treatments-xml/data/03/A4/A7/03A4A725CA4BFFEDFEC6F914FCA8FA92.xml
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<document id="611E241ED5D3C4843C663C6FE4AC4518" ID-CLB-Dataset="4845" ID-DOI="10.11646/zootaxa.502.1.1" ID-GBIF-Dataset="76aa06a8-f646-4c5a-b596-54bfd8875223" ID-ISSN="1175­5334" ID-Zenodo-Dep="5030148" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624553872186" checkinUser="felipe" docAuthor="Jameson, Mary Liz &amp; Wada, Kaoru" docDate="2004" docId="03A4A725CA4BFFEDFEC6F914FCA8FA92" docLanguage="en" docName="zt00502.pdf" docOrigin="Zootaxa 502 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.502.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Peltonotus malayensis Arrow" docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="FF9DDF5DCA52FFF6FFCEFF9DFFB5FF8C" masterDocTitle="Revision of the genus Peltonotus Burmeister (Coleoptera: Scarabaeidae: Dynastinae) from Southeastern Asia" masterLastPageNumber="66" masterPageNumber="1" pageNumber="26" updateTime="1699020591651" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DFF8077390D06783FEB984F7DB5696D7">Revision of the genus Peltonotus Burmeister (Coleoptera: Scarabaeidae: Dynastinae) from Southeastern Asia</mods:title>
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<mods:namePart id="C83B199B57AD096296D9EBA149EFEDD2">Jameson, Mary Liz</mods:namePart>
<mods:affiliation id="B40153DCAFF4E60B1689AE19DE7FD9E8">University of Nebraska State Museum, Division of Entomology, Lincoln, NE U. S. A., 68588 - 0514 mjameson 1 @ unl. edu</mods:affiliation>
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<mods:namePart id="24DEEC5C720F82B35B6626A191E3E51F">Wada, Kaoru</mods:namePart>
<mods:affiliation id="BBBC90D1989A460CD1E77194D2DE9950">Musashimurayama Daini Junior School, 1460 Midorigaoka, Musashimurayama-shi, Tokyo 208 - 0012 Japan kwada 007 @ hotmail. com</mods:affiliation>
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<mods:title id="80667EFD469E1F1857131D903D612C18">Zootaxa</mods:title>
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<mods:date id="C41D8FB0D385D87E87DE8D60B768B469">2004</mods:date>
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<treatment id="03A4A725CA4BFFEDFEC6F914FCA8FA92" ID-DOI="http://doi.org/10.5281/zenodo.5248457" ID-GBIF-Taxon="184930985" ID-Zenodo-Dep="5248457" LSID="urn:lsid:plazi:treatment:03A4A725CA4BFFEDFEC6F914FCA8FA92" httpUri="http://treatment.plazi.org/id/03A4A725CA4BFFEDFEC6F914FCA8FA92" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
<subSubSection id="C31745B8CA4BFFEFFEC6F914FDDBF92F" box="[264,622,1673,1699]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8BB21633CA4BFFEFFEC6F914FDDBF92F" blockId="25.[264,741,1673,1734]" box="[264,622,1673,1699]" pageId="25" pageNumber="26">
<heading id="D0FAA15FCA4BFFEFFEC6F914FDDBF92F" bold="true" box="[264,622,1673,1699]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<taxonomicName id="4C0D6DB0CA4BFFEFFEC6F914FDDBF92F" authority="Arrow" authorityName="Arrow" box="[264,622,1673,1699]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B979CA21CA4BFFEFFEC6F914FDDBF92F" bold="true" box="[264,622,1673,1699]" pageId="25" pageNumber="26">
<emphasis id="B979CA21CA4BFFEFFEC6F914FDA1F92F" bold="true" box="[264,532,1673,1699]" italics="true" pageId="25" pageNumber="26">Peltonotus malayensis</emphasis>
Arrow
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31745B8CA4BFFEFFEC6F931FD50F94A" box="[264,741,1708,1734]" pageId="25" pageNumber="26" type="description">
<paragraph id="8BB21633CA4BFFEFFEC6F931FD50F94A" blockId="25.[264,741,1673,1734]" box="[264,741,1708,1734]" pageId="25" pageNumber="26">
(
<figureCitation id="13360AB6CA4BFFEFFEDFF931FE37F94A" box="[273,386,1708,1734]" captionStart="FIGURES 213" captionStartId="52.[264,382,1552,1576]" captionTargetBox="[352,1234,326,1520]" captionTargetId="figure-62@52.[346,1242,326,1528]" captionTargetPageId="52" captionText="FIGURES 213. Dorsal habitus of: 24) P. fujiokai (males), 5) P. fujiokai (female), 6) P. malayensis (male), 7) P. malayensis (female), 8) P. morio (male), 9) P. morio (female), 10) P. nasutus (male), 11) P.nasutus (female), 12) P. vittatus (male), 13) P. vittatus (female)." figureDoi="http://doi.org/10.5281/zenodo.5030156" httpUri="https://zenodo.org/record/5030156/files/figure.png" pageId="25" pageNumber="26">Figs. 67</figureCitation>
,
<figureCitation id="13360AB6CA4BFFEFFE43F931FE19F94A" box="[397,428,1708,1734]" captionStart="FIGURES 1418" captionStartId="53.[264,382,1425,1449]" captionTargetBox="[340,1245,341,1398]" captionTargetId="figure-118@53.[337,1249,326,1401]" captionTargetPageId="53" captionText="FIGURES 1418. Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 14) P.karubei (apex of labrum deeply, narrowly emarginated; apex of mandible rounded laterally), 15) P. malayensis (apex of labrum broadly emarginate; apex of mandible rounded laterally), 16) P. morio (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male), 17) P. nasutus (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male), 18) P. vittatus (apex of labrum broadly emarginate; apex of mandible obliquely quadrate laterally)." figureDoi="http://doi.org/10.5281/zenodo.5030160" httpUri="https://zenodo.org/record/5030160/files/figure.png" pageId="25" pageNumber="26">15</figureCitation>
,
<figureCitation id="13360AB6CA4BFFEFFE77F931FE6CF94A" box="[441,473,1708,1734]" captionStart="FIGURES 1928" captionStartId="54.[264,382,1043,1067]" captionTargetBox="[366,1231,326,1016]" captionTargetId="figure-89@54.[355,1233,326,1019]" captionTargetPageId="54" captionText="FIGURES 1928. Mentum, dorsal view of: 19) P. deltamentum, 20) P. gracilipodus, 21) P. kyojinus, 22) P. malayensis, 23) P. podocrassus, 24) P. nasutus, 25) P. nethis, 26) P. pruinosus, 27) P. similis, 28) P. sisyrus." figureDoi="http://doi.org/10.5281/zenodo.5030164" httpUri="https://zenodo.org/record/5030164/files/figure.png" pageId="25" pageNumber="26">22</figureCitation>
,
<figureCitation id="13360AB6CA4BFFEFFE2BF931FD84F94A" box="[485,561,1708,1734]" captionStart-0="FIGURES 2930" captionStart-1="FIGURES 3136" captionStartId-0="54.[264,382,1823,1847]" captionStartId-1="55.[264,382,1495,1519]" captionTargetBox-0="[466,1118,1261,1796]" captionTargetBox-1="[342,1225,329,1468]" captionTargetId-0="figure-91@54.[464,1123,1259,1799]" captionTargetId-1="figure-95@55.[337,1249,326,1471]" captionTargetPageId-0="54" captionTargetPageId-1="55" captionText-0="FIGURES 2930. Maxilla (dorsal view) showing articulated maxillary tooth, mala with lamellate setal brush (black arrow), and form in: 29) P. deltamentum and 30) P. malayensis." captionText-1="FIGURES 3136. Detail of right forelegs of: 31) P. malayensis (showing bidentate foretibia, dorsal view), 32) P.rubripennis (showing tridentate foretibia with basal tooth very weakly developed, dorsal view), 33) P. deltamentum (showing strongly arcuate smaller claw, ventral view), 34) P. nasutus (showing form of foretarsomeres 15), 35) P.similis (showing form of foretarsomeres 15 and median protuberance of foretarsomere 5 [arrow]), 36) P. adelphosimilis (showing form of foretarsomeres 15 and apical protuberance of foretarsomere 5 [arrow])." figureDoi-0="http://doi.org/10.5281/zenodo.5030168" figureDoi-1="http://doi.org/10.5281/zenodo.5030172" httpUri-0="https://zenodo.org/record/5030168/files/figure.png" httpUri-1="https://zenodo.org/record/5030172/files/figure.png" pageId="25" pageNumber="26">3031</figureCitation>
,
<figureCitation id="13360AB6CA4BFFEFFDF3F931FD33F94A" box="[573,646,1708,1734]" captionStart="FIGURES 4345" captionStartId="58.[264,382,1739,1763]" captionTargetBox="[449,1146,330,1677]" captionTargetId="figure-45@58.[438,1148,312,1680]" captionTargetPageId="58" captionText="FIGURES 4345. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [bc]) of: 43) P. malayensis, 44) P.morio, 45) P. nasutus." figureDoi="http://doi.org/10.5281/zenodo.5030184" httpUri="https://zenodo.org/record/5030184/files/figure.png" pageId="25" pageNumber="26">43ac</figureCitation>
,
<figureCitation id="13360AB6CA4BFFEFFD5FF931FD04F94A" box="[657,689,1708,1734]" captionStart="FIGURES 5155" captionStartId="61.[264,382,1252,1276]" captionTargetBox="[345,1246,334,1218]" captionTargetId="figure-51@61.[337,1249,326,1228]" captionTargetPageId="61" captionText="FIGURES 5155. Elytral epipleuron (gray, ventral view) with position of metacoxa of: 51) P. brunnipennis, 52) P. fujiokai, 53) P. gracilipodus, P. podocrassus, P. silvanus, 54) P. kyojinus, 55) P. malayensis." figureDoi="http://doi.org/10.5281/zenodo.5030194" httpUri="https://zenodo.org/record/5030194/files/figure.png" pageId="25" pageNumber="26">55</figureCitation>
,
<figureCitation id="13360AB6CA4BFFEFFD73F931FD6EF94A" box="[701,731,1708,1734]" captionStart="FIGURE 65" captionStartId="64.[264,368,1270,1294]" captionTargetBox="[337,1251,315,1246]" captionTargetId="figure-27@64.[337,1251,315,1246]" captionTargetPageId="64" captionText="FIGURE 65. Distribution of Peltonotus species in Southeast Asia and Sundaland." figureDoi="http://doi.org/10.5281/zenodo.5030204" httpUri="https://zenodo.org/record/5030204/files/figure.png" pageId="25" pageNumber="26">65</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31745B8CA4BFFECFEC6F96FFB69FD97" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="8BB21633CA4BFFECFEC6F96FFB69FD97" blockId="25.[264,1323,1777,1901]" lastBlockId="26.[312,1323,284,539]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB1D311DCA4BFFECFEC6F96FFB69FD97" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<treatmentCitation id="0AAC3022CA4BFFEFFEC6F96FFCB5F884" author="Arrow, G. J." box="[264,768,1777,1801]" page="156" pageId="25" pageNumber="26" year="1910">
<taxonomicName id="4C0D6DB0CA4BFFEFFEC6F96FFCB5F884" authority="Arrow 1910: 155 - 156" authorityName="Arrow" authorityPageNumber="155 - 156" authorityYear="1910" box="[264,768,1777,1801]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B979CA21CA4BFFEFFEC6F96FFE49F885" box="[264,508,1778,1801]" italics="true" pageId="25" pageNumber="26">Peltonotus malayensis</emphasis>
<bibRefCitation id="EF9C6BC2CA4BFFEFFDC9F96CFCB5F884" author="Arrow, G. J." box="[519,768,1777,1800]" pageId="25" pageNumber="26" pagination="153 - 157" refId="ref31219" refString="Arrow, G. J. (1910) On the lamellicorn beetles of the genus Peltonotus with descriptions of four new species. Annals and Magazine of Natural History (Series 8), 5, 153 - 157." type="journal article" year="1910">Arrow 1910: 155156</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54B6A891CA4BFFEFFCD4F96CFC30F884" box="[794,901,1777,1800]" pageId="25" pageNumber="26" type="lectotype">Lectotype</typeStatus>
female at BMNH with label data: a) &quot;Type&quot; (round label with red border, typeface), b) &quot;Bornéo Occ. Pontianak 1903&quot; (typeface), c) &quot;ex Oberthur Coll. 1908134.&quot; (typeface), d) &quot;
<taxonomicName id="4C0D6DB0CA4BFFEFFCFAF8AEFBC8F8C6" authority=", Arrow" authorityName="Arrow" box="[820,1149,1843,1866]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="malayensis">Peltonotus malayensis, Arrow</taxonomicName>
type
<specimenCount id="9D0BDDBACA4BFFEFFB73F8ACFB64F8C9" box="[1213,1233,1841,1861]" pageId="25" pageNumber="26" type="female"></specimenCount>
&quot; (handwritten), e) &quot;
<taxonomicName id="4C0D6DB0CA4BFFEFFE0EF8CBFCB5F8E1" authority="Arrow" authorityName="Arrow" box="[448,768,1878,1901]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="malayensis">Peltonotus malayensis Arrow</taxonomicName>
<specimenCount id="9D0BDDBACA4BFFEFFCC4F8CEFCABF8EB" box="[778,798,1875,1895]" pageId="25" pageNumber="26" type="female"></specimenCount>
<typeStatus id="54B6A891CA4BFFEFFCEAF8CBFC27F8E1" box="[804,914,1878,1901]" pageId="25" pageNumber="26" type="lectotype">Lectotype</typeStatus>
&quot; (our red
<typeStatus id="54B6A891CA4BFFEFFBCFF8CBFBD6F8E1" box="[1025,1123,1878,1901]" pageId="25" pageNumber="26" type="lectotype">lectotype</typeStatus>
label).
<collectingMunicipality id="6BD68C49CA4BFFEFFB71F8CBFB5EF8E1" box="[1215,1259,1878,1901]" pageId="25" pageNumber="26">One</collectingMunicipality>
<typeStatus id="54B6A891CA4BFFECFB3AF8CBFE2FFEBF" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="paralectotype">paralectotype</typeStatus>
at MNHN with label data: a) “Bornéo Oc., Riv. Sintang,
<collectorName id="26F873E5CA48FFECFBC6FE81FBDDFEBF" box="[1032,1128,284,307]" pageId="26" pageNumber="27">F. Buffat</collectorName>
1905” (typeface), b”
<taxonomicName id="4C0D6DB0CA48FFECFE80FEA0FD3AFED8" authority="Arrow" authorityName="Arrow" box="[334,655,317,340]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="malayensis">Peltonotus malayensis Arrow</taxonomicName>
co­type
<specimenCount id="9D0BDDBACA48FFECFD3CFEA6FCB1FEC3" box="[754,772,315,335]" pageId="26" pageNumber="27" type="female"></specimenCount>
” (handwritten), c) “
<collectorName id="26F873E5CA48FFECFC11FEA0FBEAFED8" box="[991,1119,317,340]" pageId="26" pageNumber="27">G. J. Arrow</collectorName>
vidit 1910” (typeface), d) “Museum Paris ex.
<collectorName id="26F873E5CA48FFECFDBCFEC3FC87FEF9" box="[626,818,350,373]" pageId="26" pageNumber="27">Coll. R. Oberthur</collectorName>
” (typeface, yellow label), e) our
<typeStatus id="54B6A891CA48FFECFB55FEC3FA9EFEF9" box="[1179,1323,350,373]" pageId="26" pageNumber="27" type="paralectotype">paralectotype</typeStatus>
label.
<location id="8ED240E8CA48FFECFE4AFEE3FE05FE19" LSID="urn:lsid:plazi:treatment:03A4A725CA4BFFEDFEC6F914FCA8FA92:8ED240E8CA48FFECFE4AFEE3FE05FE19" box="[388,432,382,405]" municipality="One" name="One" pageId="26" pageNumber="27" stateProvince="Sumatera Utara">One</location>
additional
<typeStatus id="54B6A891CA48FFECFDE2FEE3FD09FE19" box="[556,700,382,405]" pageId="26" pageNumber="27" type="paralectotype">paralectotype</typeStatus>
from MNHN is conspecific with
<taxonomicName id="4C0D6DB0CA48FFECFBE3FEE2FB65FE1A" authorityName="Jameson and Wada" authorityYear="2004" box="[1069,1232,383,406]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="gracilipodus">
<emphasis id="B979CA21CA48FFECFBE3FEE2FB65FE1A" box="[1069,1232,383,406]" italics="true" pageId="26" pageNumber="27">P. gracilipodus</emphasis>
</taxonomicName>
and has the following label data: a) “
<collectingRegion id="49C9D8D1CA48FFECFDB8FE03FD66FE39" box="[630,723,414,437]" country="Indonesia" name="Sumatera Utara" pageId="26" pageNumber="27">Sumatra</collectingRegion>
, Siboga,
<date id="FFB330F3CA48FFECFCF6FE03FC3AFE39" box="[824,911,414,437]" pageId="26" pageNumber="27" value="1903-03">
<collectingDate id="EFF7C91BCA48FFECFCF6FE03FC3AFE39" box="[824,911,414,437]" pageId="26" pageNumber="27" value="1903-03">III.1903</collectingDate>
</date>
” (typeface), b)
<taxonomicName id="4C0D6DB0CA48FFECFBF4FE03FEC8FE5B" authority="Arrow" authorityName="Arrow" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="malayensis">Peltonotus malayensis Arrow</taxonomicName>
co­type
<specimenCount id="9D0BDDBACA48FFECFE15FE23FE5BFE5E" box="[475,494,446,466]" pageId="26" pageNumber="27" type="female"></specimenCount>
” (handwritten), c) “
<collectorName id="26F873E5CA48FFECFD08FE5DFCF7FE5B" box="[710,834,448,471]" pageId="26" pageNumber="27">G. J. Arrow</collectorName>
vidit 1910” (typeface), d) “Museum Paris ex.
<collectorName id="26F873E5CA48FFECFEF6FE7EFE49FE76" box="[312,508,483,506]" pageId="26" pageNumber="27">Coll. R. Oberthur</collectorName>
” (typeface, yellow label), e) &quot;
<taxonomicName id="4C0D6DB0CA48FFECFC87FE7EFB39FE76" authority="Arrow" authorityName="Arrow" box="[841,1164,483,506]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="malayensis">Peltonotus malayensis Arrow</taxonomicName>
<specimenCount id="9D0BDDBACA48FFECFB59FE7CFB1EFE79" box="[1175,1195,481,501]" pageId="26" pageNumber="27" type="female"></specimenCount>
<typeStatus id="54B6A891CA48FFECFB7BFE7EFA97FE76" box="[1205,1314,483,506]" pageId="26" pageNumber="27" type="lectotype">Lectotype</typeStatus>
&quot; (our yellow
<typeStatus id="54B6A891CA48FFECFE74FD99FDA9FD97" box="[442,540,516,539]" pageId="26" pageNumber="27" type="lectotype">lectotype</typeStatus>
label), f) “
<taxonomicName id="4C0D6DB0CA48FFECFD5BFD99FBD1FD97" authority="Jameson and Wada" authorityName="Jameson and Wada" authorityYear="2004" box="[661,1124,516,539]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="gracilipodus">Peltonotus gracilipodus Jameson and Wada</taxonomicName>
<typeStatus id="54B6A891CA48FFECFBA5FD99FB7BFD97" box="[1131,1230,516,539]" pageId="26" pageNumber="27" type="paratype">Paratype</typeStatus>
”.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31745B8CA48FFEDFEC6FDD1FBD6FC9A" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="8BB21633CA48FFEDFEC6FDD1FA94FE0A" blockId="26.[264,1324,588,1894]" lastBlockId="27.[264,1323,284,1310]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Description (males=11, females=7). Length
<quantity id="4CF5BBD6CA48FFECFCEDFDD1FC66FDEA" box="[803,979,588,614]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" metricValueMax="1.72" metricValueMin="1.44" pageId="26" pageNumber="27" unit="mm" value="15.8" valueMax="17.2" valueMin="14.4">14.417.2 mm</quantity>
. Widest width
<quantity id="4CF5BBD6CA48FFECFB5AFDD1FA93FDEA" box="[1172,1318,588,614]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.3" metricValueMin="7.2" pageId="26" pageNumber="27" unit="mm" value="7.75" valueMax="8.3" valueMin="7.2">7.28.3 mm</quantity>
. Color: Male (
<figureCitation id="13360AB6CA48FFECFE75FDE9FDB2FD02" box="[443,519,628,654]" captionStart="FIGURES 213" captionStartId="52.[264,382,1552,1576]" captionTargetBox="[352,1234,326,1520]" captionTargetId="figure-62@52.[346,1242,326,1528]" captionTargetPageId="52" captionText="FIGURES 213. Dorsal habitus of: 24) P. fujiokai (males), 5) P. fujiokai (female), 6) P. malayensis (male), 7) P. malayensis (female), 8) P. morio (male), 9) P. morio (female), 10) P. nasutus (male), 11) P.nasutus (female), 12) P. vittatus (male), 13) P. vittatus (female)." figureDoi="http://doi.org/10.5281/zenodo.5030156" httpUri="https://zenodo.org/record/5030156/files/figure.png" pageId="26" pageNumber="27">Fig. 6</figureCitation>
): Head, pronotum, scutellum, propygidium, pygidium, and venter shining castaneous to black; elytra reddish­brown with iridescent bloom. Female (
<figureCitation id="13360AB6CA48FFECFB1EFD01FAADFD3A" box="[1232,1304,668,694]" captionStart="FIGURES 213" captionStartId="52.[264,382,1552,1576]" captionTargetBox="[352,1234,326,1520]" captionTargetId="figure-62@52.[346,1242,326,1528]" captionTargetPageId="52" captionText="FIGURES 213. Dorsal habitus of: 24) P. fujiokai (males), 5) P. fujiokai (female), 6) P. malayensis (male), 7) P. malayensis (female), 8) P. morio (male), 9) P. morio (female), 10) P. nasutus (male), 11) P.nasutus (female), 12) P. vittatus (male), 13) P. vittatus (female)." figureDoi="http://doi.org/10.5281/zenodo.5030156" httpUri="https://zenodo.org/record/5030156/files/figure.png" pageId="26" pageNumber="27">Fig. 7</figureCitation>
): Head, pronotum, scutellum, propygidium, pygidium, and venter shining castaneous to black; elytra black with iridescent blue bloom. Head (
<figureCitation id="13360AB6CA48FFECFC46FD71FC55FC8A" box="[904,992,748,774]" captionStart="FIGURES 1418" captionStartId="53.[264,382,1425,1449]" captionTargetBox="[340,1245,341,1398]" captionTargetId="figure-118@53.[337,1249,326,1401]" captionTargetPageId="53" captionText="FIGURES 1418. Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 14) P.karubei (apex of labrum deeply, narrowly emarginated; apex of mandible rounded laterally), 15) P. malayensis (apex of labrum broadly emarginate; apex of mandible rounded laterally), 16) P. morio (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male), 17) P. nasutus (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male), 18) P. vittatus (apex of labrum broadly emarginate; apex of mandible obliquely quadrate laterally)." figureDoi="http://doi.org/10.5281/zenodo.5030160" httpUri="https://zenodo.org/record/5030160/files/figure.png" pageId="26" pageNumber="27">Fig. 15</figureCitation>
): Surface of frons with base impunctate to sparsely punctate, middle of frons to apex densely punctate; punctures ocellate, multisetigerous (less setigerous in females); short setae on disc (112 per puncture), moderately long setae (1 per puncture) adjacent to eye. Surface of clypeus densely punctate; punctures ocellate, multisetigerous (less setigerous in females); short setae on disc (112 per puncture). Clypeus laterally weakly bowed, apex truncate, corners square, beaded; bead weakly arcuate posteriorly. Labrum broadly emarginate at middle. Mandibles of male and female rounded laterally, inner apex with 2 teeth.
<collectingRegion id="49C9D8D1CA48FFECFBD1FB99FB37FB92" box="[1055,1154,1028,1054]" country="Myanmar" name="Chin" pageId="26" pageNumber="27">Mentum</collectingRegion>
(
<figureCitation id="13360AB6CA48FFECFB5DFB99FB5CFB92" box="[1171,1257,1028,1054]" captionStart="FIGURES 1928" captionStartId="54.[264,382,1043,1067]" captionTargetBox="[366,1231,326,1016]" captionTargetId="figure-89@54.[355,1233,326,1019]" captionTargetPageId="54" captionText="FIGURES 1928. Mentum, dorsal view of: 19) P. deltamentum, 20) P. gracilipodus, 21) P. kyojinus, 22) P. malayensis, 23) P. podocrassus, 24) P. nasutus, 25) P. nethis, 26) P. pruinosus, 27) P. similis, 28) P. sisyrus." figureDoi="http://doi.org/10.5281/zenodo.5030164" httpUri="https://zenodo.org/record/5030164/files/figure.png" pageId="26" pageNumber="27">Fig. 22</figureCitation>
) with apical half rounded, notched at middle; palpomere 2 dorsoventrally flattened, twice width of palpomere 1, setose; setae dense, moderately long, rufous, some curled at apex, some flattened. Maxilla (
<figureCitation id="13360AB6CA48FFECFE20FBE1FDF2FB1A" box="[494,583,1148,1174]" captionStart="FIGURES 2930" captionStartId="54.[264,382,1823,1847]" captionTargetBox="[466,1118,1261,1796]" captionTargetId="figure-91@54.[464,1123,1259,1799]" captionTargetPageId="54" captionText="FIGURES 2930. Maxilla (dorsal view) showing articulated maxillary tooth, mala with lamellate setal brush (black arrow), and form in: 29) P. deltamentum and 30) P. malayensis." figureDoi="http://doi.org/10.5281/zenodo.5030168" httpUri="https://zenodo.org/record/5030168/files/figure.png" pageId="26" pageNumber="27">Fig. 30</figureCitation>
): mala with weak lamellate setal brush; stipes with setae dense, long, not flattened, curled at apex; palpomere 2 with weak internomedial bump. Antennal club subequal to segments 27. Pronotum: Bead lacking anterior to scutellum. Surface moderately densely punctate, punctures ocellate, some multisetigerous laterally in males; setae minute. Lateral margin lacking long setae. Elytral sutural length: About 4.0 times length of scutellum. Elytron: Surface with 5 poorly developed, punctate, longitudinal striae between suture and humerus; punctures moderate in size, moderately dense. Intervals with similar sculpturing. Epipleuron of female in ventral view expanded, incised at apex; in dorsal view expansion well­developed (
<figureCitation id="13360AB6CA48FFECFC89FA21FC15FA5A" box="[839,928,1468,1494]" captionStart="FIGURES 5155" captionStartId="61.[264,382,1252,1276]" captionTargetBox="[345,1246,334,1218]" captionTargetId="figure-51@61.[337,1249,326,1228]" captionTargetPageId="61" captionText="FIGURES 5155. Elytral epipleuron (gray, ventral view) with position of metacoxa of: 51) P. brunnipennis, 52) P. fujiokai, 53) P. gracilipodus, P. podocrassus, P. silvanus, 54) P. kyojinus, 55) P. malayensis." figureDoi="http://doi.org/10.5281/zenodo.5030194" httpUri="https://zenodo.org/record/5030194/files/figure.png" pageId="26" pageNumber="27">Fig. 55</figureCitation>
). Propygidium: Surface moderately densely punctate; punctures ocellate, unisetigerous; setae moderately long, rufous. Pygidium: Surface densely punctate; punctures ocellate, mixed small and large, multisetigerous; setae short (over 15 per puncture) and moderately long (1 per puncture). Venter: Prosternal keel triangular; apex projecting anteriorly at about 70° with respect to ventral plane; apex produced to middle of protrochanter, rounded. Legs (
<figureCitation id="13360AB6CA48FFECFBEDF919FBCFF912" box="[1059,1146,1668,1694]" captionStart="FIGURES 3136" captionStartId="55.[264,382,1495,1519]" captionTargetBox="[342,1225,329,1468]" captionTargetId="figure-95@55.[337,1249,326,1471]" captionTargetPageId="55" captionText="FIGURES 3136. Detail of right forelegs of: 31) P. malayensis (showing bidentate foretibia, dorsal view), 32) P.rubripennis (showing tridentate foretibia with basal tooth very weakly developed, dorsal view), 33) P. deltamentum (showing strongly arcuate smaller claw, ventral view), 34) P. nasutus (showing form of foretarsomeres 15), 35) P.similis (showing form of foretarsomeres 15 and median protuberance of foretarsomere 5 [arrow]), 36) P. adelphosimilis (showing form of foretarsomeres 15 and apical protuberance of foretarsomere 5 [arrow])." figureDoi="http://doi.org/10.5281/zenodo.5030172" httpUri="https://zenodo.org/record/5030172/files/figure.png" pageId="26" pageNumber="27">Fig. 31</figureCitation>
): Foretibia of male bidentate; lateral margin with short, dense setae. Foretarsomere 5 of male subequal in length to tarsomeres 14; foretarsomere 34 with apices expanded, dorsal and ventral apices clothed with dense, short setae. Foreclaws of male with inner claw roundly curved, about 4 times thicker than outer claw; outer claw simply arcuate, about 1/2 the length of inner claw; empodium bulbous at base. Foreclaws of female 2/3 length of foretarsomere 5, claws angled toward venter. Meso­ and metatibial claws of male with 2 setae. Metatibia of male with apical spurs weakly curved spurs; ventral spur produced to middle of metatarsomere 1, dorsal spur produced to middle of metatarsomere 2. Parameres:
<figureCitation id="13360AB6CA49FFEDFB53FEF1FAA8FE0A" box="[1181,1309,364,390]" captionStart="FIGURES 4345" captionStartId="58.[264,382,1739,1763]" captionTargetBox="[449,1146,330,1677]" captionTargetId="figure-45@58.[438,1148,312,1680]" captionTargetPageId="58" captionText="FIGURES 4345. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [bc]) of: 43) P. malayensis, 44) P.morio, 45) P. nasutus." figureDoi="http://doi.org/10.5281/zenodo.5030184" httpUri="https://zenodo.org/record/5030184/files/figure.png" pageId="27" pageNumber="28">Fig. 43ac</figureCitation>
.
</paragraph>
<paragraph id="8BB21633CA49FFEDFEF6FE09FC8AFD12" blockId="27.[264,1323,284,1310]" pageId="27" pageNumber="28">
Diagnosis.
<taxonomicName id="4C0D6DB0CA49FFEDFE05FE09FD63FE21" box="[459,726,404,429]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B979CA21CA49FFEDFE05FE09FD63FE21" box="[459,726,404,429]" italics="true" pageId="27" pageNumber="28">Peltonotus malayensis</emphasis>
</taxonomicName>
is distinguished from other species in the genus based on the following characters: labrum broadly emarginate at middle apex; palpomere 2 dorsoventrally flattened, setose (some setae curled at apex); mala with weak lamellate setal brush; stipes with setae curled at apex; palpomere 2 with weak internomedial bump; foretibia of male bidentate with short, dense setae at lateral margin; form of male genitalia (
<figureCitation id="13360AB6CA49FFEDFEDFFDC1FE24FDFA" box="[273,401,604,630]" captionStart="FIGURES 4345" captionStartId="58.[264,382,1739,1763]" captionTargetBox="[449,1146,330,1677]" captionTargetId="figure-45@58.[438,1148,312,1680]" captionTargetPageId="58" captionText="FIGURES 4345. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [bc]) of: 43) P. malayensis, 44) P.morio, 45) P. nasutus." figureDoi="http://doi.org/10.5281/zenodo.5030184" httpUri="https://zenodo.org/record/5030184/files/figure.png" pageId="27" pageNumber="28">Fig. 43ac</figureCitation>
); epipleuron of female in ventral view expanded, incised at sternite 3, in dorsal view expansion moderately developed (
<figureCitation id="13360AB6CA49FFEDFD14FD19FC87FD12" box="[730,818,644,670]" captionStart="FIGURES 5155" captionStartId="61.[264,382,1252,1276]" captionTargetBox="[345,1246,334,1218]" captionTargetId="figure-51@61.[337,1249,326,1228]" captionTargetPageId="61" captionText="FIGURES 5155. Elytral epipleuron (gray, ventral view) with position of metacoxa of: 51) P. brunnipennis, 52) P. fujiokai, 53) P. gracilipodus, P. podocrassus, P. silvanus, 54) P. kyojinus, 55) P. malayensis." figureDoi="http://doi.org/10.5281/zenodo.5030194" httpUri="https://zenodo.org/record/5030194/files/figure.png" pageId="27" pageNumber="28">Fig. 55</figureCitation>
).
</paragraph>
<paragraph id="8BB21633CA49FFEDFEF6FD31FD41FD62" blockId="27.[264,1323,284,1310]" pageId="27" pageNumber="28">
Distribution (
<figureCitation id="13360AB6CA49FFEDFE19FD31FD84FD4A" box="[471,561,684,710]" captionStart="FIGURE 65" captionStartId="64.[264,368,1270,1294]" captionTargetBox="[337,1251,315,1246]" captionTargetId="figure-27@64.[337,1251,315,1246]" captionTargetPageId="64" captionText="FIGURE 65. Distribution of Peltonotus species in Southeast Asia and Sundaland." figureDoi="http://doi.org/10.5281/zenodo.5030204" httpUri="https://zenodo.org/record/5030204/files/figure.png" pageId="27" pageNumber="28">Fig. 65</figureCitation>
).
<collectingCountry id="F31A56A3CA49FFEDFD86FD31FD2DFD4A" box="[584,664,684,710]" name="Brunei" pageId="27" pageNumber="28">Brunei</collectingCountry>
(Borneo);
<collectingCountry id="F31A56A3CA49FFEDFCD2FD31FC34FD4A" box="[796,897,684,710]" name="Malaysia" pageId="27" pageNumber="28">Sarawak</collectingCountry>
State, Bornean
<collectingCountry id="F31A56A3CA49FFEDFB8FFD31FB05FD4A" box="[1089,1200,684,710]" name="Malaysia" pageId="27" pageNumber="28">Malaysia</collectingCountry>
; and
<collectingRegion id="49C9D8D1CA49FFEDFB3FFD31FDB6FD62" country="Indonesia" name="Kalimantan Barat" pageId="27" pageNumber="28">West Kalimantan Province</collectingRegion>
, Bornean
<collectingCountry id="F31A56A3CA49FFEDFDB4FD49FD45FD62" box="[634,752,724,750]" name="Indonesia" pageId="27" pageNumber="28">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="8BB21633CA49FFEDFEF6FD61FBD6FC9A" blockId="27.[264,1323,284,1310]" box="[312,1123,764,790]" pageId="27" pageNumber="28">
<materialsCitation id="3B651C6ECA49FFEDFEF6FD61FBEAFC9A" ID-GBIF-Occurrence="3337342321" box="[312,1119,764,790]" collectionCode="BMNH, WADA, FUJI, MLJC" pageId="27" pageNumber="28" specimenCount="18">
Locality records (
<specimenCount id="9D0BDDBACA49FFEDFDC6FD61FD1CFC9A" box="[520,681,764,790]" pageId="27" pageNumber="28" type="generic">18 specimens</specimenCount>
) from
<collectionCode id="ED1C8EF6CA49FFEDFD37FD61FCE1FC9A" box="[761,852,764,790]" pageId="27" pageNumber="28">BMNH</collectionCode>
,
<collectionCode id="ED1C8EF6CA49FFEDFCAEFD61FC0EFC9A" box="[864,955,764,790]" country="Australia" httpUri="http://grbio.org/cool/mnrx-zeat" name="Western Australia Department of Agriculture" pageId="27" pageNumber="28">WADA</collectionCode>
,
<collectionCode id="ED1C8EF6CA49FFEDFC09FD61FBB0FC9A" box="[967,1029,764,790]" pageId="27" pageNumber="28">FUJI</collectionCode>
,
<collectionCode id="ED1C8EF6CA49FFEDFBDEFD61FBEAFC9A" box="[1040,1119,764,790]" pageId="27" pageNumber="28">MLJC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31745B8CA49FFEDFEF6FCB9FCA8FA92" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BB21633CA49FFEDFEF6FCB9FAADFCB2" blockId="27.[264,1323,284,1310]" box="[312,1304,804,830]" pageId="27" pageNumber="28">
BORNEAN
<collectingCountry id="F31A56A3CA49FFEDFE03FCB9FDC5FCB2" box="[461,624,804,830]" name="Indonesia" pageId="27" pageNumber="28">INDONESIA</collectingCountry>
.
<collectingRegion id="49C9D8D1CA49FFEDFDB5FCB9FC01FCB2" box="[635,948,804,830]" country="Indonesia" name="Kalimantan Barat" pageId="27" pageNumber="28">West Kalimantan Province</collectingRegion>
(12): Mt. Bawang, Pontianak.
</paragraph>
<paragraph id="8BB21633CA49FFEDFEF6FCD1FE05FC02" blockId="27.[264,1323,284,1310]" pageId="27" pageNumber="28">
BORNEAN
<collectingCountry id="F31A56A3CA49FFEDFE1CFCD1FDDBFCEA" box="[466,622,844,870]" name="Malaysia" pageId="27" pageNumber="28">MALAYSIA</collectingCountry>
.
<collectingRegion id="49C9D8D1CA49FFEDFDB1FCD1FC9EFCEA" box="[639,811,844,870]" country="Malaysia" name="Sarawak" pageId="27" pageNumber="28">Sarawak State</collectingRegion>
(1): Mt. Dulit (at junction of Tinjar and Lejak Rivers).
</paragraph>
<paragraph id="8BB21633CA49FFEDFEF6FC01FCA8FC3A" blockId="27.[264,1323,284,1310]" box="[312,797,924,950]" pageId="27" pageNumber="28">
<collectingCountry id="F31A56A3CA49FFEDFEF6FC01FE10FC3A" box="[312,421,924,950]" name="Brunei" pageId="27" pageNumber="28">BRUNEI</collectingCountry>
(
<collectionCode id="ED1C8EF6CA49FFEDFE7AFC01FD81FC3A" box="[436,564,924,950]" pageId="27" pageNumber="28">BORNEO</collectionCode>
). Kaula Belalag (5).
</paragraph>
<paragraph id="8BB21633CA49FFEDFEF6FC59FDD0FB8A" blockId="27.[264,1323,284,1310]" pageId="27" pageNumber="28">Temporal Data. February (4), March (2), April (4), July (1), August (1), October (1), November (1), December (2).</paragraph>
<paragraph id="8BB21633CA49FFEDFEF6FB89FBB2FBF2" blockId="27.[264,1323,284,1310]" pageId="27" pageNumber="28">
Natural History. The scant knowledge of natural history of
<taxonomicName id="4C0D6DB0CA49FFEDFC20FB89FB73FBA2" box="[1006,1222,1044,1070]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B979CA21CA49FFEDFC20FB89FBDEFBA1" box="[1006,1131,1044,1069]" italics="true" pageId="27" pageNumber="28">Peltonotus</emphasis>
species
</taxonomicName>
is based on observations of
<taxonomicName id="4C0D6DB0CA49FFEDFE3FFBA1FD26FBD9" box="[497,659,1084,1109]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B979CA21CA49FFEDFE3FFBA1FD26FBD9" box="[497,659,1084,1109]" italics="true" pageId="27" pageNumber="28">P. malayensis</emphasis>
</taxonomicName>
males and females in inflorescences of aroids at night (see discussion under “Natural History” of the genus
<taxonomicName id="4C0D6DB0CA49FFEDFCB7FBF9FC43FBF1" box="[889,1014,1124,1149]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B979CA21CA49FFEDFCB7FBF9FC43FBF1" box="[889,1014,1124,1149]" italics="true" pageId="27" pageNumber="28">Peltonotus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BB21633CA49FFEDFEF6FB11FCA8FA92" blockId="27.[264,1323,284,1310]" pageId="27" pageNumber="28">
Remarks. This species is sexually dimorphic for elytral color: males have reddishbrown elytra and females have black elytra with an iridescent bloom.
<bibRefCitation id="EF9C6BC2CA49FFEDFB4DFB29FA9FFB42" author="Arrow, G. J." box="[1155,1322,1204,1230]" pageId="27" pageNumber="28" pagination="153 - 157" refId="ref31219" refString="Arrow, G. J. (1910) On the lamellicorn beetles of the genus Peltonotus with descriptions of four new species. Annals and Magazine of Natural History (Series 8), 5, 153 - 157." type="journal article" year="1910">Arrow (1910)</bibRefCitation>
described this species based on female specimens. In 1931, Arrow (
<bibRefCitation id="EF9C6BC2CA49FFEDFB8DFB41FB62FB7A" author="Arrow, G. J." box="[1091,1239,1244,1270]" pageId="27" pageNumber="28" pagination="611 - 612" refId="ref31305" refString="Arrow, G. J. (1931) Two new species of lamellicorn beetles belonging to the genus Peltonotus. Annals and Magazine of Natural History (Series 10), 8, 611 - 612." type="journal article" year="1931">Arrow 1931</bibRefCitation>
) noted that males of this species were still unknown.
</paragraph>
</subSubSection>
</treatment>
</document>