278 lines
37 KiB
XML
278 lines
37 KiB
XML
<document id="97A025611416D95A256CDD5EDD7953F6" ID-CLB-Dataset="4845" ID-DOI="10.11646/zootaxa.502.1.1" ID-GBIF-Dataset="76aa06a8-f646-4c5a-b596-54bfd8875223" ID-ISSN="11755334" ID-Zenodo-Dep="5030148" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624553872186" checkinUser="felipe" docAuthor="Jameson, Mary Liz & Wada, Kaoru" docDate="2004" docId="03A4A725CA5FFFF9FEC6FBA1FD26FBF2" docLanguage="en" docName="zt00502.pdf" docOrigin="Zootaxa 502 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.502.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Peltonotus adelphosimilis Jameson and Wada 2004, n. sp." docType="treatment" docVersion="8" lastPageNumber="16" masterDocId="FF9DDF5DCA52FFF6FFCEFF9DFFB5FF8C" masterDocTitle="Revision of the genus Peltonotus Burmeister (Coleoptera: Scarabaeidae: Dynastinae) from Southeastern Asia" masterLastPageNumber="66" masterPageNumber="1" pageNumber="14" updateTime="1699020591651" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="DFCBA9022BA744220D5954718C6459CA">
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<mods:title id="5483A397CBA36F9E2B3A1FB118FB8F71">Revision of the genus Peltonotus Burmeister (Coleoptera: Scarabaeidae: Dynastinae) from Southeastern Asia</mods:title>
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<mods:name id="5318DB81871FD0D68D60289835B3C66B" type="personal">
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<mods:role id="1EF947CC846A8BA34F02265853E30C48">
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<mods:roleTerm id="0D17699D042BAA35570051B5518F6DD7">Author</mods:roleTerm>
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<mods:namePart id="97A115FCE7A2FB2695E9B012D59A632F">Jameson, Mary Liz</mods:namePart>
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<mods:affiliation id="1923D82C6756E9F9D430AB2000E80E77">University of Nebraska State Museum, Division of Entomology, Lincoln, NE U. S. A., 68588 - 0514 mjameson 1 @ unl. edu</mods:affiliation>
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<mods:roleTerm id="81E32B3560C9A0E51DE5F93491F360DF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6278AA4AE4BFDD0DE88BA9E56D5F1197">Wada, Kaoru</mods:namePart>
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<mods:affiliation id="FC61B8D487DCEF1A07B452318C553433">Musashimurayama Daini Junior School, 1460 Midorigaoka, Musashimurayama-shi, Tokyo 208 - 0012 Japan kwada 007 @ hotmail. com</mods:affiliation>
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<mods:typeOfResource id="12BAA9B29BE71DBE2620B17F934EFFC3">text</mods:typeOfResource>
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<mods:title id="BABAE5FC0D43B83DE9342D09A9B9143E">Zootaxa</mods:title>
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<mods:part id="340BF980F62AFED8D79FEF6633F6BE42">
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<mods:date id="6843B742621A58EB197B3CB17683C609">2004</mods:date>
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<mods:number id="B10CD3958A1A343EE29489958A7CDA52">2004-05-04</mods:number>
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<mods:number id="8E97CD3E23703448978FD20E59621B56">502</mods:number>
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<mods:url id="C1CDC955489D0CD1BF0E408725F391B6">https://biotaxa.org/Zootaxa/article/view/zootaxa.502.1.1</mods:url>
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<mods:classification id="3C1EE1ADD1379A23D4F9B9471CA289A0">journal article</mods:classification>
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<treatment id="03A4A725CA5FFFF9FEC6FBA1FD26FBF2" ID-DOI="http://doi.org/10.5281/zenodo.5248445" ID-GBIF-Taxon="184930980" ID-Zenodo-Dep="5248445" LSID="urn:lsid:plazi:treatment:03A4A725CA5FFFF9FEC6FBA1FD26FBF2" httpUri="http://treatment.plazi.org/id/03A4A725CA5FFFF9FEC6FBA1FD26FBF2" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
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<subSubSection id="C31745B8CA5FFFFBFEC6FBA1FC3EFBDA" box="[264,907,1084,1110]" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="8BB21633CA5FFFFBFEC6FBA1FC3EFBDA" blockId="13.[264,907,1084,1145]" box="[264,907,1084,1110]" pageId="13" pageNumber="14">
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<heading id="D0FAA15FCA5FFFFBFEC6FBA1FC3EFBDA" bold="true" box="[264,907,1084,1110]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis id="B979CA21CA5FFFFBFEC6FBA1FC3EFBDA" bold="true" box="[264,907,1084,1110]" pageId="13" pageNumber="14">
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<taxonomicName id="4C0D6DB0CA5FFFFBFEC6FBA1FC8EFBDA" authority="Jameson and Wada" authorityName="Jameson and Wada" authorityYear="2004" box="[264,827,1084,1110]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="adelphosimilis" status="sp. nov.">
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<emphasis id="B979CA21CA5FFFFBFEC6FBA1FD8BFBDA" bold="true" box="[264,574,1084,1110]" italics="true" pageId="13" pageNumber="14">Peltonotus adelphosimilis</emphasis>
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Jameson and Wada
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24A775ACA5FFFFBFC86FBA1FC3EFBDA" box="[840,907,1084,1110]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31745B8CA5FFFF9FEC6FBC2FC26FD4A" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
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<paragraph id="8BB21633CA5FFFFBFEC6FBC2FE4EFBF5" blockId="13.[264,907,1084,1145]" box="[264,507,1119,1145]" pageId="13" pageNumber="14">
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(
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<figureCitation id="13360AB6CA5FFFFBFEDFFBC2FEC6FBF5" box="[273,371,1119,1145]" captionStart="FIGURES 31–36" captionStartId="55.[264,382,1495,1519]" captionTargetBox="[342,1225,329,1468]" captionTargetId="figure-95@55.[337,1249,326,1471]" captionTargetPageId="55" captionText="FIGURES 31–36. Detail of right forelegs of: 31) P. malayensis (showing bidentate foretibia, dorsal view), 32) P.rubripennis (showing tridentate foretibia with basal tooth very weakly developed, dorsal view), 33) P. deltamentum (showing strongly arcuate smaller claw, ventral view), 34) P. nasutus (showing form of foretarsomeres 1–5), 35) P.similis (showing form of foretarsomeres 1–5 and median protuberance of foretarsomere 5 [arrow]), 36) P. adelphosimilis (showing form of foretarsomeres 1–5 and apical protuberance of foretarsomere 5 [arrow])." figureDoi="http://doi.org/10.5281/zenodo.5030172" httpUri="https://zenodo.org/record/5030172/files/figure.png" pageId="13" pageNumber="14">Figs. 36</figureCitation>
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,
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<figureCitation id="13360AB6CA5FFFFBFEB1FBC2FE72FBF5" box="[383,455,1119,1145]" captionStart="FIGURES 37–39" captionStartId="56.[264,382,1718,1742]" captionTargetBox="[476,1110,329,1684]" captionTargetId="figure-45@56.[466,1120,326,1694]" captionTargetPageId="56" captionText="FIGURES 37–39. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 37) P. adelphosimilis, 38) P. brunnipennis, 39) P. deltomentum." figureDoi="http://doi.org/10.5281/zenodo.5030178" httpUri="https://zenodo.org/record/5030178/files/figure.png" pageId="13" pageNumber="14">37a–c</figureCitation>
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, 67)
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</paragraph>
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<paragraph id="8BB21633CA5FFFFBFEC6FB32FE37FA35" blockId="13.[264,1325,1199,1905]" pageId="13" pageNumber="14">
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Type Material.
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<materialsCitation id="3B651C6ECA5FFFFBFE0DFB32FE52FACD" ID-GBIF-Occurrence="3337342355" collectingDate="1990-10" collectionCode="WADA" collectorName="Col. No." location="Mt. Bawang" pageId="13" pageNumber="14" specimenCount="2" stateProvince="Kalimantan Barat" typeStatus="holotype">
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<typeStatus id="54B6A891CA5FFFFBFE0DFB32FD85FB45" box="[451,560,1199,1225]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
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male housed at
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<collectionCode id="ED1C8EF6CA5FFFFBFD21FB32FCF2FB45" box="[751,839,1199,1225]" country="Australia" httpUri="http://grbio.org/cool/mnrx-zeat" name="Western Australia Department of Agriculture" pageId="13" pageNumber="14">WADA</collectionCode>
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with following label data and male genitalia mounted beneath specimen: a) "
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<location id="8ED240E8CA5FFFFBFD0CFB4AFCECFB7D" LSID="urn:lsid:plazi:treatment:03A4A725CA5FFFF9FEC6FBA1FD26FBF2:8ED240E8CA5FFFFBFD0CFB4AFCECFB7D" box="[706,857,1239,1265]" name="Mt. Bawang" pageId="13" pageNumber="14" stateProvince="Kalimantan Barat">Mt. Bawang</location>
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,
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<collectingRegion id="49C9D8D1CA5FFFFBFCABFB4AFB86FB7D" box="[869,1075,1239,1265]" country="Indonesia" name="Kalimantan Barat" pageId="13" pageNumber="14">West Kalimantan</collectingRegion>
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, BORNEO,
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<date id="FFB330F3CA5FFFFBFB09FB4AFA97FB7D" box="[1223,1314,1239,1265]" pageId="13" pageNumber="14" value="1990-10">
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<collectingDate id="EFF7C91BCA5FFFFBFB09FB4AFA97FB7D" box="[1223,1314,1239,1265]" pageId="13" pageNumber="14" value="1990-10">X 1990</collectingDate>
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</date>
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" (type face), b) "Kaoru
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<collectionCode id="ED1C8EF6CA5FFFFBFDC0FB62FDD3FA95" box="[526,614,1279,1305]" country="Australia" httpUri="http://grbio.org/cool/mnrx-zeat" name="Western Australia Department of Agriculture" pageId="13" pageNumber="14">WADA</collectionCode>
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||
<collectorName id="26F873E5CA5FFFFBFDA3FB62FD65FA95" box="[621,720,1279,1305]" pageId="13" pageNumber="14">Col. No.</collectorName>
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028b" (typeface and handwritten, yellow label), c) our
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<typeStatus id="54B6A891CA5FFFFBFEF7FABAFE2AFACD" box="[313,415,1319,1345]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
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label
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</materialsCitation>
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.
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<materialsCitation id="3B651C6ECA5FFFFBFDCEFABAFECAFA35" ID-GBIF-Occurrence="3337342333" collectionCode="FUJI" collectorName="N. Nishikawa-leg." country="Indonesia" location="Mt. Bawang" pageId="13" pageNumber="14" specimenCount="2" typeStatus="allotype">
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<typeStatus id="54B6A891CA5FFFFBFDCEFABAFDD3FACD" box="[512,614,1319,1345]" pageId="13" pageNumber="14" type="allotype">Allotype</typeStatus>
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female housed at
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<collectionCode id="ED1C8EF6CA5FFFFBFC87FABAFC36FACD" box="[841,899,1319,1345]" pageId="13" pageNumber="14">FUJI</collectionCode>
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with following label data: a) "
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||
<location id="8ED240E8CA5FFFFBFB32FABAFEDFFAE5" LSID="urn:lsid:plazi:treatment:03A4A725CA5FFFF9FEC6FBA1FD26FBF2:8ED240E8CA5FFFFBFB32FABAFEDFFAE5" country="Indonesia" name="Mt. Bawang" pageId="13" pageNumber="14">Mt. Bawang</location>
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, BORNEO
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<collectingCountry id="F31A56A3CA5FFFFBFDC0FAD2FD18FAE5" box="[526,685,1359,1385]" name="Indonesia" pageId="13" pageNumber="14">INDONESIA</collectingCountry>
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1993. AUG.
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<collectorName id="26F873E5CA5FFFFBFCA2FAD2FBE7FAE5" box="[876,1106,1359,1385]" pageId="13" pageNumber="14">N. Nishikawaleg.</collectorName>
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" (type face), b) "FujiokaCollection PeltonotusT1" (typeface and handwritten, yellow label), c) our
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<typeStatus id="54B6A891CA5FFFFBFB3BFAEAFE8FFA35" pageId="13" pageNumber="14" type="allotype">allotype</typeStatus>
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label
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BB21633CA5FFFF8FEF6FA5AFDA9FB2A" blockId="13.[264,1325,1199,1905]" lastBlockId="14.[264,1324,284,1870]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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Description.
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<typeStatus id="54B6A891CA5FFFFBFE2AFA5AFDE4FA6D" box="[484,593,1479,1505]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
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male. Length
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<quantity id="4CF5BBD6CA5FFFFBFCD2FA5AFC3AFA6D" box="[796,911,1479,1505]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="13" pageNumber="14" unit="mm" value="20.3">20.3 mm</quantity>
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. Widest width
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<quantity id="4CF5BBD6CA5FFFFBFB91FA5AFB77FA6D" box="[1119,1218,1479,1505]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="13" pageNumber="14" unit="mm" value="9.4">9.4 mm</quantity>
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. Color: Clypeus, pronotum, scutellum, elytra, propygidium, pygidium, and venter dark brown; frons black or castaneous; elytra with or without iridescent bloom. Head: Surface of frons impunctate to sparsely punctate (base to middisc), densely punctate (middisc to apex); punctures simple, multisetigerous; setae short (1–20+ per puncture) and moderately long (1–2 per puncture). Surface of clypeus densely punctate; punctures simple, multisetigerous; setae short (1–20+ per puncture) and moderately long (1–2 per puncture). Clypeus laterally weakly bowed, apex truncate, corners square, beaded; bead weakly arcuate posteriorly. Labrum deeply, narrowly emarginate at middle. Mandibles rounded laterally, inner apex with 2 teeth.
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<collectingRegion id="49C9D8D1CA5FFFFBFE25F8B2FDFAF8C5" box="[491,591,1839,1865]" country="Myanmar" name="Chin" pageId="13" pageNumber="14">Mentum</collectingRegion>
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with apical half rounded, notched at middle; palpomere 2 not obviously flattened dorsoventrally, 1.5 times width of palpomere 1, sparsely setose; setae moderately long, rufous, not curled at apex, not flattened. Maxilla: mala lacking lamellate setal brush; stipes with setae sparse, long, not flattened, not curled at apex; palpomere 2 with weak internomedial bump. Antennal club subequal to segments 2–7. Pronotum: Bead lacking anterior to scutellum. Surface moderately densely punctate; punctures simple, multisetigerous laterally; setae minute (1–7 per puncture) and short (1 per puncture). Lateral margin lacking long setae. Elytral sutural length: About 4.0 times length of scutellum. Elytron: Surface with 5 poorly developed, punctate, longitudinal striae between suture and humerus; punctures moderate in size, moderately dense, some multisetigerous; setae minute (1–5 per puncture). Intervals with similar sculpturing. Propygidium: Surface moderately densely punctate with a row of setose punctures at apex; punctures simple, unisetigerous; setae short, rufous. Pygidium: Surface densely punctate and shagreened; punctures ocellate, mixed small and moderately large, multisetigerous; setae minute (1–7 per puncture) and short (1 per puncture). Venter: Prosternal keel subtriangular; apex projecting anteriorly at about 90° with respect to ventral plane; apex produced to middle of protrochanter, rounded. Legs: Foretibia of male bidentate; lateral margin with foretarsomeres 1–4 with apices expanded, dorsal and ventral apices clothed with dense, short setae; protarsomere
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<quantity id="4CF5BBD6CA5CFFF8FD06FC01FD43FC3A" box="[712,758,924,950]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="14" pageNumber="15" unit="in" value="5.0">5 in</quantity>
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male with apical protuberance (
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<figureCitation id="13360AB6CA5CFFF8FBBAFC01FB78FC3A" box="[1140,1229,924,950]" captionStart="FIGURES 31–36" captionStartId="55.[264,382,1495,1519]" captionTargetBox="[342,1225,329,1468]" captionTargetId="figure-95@55.[337,1249,326,1471]" captionTargetPageId="55" captionText="FIGURES 31–36. Detail of right forelegs of: 31) P. malayensis (showing bidentate foretibia, dorsal view), 32) P.rubripennis (showing tridentate foretibia with basal tooth very weakly developed, dorsal view), 33) P. deltamentum (showing strongly arcuate smaller claw, ventral view), 34) P. nasutus (showing form of foretarsomeres 1–5), 35) P.similis (showing form of foretarsomeres 1–5 and median protuberance of foretarsomere 5 [arrow]), 36) P. adelphosimilis (showing form of foretarsomeres 1–5 and apical protuberance of foretarsomere 5 [arrow])." figureDoi="http://doi.org/10.5281/zenodo.5030172" httpUri="https://zenodo.org/record/5030172/files/figure.png" pageId="14" pageNumber="15">Fig. 36</figureCitation>
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). Foreclaws with inner claw sickleshaped, about 4 times thicker than outer claw; outer claw simply arcuate, about 1/3 the length of inner claw; empodium bulbous at base. Meso and metatibial claws of male with 2 setae, claws angled toward venter, about 2/3 length of metatarsomere 5. Metatibia of male with apical spurs nearly straight; ventral spur produced to middle of metatarsomere 1, dorsal spur produced to middle of metatarsomere 2. Parameres:
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||
<figureCitation id="13360AB6CA5CFFF8FE57FB11FDADFB2A" box="[409,536,1164,1190]" captionStart="FIGURES 37–39" captionStartId="56.[264,382,1718,1742]" captionTargetBox="[476,1110,329,1684]" captionTargetId="figure-45@56.[466,1120,326,1694]" captionTargetPageId="56" captionText="FIGURES 37–39. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 37) P. adelphosimilis, 38) P. brunnipennis, 39) P. deltomentum." figureDoi="http://doi.org/10.5281/zenodo.5030178" httpUri="https://zenodo.org/record/5030178/files/figure.png" pageId="14" pageNumber="15">Fig. 37a–c</figureCitation>
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.
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||
</paragraph>
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||
<paragraph id="8BB21633CA5CFFF8FEF6FB29FD7BF982" blockId="14.[264,1324,284,1870]" pageId="14" pageNumber="15">
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||
<typeStatus id="54B6A891CA5CFFF8FEF6FB29FE2BFB42" box="[312,414,1204,1230]" pageId="14" pageNumber="15" type="allotype">Allotype</typeStatus>
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Female. Differs from the
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<typeStatus id="54B6A891CA5CFFF8FD28FB29FCF8FB42" box="[742,845,1204,1230]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
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||
in the following respects. Length
|
||
<quantity id="4CF5BBD6CA5CFFF8FB36FB29FE8CFB7A" metricMagnitude="-2" metricUnit="m" metricValue="1.89" pageId="14" pageNumber="15" unit="mm" value="18.9">18.9 mm</quantity>
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||
. Widest width
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<quantity id="4CF5BBD6CA5CFFF8FE3AFB41FDE7FB7A" box="[500,594,1244,1270]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="14" pageNumber="15" unit="mm" value="8.9">8.9 mm</quantity>
|
||
. Color: Head, pronotum, scutellum, propygidium, pygidium, and venter castaneous; elytra castaneous with iridescent bloom. Head: Surface of frons moderately densely punctate (base) to densely punctate (middisc to apex); punctures simple, multisetigerous; setae short (1–12 per puncture) and moderately long (1 per puncture). Pronotum: Surface moderately densely punctate, punctures simple, lacking setae laterally. Elytron: Epipleuron of female in ventral view expanded, deeply incised at sternite 3; in dorsal view, expansion welldeveloped (similar to
|
||
<figureCitation id="13360AB6CA5CFFF8FC97FA51FC1BFA6A" box="[857,942,1484,1510]" captionStart="FIGURES 61–64" captionStartId="63.[264,382,1259,1283]" captionTargetBox="[427,1142,361,1231]" captionTargetId="figure-43@63.[419,1168,361,1236]" captionTargetPageId="63" captionText="FIGURES 61–64. Elytral epipleuron (gray, ventral view) with position of metacoxa of: 61) P. similis, 62) P. sisyrus, 63) P. suehirogarus, 64a–b) P. vittatus." figureDoi="http://doi.org/10.5281/zenodo.5030200" httpUri="https://zenodo.org/record/5030200/files/figure.png" pageId="14" pageNumber="15">Fig. 61</figureCitation>
|
||
). Legs: Foreclaws 1/2 length of foretarsomere 5, angled toward venter.
|
||
</paragraph>
|
||
<paragraph id="8BB21633CA5CFFF9FEF6F981FB01FE72" blockId="14.[264,1324,284,1870]" lastBlockId="15.[264,1323,284,1150]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
|
||
Diagnosis. Based on shared characters,
|
||
<taxonomicName id="4C0D6DB0CA5CFFF8FCD2F981FC52F9B9" authorityName="Jameson and Wada" authorityYear="2004" box="[796,999,1564,1589]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="adelphosimilis">
|
||
<emphasis id="B979CA21CA5CFFF8FCD2F981FC52F9B9" box="[796,999,1564,1589]" italics="true" pageId="14" pageNumber="15">P. adelphosimilis</emphasis>
|
||
</taxonomicName>
|
||
most closely resembles
|
||
<taxonomicName id="4C0D6DB0CA5CFFF8FADBF981FEE1F9D1" authorityName="Arrow. A" authorityYear="1931" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B979CA21CA5CFFF8FADBF981FEE1F9D1" italics="true" pageId="14" pageNumber="15">P. similis</emphasis>
|
||
</taxonomicName>
|
||
. The two species are separated based on the form of the labrum (deeply, narrowly emarginate in
|
||
<taxonomicName id="4C0D6DB0CA5CFFF8FE7DF9F1FDC8F909" authorityName="Jameson and Wada" authorityYear="2004" box="[435,637,1644,1669]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="adelphosimilis">
|
||
<emphasis id="B979CA21CA5CFFF8FE7DF9F1FDC8F909" box="[435,637,1644,1669]" italics="true" pageId="14" pageNumber="15">P. adelphosimilis</emphasis>
|
||
</taxonomicName>
|
||
[e.g.,
|
||
<figureCitation id="13360AB6CA5CFFF8FD05F9F1FC91F90A" box="[715,804,1644,1670]" captionStart="FIGURES 14–18" captionStartId="53.[264,382,1425,1449]" captionTargetBox="[340,1245,341,1398]" captionTargetId="figure-118@53.[337,1249,326,1401]" captionTargetPageId="53" captionText="FIGURES 14–18. Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 14) P.karubei (apex of labrum deeply, narrowly emarginated; apex of mandible rounded laterally), 15) P. malayensis (apex of labrum broadly emarginate; apex of mandible rounded laterally), 16) P. morio (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male), 17) P. nasutus (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male), 18) P. vittatus (apex of labrum broadly emarginate; apex of mandible obliquely quadrate laterally)." figureDoi="http://doi.org/10.5281/zenodo.5030160" httpUri="https://zenodo.org/record/5030160/files/figure.png" pageId="14" pageNumber="15">Fig. 14</figureCitation>
|
||
]; broadly emarginate in
|
||
<taxonomicName id="4C0D6DB0CA5CFFF8FB84F9F1FB00F909" authorityName="Arrow. A" authorityYear="1931" box="[1098,1205,1644,1669]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B979CA21CA5CFFF8FB84F9F1FB00F909" box="[1098,1205,1644,1669]" italics="true" pageId="14" pageNumber="15">P. similis</emphasis>
|
||
</taxonomicName>
|
||
), form of foretarsomere
|
||
<quantity id="4CF5BBD6CA5CFFF8FE78F909FE52F922" box="[438,487,1684,1710]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="14" pageNumber="15" unit="in" value="5.0">5 in</quantity>
|
||
males (with apical protuberance in
|
||
<taxonomicName id="4C0D6DB0CA5CFFF8FC6CF909FBDBF921" authorityName="Jameson and Wada" authorityYear="2004" box="[930,1134,1684,1709]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="adelphosimilis">
|
||
<emphasis id="B979CA21CA5CFFF8FC6CF909FBDBF921" box="[930,1134,1684,1709]" italics="true" pageId="14" pageNumber="15">P. adelphosimilis</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="13360AB6CA5CFFF8FB4EF909FB6BF922" box="[1152,1246,1684,1710]" captionStart="FIGURES 31–36" captionStartId="55.[264,382,1495,1519]" captionTargetBox="[342,1225,329,1468]" captionTargetId="figure-95@55.[337,1249,326,1471]" captionTargetPageId="55" captionText="FIGURES 31–36. Detail of right forelegs of: 31) P. malayensis (showing bidentate foretibia, dorsal view), 32) P.rubripennis (showing tridentate foretibia with basal tooth very weakly developed, dorsal view), 33) P. deltamentum (showing strongly arcuate smaller claw, ventral view), 34) P. nasutus (showing form of foretarsomeres 1–5), 35) P.similis (showing form of foretarsomeres 1–5 and median protuberance of foretarsomere 5 [arrow]), 36) P. adelphosimilis (showing form of foretarsomeres 1–5 and apical protuberance of foretarsomere 5 [arrow])." figureDoi="http://doi.org/10.5281/zenodo.5030172" httpUri="https://zenodo.org/record/5030172/files/figure.png" pageId="14" pageNumber="15">Fig. 36</figureCitation>
|
||
]; with median protuberance in
|
||
<taxonomicName id="4C0D6DB0CA5CFFF8FDE2F921FD2DF959" authorityName="Arrow. A" authorityYear="1931" box="[556,664,1724,1749]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B979CA21CA5CFFF8FDE2F921FD2DF959" box="[556,664,1724,1749]" italics="true" pageId="14" pageNumber="15">P. similis</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="13360AB6CA5CFFF8FD67F921FCB0F95A" box="[681,773,1724,1750]" captionStart="FIGURES 31–36" captionStartId="55.[264,382,1495,1519]" captionTargetBox="[342,1225,329,1468]" captionTargetId="figure-95@55.[337,1249,326,1471]" captionTargetPageId="55" captionText="FIGURES 31–36. Detail of right forelegs of: 31) P. malayensis (showing bidentate foretibia, dorsal view), 32) P.rubripennis (showing tridentate foretibia with basal tooth very weakly developed, dorsal view), 33) P. deltamentum (showing strongly arcuate smaller claw, ventral view), 34) P. nasutus (showing form of foretarsomeres 1–5), 35) P.similis (showing form of foretarsomeres 1–5 and median protuberance of foretarsomere 5 [arrow]), 36) P. adelphosimilis (showing form of foretarsomeres 1–5 and apical protuberance of foretarsomere 5 [arrow])." figureDoi="http://doi.org/10.5281/zenodo.5030172" httpUri="https://zenodo.org/record/5030172/files/figure.png" pageId="14" pageNumber="15">Fig. 35</figureCitation>
|
||
]), and form of the female epipleuron (incised area round in
|
||
<taxonomicName id="4C0D6DB0CA5CFFF8FE7DF979FDC8F971" authorityName="Jameson and Wada" authorityYear="2004" box="[435,637,1764,1789]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="adelphosimilis">
|
||
<emphasis id="B979CA21CA5CFFF8FE7DF979FDC8F971" box="[435,637,1764,1789]" italics="true" pageId="14" pageNumber="15">P. adelphosimilis</emphasis>
|
||
</taxonomicName>
|
||
; ovate in
|
||
<taxonomicName id="4C0D6DB0CA5CFFF8FD34F979FCD3F971" authorityName="Arrow. A" authorityYear="1931" box="[762,870,1764,1789]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B979CA21CA5CFFF8FD34F979FCD3F971" box="[762,870,1764,1789]" italics="true" pageId="14" pageNumber="15">P. similis</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="13360AB6CA5CFFF8FCB6F979FC60F972" box="[888,981,1764,1790]" captionStart="FIGURES 61–64" captionStartId="63.[264,382,1259,1283]" captionTargetBox="[427,1142,361,1231]" captionTargetId="figure-43@63.[419,1168,361,1236]" captionTargetPageId="63" captionText="FIGURES 61–64. Elytral epipleuron (gray, ventral view) with position of metacoxa of: 61) P. similis, 62) P. sisyrus, 63) P. suehirogarus, 64a–b) P. vittatus." figureDoi="http://doi.org/10.5281/zenodo.5030200" httpUri="https://zenodo.org/record/5030200/files/figure.png" pageId="14" pageNumber="15">Fig. 61</figureCitation>
|
||
]). The following characters will separate
|
||
<taxonomicName id="4C0D6DB0CA5CFFF8FE60F891FDCFF8A9" authorityName="Jameson and Wada" authorityYear="2004" box="[430,634,1804,1829]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="adelphosimilis">
|
||
<emphasis id="B979CA21CA5CFFF8FE60F891FDCFF8A9" box="[430,634,1804,1829]" italics="true" pageId="14" pageNumber="15">P. adelphosimilis</emphasis>
|
||
</taxonomicName>
|
||
from other
|
||
<taxonomicName id="4C0D6DB0CA5CFFF8FCDAF891FC43F8AA" box="[788,1014,1804,1830]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B979CA21CA5CFFF8FCDAF891FC24F8A9" box="[788,913,1804,1829]" italics="true" pageId="14" pageNumber="15">Peltonotus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
: mandibles of male and female rounded laterally, inner apex with 2 teeth; mentum with apical half rounded, palpomere 2 not obviously flattened or greatly enlarged and lacking curled setae; mala lacking lamellate setal brush; stipes with setae sparse and not curled at apex; form of female epipleuron (dorsal and ventral views); elytra with multisetigerous punctures; foretarsomere 5 of male subequal in length to tarsomere 5 with medial protuberance; foretarsomeres 1–4 with apices expanded, dorsal and ventral apices clothed with dense, short setae; dorsal metatibial spur produced to middle of metatarsomere 2, nearly straight in males.
|
||
</paragraph>
|
||
<paragraph id="8BB21633CA5DFFF9FEF6FD91FD4CFDFA" blockId="15.[264,1323,284,1150]" pageId="15" pageNumber="16">
|
||
Etymology. The species epithet,
|
||
<taxonomicName id="4C0D6DB0CA5DFFF9FD06FD91FCC6FDA9" authorityName="Jameson and Wada" authorityYear="2004" box="[712,883,524,549]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="adelphosimilis">
|
||
<emphasis id="B979CA21CA5DFFF9FD06FD91FCC6FDA9" box="[712,883,524,549]" italics="true" pageId="15" pageNumber="16">adelphosimilis</emphasis>
|
||
</taxonomicName>
|
||
, is derived from the Greek word for near kinship, twin, or brother (
|
||
<emphasis id="B979CA21CA5DFFF9FDB2FDA9FD53FDC1" box="[636,742,564,589]" italics="true" pageId="15" pageNumber="16">adelphos</emphasis>
|
||
). The combination,
|
||
<emphasis id="B979CA21CA5DFFF9FC2BFDA9FB3DFDC1" box="[997,1160,564,589]" italics="true" pageId="15" pageNumber="16">adephosimilis</emphasis>
|
||
, refers to the near kinship of this species with
|
||
<taxonomicName id="4C0D6DB0CA5DFFF9FD46FDC1FD44FDF9" authorityName="Arrow. A" authorityYear="1931" box="[648,753,604,629]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B979CA21CA5DFFF9FD46FDC1FD44FDF9" box="[648,753,604,629]" italics="true" pageId="15" pageNumber="16">P. similis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB21633CA5DFFF9FEF6FD19FBC3FD12" blockId="15.[264,1323,284,1150]" box="[312,1142,644,670]" pageId="15" pageNumber="16">
|
||
Distribution (Fig. 67).
|
||
<collectingRegion id="49C9D8D1CA5DFFF9FD87FD19FC33FD12" box="[585,902,644,670]" country="Indonesia" name="Kalimantan Barat" pageId="15" pageNumber="16">West Kalimantan Province</collectingRegion>
|
||
, Bornean
|
||
<collectingCountry id="F31A56A3CA5DFFF9FC32FD19FBC7FD12" box="[1020,1138,644,670]" name="Indonesia" pageId="15" pageNumber="16">Indonesia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB21633CA5DFFF9FEF6FD31FC26FD4A" blockId="15.[264,1323,284,1150]" box="[312,915,684,710]" pageId="15" pageNumber="16">
|
||
<materialsCitation id="3B651C6ECA5DFFF9FEF6FD31FC25FD4A" ID-GBIF-Occurrence="3337342346" box="[312,912,684,710]" collectionCode="FUJI, WADA" pageId="15" pageNumber="16" specimenCount="2">
|
||
Locality records (
|
||
<specimenCount id="9D0BDDBACA5DFFF9FDC9FD31FD2FFD4A" box="[519,666,684,710]" pageId="15" pageNumber="16" type="generic">2 specimens</specimenCount>
|
||
) from
|
||
<collectionCode id="ED1C8EF6CA5DFFF9FD24FD31FC9DFD4A" box="[746,808,684,710]" pageId="15" pageNumber="16">FUJI</collectionCode>
|
||
,
|
||
<collectionCode id="ED1C8EF6CA5DFFF9FCFAFD31FC25FD4A" box="[820,912,684,710]" country="Australia" httpUri="http://grbio.org/cool/mnrx-zeat" name="Western Australia Department of Agriculture" pageId="15" pageNumber="16">WADA</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31745B8CA5DFFF9FEF6FD49FD26FBF2" pageId="15" pageNumber="16" type="materials_examined">
|
||
<paragraph id="8BB21633CA5DFFF9FEF6FD49FBCBFD62" blockId="15.[264,1323,284,1150]" box="[312,1150,724,750]" pageId="15" pageNumber="16">
|
||
BORNEAN
|
||
<collectingCountry id="F31A56A3CA5DFFF9FE03FD49FDC5FD62" box="[461,624,724,750]" name="Indonesia" pageId="15" pageNumber="16">INDONESIA</collectingCountry>
|
||
.
|
||
<collectingRegion id="49C9D8D1CA5DFFF9FDB5FD49FC01FD62" box="[635,948,724,750]" country="Indonesia" name="Kalimantan Barat" pageId="15" pageNumber="16">West Kalimantan Province</collectingRegion>
|
||
(2): Mt Bawang.
|
||
</paragraph>
|
||
<paragraph id="8BB21633CA5DFFF9FEF6FD61FCADFC9A" blockId="15.[264,1323,284,1150]" box="[312,792,764,790]" pageId="15" pageNumber="16">Temporal Data. August (1), October (1).</paragraph>
|
||
<paragraph id="8BB21633CA5DFFF9FEF6FCB9FD26FBF2" blockId="15.[264,1323,284,1150]" pageId="15" pageNumber="16">
|
||
Remarks. Based on shared characters,
|
||
<taxonomicName id="4C0D6DB0CA5DFFF9FCC3FCB9FCC2FCB1" authorityName="Arrow. A" authorityYear="1931" box="[781,887,804,829]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B979CA21CA5DFFF9FCC3FCB9FCC2FCB1" box="[781,887,804,829]" italics="true" pageId="15" pageNumber="16">P. similis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0D6DB0CA5DFFF9FC7BFCB9FBC8FCB1" authorityName="Jameson and Wada" authorityYear="2004" box="[949,1149,804,830]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="57" phylum="Arthropoda" rank="species" species="adelphosimilis">
|
||
P.
|
||
<emphasis id="B979CA21CA5DFFF9FC1CFCB9FBC8FCB1" box="[978,1149,804,829]" italics="true" pageId="15" pageNumber="16">adelphosimilis</emphasis>
|
||
</taxonomicName>
|
||
are sister species. They differ in only three character states (form of the male foretarsomere 5, form of the labrum, and form of the female epipleuron). We considered that the character states observed in
|
||
<taxonomicName id="4C0D6DB0CA5DFFF9FE6CFC01FDC5FC39" authorityName="Jameson and Wada" authorityYear="2004" box="[418,624,924,949]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="adelphosimilis">
|
||
<emphasis id="B979CA21CA5DFFF9FE6CFC01FDC5FC39" box="[418,624,924,949]" italics="true" pageId="15" pageNumber="16">P. adelphosimilis</emphasis>
|
||
</taxonomicName>
|
||
may be intraspecific variation, but the two species also appear to be geographically isolated:
|
||
<taxonomicName id="4C0D6DB0CA5DFFF9FD72FC59FC90FC51" authorityName="Arrow. A" authorityYear="1931" box="[700,805,964,989]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B979CA21CA5DFFF9FD72FC59FC90FC51" box="[700,805,964,989]" italics="true" pageId="15" pageNumber="16">P. similis</emphasis>
|
||
</taxonomicName>
|
||
occurs in the Crocker Range in
|
||
<collectingRegion id="49C9D8D1CA5DFFF9FB55FC59FB56FC52" box="[1179,1251,964,990]" country="Malaysia" name="Sabah" pageId="15" pageNumber="16">Sabah</collectingRegion>
|
||
while
|
||
<taxonomicName id="4C0D6DB0CA5DFFF9FEC6FC71FE66FB89" authorityName="Jameson and Wada" authorityYear="2004" box="[264,467,1004,1029]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="adelphosimilis">
|
||
<emphasis id="B979CA21CA5DFFF9FEC6FC71FE66FB89" box="[264,467,1004,1029]" italics="true" pageId="15" pageNumber="16">P. adelphosimilis</emphasis>
|
||
</taxonomicName>
|
||
occurs in Boven Kapuas Mountains in
|
||
<collectingRegion id="49C9D8D1CA5DFFF9FC76FC71FB3FFB8A" box="[952,1162,1004,1030]" country="Indonesia" name="Kalimantan Barat" pageId="15" pageNumber="16">West Kalimantan</collectingRegion>
|
||
. Geographic isolation in addition to the character state differences led us to the conclusion that these are two distinct taxa. Additional specimens of both
|
||
<taxonomicName id="4C0D6DB0CA5DFFF9FC82FBA1FC02FBD9" authorityName="Arrow. A" authorityYear="1931" box="[844,951,1084,1109]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B979CA21CA5DFFF9FC82FBA1FC02FBD9" box="[844,951,1084,1109]" italics="true" pageId="15" pageNumber="16">P. similis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0D6DB0CA5DFFF9FC3DFBA1FB09FBD9" authorityName="Jameson and Wada" authorityYear="2004" box="[1011,1212,1084,1109]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="adelphosimilis">
|
||
<emphasis id="B979CA21CA5DFFF9FC3DFBA1FB09FBD9" box="[1011,1212,1084,1109]" italics="true" pageId="15" pageNumber="16">P. adelphosimilis</emphasis>
|
||
</taxonomicName>
|
||
will corroborate or refute this hypothesis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |