518,1537]" pageId="15" pageNumber="244">
<collectingCountry id="F34D76437A21FFB1FBDC04C630A6B108" box="[1109,1175,1518,1537]" name="India" pageId="15" pageNumber="244">INDIA</collectingCountry>
:
<collectingRegion id="499EF8317A21FFB1FB2904C630D4B2F6" box="[1184,1253,1518,1537]" country="India" name="Kerala" pageId="15" pageNumber="244">KERALA</collectingRegion>
:
</emphasis>
1 ♂ (
<collectionCode id="ED4BAE167A21FFB1FA9204C63156B108" box="[1307,1383,1518,1537]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="15" pageNumber="244" type="Museum">NHMW</collectionCode>
): Shoranur, bank of Ponnani river,
<date id="FFE410137A21FFB1FBB1072030BAB112" box="[1080,1163,1544,1563]" pageId="15" pageNumber="244" value="1994-01-31">31.i.1994</date>
, lgt. Z. Kejval.
</materialsCitation>
<materialsCitation id="3B323C8E7A21FFB1FA9B0720302CB158" ID-GBIF-Occurrence="3756360352" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FA9B07203151B112" bold="true" box="[1298,1376,1544,1563]" pageId="15" pageNumber="244">
<collectingCountry id="F34D76437A21FFB1FA9B0720316DB112" box="[1298,1372,1544,1563]" name="Nepal" pageId="15" pageNumber="244">NEPAL</collectingCountry>
:
</emphasis>
1 ♂,
<specimenCount id="9D5CFD5A7A21FFB1FA1807203759B13F" pageId="15" pageNumber="244" type="generic">2 spec.</specimenCount>
(
<collectionCode id="ED4BAE167A21FFB1FCFB070B379EB13F" box="[882,943,1571,1590]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="15" pageNumber="244">SMNS</collectionCode>
,
<collectionCode id="ED4BAE167A21FFB1FC31070B37C8B13F" box="[952,1017,1571,1590]" country="Czech Republic" name="National Museum Prague" pageId="15" pageNumber="244">NMPC</collectionCode>
):
<collectingRegion id="499EF8317A21FFB1FB8F070B3067B13F" box="[1030,1110,1571,1590]" country="Nepal" name="Narayani" pageId="15" pageNumber="244">Narayani</collectingRegion>
, Sauraha,
<collectingRegion id="499EF8317A21FFB1FB27070B30EDB13F" box="[1198,1244,1571,1590]" country="Nepal" name="Rapti" pageId="15" pageNumber="244">Rapti</collectingRegion>
River bank, light trap,
<date id="FFE410137A21FFB1FCB5071637A9B158" box="[828,920,1598,1617]" pageId="15" pageNumber="244" value="2000-04-18">18.iv.2000</date>
, lgt. A. Weigel.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34065587A21FFAFFCB50741350AB6BC" lastPageId="17" lastPageNumber="246" pageId="15" pageNumber="244" type="description">
<paragraph id="8BE536D37A21FFAFFCB50741350AB6BC" blockId="15.[828,1437,1641,2080]" lastBlockId="17.[151,760,190,789]" lastPageId="17" lastPageNumber="246" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FCB507413018B189" bold="true" box="[828,1065,1641,1664]" pageId="15" pageNumber="244">
Redescription.
<emphasis id="B92EEAC17A21FFB1FC7907413018B189" bold="true" box="[1008,1065,1641,1664]" italics="true" pageId="15" pageNumber="244">Body</emphasis>
</emphasis>
length
<quantity id="4CA29B367A21FFB1FB0D074130EFB189" box="[1156,1246,1641,1664]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="244" unit="mm" value="1.3">1.3 mm</quantity>
, maximum body width
<quantity id="4CA29B367A21FFB1FCF607A137E5B1A9" box="[895,980,1673,1696]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="244" unit="mm" value="0.9">0.9 mm</quantity>
. Head and labrum black; pronotum uniformly yellowish; elytra uniformly yellowish; legs yellowish.
<emphasis id="B92EEAC17A21FFB1FCB507E13747B1E9" bold="true" box="[828,886,1737,1760]" italics="true" pageId="15" pageNumber="244">Head</emphasis>
with weak granulate microsculpture on interstices; punctation sparse, each puncture bearing pointed seta. Eyes separated by 4.0× the width of one eye in dorsal view.
<emphasis id="B92EEAC17A21FFB1FCB506013798B049" bold="true" box="[828,937,1833,1856]" italics="true" pageId="15" pageNumber="244">Pronotum</emphasis>
with sparse setiferous punctation similar to that on head; interstices with weak granulate microsculpture.
<emphasis id="B92EEAC17A21FFB1FCB50641374EB089" bold="true" box="[828,895,1897,1920]" italics="true" pageId="15" pageNumber="244">Elytra</emphasis>
with 10 striae sharply impressed except anteromedially (near scutellar shield) where neither striae nor serial punctures are visible; intervals weakly convex at midlength and near apex; interval punctation sparse, setiferous; interstices without microsculpture.
<emphasis id="B92EEAC17A21FFB1FBF706C130D6BF09" bold="true" box="[1150,1255,2025,2048]" italics="true" pageId="15" pageNumber="244">Aedeagus</emphasis>
(
<figureCitation id="13612A567A21FFB1FB7C06C13107BF09" box="[1269,1334,2025,2048]" captionStart="Fig" captionStartId="16.[151,182,1976,1995]" captionTargetBox="[204,1420,202,1944]" captionTargetId="figure-108@16.[162,1439,189,1956]" captionTargetPageId="16" captionText="Fig. 8. Male genitalia of Thysanarthria species, holotypes.AE T. chui sp. nov.(AC whole aedeagus in dorsal, lateral and ventral view; DE detail of median lobe and parameres in dorsal and ventral view). FJ T. hongsonensis Hebauer, 2001 (FH whole aedeagus in dorsal, lateral and ventral view; IJ details of parameres and median lobe in dorsal and ventral view). KO T. madurensis Hebauer, 2001 (KM whole aedeagus in dorsal, lateral and ventral view; NO details of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488940" httpUri="https://zenodo.org/record/4488940/files/figure.png" pageId="15" pageNumber="244">Figs 8</figureCitation>
KO)
<quantity id="4CA29B367A21FFB1FAF206C13754BF29" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="244" unit="mm" value="0.4">0.4 mm</quantity>
long. Phallobase moderately wide at base of parameres, c. as wide as paramares combined, gradually narrowing towards basal fifth, base abruptly widened, arcuate in lateral view. Parameres narrow and elongate, gradually narrowing towards a pointed apex, apices slightly divergent, outer face of parameres arcuate. Median lobe narrow, not reaching apex of parameres, with a pair of pointed lobes subapically, apex cut off between these projections; gonopore transversely oval, subapical.
</paragraph>
</subSubSection>
<caption id="DF25665B7A3EFFAEFF1E069037E8BF12" ID-DOI="http://doi.org/10.5281/zenodo.4488940" ID-Zenodo-Dep="4488940" httpUri="https://zenodo.org/record/4488940/files/figure.png" pageId="16" pageNumber="245" startId="16.[151,182,1976,1995]" targetBox="[204,1420,202,1944]" targetPageId="16">
<paragraph id="8BE536D37A3EFFAEFF1E069037E8BF12" blockId="16.[151,1437,1976,2075]" pageId="16" pageNumber="245">
Fig. 8. Male genitalia of
<taxonomicName id="4C5A4D507A3EFFAEFEEC069035EEB0C2" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[357,479,1976,1995]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A3EFFAEFEEC069035EEB0C2" box="[357,479,1976,1995]" italics="true" pageId="16" pageNumber="245">Thysanarthria</emphasis>
</taxonomicName>
species, holotypes.AE
<taxonomicName id="4C5A4D507A3EFFAEFD48069036CBB0C2" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[705,762,1976,1995]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="245" phylum="Arthropoda" rank="species" species="chui" status="sp. nov.">
<emphasis id="B92EEAC17A3EFFAEFD48069036CBB0C2" box="[705,762,1976,1995]" italics="true" pageId="16" pageNumber="245">T. chui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A3EFFAEFD7706903771B0C2" box="[766,832,1976,1995]" pageId="16" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
(AC whole aedeagus in dorsal, lateral and ventral view; DE detail of median lobe and parameres in dorsal and ventral view). FJ
<taxonomicName id="4C5A4D507A3EFFAEFD4C06FB37E1B0EF" authority="Hebauer, 2001" authorityName="Hebauer" authorityYear="2001" box="[709,976,2003,2022]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="245" phylum="Arthropoda" rank="species" species="hongsonensis">
<emphasis id="B92EEAC17A3EFFAEFD4C06FB377CB0EF" box="[709,845,2003,2022]" italics="true" pageId="16" pageNumber="245">T. hongsonensis</emphasis>
<bibRefCitation id="EFCB4B227A3EFFAEFCDA06FB37E1B0EF" author="Hebauer" box="[851,976,2003,2022]" firstAuthor="Hebauer" pageId="16" pageNumber="245" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">Hebauer, 2001</bibRefCitation>
</taxonomicName>
(FH whole aedeagus in dorsal, lateral and ventral view; IJ details of parameres and median lobe in dorsal and ventral view). KO
<taxonomicName id="4C5A4D507A3EFFAEFCFD06C6305EBF08" authority="Hebauer, 2001" authorityName="Hebauer" authorityYear="2001" box="[884,1135,2030,2049]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="245" phylum="Arthropoda" rank="species" species="madurensis">
<emphasis id="B92EEAC17A3EFFAEFCFD06C637DDBF08" box="[884,1004,2030,2049]" italics="true" pageId="16" pageNumber="245">T. madurensis</emphasis>
<bibRefCitation id="EFCB4B227A3EFFAEFC7806C6305EBF08" author="Hebauer" box="[1009,1135,2030,2049]" firstAuthor="Hebauer" pageId="16" pageNumber="245" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">Hebauer, 2001</bibRefCitation>
</taxonomicName>
(KM whole aedeagus in dorsal, lateral and ventral view; NO details of median lobe and parameres in dorsal and ventral view).
</paragraph>
</caption>
<subSubSection id="C34065587A3FFFAFFF1E009636AEB53C" pageId="17" pageNumber="246" type="diagnosis">
<paragraph id="8BE536D37A3FFFAFFF1E009636AEB53C" blockId="17.[151,760,190,789]" pageId="17" pageNumber="246">
<emphasis id="B92EEAC17A3FFFAFFF1E009635B8B6DC" bold="true" box="[151,393,446,469]" pageId="17" pageNumber="246">Differential diagnosis.</emphasis>
In the tiny body size, dorsal surface without microsculpture and narrowly elongate pointed parameres,
<taxonomicName id="4C5A4D507A3FFFAFFF6A00D63545B51C" authorityName="Hebauer" authorityYear="2001" box="[227,372,510,533]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="madurensis">
<emphasis id="B92EEAC17A3FFFAFFF6A00D63545B51C" box="[227,372,510,533]" italics="true" pageId="17" pageNumber="246">T. madurensis</emphasis>
</taxonomicName>
only resembles
<taxonomicName id="4C5A4D507A3FFFAFFD9700D636F4B51C" authorityName="Hebauer" authorityYear="2001" box="[542,709,510,533]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="hongsonensis">
<emphasis id="B92EEAC17A3FFFAFFD9700D636F4B51C" box="[542,709,510,533]" italics="true" pageId="17" pageNumber="246">T. hongsonensis</emphasis>
</taxonomicName>
. See under the latter species for diagnostic charactares.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3FFFAFFF1E031634D8B57C" pageId="17" pageNumber="246" type="biology_ecology">
<paragraph id="8BE536D37A3FFFAFFF1E031634D8B57C" blockId="17.[151,760,190,789]" pageId="17" pageNumber="246">
<emphasis id="B92EEAC17A3FFFAFFF1E031634C1B55C" bold="true" box="[151,240,574,597]" pageId="17" pageNumber="246">Biology.</emphasis>
Unknown, the Nepalese specimens were collected at light.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3FFFAFFF1E0356350EB41C" pageId="17" pageNumber="246" type="distribution">
<paragraph id="8BE536D37A3FFFAFFF1E0356350EB41C" blockId="17.[151,760,190,789]" pageId="17" pageNumber="246">
<emphasis id="B92EEAC17A3FFFAFFF1E03563517B59C" bold="true" box="[151,294,638,661]" pageId="17" pageNumber="246">Distribution.</emphasis>
The species is known from three very distant localities (
<collectingCountry id="F34D76437A3FFFAFFE8B03B6355BB5BC" box="[258,362,670,693]" name="Sri Lanka" pageId="17" pageNumber="246">Sri Lanka</collectingCountry>
, southern
<collectingCountry id="F34D76437A3FFFAFFE5B03B63639B5BC" box="[466,520,670,693]" name="India" pageId="17" pageNumber="246">India</collectingCountry>
and
<collectingCountry id="F34D76437A3FFFAFFDBE03B6364BB5BC" box="[567,634,670,693]" name="Nepal" pageId="17" pageNumber="246">Nepal</collectingCountry>
). If the label data especially of the Nepalese specimens are correct, the species is likely widely distributed in the Indian peninsula, but overlooked.
</paragraph>
</subSubSection>
</treatment>
</document>