302 lines
34 KiB
XML
302 lines
34 KiB
XML
<document id="4556F1AC2F108CDE9D43A57E73C7C134" ID-DOI="10.11646/zootaxa.1423.1.6" ID-ISSN="11755334" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625625910432" checkinUser="felipe" docAuthor="ASSING, VOLKER" docDate="2007" docId="039C87E3FF8BFF94FF64FA79FA16FA16" docLanguage="en" docName="zt01423p066.pdf.imf" docOrigin="Zootaxa 1423 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.1423.1.6" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Arpedium ludgeri ASSING, 2007, sp. n." docType="treatment" docVersion="2" lastPageNumber="65" masterDocId="FFA5FF9BFF8BFF96FFF3FFDBFF8AFFE9" masterDocTitle="A new species of Arpedium from Kyrgyzstan (Coleoptera: Staphylinidae, Omaliinae)" masterLastPageNumber="66" masterPageNumber="63" pageNumber="63" updateTime="1698993236489" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="B3FF07A9CF5DBA6CEDAD0F682B3DA86C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="18A98DDEF3F5523EA9FDCF84E5CD3D5B">
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<mods:title id="FE6A97D03CFFA0E3EEBBB6D625D78EA6">A new species of Arpedium from Kyrgyzstan (Coleoptera: Staphylinidae, Omaliinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="1FB2752003FC467D4242DCD736A85C63" type="personal">
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<mods:role id="EEF947314B5407071304741D568427C4">
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<mods:roleTerm id="A5371E703BC9FE04E1026AEBB0D1DB99">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DBEC92517395F5646B00B8C88FD5B880">ASSING, VOLKER</mods:namePart>
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<mods:typeOfResource id="E7B7653C4C5907B31B027C80E2C7DC0A">text</mods:typeOfResource>
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<mods:titleInfo id="BE369DF7779311679E67CB46A1D5BD03">
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<mods:title id="AA4B6BAF7AB11AF1FE6AA47C0C8E1201">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="51C4334CD2B07FC65E2D08639BF6DA17">
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<mods:date id="E753F637ABD70E107E55F2E93AB74BEA">2007</mods:date>
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<mods:detail id="80B73CFD2B72D253CE918CE065CD15A8" type="pubDate">
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<mods:number id="D3A66F08BD316FA57D092899EE333F0E">2007-03-12</mods:number>
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<mods:detail id="C093C15AED2F6CBF577A3AF731AC58F6" type="volume">
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<mods:number id="11B0C5C4A6E47037BEDC30A68EB307CF">1423</mods:number>
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</mods:detail>
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<mods:detail id="93AD9269A2B59E8361C691459CB372B0" type="issue">
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<mods:number id="14F3EF82A4DF4C850CF57CB3DF1EE508">1</mods:number>
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<mods:start id="388F1D400654556E446FF0CA942B1547">63</mods:start>
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<mods:end id="F0B295822EB38A4F979A575EEBFE344F">66</mods:end>
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<mods:location id="D60DC0871A8EEC8FB4F1DBB10990B103">
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<mods:url id="59D64344E097DE472F1F0D9CCFABE711">http://dx.doi.org/10.11646/zootaxa.1423.1.6</mods:url>
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<mods:classification id="D1502F2238172EAA93BADC97E2C6CAE0">journal article</mods:classification>
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<mods:identifier id="D9D6C0EBDAA8D08347570C9AD61BE8B6" type="DOI">10.11646/zootaxa.1423.1.6</mods:identifier>
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<mods:identifier id="4EBEF70B1B3044D1881DD72DA4655EF2" type="ISSN">11755334</mods:identifier>
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</mods:mods>
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<treatment id="039C87E3FF8BFF94FF64FA79FA16FA16" LSID="urn:lsid:plazi:treatment:039C87E3FF8BFF94FF64FA79FA16FA16" httpUri="http://treatment.plazi.org/id/039C87E3FF8BFF94FF64FA79FA16FA16" lastPageId="2" lastPageNumber="65" pageId="0" pageNumber="63">
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<subSubSection id="C32F657EFF8BFF96FF64FA79FE9DFA36" pageId="0" pageNumber="63" type="nomenclature">
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<paragraph id="8B8A36F5FF8BFF96FF64FA79FE3CFA55" blockId="0.[151,438,1442,1503]" box="[151,438,1442,1468]" pageId="0" pageNumber="63">
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<heading id="D0C28199FF8BFF96FF64FA79FE3CFA55" bold="true" box="[151,438,1442,1468]" fontSize="11" level="1" pageId="0" pageNumber="63" reason="1">
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<emphasis id="B941EAE7FF8BFF96FF64FA79FE3CFA55" bold="true" box="[151,438,1442,1468]" pageId="0" pageNumber="63">
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<taxonomicName id="4C354D76FF8BFF96FF64FA79FEE7FA55" ID-CoL="67T5S" authorityName="Assing" authorityYear="2007" box="[151,365,1442,1468]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="ludgeri" status="sp. nov.">
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<emphasis id="B941EAE7FF8BFF96FF64FA79FEE7FA55" bold="true" box="[151,365,1442,1468]" italics="true" pageId="0" pageNumber="63">Arpedium ludgeri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A272579CFF8BFF96FE87FA79FE3CFA55" box="[372,438,1442,1468]" pageId="0" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8A36F5FF8BFF96FF64FA1EFE9DFA36" blockId="0.[151,438,1442,1503]" box="[151,279,1477,1503]" pageId="0" pageNumber="63">
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(Figs.
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<docIssue id="CF56491AFF8BFF96FF11FA1EFF66FA36" box="[226,236,1477,1503]" pageId="0" pageNumber="63">1</docIssue>
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–9)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F657EFF8BFF96FF64F9CFFD3DF998" pageId="0" pageNumber="63" type="materials_examined">
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<paragraph id="8B8A36F5FF8BFF96FF64F9CFFD3DF998" blockId="0.[151,1437,1556,2031]" pageId="0" pageNumber="63">
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Type material.
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<materialsCitation id="3B5D3CA8FF8BFF96FEA0F9CFFA13F9A6" collectingDate="2005-07-22" collectionCode="V" collectorName="L. Schmidt & Assing" country="Kyrgyzstan" elevation="3775" latitude="41.946667" location="Issyk-Kul" longLatPrecision="19" longitude="77.36917" municipality="Torsor" pageId="0" pageNumber="63" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
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<emphasis id="B941EAE7FF8BFF96FEA0F9CFFE31F9C5" bold="true" box="[339,443,1556,1580]" pageId="0" pageNumber="63">
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<typeStatus id="548E8857FF8BFF96FEA0F9CFFE31F9C5" box="[339,443,1556,1580]" pageId="0" pageNumber="63" type="holotype">Holotype</typeStatus>
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</emphasis>
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<specimenCount id="9D33FD7CFF8BFF96FE39F9CDFE56F9CF" box="[458,476,1558,1574]" pageId="0" pageNumber="63" type="male">♂</specimenCount>
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: "
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<collectingCountry id="F3227665FF8BFF96FDF8F9CEFD01F9C5" box="[523,651,1557,1580]" name="Kyrgyzstan" pageId="0" pageNumber="63">Kyrgyzstan</collectingCountry>
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,
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<location id="8EEA602EFF8BFF96FD6DF9CEFC80F9C5" LSID="urn:lsid:plazi:treatment:039C87E3FF8BFF94FF64FA79FA16FA16:8EEA602EFF8BFF96FD6DF9CEFC80F9C5" box="[670,778,1557,1580]" country="Kyrgyzstan" latitude="41.946667" longLatPrecision="19" longitude="77.36917" municipality="Torsor" name="Issyk-Kul" pageId="0" pageNumber="63">Issyk-Kul</location>
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, Terskej-A[latau].,
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<collectingMunicipality id="6BEEAC8FFF8BFF96FC1BF9CEFBA7F9C5" box="[1000,1069,1557,1580]" pageId="0" pageNumber="63">Torsor</collectingMunicipality>
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pass,
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<date id="FF8B1035FF8BFF96FB73F9CEFA8EF9C5" box="[1152,1284,1557,1580]" pageId="0" pageNumber="63" value="2005-07-22">
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<collectingDate id="EFCFE9DDFF8BFF96FB73F9CEFA8EF9C5" box="[1152,1284,1557,1580]" pageId="0" pageNumber="63" value="2005-07-22">22.VII.2005</collectingDate>
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</date>
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,
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<geoCoordinate id="EE015032FF8BFF96FAEBF9CEFA12F9C5" box="[1304,1432,1557,1580]" degrees="41" direction="north" minutes="56" orientation="latitude" pageId="0" pageNumber="63" precision="15" seconds="48" value="41.946667">41°56'48"N</geoCoordinate>
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,
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<geoCoordinate id="EE015032FF8BFF96FF64F9E3FE9EF9A6" box="[151,276,1592,1615]" degrees="77" direction="east" minutes="22" orientation="longitude" pageId="0" pageNumber="63" precision="15" seconds="09" value="77.36917">77°22'09"E</geoCoordinate>
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,
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<quantity id="4CCD9B10FF8BFF96FED2F9E3FE30F9A6" box="[289,442,1592,1615]" metricMagnitude="3" metricUnit="m" metricValue="3.775" metricValueMax="3.85" metricValueMin="3.7" pageId="0" pageNumber="63" unit="m" value="3775.0" valueMax="3850.0" valueMin="3700.0">
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<elevation id="0018D1C6FF8BFF96FED2F9E3FE30F9A6" box="[289,442,1592,1615]" metricMagnitude="3" metricUnit="m" metricValue="3.775" metricValueMax="3.85" metricValueMin="3.7" pageId="0" pageNumber="63" unit="m" value="3775.0" valueMax="3850.0" valueMin="3700.0">3700–3850 m</elevation>
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</quantity>
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, l.
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<collectorName id="26C05323FF8BFF96FE2DF9E3FDDDF9A6" box="[478,599,1592,1615]" pageId="0" pageNumber="63">L. Schmidt</collectorName>
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/
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<typeStatus id="548E8857FF8BFF96FD82F9E3FD6AF9A6" box="[625,736,1592,1615]" pageId="0" pageNumber="63" type="holotype">Holotypus</typeStatus>
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<specimenCount id="9D33FD7CFF8BFF96FD19F9E2FD76F9A0" box="[746,764,1593,1609]" pageId="0" pageNumber="63" type="male">♂</specimenCount>
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<taxonomicName id="4C354D76FF8BFF96FCFDF9E3FC47F9A6" authorityName="Assing" authorityYear="2007" box="[782,973,1592,1615]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="ludgeri" status="sp. nov.">
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<emphasis id="B941EAE7FF8BFF96FCFDF9E3FC47F9A6" box="[782,973,1592,1615]" italics="true" pageId="0" pageNumber="63">Arpedium ludgeri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A272579CFF8BFF96FC25F9E3FB98F9A6" box="[982,1042,1592,1615]" pageId="0" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
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det
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<collectionCode id="ED24AE30FF8BFF96FBB6F9E3FBD2F9A6" box="[1093,1112,1592,1615]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="0" pageNumber="63" type="Museum">V</collectionCode>
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||
. Assing 2006 (coll.
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<collectorName id="26C05323FF8BFF96FAC4F9E3FA02F9A6" box="[1335,1416,1592,1615]" pageId="0" pageNumber="63">Assing</collectorName>
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||
)"
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B5D3CA8FF8BFF96FF64F981FD39F998" box="[151,691,1626,1650]" collectingDate="2005-07-22" collectionCode="V" collectorName="Assing" country="Kyrgyzstan" elevation="3775" latitude="41.946667" location="Issyk-Kul" longLatPrecision="19" longitude="77.36917" municipality="Torsor" pageId="0" pageNumber="63" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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||
<emphasis id="B941EAE7FF8BFF96FF64F981FF75F99B" bold="true" box="[151,255,1626,1650]" pageId="0" pageNumber="63">
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||
<typeStatus id="548E8857FF8BFF96FF64F981FF75F99B" box="[151,255,1626,1650]" pageId="0" pageNumber="63" type="paratype">Paratype</typeStatus>
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||
</emphasis>
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||
<specimenCount id="9D33FD7CFF8BFF96FEF6F980FE9DF982" box="[261,279,1627,1643]" pageId="0" pageNumber="63" type="male">♂</specimenCount>
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||
: same data as holotype (coll.
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||
<collectorName id="26C05323FF8BFF96FDAEF981FD26F998" box="[605,684,1626,1649]" pageId="0" pageNumber="63">Assing</collectorName>
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)
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</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32F657EFF8BFF97FF34F9A7FEE9F85A" lastPageId="1" lastPageNumber="64" pageId="0" pageNumber="63" type="description">
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<paragraph id="8B8A36F5FF8BFF96FF34F9A7FC99F8F6" blockId="0.[151,1437,1556,2031]" pageId="0" pageNumber="63">
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||
<emphasis id="B941EAE7FF8BFF96FF34F9A7FED8F97D" bold="true" box="[199,338,1660,1684]" inLineHeading="true" pageId="0" pageNumber="63" reason="1">Description.</emphasis>
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Measurements (mm) and ratios (
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<typeStatus id="548E8857FF8BFF96FD32F9A6FCA9F97D" box="[705,803,1661,1684]" pageId="0" pageNumber="63" type="holotype">holotype</typeStatus>
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,
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<typeStatus id="548E8857FF8BFF96FCC3F9A6FC1BF97D" box="[816,913,1661,1684]" pageId="0" pageNumber="63" type="paratype">paratype</typeStatus>
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): total body length: 4.5, 5.0; length of antenna: 1.37, 1.48; head width (HW): 0.62, 0.68; width of pronotum (PW): 0.83, 0.86; length of pronotum (PL): 0.57, 0.62; length of elytra at suture from apex of scutellum to posterior margin (EL): 0.62, 0.76; combined width of elytra: 1.01, 1.09; length of metatibia (TiL): 0.69, 0.72; length of metatarsus without claws (TaL): 0.36, 0.39; PW/HW: 1.34, 1.27; PW/PL: 1.45, 1.39; EL/PL: 1.08, 1.22; TaL/TiL: 0.52, 0.54.
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</paragraph>
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<paragraph id="8B8A36F5FF8BFF96FF34F8F1FBA0F806" blockId="0.[151,1437,1556,2031]" pageId="0" pageNumber="63">Habitus as in Fig. 1. Head with moderately fine puncturation, interstices on frons on average approximately as wide as diameter of punctures; pubescence moderately long, pale, and suberect; dorsal surface without distinct microsculpture; in front of ocelli with oblong and sharply delimited impressions. Eyes moderately small, distinctly projecting from lateral contours of head, slightly longer than postocular region in dorsal view. Maxillary palpus with apical palpomere 1.5–1.8 times as long as preapical palpomere. Antenna rather short; antennomeres IV–VI less than twice as long as wide; VII weakly oblong; VIII–IX approximately as wide as long; X weakly transverse (Fig. 4).</paragraph>
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<paragraph id="8B8A36F5FF8AFF97FF34FF43FE7AFED0" blockId="1.[151,1436,152,417]" pageId="1" pageNumber="64">
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<materialsCitation id="3B5D3CA8FF8AFF97FF34FF43FE66FED0" collectionCode="PW, PL" pageId="1" pageNumber="64" specimenCount="2" typeStatus="holotype">
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Pronotum distinctly transverse and wider than head (see measurements, ratios
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<collectionCode id="ED24AE30FF8AFF97FBE3FF43FBB2FF46" box="[1040,1080,152,175]" country="Thailand" name="Paleontological Collections" pageId="1" pageNumber="64">PW</collectionCode>
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||
/
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||
<collectionCode id="ED24AE30FF8AFF97FBB3FF43FBD5FF46" box="[1088,1119,152,175]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="1" pageNumber="64" type="Museum">PL</collectionCode>
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and
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<collectionCode id="ED24AE30FF8AFF97FB67FF43FB36FF46" box="[1172,1212,152,175]" country="Thailand" name="Paleontological Collections" pageId="1" pageNumber="64">PW</collectionCode>
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/HW, and Figs. 2–3), widest in anterior half; posterior angles obtuse, shortly rounded; posterior lateral area more or less distinctly depressed; puncturation variable, in
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<typeStatus id="548E8857FF8AFF97FE55FF06FD89FF1D" box="[422,515,221,244]" pageId="1" pageNumber="64" type="holotype">holotype</typeStatus>
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moderately pronounced, in
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<typeStatus id="548E8857FF8AFF97FCC7FF06FC1AFF1D" box="[820,912,221,244]" pageId="1" pageNumber="64" type="paratype">paratype</typeStatus>
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distinctly coarser and well-delimited, interstices on average as wide as or slightly narrower than diameter of punctures (Figs. 2–3); pubescence moderately long, pale, and suberect; microsculpture absent
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</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8B8A36F5FF8AFF97FF34FE9EFA8AFE48" blockId="1.[151,1436,152,417]" pageId="1" pageNumber="64">Elytra comparatively short, slightly longer than pronotum (see measurements and ratio EL/PL), and distinctly dilated caudad; puncturation distinct, but ill-defined; pubescence similar to that of pronotum; interstices without microsculpture. Hind wings reduced. Metatarsus approximately half the length of metatibia (see ratio TaL/TiL).</paragraph>
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<paragraph id="8B8A36F5FF8AFF97FF64F9BFFBF2F97F" blockId="1.[151,1436,1636,1686]" pageId="1" pageNumber="64">
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||
<emphasis id="B941EAE7FF8AFF97FF64F9BFFEB9F990" bold="true" box="[151,307,1636,1657]" pageId="1" pageNumber="64">FIGURES 1–9.</emphasis>
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||
<taxonomicName id="4C354D76FF8AFF97FEC9F9BFFE6EF990" authorityName="Assing" authorityYear="2007" box="[314,484,1636,1657]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="ludgeri">
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||
<emphasis id="B941EAE7FF8AFF97FEC9F9BFFE6EF990" box="[314,484,1636,1657]" italics="true" pageId="1" pageNumber="64">Arpedium ludgeri</emphasis>
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</taxonomicName>
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||
(3, 7, 9:
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<typeStatus id="548E8857FF8AFF97FDB3F9BFFD1DF990" box="[576,663,1636,1657]" pageId="1" pageNumber="64" type="holotype">holotype</typeStatus>
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||
; 1–2, 4–6, 8:
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<typeStatus id="548E8857FF8AFF97FCEDF9BFFCFCF990" box="[798,886,1636,1657]" pageId="1" pageNumber="64" type="paratype">paratype</typeStatus>
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): 1—habitus; 2–3—forebody; 4—antenna; 5—abdomen; 6–7—aedeagus; 8–9—apical portion of aedeagus. Scale bars: 1–3, 5: 1.0 mm; 4:
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<quantity id="4CCD9B10FF8AFF97FC52F95AFC67F97F" box="[929,1005,1665,1686]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="64" unit="mm" value="0.5">0.5 mm</quantity>
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; 6–9:
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<quantity id="4CCD9B10FF8AFF97FBD4F95AFBF9F97F" box="[1063,1139,1665,1686]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="64" unit="mm" value="0.1">0.1 mm</quantity>
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.
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</paragraph>
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<paragraph id="8B8A36F5FF8AFF97FF34F934FF3EF887" blockId="1.[151,1436,1775,1971]" pageId="1" pageNumber="64">Abdomen approximately 1.1 times as wide as elytra, widest at segments IV/V; puncturation moderately fine and conspicuously dense; pubescence more or less depressed, darker and shorter than that of forebody; all segments with pronounced microsculpture and almost matt; posterior margin of tergite VII with narrow rudiment of palisade fringe (Fig. 5).</paragraph>
|
||
<paragraph id="8B8A36F5FF8AFF97FF34F8A2FEE9F85A" blockId="1.[151,1436,1775,1971]" pageId="1" pageNumber="64">Male: aedeagus as in Figs. 6–9, with semitransparent internal tube; parameres apically not distinctly dilated and with 2 setae (Figs. 8–9).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F657EFF89FF94FF34FF4CFD3DFF38" pageId="2" pageNumber="65" type="etymology">
|
||
<paragraph id="8B8A36F5FF89FF94FF34FF4CFD3DFF38" blockId="2.[151,1437,151,1423]" pageId="2" pageNumber="65">
|
||
<emphasis id="B941EAE7FF89FF94FF34FF4CFECDFF46" bold="true" box="[199,327,151,175]" inLineHeading="true" pageId="2" pageNumber="65" reason="1">Etymology.</emphasis>
|
||
The species is dedicated to the collector of the
|
||
<typeStatus id="548E8857FF89FF94FCBFFF43FC0DFF46" box="[844,903,152,175]" pageId="2" pageNumber="65">types</typeStatus>
|
||
, Ludger Schmidt (Hannover), in gratitude for the gift of his staphylinid by-catches from
|
||
<collectingCountry id="F3227665FF89FF94FDC6FF61FD39FF38" box="[565,691,186,209]" name="Kyrgyzstan" pageId="2" pageNumber="65">Kyrgyzstan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F657EFF89FF94FF34FF07FBA4FAA0" pageId="2" pageNumber="65" type="discussion">
|
||
<paragraph id="8B8A36F5FF89FF94FF34FF07FC09FD15" blockId="2.[151,1437,151,1423]" pageId="2" pageNumber="65">
|
||
<emphasis id="B941EAE7FF89FF94FF34FF07FDDEFF1D" bold="true" box="[199,596,220,244]" inLineHeading="true" pageId="2" pageNumber="65" reason="1">Comments on generic assignment.</emphasis>
|
||
<taxonomicName id="4C354D76FF89FF94FDAEFF06FC96FF1D" authorityName="Assing" authorityYear="2007" box="[605,796,221,244]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="ludgeri">
|
||
<emphasis id="B941EAE7FF89FF94FDAEFF06FC96FF1D" box="[605,796,221,244]" italics="true" pageId="2" pageNumber="65">Arpedium ludgeri</emphasis>
|
||
</taxonomicName>
|
||
combines characters typically observed in
|
||
<taxonomicName id="4C354D76FF89FF94FB09FF06FAE9FF1D" authorityName="Erichson" authorityYear="1839" box="[1274,1379,221,244]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAE7FF89FF94FB09FF06FAE9FF1D" box="[1274,1379,221,244]" italics="true" pageId="2" pageNumber="65">Arpedium</emphasis>
|
||
</taxonomicName>
|
||
with those of
|
||
<taxonomicName id="4C354D76FF89FF94FF07FEDBFEF1FEFE" authorityName="Reitter" authorityYear="1909" box="[244,379,256,279]" class="Insecta" family="Staphylinidae" genus="Eucnecosum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAE7FF89FF94FF07FEDBFEF1FEFE" box="[244,379,256,279]" italics="true" pageId="2" pageNumber="65">Eucnecosum</emphasis>
|
||
</taxonomicName>
|
||
, thus rendering the separation of these taxa on the generic level doubtful. Based on the absence of microsculpture of the forebody, the long apical palpomere of the maxillary palpus (similar to the condition in
|
||
<taxonomicName id="4C354D76FF89FF94FAC1FEF9FE99FEB5" authority="Fauvel" authorityName="Fauvel" authorityYear="1878" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="cribratum">
|
||
<emphasis id="B941EAE7FF89FF94FAC1FEF9FF35FEB5" italics="true" pageId="2" pageNumber="65">A. cribratum</emphasis>
|
||
Fauvel
|
||
</taxonomicName>
|
||
), the shape of the impressions in front of the ocelli, as well as the rather coarse puncturation of the head and pronotum, the new species would have to be placed in
|
||
<taxonomicName id="4C354D76FF89FF94FD07FEB3FCD7FE96" authorityName="Erichson" authorityYear="1839" box="[756,861,360,383]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAE7FF89FF94FD07FEB3FCD7FE96" box="[756,861,360,383]" italics="true" pageId="2" pageNumber="65">Arpedium</emphasis>
|
||
</taxonomicName>
|
||
, whereas in size, shape and length of the wings, the presence of distinct pubescence on the forebody, and the relatively short antennae it is more similar to species currently attributed to
|
||
<taxonomicName id="4C354D76FF89FF94FEECFE76FE2CFE2D" authorityName="Reitter" authorityYear="1909" box="[287,422,429,452]" class="Insecta" family="Staphylinidae" genus="Eucnecosum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAE7FF89FF94FEECFE76FE2CFE2D" box="[287,422,429,452]" italics="true" pageId="2" pageNumber="65">Eucnecosum</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, a comparison of several species of
|
||
<taxonomicName id="4C354D76FF89FF94FC6DFE76FB8DFE2D" authorityName="Erichson" authorityYear="1839" box="[926,1031,429,452]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAE7FF89FF94FC6DFE76FB8DFE2D" box="[926,1031,429,452]" italics="true" pageId="2" pageNumber="65">Arpedium</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C354D76FF89FF94FBC8FE76FB48FE2D" authorityName="Reitter" authorityYear="1909" box="[1083,1218,429,452]" class="Insecta" family="Staphylinidae" genus="Eucnecosum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAE7FF89FF94FBC8FE76FB48FE2D" box="[1083,1218,429,452]" italics="true" pageId="2" pageNumber="65">Eucnecosum</emphasis>
|
||
</taxonomicName>
|
||
yielded no convincing evidence that both are distinct on the generic level. The characters indicated in the literature (see introduction) to distinguish the two taxa (e. g. presence/absence of microsculpture, length and density of pubescence, length of antennae, relative length of palpomeres) do not seem to justify such a distinction. Finally, I have been unable to appreciate the presence/absence of a subocular ridge as a useful character. This ridge is more or less pronounced in all
|
||
<taxonomicName id="4C354D76FF89FF94FB38FDE3FABEFDA6" authorityName="Erichson" authorityYear="1839" box="[1227,1332,568,591]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAE7FF89FF94FB38FDE3FABEFDA6" box="[1227,1332,568,591]" italics="true" pageId="2" pageNumber="65">Arpedium</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C354D76FF89FF94FA99FDE3FEC6FD98" class="Insecta" family="Staphylinidae" genus="Eucnecosum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B941EAE7FF89FF94FA99FDE3FF7EFD98" italics="true" pageId="2" pageNumber="65">Eucnecosum</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, but can be seen in ventral aspect only in those species with rather large eyes; in species with eyes of reduced size it is usually covered by the (more or less bulging) subocular portion of the head and can be seen only in lateral aspect. In conclusion, there is little doubt that
|
||
<taxonomicName id="4C354D76FF89FF94FD40FD7BFCB0FD5E" authorityName="Reitter" authorityYear="1909" box="[691,826,672,695]" class="Insecta" family="Staphylinidae" genus="Eucnecosum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAE7FF89FF94FD40FD7BFCB0FD5E" box="[691,826,672,695]" italics="true" pageId="2" pageNumber="65">Eucnecosum</emphasis>
|
||
</taxonomicName>
|
||
will eventually have to be treated as a subgenus again or may even have to be synonymised with
|
||
<taxonomicName id="4C354D76FF89FF94FDB6FD19FD24FD30" authorityName="Erichson" authorityYear="1839" box="[581,686,706,729]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAE7FF89FF94FDB6FD19FD24FD30" box="[581,686,706,729]" italics="true" pageId="2" pageNumber="65">Arpedium</emphasis>
|
||
</taxonomicName>
|
||
. Such changes, however, should be based on a thorough phylogenetic study of these taxa, which is not within the scope of the present paper.
|
||
</paragraph>
|
||
<paragraph id="8B8A36F5FF89FF94FF34FCDCFE86FCA8" blockId="2.[151,1437,151,1423]" pageId="2" pageNumber="65">
|
||
<emphasis id="B941EAE7FF89FF94FF34FCDCFE2FFCF6" bold="true" box="[199,421,775,799]" inLineHeading="true" pageId="2" pageNumber="65" reason="1">Comparative notes.</emphasis>
|
||
The new species is distinguished from other Palaearctic species of
|
||
<taxonomicName id="4C354D76FF89FF94FB8BFCD3FB6BFCF6" authorityName="Erichson" authorityYear="1839" box="[1144,1249,776,799]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAE7FF89FF94FB8BFCD3FB6BFCF6" box="[1144,1249,776,799]" italics="true" pageId="2" pageNumber="65">Arpedium</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C354D76FF89FF94FAE6FCD3FA16FCF6" authorityName="Reitter" authorityYear="1909" box="[1301,1436,776,799]" class="Insecta" family="Staphylinidae" genus="Eucnecosum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAE7FF89FF94FAE6FCD3FA16FCF6" box="[1301,1436,776,799]" italics="true" pageId="2" pageNumber="65">Eucnecosum</emphasis>
|
||
</taxonomicName>
|
||
as follows:
|
||
</paragraph>
|
||
<paragraph id="8B8A36F5FF89FF94FF34FC96FBF2FBF8" blockId="2.[151,1437,151,1423]" pageId="2" pageNumber="65">
|
||
From
|
||
<taxonomicName id="4C354D76FF89FF94FEFBFC96FE0CFC8D" baseAuthorityName="Gravenhorst" baseAuthorityYear="1806" box="[264,390,845,868]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="quadrum">
|
||
<emphasis id="B941EAE7FF89FF94FEFBFC96FE0CFC8D" box="[264,390,845,868]" italics="true" pageId="2" pageNumber="65">A. quadrum</emphasis>
|
||
</taxonomicName>
|
||
by much smaller size, darker coloration especially of pronotum and elytra, less glossy, as well as more densely and distinctly pubescent forebody, reduced hind wings, shorter tibiae and tarsi, the much shorter antennae with less slender antennomeres, stouter maxillary palpi with relatively longer apical palpomeres, much smaller eyes, a distinctly more transverse and posteriorly more distinctly narrowed pronotum, distinctly finer and less well-defined puncturation of pronotum and elytra, much shorter and posteriorly more strongly dilated elytra, coarser and denser puncturation of the abdomen, and by the apically not distinctly dilated parameres of the aedeagus;
|
||
</paragraph>
|
||
<paragraph id="8B8A36F5FF89FF94FF34FBC6FCF3FB56" blockId="2.[151,1437,151,1423]" pageId="2" pageNumber="65">
|
||
From
|
||
<taxonomicName id="4C354D76FF89FF94FEFEFBC6FE2AFBDD" authorityName="Coiffait" authorityYear="1982" box="[269,416,1053,1076]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="nepalicum">
|
||
<emphasis id="B941EAE7FF89FF94FEFEFBC6FE2AFBDD" box="[269,416,1053,1076]" italics="true" pageId="2" pageNumber="65">A. nepalicum</emphasis>
|
||
</taxonomicName>
|
||
[characters taken from description by
|
||
<bibRefCitation id="EFA44B04FF89FF94FCACFBC6FB83FBDD" author="Coiffait, H." box="[863,1033,1053,1076]" pageId="2" pageNumber="65" pagination="21 - 179" refId="ref1985" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana biologica, 62 (1981), 21 - 179." type="journal article" year="1982">Coiffait (1982)</bibRefCitation>
|
||
], whose male sexual characters are unknown, by distinctly larger size (
|
||
<taxonomicName id="4C354D76FF89FF94FDFEFB9BFD13FBBE" authorityName="Coiffait" authorityYear="1982" box="[525,665,1088,1111]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="nepalicum">
|
||
<emphasis id="B941EAE7FF89FF94FDFEFB9BFD13FBBE" box="[525,665,1088,1111]" italics="true" pageId="2" pageNumber="65">A. nepalicum</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<quantity id="4CCD9B10FF89FF94FD54FB9BFD71FBBE" box="[679,763,1088,1111]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="65" unit="mm" value="2.5">2.5 mm</quantity>
|
||
), darker coloration (
|
||
<taxonomicName id="4C354D76FF89FF94FC20FB9BFBD5FBBE" authorityName="Coiffait" authorityYear="1982" box="[979,1119,1088,1111]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="nepalicum">
|
||
<emphasis id="B941EAE7FF89FF94FC20FB9BFBD5FBBE" box="[979,1119,1088,1111]" italics="true" pageId="2" pageNumber="65">A. nepalicum</emphasis>
|
||
</taxonomicName>
|
||
: whole body reddish yellow), more pronounced posterior angles of the pronotum (in
|
||
<taxonomicName id="4C354D76FF89FF94FD2CFBB8FCE1FB93" authorityName="Coiffait" authorityYear="1982" box="[735,875,1123,1146]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="nepalicum">
|
||
<emphasis id="B941EAE7FF89FF94FD2CFBB8FCE1FB93" box="[735,875,1123,1146]" italics="true" pageId="2" pageNumber="65">A. nepalicum</emphasis>
|
||
</taxonomicName>
|
||
broadly rounded), reduced hind wings (in
|
||
<taxonomicName id="4C354D76FF89FF94FAC0FBB8FF49FB75" authorityName="Coiffait" authorityYear="1982" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="nepalicum">
|
||
<emphasis id="B941EAE7FF89FF94FAC0FBB8FF49FB75" italics="true" pageId="2" pageNumber="65">A. nepalicum</emphasis>
|
||
</taxonomicName>
|
||
almost reaching apex of abdomen), denser puncturation and the more pronounced microreticulation of the abdomen (in
|
||
<taxonomicName id="4C354D76FF89FF94FF48FB73FECDFB56" authorityName="Coiffait" authorityYear="1982" box="[187,327,1192,1215]" class="Insecta" family="Staphylinidae" genus="Arpedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="nepalicum">
|
||
<emphasis id="B941EAE7FF89FF94FF48FB73FECDFB56" box="[187,327,1192,1215]" italics="true" pageId="2" pageNumber="65">A. nepalicum</emphasis>
|
||
</taxonomicName>
|
||
with shallow transverse microsculpture and glossy);
|
||
</paragraph>
|
||
<paragraph id="8B8A36F5FF89FF94FF34FB11FBA4FAA0" blockId="2.[151,1437,151,1423]" pageId="2" pageNumber="65">
|
||
From the species currently attributed to
|
||
<taxonomicName id="4C354D76FF89FF94FD9DFB10FD7FFB0B" authorityName="Reitter" authorityYear="1909" box="[622,757,1227,1250]" class="Insecta" family="Staphylinidae" genus="Eucnecosum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAE7FF89FF94FD9DFB10FD7FFB0B" box="[622,757,1227,1250]" italics="true" pageId="2" pageNumber="65">Eucnecosum</emphasis>
|
||
</taxonomicName>
|
||
by the absence of microsculpture on the head and pronotum, by the longer impressions in front of the ocelli, by the stouter maxillary palpus with a relatively longer apical palpomere, by the distinctly more transverse, less convex (cross-section), and posteriorly more distinctly narrowed pronotum, the coarser puncturation of the pronotum, and by the denser puncturation of the abdomen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F657EFF89FF94FF34FA8FFA16FA16" pageId="2" pageNumber="65" type="distribution">
|
||
<paragraph id="8B8A36F5FF89FF94FF34FA8FFBD2FA66" blockId="2.[151,1437,151,1423]" pageId="2" pageNumber="65">
|
||
<emphasis id="B941EAE7FF89FF94FF34FA8FFD8CFA85" bold="true" box="[199,518,1364,1388]" inLineHeading="true" pageId="2" pageNumber="65" reason="1">Distribution and bionomics.</emphasis>
|
||
The
|
||
<typeStatus id="548E8857FF89FF94FDCDFA8EFDE6FA85" box="[574,620,1365,1388]" pageId="2" pageNumber="65">type</typeStatus>
|
||
locality is situated in eastern
|
||
<collectingCountry id="F3227665FF89FF94FC5FFA8EFBA3FA85" box="[940,1065,1365,1388]" name="Kyrgyzstan" pageId="2" pageNumber="65">Kyrgyzstan</collectingCountry>
|
||
, to the south of the Issyk-kul lake. The specimens were found under stones in stony grassland at an altitude of
|
||
<quantity id="4CCD9B10FF89FF94FC4EFAA3FBDEFA66" box="[957,1108,1400,1423]" metricMagnitude="3" metricUnit="m" metricValue="3.775" metricValueMax="3.85" metricValueMin="3.7" pageId="2" pageNumber="65" unit="m" value="3775.0" valueMax="3850.0" valueMin="3700.0">3700–3850 m</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8A36F5FF89FF94FB47FA19FA16FA16" blockId="2.[620,1436,1474,1535]" pageId="2" pageNumber="65">
|
||
VOLKER ASSING
|
||
<emphasis id="B941EAE7FF89FF94FD9FFA31FA16FA16" box="[620,1436,1514,1535]" italics="true" pageId="2" pageNumber="65">
|
||
Gabelsbergerstr. 2, D-30163 Hannover,
|
||
<collectingCountry id="F3227665FF89FF94FC1EFA31FBC2FA16" box="[1005,1096,1514,1535]" name="Germany" pageId="2" pageNumber="65">Germany</collectingCountry>
|
||
. E-mail: vassing.hann@t-online.de
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |