259 lines
30 KiB
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259 lines
30 KiB
XML
<document ID-DOI="10.11646/zootaxa.5278.2.12" ID-ISSN="1175-5326" ID-Zenodo-Dep="7906063" ID-ZooBank="FAF52B55-44A7-4B6C-BCA0-483BE43D1BC2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683528846200" checkinUser="plazi" docAuthor="Leschen, Richard A. B. & Gimmel, Matthew L." docDate="2023" docId="03F38795FFC1596C65FEFD78FD68FD45" docLanguage="en" docName="zootaxa.5278.2.12.pdf" docOrigin="Zootaxa 5278 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.2.12" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sternodea arcana Leschen and Gimmel 2023, new species" docType="treatment" docVersion="2" lastPageNumber="398" masterDocId="FFCAFFEDFFC0596E6569FFF6FFF0FFB4" masterDocTitle="Sternodea arcana, new species (Coleoptera: Cryptophagidae: Caenoscelini): part of the Nearctic fauna?" masterLastPageNumber="400" masterPageNumber="396" pageNumber="397" updateTime="1683529220222" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
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<mods:title>Sternodea arcana, new species (Coleoptera: Cryptophagidae: Caenoscelini): part of the Nearctic fauna?</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Leschen, Richard A. B.</mods:namePart>
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<mods:affiliation>Manaaki Whenua Landcare Research, New Zealand Arthropod Collection, Private Bag 92170, Auckland, New Zealand. leschenr @ landcareresearch. co. nz</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gimmel, Matthew L.</mods:namePart>
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<mods:affiliation>Department of Invertebrate Zoology, Santa Barbara Museum of Natural History, 2559 Puesta del Sol Rd, Santa Barbara, CA 93105, USA.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-05</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5278</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>396</mods:start>
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<mods:end>400</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.2.12</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5278.2.12</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7906063</mods:identifier>
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<mods:identifier type="ZooBank">FAF52B55-44A7-4B6C-BCA0-483BE43D1BC2</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7906065" ID-Zenodo-Dep="7906065" LSID="urn:lsid:plazi:treatment:03F38795FFC1596C65FEFD78FD68FD45" httpUri="http://treatment.plazi.org/id/03F38795FFC1596C65FEFD78FD68FD45" lastPageId="2" lastPageNumber="398" pageId="1" pageNumber="397">
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<subSubSection box="[151,797,654,681]" pageId="1" pageNumber="397" type="nomenclature">
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<paragraph blockId="1.[151,797,654,717]" box="[151,797,654,681]" pageId="1" pageNumber="397">
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<heading bold="true" box="[151,797,654,681]" fontSize="11" level="1" pageId="1" pageNumber="397" reason="1">
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<emphasis bold="true" box="[151,797,654,681]" pageId="1" pageNumber="397">
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<taxonomicName authority="Leschen and Gimmel" authorityName="Leschen and Gimmel" authorityYear="2023" box="[151,640,654,681]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="arcana" status="new species">
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<emphasis bold="true" box="[151,366,654,681]" italics="true" pageId="1" pageNumber="397">Sternodea arcana</emphasis>
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Leschen and Gimmel
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</taxonomicName>
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,
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<taxonomicNameLabel box="[653,797,654,681]" pageId="1" pageNumber="397" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,274,690,717]" pageId="1" pageNumber="397" type="description">
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<paragraph blockId="1.[151,797,654,717]" box="[151,274,690,717]" pageId="1" pageNumber="397">
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(
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||
<figureCitation box="[159,265,690,717]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1502,1526]" captionTargetBox="[152,1434,778,1477]" captionTargetId="figure-281@2.[151,1435,777,1478]" captionTargetPageId="2" captionText="FIGURES 1–5. Sternodea arcana, new species, holotype. 1) Antenna; 2) Pronotum, dorsal view showing punctation in anterior portion and transverse punctate-crenulate line; 3) Prosternal process, ventral view (not to scale); 4) Aedeagus, dorsal view; 5) Aedeagus, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906067" httpUri="https://zenodo.org/record/7906067/files/figure.png" pageId="1" pageNumber="397">Figs. 1–5</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="1" pageNumber="397" type="materials_examined">
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<paragraph blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="397">
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<emphasis bold="true" box="[151,334,762,789]" pageId="1" pageNumber="397">Type specimen.</emphasis>
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<materialsCitation collectionCode="CNCI" collectorName="Monroe Co. Upper Matacumba Key & S. Peck" country="Iceland" county="Monroe Co." determinerName="C. Johnson" location="Upper Matacumba Key" pageId="1" pageNumber="397" specimenCount="2" typeStatus="holotype">
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<typeStatus box="[341,459,763,789]" pageId="1" pageNumber="397">HOLOTYPE</typeStatus>
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, male (
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<collectionCode box="[543,615,763,789]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="1" pageNumber="397">CNCI</collectionCode>
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), labeled: “FLA.
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<collectorName box="[810,1217,763,789]" pageId="1" pageNumber="397">
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<collectingCounty box="[810,946,763,789]" pageId="1" pageNumber="397">Monroe Co.</collectingCounty>
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<location LSID="urn:lsid:plazi:treatment:03F38795FFC1596C65FEFD78FD68FD45:8E856058FFC1596F66D1FD0DFB31FCA1" box="[952,1217,763,789]" country="Iceland" county="Monroe Co." name="Upper Matacumba Key" pageId="1" pageNumber="397">Upper Matacumba Key</location>
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</collectorName>
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,
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<collectingCountry box="[1228,1268,763,789]" name="Iceland" pageId="1" pageNumber="397">Isla</collectingCountry>
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Moranda, Ber. 225, 8.VIII.74,
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<collectorName box="[325,409,799,825]" pageId="1" pageNumber="397">S. Peck</collectorName>
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//
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<taxonomicName authority="Leschen & Gimmel, 2023" authorityName="Leschen & Gimmel" authorityYear="2023" box="[439,676,799,825]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="floridensis" status="sp. n.">Sternodea floridensis</taxonomicName>
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<taxonomicNameLabel box="[683,743,799,825]" pageId="1" pageNumber="397" rank="species">sp. n.</taxonomicNameLabel>
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det.
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<determinerName box="[798,923,799,825]" pageId="1" pageNumber="397">C. Johnson</determinerName>
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1992 [partly handwritten, unpublished name] //
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<typeStatus box="[174,322,835,861]" pageId="1" pageNumber="397">HOLOTYPE</typeStatus>
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<taxonomicName authority="Leschen & Gimmel, 2023" authorityName="Leschen & Gimmel" authorityYear="2023" box="[329,527,835,861]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="arcana" status="new species">
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<emphasis box="[329,527,835,861]" italics="true" pageId="1" pageNumber="397">Sternodea arcana</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[541,679,835,861]" pageId="1" pageNumber="397" rank="species">new species</taxonomicNameLabel>
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, desig. by Leschen & Gimmel 2023 [red label]”.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="397" type="diagnosis">
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<paragraph blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="397">
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<emphasis bold="true" box="[199,322,870,896]" pageId="1" pageNumber="397">Diagnosis.</emphasis>
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<taxonomicName authorityName="Leschen & Gimmel" authorityYear="2023" box="[331,532,871,897]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="arcana">
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<emphasis box="[331,532,871,897]" italics="true" pageId="1" pageNumber="397">Sternodea arcana</emphasis>
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</taxonomicName>
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is the sole member of the genus in the
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<collectingCountry box="[995,1148,871,897]" name="United States of America" pageId="1" pageNumber="397">United States</collectingCountry>
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(assuming the label data to be correct) and can be distinguished from members of the widespread
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<taxonomicName box="[987,1122,907,933]" class="Insecta" family="Cryptophagidae" genus="Caenoscelis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="genus">
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<emphasis box="[987,1122,907,933]" italics="true" pageId="1" pageNumber="397">Caenoscelis</emphasis>
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</taxonomicName>
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by having a short, rounded body and antennal club with two antennomeres (
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<figureCitation box="[692,756,943,969]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1502,1526]" captionTargetBox="[152,1434,778,1477]" captionTargetId="figure-281@2.[151,1435,777,1478]" captionTargetPageId="2" captionText="FIGURES 1–5. Sternodea arcana, new species, holotype. 1) Antenna; 2) Pronotum, dorsal view showing punctation in anterior portion and transverse punctate-crenulate line; 3) Prosternal process, ventral view (not to scale); 4) Aedeagus, dorsal view; 5) Aedeagus, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906067" httpUri="https://zenodo.org/record/7906067/files/figure.png" pageId="1" pageNumber="397">Fig. 1</figureCitation>
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||
) (body elongate, parallel-sided, and antennal club with three antennomeres in
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<taxonomicName box="[349,484,979,1005]" class="Insecta" family="Cryptophagidae" genus="Caenoscelis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="genus">
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<emphasis box="[349,484,979,1005]" italics="true" pageId="1" pageNumber="397">Caenoscelis</emphasis>
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</taxonomicName>
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) and the more similar
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<taxonomicName box="[754,871,979,1005]" class="Insecta" family="Cryptophagidae" genus="Renodesta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="genus">
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<emphasis box="[754,871,979,1005]" italics="true" pageId="1" pageNumber="397">Renodesta</emphasis>
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</taxonomicName>
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, known from
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<collectingRegion box="[1039,1155,979,1005]" country="United States of America" name="California" pageId="1" pageNumber="397">California</collectingRegion>
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, by the antenna with 11 antennomeres and antennal club with two antennomeres (
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<figureCitation box="[796,861,1015,1041]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1502,1526]" captionTargetBox="[152,1434,778,1477]" captionTargetId="figure-281@2.[151,1435,777,1478]" captionTargetPageId="2" captionText="FIGURES 1–5. Sternodea arcana, new species, holotype. 1) Antenna; 2) Pronotum, dorsal view showing punctation in anterior portion and transverse punctate-crenulate line; 3) Prosternal process, ventral view (not to scale); 4) Aedeagus, dorsal view; 5) Aedeagus, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906067" httpUri="https://zenodo.org/record/7906067/files/figure.png" pageId="1" pageNumber="397">Fig. 1</figureCitation>
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) (antenna with 10 antennomeres and antennal club with one antennomere in
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<taxonomicName box="[436,553,1051,1077]" class="Insecta" family="Cryptophagidae" genus="Renodesta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="genus">
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<emphasis box="[436,553,1051,1077]" italics="true" pageId="1" pageNumber="397">Renodesta</emphasis>
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</taxonomicName>
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).
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</paragraph>
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<paragraph blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="397">
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With respect to the described species from the Palearctic (
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<bibRefCitation author="Reitter, E." box="[853,998,1087,1113]" pageId="1" pageNumber="397" pagination="1 - 56" refId="ref3148" refString="Reitter, E. (1888) Bestimmungs-Tabellen der europaischen Coleopteren. XVI. Heft. Enthaltend die Familien: Erotylidae und Cryptophagidae. Verhandlungen des Naturforschenden Vereins in Brunn, 26, 1 - 56. [1887]" type="journal article" year="1888">Reitter 1888</bibRefCitation>
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;
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<bibRefCitation author="Lyubarsky, G. Y." box="[1011,1196,1087,1113]" pageId="1" pageNumber="397" pagination="53 - 62" refId="ref3058" refString="Lyubarsky, G. Y. (1987) Review of species of the genus Sternodea Reitter, 1875 (Coleoptera: Cryptophagidae: Atomariinae). Biulleten' Moskovskogo Obshchestva Ispytatelei Prirody, Otdel Biologicheskii, 92, 53 - 62." type="journal article" year="1987">Lyubarsky 1987</bibRefCitation>
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;
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<bibRefCitation author="Johnson, C. & Otero, J. C. & Leschen, R. A. B." box="[1208,1431,1087,1113]" pageId="1" pageNumber="397" refId="ref2855" refString="Johnson, C., Otero, J. C. & Leschen, R. A. B. (2007) Family Cryptophagidae. In: Lobl, I. & Smetana, A. (Eds.), A Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 513 - 531." type="book" year="2007">
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Johnson
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<emphasis box="[1308,1365,1087,1113]" italics="true" pageId="1" pageNumber="397">et al.</emphasis>
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2007
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</bibRefCitation>
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||
;
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<bibRefCitation author="Johnson, C." box="[151,311,1123,1149]" pageId="1" pageNumber="397" pagination="267 - 272" refId="ref2741" refString="Johnson, C. (2012) A new Himalayan Sternodea species, together with notes and corrections on Palaearctic Cryptophagidae (Coleoptera). Entomologists Gazette, 63 (4), 267 - 272." type="journal article" year="2012">Johnson 2012</bibRefCitation>
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;
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<bibRefCitation author="Sasaji, H." box="[322,457,1123,1149]" pageId="1" pageNumber="397" pagination="21 - 63" refId="ref3188" refString="Sasaji, H. (1984) Contribution to the taxonomy of the superfamily Cucujoidea (Coleoptera) of Japan and her adjacent districts, II. Memoirs of the Faculty of Education, Fukui University Series II (Natural Science), 34, 21 - 63." type="journal article" year="1984">Sasaji 1984</bibRefCitation>
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),
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<taxonomicName authorityName="Leschen & Gimmel" authorityYear="2023" box="[477,584,1123,1149]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="arcana">
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<emphasis box="[477,584,1123,1149]" italics="true" pageId="1" pageNumber="397">S. arcana</emphasis>
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</taxonomicName>
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can be distinguished from
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<taxonomicName authority="Reitter" authorityName="Reitter" authorityYear="1888" box="[892,1074,1123,1149]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="grilati">
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<emphasis box="[892,991,1123,1149]" italics="true" pageId="1" pageNumber="397">S. grilati</emphasis>
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Reitter
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</taxonomicName>
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and
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<taxonomicName authority="Reitter" authorityName="Reitter" authorityYear="1875" box="[1131,1316,1123,1149]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="raddei">
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<emphasis box="[1131,1232,1123,1149]" italics="true" pageId="1" pageNumber="397">S. raddei</emphasis>
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Reitter
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||
</taxonomicName>
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by having dense and coarse punctation on the pronotum (
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<figureCitation box="[684,753,1159,1185]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1502,1526]" captionTargetBox="[152,1434,778,1477]" captionTargetId="figure-281@2.[151,1435,777,1478]" captionTargetPageId="2" captionText="FIGURES 1–5. Sternodea arcana, new species, holotype. 1) Antenna; 2) Pronotum, dorsal view showing punctation in anterior portion and transverse punctate-crenulate line; 3) Prosternal process, ventral view (not to scale); 4) Aedeagus, dorsal view; 5) Aedeagus, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906067" httpUri="https://zenodo.org/record/7906067/files/figure.png" pageId="1" pageNumber="397">Fig. 2</figureCitation>
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) and sparse and fine punctation on the elytra (both surfaces coarsely [
|
||
<taxonomicName authorityName="Reitter" authorityYear="1888" box="[262,361,1195,1221]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="grilati">
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||
<emphasis box="[262,361,1195,1221]" italics="true" pageId="1" pageNumber="397">S. grilati</emphasis>
|
||
</taxonomicName>
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] or finely [
|
||
<taxonomicName authorityName="Reitter" authorityYear="1875" box="[492,594,1195,1221]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="raddei">
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||
<emphasis box="[492,594,1195,1221]" italics="true" pageId="1" pageNumber="397">S. raddei</emphasis>
|
||
</taxonomicName>
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||
] punctate in the other species), and from
|
||
<taxonomicName authority="Reitter" authorityName="Reitter" authorityYear="1875" box="[1068,1253,1195,1221]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="baudii">
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||
<emphasis box="[1068,1169,1195,1221]" italics="true" pageId="1" pageNumber="397">S. baudii</emphasis>
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Reitter
|
||
</taxonomicName>
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||
and
|
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<taxonomicName authority="Sasaji" authorityName="Sasaji" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="japonica">
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||
<emphasis box="[1309,1436,1195,1221]" italics="true" pageId="1" pageNumber="397">S. japonica</emphasis>
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Sasaji
|
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</taxonomicName>
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by the presence of a transverse punctate-crenulate line on the pronotum (
|
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<figureCitation box="[1064,1133,1231,1257]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1502,1526]" captionTargetBox="[152,1434,778,1477]" captionTargetId="figure-281@2.[151,1435,777,1478]" captionTargetPageId="2" captionText="FIGURES 1–5. Sternodea arcana, new species, holotype. 1) Antenna; 2) Pronotum, dorsal view showing punctation in anterior portion and transverse punctate-crenulate line; 3) Prosternal process, ventral view (not to scale); 4) Aedeagus, dorsal view; 5) Aedeagus, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906067" httpUri="https://zenodo.org/record/7906067/files/figure.png" pageId="1" pageNumber="397">Fig. 2</figureCitation>
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) (transverse crenulate line absent in those two species). In
|
||
<taxonomicName authorityName="Leschen & Gimmel" authorityYear="2023" box="[515,622,1267,1293]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="arcana">
|
||
<emphasis box="[515,622,1267,1293]" italics="true" pageId="1" pageNumber="397">S. arcana</emphasis>
|
||
</taxonomicName>
|
||
, the prosternal process is distinctly dilated posteriorly (
|
||
<figureCitation box="[1253,1321,1267,1293]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1502,1526]" captionTargetBox="[152,1434,778,1477]" captionTargetId="figure-281@2.[151,1435,777,1478]" captionTargetPageId="2" captionText="FIGURES 1–5. Sternodea arcana, new species, holotype. 1) Antenna; 2) Pronotum, dorsal view showing punctation in anterior portion and transverse punctate-crenulate line; 3) Prosternal process, ventral view (not to scale); 4) Aedeagus, dorsal view; 5) Aedeagus, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906067" httpUri="https://zenodo.org/record/7906067/files/figure.png" pageId="1" pageNumber="397">Fig. 3</figureCitation>
|
||
), while
|
||
<taxonomicName authority="Johnson" authorityName="Johnson" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="1" pageNumber="397">S. franzi</emphasis>
|
||
Johnson
|
||
</taxonomicName>
|
||
has the prosternal process slightly dilated posteriorly and the process is parallel-sided in
|
||
<taxonomicName box="[1341,1436,1303,1329]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="lederi">
|
||
<emphasis box="[1341,1436,1303,1329]" italics="true" pageId="1" pageNumber="397">S. lederi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[199,275,1339,1365]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="miki">
|
||
<emphasis box="[199,275,1339,1365]" italics="true" pageId="1" pageNumber="397">S. miki</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="2" lastPageNumber="398" pageId="1" pageNumber="397" type="description">
|
||
<paragraph blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="397">
|
||
<emphasis bold="true" box="[199,345,1374,1400]" pageId="1" pageNumber="397">Description.</emphasis>
|
||
Length
|
||
<quantity box="[436,537,1375,1401]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="1" pageNumber="397" unit="mm" value="1.29">1.29 mm</quantity>
|
||
, breadth
|
||
<quantity box="[638,740,1375,1401]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="397" unit="mm" value="0.82">0.82 mm</quantity>
|
||
. Body short-oval. Color of body dark reddish-brown, antennae and legs lighter. Dorsal setae uniformly golden, pronotum and elytra with long, decumbent, suberect, curved setae; microsculpture absent. Ventral surfaces with recumbent setae, about the same length compared with those on dorsum; prosternal process with sparse, short, curved setae along margins; punctation dense on most surfaces, less impressed and sparser on elytra and abdominal ventrites 2–5; abdominal ventrite 1 with punctures about as dense as on metaventrite, ventrite 5 bearing a patch of posteriorly-directed setae and sensory pegs.
|
||
</paragraph>
|
||
<paragraph blockId="1.[151,1437,762,2013]" lastBlockId="2.[151,1437,151,753]" lastPageId="2" lastPageNumber="398" pageId="1" pageNumber="397">
|
||
Antenna (
|
||
<figureCitation box="[308,372,1591,1617]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1502,1526]" captionTargetBox="[152,1434,778,1477]" captionTargetId="figure-281@2.[151,1435,777,1478]" captionTargetPageId="2" captionText="FIGURES 1–5. Sternodea arcana, new species, holotype. 1) Antenna; 2) Pronotum, dorsal view showing punctation in anterior portion and transverse punctate-crenulate line; 3) Prosternal process, ventral view (not to scale); 4) Aedeagus, dorsal view; 5) Aedeagus, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906067" httpUri="https://zenodo.org/record/7906067/files/figure.png" pageId="1" pageNumber="397">Fig. 1</figureCitation>
|
||
) with 11 antennomeres; antennomere 3 shorter than 4 and 5 combined, lengths of antennomeres 4 and 5 equal and slightly longer than antennomeres 6 and 8, antennomere 9 slightly wider than antennomere 8, antennomere 11 asymmetrical and twice the length of antennomere 10. Eye with 10 ommatidia. Pronotum (
|
||
<figureCitation box="[1360,1428,1663,1689]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1502,1526]" captionTargetBox="[152,1434,778,1477]" captionTargetId="figure-281@2.[151,1435,777,1478]" captionTargetPageId="2" captionText="FIGURES 1–5. Sternodea arcana, new species, holotype. 1) Antenna; 2) Pronotum, dorsal view showing punctation in anterior portion and transverse punctate-crenulate line; 3) Prosternal process, ventral view (not to scale); 4) Aedeagus, dorsal view; 5) Aedeagus, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906067" httpUri="https://zenodo.org/record/7906067/files/figure.png" pageId="1" pageNumber="397">Fig. 2</figureCitation>
|
||
) with length/width ratio ~0.61; front margin weakly procurved and anterior angles projecting; lateral carina lacking a distinct bead (absent at base), margin irregularly and weakly crenulate; posterior margin sinuate, with a weak scutellar lobe; setae long, regular and curved; with slightly procurved transverse punctate-crenulate line present at middle third, loosely extended by a row of larger punctures that end near the posterior margin; punctation relatively uniform over most of disc, punctures separated by about 1.3 diameters, with smooth and shiny interspaces, separated by about 2–4 diameters and finer behind transverse line. Hypomeron densely punctate, punctures separated by about 1.3 diameters. Prosternal process (
|
||
<figureCitation box="[551,622,1915,1941]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1502,1526]" captionTargetBox="[152,1434,778,1477]" captionTargetId="figure-281@2.[151,1435,777,1478]" captionTargetPageId="2" captionText="FIGURES 1–5. Sternodea arcana, new species, holotype. 1) Antenna; 2) Pronotum, dorsal view showing punctation in anterior portion and transverse punctate-crenulate line; 3) Prosternal process, ventral view (not to scale); 4) Aedeagus, dorsal view; 5) Aedeagus, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906067" httpUri="https://zenodo.org/record/7906067/files/figure.png" pageId="1" pageNumber="397">Fig. 3</figureCitation>
|
||
) elevated,
|
||
<quantity box="[753,859,1915,1941]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="397" unit="mm" value="0.24">0.24 mm</quantity>
|
||
, anteriorly acute with a short longitudinal carina, strongly dilated posteriorly, about 2.0× longer than width at posterior margin, posterior margin evenly arcuate, marginal bead present. Elytra with sides rounded near base, about 1.05× longer than combined width, about 1.15× wider than pronotum at greatest width (at basal one-fourth), about 2.22× longer than pronotum at greatest length; setae dual, rows of suberect curved setae (approximately
|
||
<quantity box="[785,885,187,213]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="398" unit="mm" value="0.08">0.08 mm</quantity>
|
||
) alternating with rows of more adpressed setae of about similar lengths; punctation less dense and less impressed than on pronotum, punctures one-fourth to one-half the diameter of those on pronotum, denser near basal margin and separated by 1–2 puncture diameters, sparser on disc. Abdominal ventrite 1 with moderately broad intercoxal process, femoral lines and median carina absent, bead broadened near base of intercoxal process.
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,753]" pageId="2" pageNumber="398">
|
||
Aedeagus (
|
||
<figureCitation box="[327,436,366,393]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1502,1526]" captionTargetBox="[152,1434,778,1477]" captionTargetId="figure-281@2.[151,1435,777,1478]" captionTargetPageId="2" captionText="FIGURES 1–5. Sternodea arcana, new species, holotype. 1) Antenna; 2) Pronotum, dorsal view showing punctation in anterior portion and transverse punctate-crenulate line; 3) Prosternal process, ventral view (not to scale); 4) Aedeagus, dorsal view; 5) Aedeagus, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906067" httpUri="https://zenodo.org/record/7906067/files/figure.png" pageId="2" pageNumber="398">Figs. 4, 5</figureCitation>
|
||
) with tegminal strut absent, tegminal arms broad and fused at middle without a suture; parameres separate and partially fused to phallobase, thick and orbiculate, each with an inner dorsal cavity, apically truncate, length about 1.2× longer than width, apices unisetose; basipenis 3.3× longer than distipenis, with a welldeveloped median keel on ventral surface about half the length distipenis; distipenis relatively short (about 1.2× longer than wide), internal sclerites symmetrical; internal sac (broken) lacking sclerites; straight, not kinked.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="398" type="discussion">
|
||
<paragraph blockId="2.[151,1437,151,753]" pageId="2" pageNumber="398">
|
||
<emphasis bold="true" box="[199,315,547,573]" pageId="2" pageNumber="398">Remarks.</emphasis>
|
||
This species is known only from the male
|
||
<typeStatus box="[788,890,547,573]" pageId="2" pageNumber="398">holotype</typeStatus>
|
||
. The abdomen (mounted dorsally), the terminalia and the aedeagus were mounted separately from the remaining body on the card with a male symbol. It is missing much of the protarsomeres (the right one absent and tarsomeres 4 and 5 are missing from the left), the left hind leg, and the anterior portion of the internal sac was also broken.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="398" type="etymology">
|
||
<paragraph blockId="2.[151,1437,151,753]" pageId="2" pageNumber="398">
|
||
<emphasis bold="true" box="[199,334,691,717]" pageId="2" pageNumber="398">Etymology.</emphasis>
|
||
The specific epithet is from the Latin adjective
|
||
<taxonomicName authorityName="Leschen & Gimmel" authorityYear="2023" box="[900,978,691,716]" class="Insecta" family="Cryptophagidae" genus="Sternodea" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="398" phylum="Arthropoda" rank="species" species="arcana">
|
||
<emphasis box="[900,978,691,716]" italics="true" pageId="2" pageNumber="398">arcana</emphasis>
|
||
</taxonomicName>
|
||
, meaning “mysterious”, alluding to the mysterious geographic origin of the
|
||
<typeStatus box="[558,660,727,753]" pageId="2" pageNumber="398">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |