644 lines
73 KiB
XML
644 lines
73 KiB
XML
<document id="89E6F8F5D05344336BCFC655851E8946" ID-CLB-Dataset="26665" ID-DOI="10.11646/zootaxa.4612.1.3" ID-GBIF-Dataset="43d2d964-7ece-4f81-be1c-169c0592de90" ID-ISSN="1175-5326" ID-Zenodo-Dep="3233614" ID-ZooBank="6751A2A1-2438-4759-B14C-4639595D9F82" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1559035746592" checkinUser="plazi" docAuthor="Haber, William A." docDate="2019" docId="039C87D70659FFF6FF66298CFD15E896" docLanguage="en" docName="zootaxa.4612.1.3.pdf" docOrigin="Zootaxa 4612 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gynacantha vargasi Haber 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="70" masterDocId="FFA5FFAF0658FFFCFFF12A59FFF5EA65" masterDocTitle="Gynacantha vargasi (Odonata: Anisoptera: Aeshnidae) sp. nov. from Costa Rica" masterLastPageNumber="70" masterPageNumber="58" pageNumber="58" updateTime="1698732123088" updateUser="ExternalLinkService">
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<mods:titleInfo id="68B2477D0C671EE7D81556EBCDC96E94">
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<mods:title id="BE709D8F2A60C99C69E8395C6F9D8C0E">Gynacantha vargasi (Odonata: Anisoptera: Aeshnidae) sp. nov. from Costa Rica</mods:title>
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<mods:name id="AB11165AA9594978B7EC79651F6D3925" type="personal">
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<mods:namePart id="7825310673A8E8EADA2F99FBFF8F0A39">Haber, William A.</mods:namePart>
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<mods:typeOfResource id="5B90A31C8FA460BB7832E711216EBAEF">text</mods:typeOfResource>
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<mods:title id="C3A3620E5FD9A8310FFAEC622FB47541">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F98F000B99F17AA65F91D0BA1B508BE7">
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<mods:date id="BA38028319826FC8778AD51AB05593B6">2019</mods:date>
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<mods:detail id="141B165D6D89104E611EFA36724AC5AB" type="pubDate">
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<mods:number id="7138E511D4FD6E884C1AD48F075F7964">2019-05-28</mods:number>
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<mods:number id="F863DB366C327B7721AFD026BCCF1934">4612</mods:number>
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<mods:detail id="ADB9A378222CE2739F9E7C988B23A091" type="issue">
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<mods:number id="49772716CBF0F47949515A3DAA18723D">1</mods:number>
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<mods:identifier id="F770E5E7FBF0882EC2C1CF2F03DC92CE" type="DOI">10.11646/zootaxa.4612.1.3</mods:identifier>
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<mods:identifier id="9E6D9E7DCAC6A02012FEB0B84375377F" type="GBIF-Dataset">43d2d964-7ece-4f81-be1c-169c0592de90</mods:identifier>
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<mods:identifier id="E59D4D934737D5B5C79FFEAFBBF96127" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3422BDE202FC858CDA6950241599CA55" type="Zenodo-Dep">3233614</mods:identifier>
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</mods:mods>
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<treatment id="039C87D70659FFF6FF66298CFD15E896" ID-GBIF-Taxon="157292249" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896" httpUri="http://treatment.plazi.org/id/039C87D70659FFF6FF66298CFD15E896" lastPageId="10" pageId="1">
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<subSubSection id="C32F654A0659FFFDFF66298CFEFEEE76" pageId="1" type="nomenclature">
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<paragraph id="8B8A36C10659FFFDFF66298CFE05E995" blockId="1.[151,496,981,1043]" box="[151,496,981,1008]" pageId="1">
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<heading id="D0C281AD0659FFFDFF66298CFE05E995" bold="true" box="[151,496,981,1008]" fontSize="11" level="1" pageId="1" reason="1">
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<emphasis id="B941EAD30659FFFDFF66298CFE05E995" bold="true" box="[151,496,981,1008]" pageId="1">
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||
<taxonomicName id="4C354D420659FFFDFF66298CFE7EE98A" ID-CoL="9X2VK" authority="Haber, 2019" authorityName="Haber" authorityYear="2019" box="[151,395,981,1008]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="1" phylum="Arthropoda" rank="species" species="vargasi" status="sp. nov.">
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<emphasis id="B941EAD30659FFFDFF66298CFE7EE98A" bold="true" box="[151,395,981,1008]" italics="true" pageId="1">Gynacantha vargasi</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A27257A80659FFFDFE63298FFE05E995" box="[402,496,982,1008]" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8B8A36C10659FFFDFF6629A0FEFEEE76" blockId="1.[151,496,981,1043]" box="[151,267,1017,1043]" pageId="1">(Figs 1–7)</paragraph>
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</subSubSection>
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<subSubSection id="C32F654A0659FFFDFF662E1BFC61EF6A" pageId="1" type="materials_examined">
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<paragraph id="8B8A36C10659FFFDFF662E1BFA7CEE1A" blockId="1.[151,1437,1089,2016]" pageId="1">
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<materialsCitation id="3B5D3C9C0659FFFDFF662E1BFA7CEE1A" ID-GBIF-Occurrence="2244393148" collectingDate="2009-06-23" collectionCode="INBIO-MNCR" collectorName="Juan Chavez & Ronald Vargas & William Haber" country="Costa Rica" county="Male" elevation="850" location="Reserva Las Brisas" municipality="Provincia" pageId="1" specimenCount="1" stateProvince="Limon" typeStatus="holotype">
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||
<emphasis id="B941EAD30659FFFDFF662E1BFEA4EE39" bold="true" box="[151,337,1090,1116]" pageId="1">
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||
<typeStatus id="548E88630659FFFDFF662E1BFEF3EE39" box="[151,262,1090,1116]" pageId="1" type="holotype">Holotype</typeStatus>
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||
.
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||
<collectingCounty id="62EB4E4D0659FFFDFEE52E1BFEA4EE39" box="[276,337,1090,1116]" pageId="1">Male</collectingCounty>
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||
</emphasis>
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||
(
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||
<collectionCode id="ED24AE040659FFFDFE912E18FDE5EE3E" box="[352,528,1089,1115]" pageId="1">INBIO–MNCR</collectionCode>
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<emphasis id="B941EAD30659FFFDFDE02E18FDD4EE39" bold="true" box="[529,545,1089,1116]" pageId="1">),</emphasis>
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<collectingCountry id="F32276510659FFFDFDDB2E18FD24EE3E" box="[554,721,1089,1115]" name="Costa Rica" pageId="1">COSTA RICA</collectingCountry>
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||
:
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<collectingMunicipality id="6BEEACBB0659FFFDFD2F2E18FCBCEE3E" box="[734,841,1089,1115]" pageId="1">Provincia</collectingMunicipality>
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||
<collectingRegion id="49F1F8230659FFFDFCA32E18FC55EE3E" box="[850,928,1089,1115]" country="Costa Rica" name="Limon" pageId="1">Limón</collectingRegion>
|
||
,
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||
<locationDeviation id="9A382FE30659FFFDFC5D2E18FB5AEE39" box="[940,1199,1089,1116]" location="Pocora" pageId="1">
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||
<quantity id="4CCD9B240659FFFDFC5D2E18FC01EE3E" box="[940,1012,1089,1115]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="1" unit="km" value="11.0">11 km</quantity>
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||
SSW of Pocora
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||
</locationDeviation>
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||
,
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||
<location id="8EEA601A0659FFFDFB4A2E1BFA6DEE3E" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896:8EEA601A0659FFFDFB4A2E1BFA6DEE3E" box="[1211,1432,1089,1116]" country="Costa Rica" county="Male" municipality="Provincia" name="Reserva Las Brisas" pageId="1" stateProvince="Limon">Reserva Las Brisas</location>
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,
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<location id="8EEA601A0659FFFDFF662E3CFED9EE1A" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896:8EEA601A0659FFFDFF662E3CFED9EE1A" box="[151,300,1125,1151]" country="Costa Rica" county="Male" municipality="Provincia" name="Rio Cuarenta" pageId="1" stateProvince="Limon">Río Cuarenta</location>
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||
y
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<location id="8EEA601A0659FFFDFEB92E3CFE7EEE1A" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896:8EEA601A0659FFFDFEB92E3CFE7EEE1A" box="[328,395,1125,1151]" country="Costa Rica" county="Male" municipality="Provincia" name="Cinco" pageId="1" stateProvince="Limon">Cinco</location>
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||
(10.07, -83.63,
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||
<elevation id="0018D1F20659FFFDFDCF2E3CFD5BEEE5" box="[574,686,1125,1152]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="1" unit="m" value="850.0">
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||
<quantity id="4CCD9B240659FFFDFDCF2E3CFD70EEE5" box="[574,645,1125,1152]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="1" unit="m" value="850.0">850 m</quantity>
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asl
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</elevation>
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),
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<collectingDate id="EFCFE9E90659FFFDFD332E3CFCAEEE1A" box="[706,859,1125,1152]" pageId="1" value="2009-06-23">
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<date id="FF8B10010659FFFDFD332E3CFCAEEE1A" box="[706,859,1125,1152]" pageId="1" value="2009-06-23">23 June 2009</date>
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</collectingDate>
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||
,
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<collectorName id="26C053170659FFFDFC972E3FFC02EE1A" box="[870,1015,1125,1152]" pageId="1">Juan Chávez</collectorName>
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||
,
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||
<collectorName id="26C053170659FFFDFBF32E3CFB5DEEE5" box="[1026,1192,1125,1152]" pageId="1">Ronald Vargas</collectorName>
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||
,
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||
<collectorName id="26C053170659FFFDFB422E3CFAADEE1A" box="[1203,1368,1125,1151]" pageId="1">William Haber</collectorName>
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||
leg.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8B8A36C10659FFFDFF362ED3FC61EF6A" blockId="1.[151,1437,1089,2016]" pageId="1">
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||
<materialsCitation id="3B5D3C9C0659FFFDFF362ED3FDC5EE8E" ID-GBIF-Occurrence="2244393151" collectingDate="2003-04-19" collectionCode="WH, UCMS" collectorName="Ronald Vargas & Female" country="Costa Rica" county="Male" elevation="500" location="Cascante Refuge" municipality="Braulio Carrillo National Park" pageId="1" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
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||
<emphasis id="B941EAD30659FFFDFF362ED3FE78EEC1" bold="true" box="[199,397,1162,1188]" pageId="1">
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||
<typeStatus id="548E88630659FFFDFF362ED3FEB4EEC1" box="[199,321,1162,1188]" pageId="1" type="paratype">Paratypes</typeStatus>
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||
.
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||
<collectingCounty id="62EB4E4D0659FFFDFEA12ED3FE78EEC1" box="[336,397,1162,1188]" pageId="1">Male</collectingCounty>
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||
</emphasis>
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||
(
|
||
<collectionCode id="ED24AE040659FFFDFE6F2ED0FE27EEC6" box="[414,466,1161,1187]" pageId="1">WH</collectionCode>
|
||
),
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||
<collectingCountry id="F32276510659FFFDFE192ED0FD64EEC6" box="[488,657,1161,1187]" name="Costa Rica" pageId="1">COSTA RICA</collectingCountry>
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||
:
|
||
<collectingRegion id="49F1F8230659FFFDFD6E2ED0FC85EEC6" box="[671,880,1161,1187]" country="Costa Rica" name="Heredia" pageId="1">Provincia Heredia</collectingRegion>
|
||
,
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||
<locationDeviation id="9A382FE30659FFFDFC8C2ED0FB85EEC6" box="[893,1136,1161,1188]" location="La Virgen" pageId="1">
|
||
<quantity id="4CCD9B240659FFFDFC8C2ED0FC33EEC6" box="[893,966,1161,1187]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="1" unit="km" value="11.0">11 km</quantity>
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||
SE La Virgen
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||
</locationDeviation>
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||
,
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||
<collectingMunicipality id="6BEEACBB0659FFFDFB8C2ED0FF38EEA2" pageId="1">Braulio Carrillo National Park</collectingMunicipality>
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||
, vicinity
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||
<location id="8EEA601A0659FFFDFEC92EF4FE0FEEA2" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896:8EEA601A0659FFFDFEC92EF4FE0FEEA2" box="[312,506,1197,1223]" country="Costa Rica" county="Male" municipality="Braulio Carrillo National Park" name="Cascante Refuge" pageId="1" stateProvince="Heredia">Cascante Refuge</location>
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||
, east side of
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||
<location id="8EEA601A0659FFFDFD602EF4FD00EEA2" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896:8EEA601A0659FFFDFD602EF4FD00EEA2" box="[657,757,1197,1223]" country="Costa Rica" county="Male" municipality="Braulio Carrillo National Park" name="Rio Peje" pageId="1" stateProvince="Heredia">Río Peje</location>
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||
,
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||
<location id="8EEA601A0659FFFDFCF32EF4FC40EEA2" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896:8EEA601A0659FFFDFCF32EF4FC40EEA2" box="[770,949,1197,1223]" country="Costa Rica" county="Male" municipality="Braulio Carrillo National Park" name="Quebrada Lajas" pageId="1" stateProvince="Heredia">Quebrada Lajas</location>
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||
(10.33, -84.07,
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||
<elevation id="0018D1F20659FFFDFB9C2EF4FB15EEAD" box="[1133,1248,1197,1224]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" unit="m" value="500.0">
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||
<quantity id="4CCD9B240659FFFDFB9C2EF4FB43EEAD" box="[1133,1206,1197,1224]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" unit="m" value="500.0">500 m</quantity>
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||
asl
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||
</elevation>
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||
),
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||
<collectingDate id="EFCFE9E90659FFFDFB072EF4FA6DEEA2" box="[1270,1432,1197,1223]" pageId="1" value="2003-04-19">
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||
<date id="FF8B10010659FFFDFB072EF4FA6DEEA2" box="[1270,1432,1197,1223]" pageId="1" value="2003-04-19">19 April 2003</date>
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||
</collectingDate>
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||
,
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||
<collectorName id="26C053170659FFFDFF662E88FECFEE89" box="[151,314,1233,1260]" pageId="1">Ronald Vargas</collectorName>
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||
leg.
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||
<collectorName id="26C053170659FFFDFE9F2E8BFE30EE89" box="[366,453,1234,1260]" pageId="1">
|
||
<emphasis id="B941EAD30659FFFDFE9F2E8BFE30EE89" bold="true" box="[366,453,1234,1260]" pageId="1">Female</emphasis>
|
||
</collectorName>
|
||
(
|
||
<collectionCode id="ED24AE040659FFFDFE222E88FDDDEE8E" box="[467,552,1233,1259]" country="USA" httpUri="http://grbio.org/cool/gg3r-fyna" name="University of Connecticut Biodiversity Research Collections" pageId="1" type="University or college">UCMS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B5D3C9C0659FFFDFDCB2E88FC61EF6A" ID-GBIF-Occurrence="2244393149" collectingDate="2006-08-23" collectorName="Ronald Vargas" country="Costa Rica" county="Provincia" elevation="850" location="Rio Cuarenta" municipality="Reserva Las Brisas" pageId="1" specimenCount="1" stateProvince="Limon" typeStatus="paratype">
|
||
<collectingCounty id="62EB4E4D0659FFFDFDCB2E88FD50EE8E" box="[570,677,1233,1259]" pageId="1">Provincia</collectingCounty>
|
||
<collectingRegion id="49F1F8230659FFFDFD5A2E88FD0CEE8E" box="[683,761,1233,1259]" country="Costa Rica" name="Limon" pageId="1">Limón</collectingRegion>
|
||
,
|
||
<locationDeviation id="9A382FE30659FFFDFCF22E88FBE8EE8E" box="[771,1053,1233,1259]" location="Siquirres" pageId="1">
|
||
<quantity id="4CCD9B240659FFFDFCF22E88FCBCEE8E" box="[771,841,1233,1259]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="1" unit="km" value="14.0">14 km</quantity>
|
||
WSW of Siquirres
|
||
</locationDeviation>
|
||
,
|
||
<collectingMunicipality id="6BEEACBB0659FFFDFBD62E8BFB0AEE8E" box="[1063,1279,1233,1260]" pageId="1">Reserva Las Brisas</collectingMunicipality>
|
||
,
|
||
<location id="8EEA601A0659FFFDFAF82E88FA69EE8E" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896:8EEA601A0659FFFDFAF82E88FA69EE8E" box="[1289,1436,1233,1259]" country="Costa Rica" county="Provincia" municipality="Reserva Las Brisas" name="Rio Cuarenta" pageId="1" stateProvince="Limon">Río Cuarenta</location>
|
||
y
|
||
<location id="8EEA601A0659FFFDFF5D2EACFF1AEF6A" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896:8EEA601A0659FFFDFF5D2EACFF1AEF6A" box="[172,239,1269,1295]" country="Costa Rica" county="Provincia" municipality="Reserva Las Brisas" name="Cinco" pageId="1" stateProvince="Limon">Cinco</location>
|
||
(10.07, -83.63,
|
||
<elevation id="0018D1F20659FFFDFE522EACFE18EF75" box="[419,493,1269,1296]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="1" unit="m" value="850.0">
|
||
<quantity id="4CCD9B240659FFFDFE522EACFE18EF75" box="[419,493,1269,1296]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="1" unit="m" value="850.0">850 m</quantity>
|
||
</elevation>
|
||
),
|
||
<collectingDate id="EFCFE9E90659FFFDFDF02EACFD40EF6A" box="[513,693,1269,1296]" pageId="1" value="2006-08-23">
|
||
<date id="FF8B10010659FFFDFDF02EACFD40EF6A" box="[513,693,1269,1296]" pageId="1" value="2006-08-23">23 August 2006</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C053170659FFFDFD312EACFC91EF75" box="[704,868,1269,1296]" pageId="1">Ronald Vargas</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F654A0659FFFDFF362F40FAE7EF32" pageId="1" type="description">
|
||
<paragraph id="8B8A36C10659FFFDFF362F40FAE7EF32" blockId="1.[151,1437,1089,2016]" pageId="1">
|
||
<materialsCitation id="3B5D3C9C0659FFFDFF362F40FAE7EF32" ID-GBIF-Occurrence="2244393150" collectingDate="2004-11-20" collectionCode="INBIO-MNCR" collectorName="Eladio Cruz" country="Costa Rica" county="Female" elevation="800" location="Monteverde Cloud Forest Biological Reserve" municipality="Provincia" pageId="1" specimenCount="1" stateProvince="Alajuela" typeStatus="allotype">
|
||
<typeStatus id="548E88630659FFFDFF362F40FED9EF51" box="[199,300,1305,1332]" pageId="1" type="allotype">
|
||
<emphasis id="B941EAD30659FFFDFF362F40FED9EF51" bold="true" box="[199,300,1305,1332]" pageId="1">Allotype</emphasis>
|
||
</typeStatus>
|
||
.
|
||
<collectingCounty id="62EB4E4D0659FFFDFEC82F43FE65EF51" box="[313,400,1306,1332]" pageId="1">
|
||
<emphasis id="B941EAD30659FFFDFEC82F43FE65EF51" bold="true" box="[313,400,1306,1332]" pageId="1">Female</emphasis>
|
||
</collectingCounty>
|
||
(
|
||
<collectionCode id="ED24AE040659FFFDFE6C2F40FDB4EF56" box="[413,577,1305,1331]" pageId="1">INBIO–MNCR</collectionCode>
|
||
),
|
||
<collectingCountry id="F32276510659FFFDFDA02F40FD12EF56" box="[593,743,1305,1331]" name="Costa Rica" pageId="1">COSTA RICA</collectingCountry>
|
||
:
|
||
<collectingMunicipality id="6BEEACBB0659FFFDFD1E2F40FCBBEF56" box="[751,846,1305,1331]" pageId="1">Provincia</collectingMunicipality>
|
||
<collectingRegion id="49F1F8230659FFFDFCA02F40FC52EF56" box="[849,935,1305,1331]" country="Costa Rica" name="Alajuela" pageId="1">Alajuela</collectingRegion>
|
||
,
|
||
<location id="8EEA601A0659FFFDFC5F2F40FA9BEF51" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896:8EEA601A0659FFFDFC5F2F40FA9BEF51" box="[942,1390,1305,1332]" country="Costa Rica" county="Female" municipality="Provincia" name="Monteverde Cloud Forest Biological Reserve" pageId="1" stateProvince="Alajuela">Monteverde Cloud Forest Biological Reserve</location>
|
||
,
|
||
<location id="8EEA601A0659FFFDFA862F40FE9FEF32" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896:8EEA601A0659FFFDFA862F40FE9FEF32" country="Costa Rica" county="Female" municipality="Provincia" name="Rio Penas Blancas Valley" pageId="1" stateProvince="Alajuela">Río Peñas Blancas Valley</location>
|
||
,
|
||
<location id="8EEA601A0659FFFDFE852F64FDF8EF32" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896:8EEA601A0659FFFDFE852F64FDF8EF32" box="[372,525,1341,1367]" country="Costa Rica" county="Female" municipality="Provincia" name="Refugio Eladio" pageId="1" stateProvince="Alajuela">Refugio Eladio</location>
|
||
,
|
||
<location id="8EEA601A0659FFFDFDE42F64FD12EF32" LSID="urn:lsid:plazi:treatment:039C87D70659FFF6FF66298CFD15E896:8EEA601A0659FFFDFDE42F64FD12EF32" box="[533,743,1341,1367]" country="Costa Rica" county="Female" municipality="Provincia" name="Quebrada Pavoncillo" pageId="1" stateProvince="Alajuela">Quebrada Pavoncillo</location>
|
||
(10.31, -84.72),
|
||
<elevation id="0018D1F20659FFFDFC7B2F64FC39EF3D" box="[906,972,1341,1368]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" unit="m" value="800.0">
|
||
<quantity id="4CCD9B240659FFFDFC7B2F64FC39EF3D" box="[906,972,1341,1368]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" unit="m" value="800.0">800 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFCFE9E90659FFFDFC242F64FB93EF32" box="[981,1126,1341,1367]" pageId="1" value="2004-11-20">
|
||
<date id="FF8B10010659FFFDFC242F64FB93EF32" box="[981,1126,1341,1367]" pageId="1" value="2004-11-20">20 NOV 2004</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C053170659FFFDFB9E2F64FB12EF32" box="[1135,1255,1341,1367]" pageId="1">Eladio Cruz</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F654A0659FFFDFF362F3BFEA8EF8D" pageId="1" type="etymology">
|
||
<paragraph id="8B8A36C10659FFFDFF362F3BFEA8EF8D" blockId="1.[151,1437,1089,2016]" pageId="1">
|
||
<emphasis id="B941EAD30659FFFDFF362F3BFEBBEF19" bold="true" box="[199,334,1378,1404]" pageId="1">Etymology.</emphasis>
|
||
This species is named in honor of Ronald Vargas Castro, one of Costa Rica’s great naturalists, a trusted companion who has accompanied me on many
|
||
<taxonomicName id="4C354D420659FFFDFD062FDCFCA3EFFA" authorityName="Fabricius" authorityYear="1793" box="[759,854,1413,1439]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||
expeditions in
|
||
<collectingCountry id="F32276510659FFFDFBF12FDCFB8CEFFA" box="[1024,1145,1413,1439]" name="Costa Rica" pageId="1">Costa Rica</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32276510659FFFDFB5F2FDCFAFEEFFA" box="[1198,1291,1413,1439]" name="Ecuador" pageId="1">Ecuador</collectingCountry>
|
||
and who has collected numerous rare and undescribed species of
|
||
<taxonomicName id="4C354D420659FFFDFD1A2FF0FCB8EFA6" authorityName="Fabricius" authorityYear="1793" box="[747,845,1449,1475]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||
, including two of the four known specimens of this new
|
||
<taxonomicName id="4C354D420659FFFDFF3C2F94FEA3EF82" authorityName="Rambur" authorityYear="1842" box="[205,342,1485,1511]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAD30659FFFDFF3C2F94FEA3EF82" box="[205,342,1485,1511]" italics="true" pageId="1">Gynacantha</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F654A0659FFFBFF362FA8FAD9ED62" lastPageId="7" pageId="1" type="description">
|
||
<paragraph id="8B8A36C10659FFFDFF362FA8FE10EC69" blockId="1.[151,1437,1089,2016]" box="[199,485,1521,1548]" pageId="1">
|
||
<emphasis id="B941EAD30659FFFDFF362FA8FE2BEC69" bold="true" box="[199,478,1521,1548]" pageId="1">
|
||
Description of
|
||
<typeStatus id="548E88630659FFFDFE862FABFE2BEC69" box="[375,478,1522,1548]" pageId="1" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8A36C10659FFFDFF362C4CFE72EC81" blockId="1.[151,1437,1089,2016]" pageId="1">
|
||
<emphasis id="B941EAD30659FFFDFF362C4CFEF1EC4A" box="[199,260,1557,1583]" italics="true" pageId="1">Head</emphasis>
|
||
. Maximum width 8.8, eyes dark brown (dark green in life), labium light orange-brown, unmarked; underside of head cream in outer half, shading to black on inner half; labrum cream, unmarked; anteclypeus gray-green; postclypeus blue-gray with two ill-defined dull orange spots bordering the center; antefrons light brown shading to dark brown at upper margin; postfrons brown with a black spot centered on anterior side about 1/2 width of frons, no T-spot stem present; vertex shiny black; antenna dark brown; eyes dark green and black in life; vertex black; occipital triangle green.
|
||
</paragraph>
|
||
<paragraph id="8B8A36C10659FFFDFF362CB4FA87EDDE" blockId="1.[151,1437,1089,2016]" pageId="1">
|
||
<emphasis id="B941EAD30659FFFDFF362CB4FEEDED62" box="[199,280,1773,1799]" italics="true" pageId="1">Thorax</emphasis>
|
||
. Prothorax with forelobe light brown; midlobe light brown with darker brown and black markings; hindlobe black in middle 1/3, light brown laterally (Figs. 1a, 2a). Pterothorax dark brown with lime green markings; mesepisternum with a narrow collar stripe and an arching antehumeral stripe, free at each end; mesepimeron with a complete, nearly straight and uniform stripe in center; metepisternum mostly brown with a small green mark anterior to spiracle; metepimeron with a broad stripe covering most of surface, but separated from upper and lower sutures; mesinfraepisternum and metinfraepisternum light brown; metasternum brown with some pale shading.
|
||
</paragraph>
|
||
<paragraph id="8B8A36C10659FFF8FF362D9FFDDFED68" blockId="1.[151,1437,1089,2016]" lastBlockId="4.[151,1437,1743,2021]" lastPageId="4" pageId="1">
|
||
<emphasis id="B941EAD30659FFFDFF362D9FFF09EDBA" box="[199,252,1990,2015]" italics="true" pageId="1">Legs</emphasis>
|
||
. Fore femur dark brown at base, shading to black in distal half, inner surface yellow on basal half, tibia
|
||
<emphasis id="B941EAD3065AFFFEFF662D81FED4ED96" bold="true" box="[151,289,2008,2035]" pageId="2">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C354D42065AFFFEFED62D80FDFFED96" authority="Haber, 2019" authorityName="Haber" authorityYear="2019" box="[295,522,2009,2035]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="2" phylum="Arthropoda" rank="species" species="vargasi" status="sp. nov.">
|
||
<emphasis id="B941EAD3065AFFFEFED62D80FDFFED96" box="[295,522,2009,2035]" italics="true" pageId="2">Gynacantha vargasi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27257A8065AFFFEFDE02D83FD9DED91" box="[529,616,2010,2036]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
|
||
a)
|
||
<typeStatus id="548E8863065AFFFEFD7A2D80FD18ED96" box="[651,749,2009,2035]" pageId="2" type="holotype">holotype</typeStatus>
|
||
male, b)
|
||
<typeStatus id="548E8863065AFFFEFCA72D80FC45ED96" box="[854,944,2009,2035]" pageId="2" type="allotype">allotype</typeStatus>
|
||
female. and tarsus black; mid and hind femora orange-brown basally, shading to black in distal third; tibia black with dorsal side yellow in basal 4/5; tarsi black.
|
||
</paragraph>
|
||
<caption id="DF4A6649065BFFFFFF662DC1FF0AED9C" httpUri="https://zenodo.org/record/3233616/files/figure.png" pageId="3" startId="3.[151,250,1944,1969]" targetBox="[196,1391,201,1883]" targetPageId="3">
|
||
<paragraph id="8B8A36C1065BFFFFFF662DC1FF0AED9C" blockId="3.[151,1437,1944,2041]" pageId="3">
|
||
<emphasis id="B941EAD3065BFFFFFF662DC1FEE6EDD4" bold="true" box="[151,275,1944,1969]" pageId="3">FIGURE 2.</emphasis>
|
||
Images of head and thorax of
|
||
<taxonomicName id="4C354D42065BFFFFFDB82DC1FCE2EDD5" authorityName="Haber" authorityYear="2019" box="[585,791,1944,1968]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="3" phylum="Arthropoda" rank="species" species="vargasi">
|
||
<emphasis id="B941EAD3065BFFFFFDB82DC1FCE2EDD5" box="[585,791,1944,1968]" italics="true" pageId="3">Gynacantha vargasi</emphasis>
|
||
</taxonomicName>
|
||
and similarly patterned species from Costa Rica.
|
||
<taxonomicName id="4C354D42065BFFFFFAE02DC0FA8EEDD5" authorityName="Haber" authorityYear="2019" box="[1297,1403,1945,1968]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="3" phylum="Arthropoda" rank="species" species="vargasi">
|
||
<emphasis id="B941EAD3065BFFFFFAE02DC0FA8EEDD5" box="[1297,1403,1945,1968]" italics="true" pageId="3">G. vargasi</emphasis>
|
||
</taxonomicName>
|
||
: a) male, b) female;
|
||
<taxonomicName id="4C354D42065BFFFFFEB22DE4FE41EDB1" authorityName="Karsch" authorityYear="1891" box="[323,436,1980,2004]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="3" phylum="Arthropoda" rank="species" species="caudata">
|
||
<emphasis id="B941EAD3065BFFFFFEB22DE4FE41EDB1" box="[323,436,1980,2004]" italics="true" pageId="3">G. caudata</emphasis>
|
||
</taxonomicName>
|
||
: c) male, d) female;
|
||
<taxonomicName id="4C354D42065BFFFFFD772DE5FC86EDB1" authorityName="Calvert" authorityYear="1905" box="[646,883,1980,2004]" class="Insecta" family="Aeshnidae" genus="Aeshna" kingdom="Animalia" order="Odonata" pageId="3" phylum="Arthropoda" rank="species" species="williamsoniana">
|
||
<emphasis id="B941EAD3065BFFFFFD772DE5FC86EDB1" box="[646,883,1980,2004]" italics="true" pageId="3">Aeshna williamsoniana</emphasis>
|
||
</taxonomicName>
|
||
: e) male, f) female;
|
||
<taxonomicName id="4C354D42065BFFFFFBB32DE5FAC9EDB1" baseAuthorityName="Brauer" baseAuthorityYear="1865" box="[1090,1340,1980,2004]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="3" phylum="Arthropoda" rank="species" species="cornigera">
|
||
<emphasis id="B941EAD3065BFFFFFBB32DE5FAC9EDB1" box="[1090,1340,1980,2004]" italics="true" pageId="3">Rhionaeschna cornigera</emphasis>
|
||
</taxonomicName>
|
||
: g) male, h) female.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4A6649065CFFF8FF662CD1FBD6ECC4" box="[151,1059,1672,1697]" httpUri="https://zenodo.org/record/3233618/files/figure.png" pageId="4" startId="4.[151,250,1672,1697]" targetBox="[164,1409,176,1571]" targetPageId="4">
|
||
<paragraph id="8B8A36C1065CFFF8FF662CD1FBD6ECC4" blockId="4.[151,1059,1672,1697]" box="[151,1059,1672,1697]" pageId="4">
|
||
<emphasis id="B941EAD3065CFFF8FF662CD1FEE6ECC4" bold="true" box="[151,275,1672,1697]" pageId="4">FIGURE 3.</emphasis>
|
||
Wings of
|
||
<taxonomicName id="4C354D42065CFFF8FE8D2CD1FDBCECC5" authority="Haber, 2019" authorityName="Haber" authorityYear="2019" box="[380,585,1672,1696]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="4" phylum="Arthropoda" rank="species" species="vargasi" status="sp. nov.">
|
||
<emphasis id="B941EAD3065CFFF8FE8D2CD1FDBCECC5" box="[380,585,1672,1696]" italics="true" pageId="4">Gynacantha vargasi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27257A8065CFFF8FDBE2CD0FD6BECC5" box="[591,670,1673,1696]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
: a) holotype male, b) allotype female.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8A36C1065CFFF8FF362D4EFC65ED80" blockId="4.[151,1437,1743,2021]" pageId="4">
|
||
<emphasis id="B941EAD3065CFFF8FF362D4EFEFEED55" box="[199,267,1815,1840]" italics="true" pageId="4">Wings</emphasis>
|
||
. (data for left wing precede those for right wing) Transparent with slight brown tint, veins brown, FW Ax (left, right) 27, 26 with 1 and 9 thickened; Px 23, 23; HW Ax 21, 22 with 1 and 10 thickened; Px 23, 21; poststigmatal cross veins in FW 6, 5; in HW 7, 4; FW triangle with (left, right) 6, 6 cells; subtriangle 3, 2 cells; supratriangle 6, 7 crossveins; median space free; cubito-anal crossveins 8, 8. HW triangle with 6, 6 cells, subtriangle with 2, 2 cells, supratriangle with 5, 5 crossveins; median space free; cubito-anal crossveins 6, 6; anal loop with 13, 11 cells; anal loop separated from posterior margin by 1, 2 rows of cells (
|
||
<figureCitation id="130E2A44065CFFF8FCC02D92FC76ED80" box="[817,899,1995,2021]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1697]" captionTargetBox="[164,1409,176,1571]" captionTargetId="figure@4.[164,1423,166,1593]" captionTargetPageId="4" captionText="FIGURE 3. Wings of Gynacantha vargasi sp. nov.: a) holotype male, b) allotype female." httpUri="https://zenodo.org/record/3233618/files/figure.png" pageId="4">Fig. 3a</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8A36C1065DFFFAFF362AC0FDC9EDDB" blockId="5.[151,1436,153,359]" lastBlockId="6.[151,1437,1920,2019]" lastPageId="6" pageId="5">
|
||
<emphasis id="B941EAD3065DFFF9FF362AC0FEC4EAD6" box="[199,305,153,179]" italics="true" pageId="5">Abdomen</emphasis>
|
||
. S1 red-brown, shading to dark brown on dorsum; S2 with dorsum and sides black, a narrow blue stripe on midline; green on side at base, green bordering transverse carina, a blue patch above auricle, a wide blue stripe bordering posterior margin, auricle black with a blue patch on dorsum, ventrally bright yellow-green with black edges; 5–6 stout recurved teeth on posterior edge (
|
||
<figureCitation id="130E2A44065DFFF9FCD82B5CFC8BEB45" box="[809,894,261,288]" captionStart="FIGURE 6" captionStartId="8.[151,250,1918,1943]" captionTargetBox="[187,1421,287,1872]" captionTargetId="figure@8.[187,1421,302,1872]" captionTargetPageId="8" captionText="FIGURE 6. Abdominal segments 1−2 (ventral view) of male Gynacantha species showing secondary genitalia and auricles: a) G. vargasi, male holotype; b) G. caudata, Prov. Guanacaste, Costa Rica; c) G. tenuis, Dept. Loreto, Peru; and d) G. tibiata, Prov. Limón, Costa Rica." httpUri="https://zenodo.org/record/3233624/files/figure.png" pageId="5">Fig. 6a</figureCitation>
|
||
); 5, 8 minute black denticles bordering ventral carina, but not in a line (
|
||
<figureCitation id="130E2A44065DFFF9FE5D2B70FE0BEB21" box="[428,510,297,324]" captionStart="FIGURE 6" captionStartId="8.[151,250,1918,1943]" captionTargetBox="[187,1421,287,1872]" captionTargetId="figure@8.[187,1421,302,1872]" captionTargetPageId="8" captionText="FIGURE 6. Abdominal segments 1−2 (ventral view) of male Gynacantha species showing secondary genitalia and auricles: a) G. vargasi, male holotype; b) G. caudata, Prov. Guanacaste, Costa Rica; c) G. tenuis, Dept. Loreto, Peru; and d) G. tibiata, Prov. Limón, Costa Rica." httpUri="https://zenodo.org/record/3233624/files/figure.png" pageId="5">Fig. 6a</figureCitation>
|
||
). S3 greatly narrowed both laterally and dorsally (Figs. 1a, 5a, 6a); basal half redbrown, shading to black on posterior half; lower side with a blue patch at base, an inconspicuous green line border ing transverse carina on posterior side, two lateral blue spots at apex; S4−10 black with S4−8 having inconspicuous blue spots and streaks on lower sides.
|
||
</paragraph>
|
||
<caption id="DF4A6649065DFFF9FF662D4BFBD5EDDE" httpUri="https://zenodo.org/record/3233620/files/figure.png" pageId="5" startId="5.[151,250,1810,1835]" targetBox="[176,1406,426,1668]" targetPageId="5">
|
||
<paragraph id="8B8A36C1065DFFF9FF662D4BFBD5EDDE" blockId="5.[151,1436,1810,1979]" pageId="5">
|
||
<emphasis id="B941EAD3065DFFF9FF662D4BFEE7ED4E" bold="true" box="[151,274,1810,1835]" pageId="5">FIGURE 4.</emphasis>
|
||
Abdominal appendages of
|
||
<taxonomicName id="4C354D42065DFFF9FDD72D4AFD57ED4E" authorityName="Rambur" authorityYear="1842" box="[550,674,1811,1835]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAD3065DFFF9FDD72D4AFD57ED4E" box="[550,674,1811,1835]" italics="true" pageId="5">Gynacantha</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<taxonomicName id="4C354D42065DFFF9FD0D2D4AFC3DED4F" authorityName="Haber" authorityYear="2019" box="[764,968,1811,1835]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" phylum="Arthropoda" rank="species" species="vargasi">
|
||
<emphasis id="B941EAD3065DFFF9FD0D2D4AFC3DED4F" box="[764,968,1811,1835]" italics="true" pageId="5">Gynacantha vargasi</emphasis>
|
||
</taxonomicName>
|
||
male holotype: a) dorsal view; b) mediodorsal view; c) external lateral view; d) cross section of right cercus at center of dorsal lobe, posterior view.
|
||
<taxonomicName id="4C354D42065DFFF9FB542D6EFA90ED2B" baseAuthorityName="Martin" baseAuthorityYear="1909" box="[1189,1381,1847,1871]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" phylum="Arthropoda" rank="species" species="tenuis">
|
||
<emphasis id="B941EAD3065DFFF9FB542D6EFA90ED2B" box="[1189,1381,1847,1871]" italics="true" pageId="5">Gynacantha tenuis</emphasis>
|
||
</taxonomicName>
|
||
male (Dept. Loreto, Peru): e) cross section of right cercus at middle of dorsal lobe.
|
||
<taxonomicName id="4C354D42065DFFF9FC4F2D02FB61ED16" authorityName="Karsch" authorityYear="1891" box="[958,1172,1883,1907]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" phylum="Arthropoda" rank="species" species="caudata">
|
||
<emphasis id="B941EAD3065DFFF9FC4F2D02FB61ED16" box="[958,1172,1883,1907]" italics="true" pageId="5">Gynacantha caudata</emphasis>
|
||
</taxonomicName>
|
||
(Prov. Guanacaste, Costa Rica): f) cross section of right cercus at center of dorsal lobe, posterior view.
|
||
<taxonomicName id="4C354D42065DFFF9FC662D26FB96EDF3" authorityName="Haber" authorityYear="2019" box="[919,1123,1919,1943]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" phylum="Arthropoda" rank="species" species="vargasi">
|
||
<emphasis id="B941EAD3065DFFF9FC662D26FB96EDF3" box="[919,1123,1919,1943]" italics="true" pageId="5">Gynacantha vargasi</emphasis>
|
||
</taxonomicName>
|
||
, female allotype: g) abdominal segments 8−10, lateral view; h) forked process on abdominal segment 10, posterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4A6649065EFFFAFF662D4FFDB0ED36" httpUri="https://zenodo.org/record/3233622/files/figure.png" pageId="6" startId="6.[151,250,1814,1839]" targetBox="[164,1423,168,1757]" targetPageId="6">
|
||
<paragraph id="8B8A36C1065EFFFAFF662D4FFDB0ED36" blockId="6.[151,1436,1814,1875]" pageId="6">
|
||
<emphasis id="B941EAD3065EFFFAFF662D4FFEE0ED4B" bold="true" box="[151,277,1814,1839]" pageId="6">FIGURE 5.</emphasis>
|
||
<emphasis id="B941EAD3065EFFFAFEEC2D4FFE06ED4B" box="[285,499,1814,1838]" italics="true" pageId="6">
|
||
<taxonomicName id="4C354D42065EFFFAFEEC2D4FFE1AED4B" authorityName="Haber" authorityYear="2019" box="[285,495,1814,1838]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="6" phylum="Arthropoda" rank="species" species="vargasi">Gynacantha vargasi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
abdominal segments 1−3: a) male holotype, lateral view; b) female paratype (Las Brisas), lateral view; c) same female, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8A36C1065EFFFBFF362D90FBD1EABD" blockId="6.[151,1437,1920,2019]" lastBlockId="7.[151,1437,153,2015]" lastPageId="7" pageId="6">
|
||
<emphasis id="B941EAD3065EFFFAFF362D90FE53ED86" box="[199,422,1993,2019]" italics="true" pageId="6">Caudal appendages</emphasis>
|
||
. Left cercus detached (Fig. 1a). Appendages black; cercus 5.5; epiproct 2.6, straight in api- cal 2/3, apex minutely emarginate in dorsal view. Cercus with medial margin slightly decumbent medially, and with an asymmetric distal dorsal lobe angled medially to plane of cercus (
|
||
<figureCitation id="130E2A44065FFFFBFC682AE4FC28EAB2" box="[921,989,189,215]" captionStart="FIGURE 4" captionStartId="5.[151,250,1810,1835]" captionTargetBox="[176,1406,426,1668]" captionTargetId="figure@5.[151,1436,386,1703]" captionTargetPageId="5" captionText="FIGURE 4. Abdominal appendages of Gynacantha species. Gynacantha vargasi male holotype: a) dorsal view; b) mediodorsal view; c) external lateral view; d) cross section of right cercus at center of dorsal lobe, posterior view. Gynacantha tenuis male (Dept. Loreto, Peru): e) cross section of right cercus at middle of dorsal lobe. Gynacantha caudata (Prov. Guanacaste, Costa Rica): f) cross section of right cercus at center of dorsal lobe, posterior view. Gynacantha vargasi, female allotype: g) abdominal segments 8−10, lateral view; h) forked process on abdominal segment 10, posterior view." httpUri="https://zenodo.org/record/3233620/files/figure.png" pageId="7">Fig. 4</figureCitation>
|
||
, a−d).
|
||
</paragraph>
|
||
<paragraph id="8B8A36C1065FFFFBFF362AB8FC6DEB7A" blockId="7.[151,1437,153,2015]" pageId="7">Measurements (mm). Total length without cerci 61; abdomen 47, cerci 5.5; FW 44; HW 43 with width at nodus 12; pterostigma length on costal vein 3.5, 3.4 in FW; 3.2, 3.1 in HW.</paragraph>
|
||
<paragraph id="8B8A36C1065FFFFBFF362B71FE28EB26" blockId="7.[151,1437,153,2015]" box="[199,477,296,323]" pageId="7">
|
||
<emphasis id="B941EAD3065FFFFBFF362B71FE28EB26" bold="true" box="[199,477,296,323]" pageId="7">
|
||
Description of
|
||
<typeStatus id="548E8863065FFFFBFE862B70FE2DEB26" box="[375,472,297,323]" pageId="7" type="allotype">allotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8A36C1065FFFFBFF362B14FCADEBB6" blockId="7.[151,1437,153,2015]" pageId="7">
|
||
<emphasis id="B941EAD3065FFFFBFF362B14FEF1EB02" box="[199,260,333,359]" italics="true" pageId="7">Head</emphasis>
|
||
. Maximum width 9.3 mm. Eyes dark brown (not seen alive); labium dull orange-brown shading to cream posteriorly; underside of head cream-brown bordering eye margins, shading to black in center; labrum orangebrown, unmarked; ante- and postclypeus light brown; antefrons orange-brown, postfrons brown with anterior half black, no T-stem; vertex black; occipital triangle yellow-green.
|
||
</paragraph>
|
||
<paragraph id="8B8A36C1065FFFFBFF362B84FD57E8AA" blockId="7.[151,1437,153,2015]" pageId="7">
|
||
<emphasis id="B941EAD3065FFFFBFF362B84FEEDEB92" box="[199,280,477,503]" italics="true" pageId="7">Thorax</emphasis>
|
||
. Prothorax forelobe with light brown margins, darker brown in middle; midlobe and hindlobe light brown without darker markings. Pterothorax ground color dark brown dorsally shading to light orange-brown ventrally; venter unmarked; pale markings yellow-green, duller and wider than in male; mesepisternum with dorsal carina all brown; with a slender collar stripe; antehumeral stripe arched upward at middle; mesepimeron with a broad central stripe, narrowed posteriorly; metepisternum all brown; metepimeron with a broad complete stripe covering most of sclerite, free at each end and not reaching either suture; mesinfraepisternum brown with weak green shading; metinfraepisternum brown and unmarked.
|
||
</paragraph>
|
||
<paragraph id="8B8A36C1065FFFFBFF362883FD4EE95E" blockId="7.[151,1437,153,2015]" pageId="7">
|
||
<emphasis id="B941EAD3065FFFFBFF362883FF09E896" box="[199,252,730,755]" italics="true" pageId="7">Legs</emphasis>
|
||
. Fore femur orange-brown in basal half, shading to black in apical half, inner surface cream on basal 2/3, tibia and tarsus black; mid and hind femora orange-brown shading to black in apical 1/4 to 1/3, tibia dark brown with dorsal side tan, apical 1/4 black; tarsi black.
|
||
</paragraph>
|
||
<paragraph id="8B8A36C1065FFFFBFF36291FFF36EE39" blockId="7.[151,1437,153,2015]" pageId="7">
|
||
<emphasis id="B941EAD3065FFFFBFF36291FFEFEE93A" box="[199,267,838,863]" italics="true" pageId="7">Wings</emphasis>
|
||
. (data for left wing precede those for right wing) Distal half of right FW missing (Fig. 1b). Distinctly tinted brown with veins brown to dark brown; pterostigma light brown; Ax of FW 31, with 1 and 10 thickened; Px 26; tip of right wing missing. In HW Ax 23, 21 with 1 and 9 thickened; Px 26, 27; HW triangle with 6 plus 2 tiny cells, 6 cells; subtriangle with 2, 2 cells, supratriangle with 7, 7 crossveins; anal loop with 12, 13 cells; anal loop separated from posterior margin by 1, 2 cell rows. A subcostal crossvein present anterior to first Ax in left FW and left HW. FW triangle with 8, 8 cells; subtriangle with 2, 2 cells; supratriangle with 9, 9 crossveins, cubito-anal crossveins
|
||
<quantity id="4CCD9B24065FFFFBFF282E44FED4EE52" box="[217,289,1053,1079]" metricMagnitude="-1" metricUnit="m" metricValue="1.9558" pageId="7" unit="in" value="7.7">7, 7 in</quantity>
|
||
FW
|
||
<quantity id="4CCD9B24065FFFFBFEA92E44FE6AEE52" box="[344,415,1053,1079]" metricMagnitude="-1" metricUnit="m" metricValue="1.6764" pageId="7" unit="in" value="6.6">6, 6 in</quantity>
|
||
HW; crossveins between arculus and branching of R
|
||
<quantity id="4CCD9B24065FFFFBFC0B2E44FBA8EE52" box="[1018,1117,1052,1079]" metricMagnitude="-1" metricUnit="m" metricValue="4.102099999999999" pageId="7" unit="in" value="16.15">16, 15 in</quantity>
|
||
FW
|
||
<quantity id="4CCD9B24065FFFFBFB652E44FB03EE52" box="[1172,1270,1053,1079]" metricMagnitude="-1" metricUnit="m" metricValue="2.82194" pageId="7" unit="in" value="11.11">11, 11 in</quantity>
|
||
HW (Figs. 1b, 3b).
|
||
</paragraph>
|
||
<paragraph id="8B8A36C1065FFFFBFF362E3CFCD7EEA2" blockId="7.[151,1437,153,2015]" pageId="7">
|
||
<emphasis id="B941EAD3065FFFFBFF362E3CFEC4EE1A" box="[199,305,1125,1151]" italics="true" pageId="7">Abdomen</emphasis>
|
||
. Cerci broken near base and missing. S1 orange-brown; S2 black above lateral carina, greenish below, extensive blue shading on posterior half above lateral carina; S3 brown with a blue basal patch on side; S4−10 dark brown shading to black posteriorly (
|
||
<figureCitation id="130E2A44065FFFFBFDDC2EF4FD69EEA2" box="[557,668,1197,1223]" captionStart="FIGURE 4" captionStartId="5.[151,250,1810,1835]" captionTargetBox="[176,1406,426,1668]" captionTargetId="figure@5.[151,1436,386,1703]" captionTargetPageId="5" captionText="FIGURE 4. Abdominal appendages of Gynacantha species. Gynacantha vargasi male holotype: a) dorsal view; b) mediodorsal view; c) external lateral view; d) cross section of right cercus at center of dorsal lobe, posterior view. Gynacantha tenuis male (Dept. Loreto, Peru): e) cross section of right cercus at middle of dorsal lobe. Gynacantha caudata (Prov. Guanacaste, Costa Rica): f) cross section of right cercus at center of dorsal lobe, posterior view. Gynacantha vargasi, female allotype: g) abdominal segments 8−10, lateral view; h) forked process on abdominal segment 10, posterior view." httpUri="https://zenodo.org/record/3233620/files/figure.png" pageId="7">
|
||
Fig.
|
||
<quantity id="4CCD9B24065FFFFBFD902EF4FD75EEA2" box="[609,640,1197,1223]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="7" unit="g" value="4.0">4g</quantity>
|
||
, h
|
||
</figureCitation>
|
||
;
|
||
<figureCitation id="130E2A44065FFFFBFD592EF4FCE1EEA2" box="[680,788,1196,1223]" captionStart="FIGURE 5" captionStartId="6.[151,250,1814,1839]" captionTargetBox="[164,1423,168,1757]" captionTargetId="figure@6.[164,1423,168,1757]" captionTargetPageId="6" captionText="FIGURE 5. Gynacantha vargasi, abdominal segments 1−3: a) male holotype, lateral view; b) female paratype (Las Brisas), lateral view; c) same female, ventral view." httpUri="https://zenodo.org/record/3233622/files/figure.png" pageId="7">Fig. 5b, c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8A36C1065FFFFBFF362E88FC44EF6A" blockId="7.[151,1437,153,2015]" pageId="7">Measurements (mm). Total length without cerci 64 (cerci broken); abdomen 50; FW 48; HW 49, 49, width at nodus 13. Pterostigma length on costal vein 3.8 in FW; 3.5, 3.6 in HW.</paragraph>
|
||
<paragraph id="8B8A36C1065FFFFBFF362F41FA90EC6E" blockId="7.[151,1437,153,2015]" pageId="7">
|
||
<emphasis id="B941EAD3065FFFFBFF362F41FEBFEF56" bold="true" box="[199,330,1304,1331]" pageId="7">Variations.</emphasis>
|
||
Both males are mature. The Río Peje male, with wings slightly darker, appears to be older than the
|
||
<typeStatus id="548E8863065FFFFBFF662F64FF0CEF32" box="[151,249,1341,1367]" pageId="7" type="holotype">holotype</typeStatus>
|
||
male and comparable in age to the two females. It is slightly smaller, with head width, abdomen, and FW 8.7, 46, and 43, respectively, and the anal loop has only 9 and 10 cells. The color pattern elements are the same in both. The labrum and face are darker orange and shading more to blue in the La Virgen male. The cercus apex of the
|
||
<typeStatus id="548E8863065FFFFBFF662FF0FF0CEFA6" box="[151,249,1449,1475]" pageId="7" type="holotype">holotype</typeStatus>
|
||
is twisted about 60
|
||
<superScript id="7C409B89065FFFFBFE212FF1FE2DEFD3" attach="left" box="[464,472,1448,1462]" fontSize="6" pageId="7">o</superScript>
|
||
clockwise; those of the Río Peje male are turned about 20
|
||
<superScript id="7C409B89065FFFFBFBAD2FF1FB91EFD3" attach="none" box="[1116,1124,1448,1462]" fontSize="6" pageId="7">o</superScript>
|
||
. Because the twisted cercus is evident in the fresh
|
||
<typeStatus id="548E8863065FFFFBFE692F94FE0FEF82" box="[408,506,1485,1511]" pageId="7" type="holotype">holotype</typeStatus>
|
||
male (Fig. 1a), scanned before preservation and shows the same orientation after drying, this state appears to be natural. The Río Peje male was preserved in acetone and dried before analysis.
|
||
</paragraph>
|
||
<paragraph id="8B8A36C1065FFFFBFF362C4CFCDEEC12" blockId="7.[151,1437,153,2015]" pageId="7">Both females are mature with the wings shaded brown. The color patterns are without significant variation but duller overall than in the males. The Las Brisas female is smaller, with head width, abdomen, and FW 9.1, 51, and 46, respectively and the anal loop has only 11 and 10 cells.</paragraph>
|
||
<paragraph id="8B8A36C1065FFFFBFF362CD8FAD9ED62" blockId="7.[151,1437,153,2015]" pageId="7">The separation of the anal loop from the posterior wing margin is variable – the normal state appears to be two rows of cells between the loop and the margin as in the Las Brisas female. The other female and both males have at least one elongated cell bordered by two cell rows joining the loop and margin in each wing. A crossvein basal to the first thickened antenodal vein appears only in one FW and one HW of the Río Peñas Blancas female.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F654A065FFFF6FF362D49FB81EAB2" lastPageId="10" pageId="7" type="diagnosis">
|
||
<paragraph id="8B8A36C1065FFFFBFF362D49FE9EEDDE" blockId="7.[151,1437,153,2015]" pageId="7">
|
||
<emphasis id="B941EAD3065FFFFBFF362D49FEB7ED4E" bold="true" box="[199,322,1808,1835]" pageId="7">Diagnosis.</emphasis>
|
||
Males of
|
||
<taxonomicName id="4C354D42065FFFFBFE442D48FD6CED4E" authorityName="Haber" authorityYear="2019" box="[437,665,1809,1835]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" phylum="Arthropoda" rank="species" species="vargasi">
|
||
<emphasis id="B941EAD3065FFFFBFE442D48FD6CED4E" box="[437,665,1809,1835]" italics="true" pageId="7">Gynacantha vargasi</emphasis>
|
||
</taxonomicName>
|
||
can be separated from all other species by the bright green thoracic stripes, mid and hind legs with femora orange-brown tipped with black, and dorsal side of the middle and hind tibia yellow; the length of the epiproct at about half the length of cercus, and the unique angled distal lobe of the cercus. Females are distinct in the T-mark on the frons lacking a stem, the color patterns of the thorax, and the yellow to tan mid and hind tibia.
|
||
</paragraph>
|
||
<paragraph id="8B8A36C1065FFFF6FF362D9CFB81EAB2" blockId="7.[151,1437,153,2015]" lastBlockId="10.[151,1437,153,755]" lastPageId="10" pageId="7">
|
||
Within
|
||
<collectingCountry id="F3227651065FFFFBFEEA2D9CFE6CEDBA" box="[283,409,1989,2015]" name="Costa Rica" pageId="7">Costa Rica</collectingCountry>
|
||
, the lime-green lateral thoracic stripes are similar in size and shape to those of
|
||
<taxonomicName id="4C354D42065FFFF4FAE22D9CFE7BEAD6" authority="Karsch, 1891" authorityName="Karsch" authorityYear="1891" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" lastPageId="8" order="Odonata" pageId="7" phylum="Arthropoda" rank="species" species="caudata">
|
||
<emphasis id="B941EAD3065FFFF4FAE22D9CFF04EAD6" italics="true" lastPageId="8" pageId="7">Gynacantha caudata</emphasis>
|
||
Karsch, 1891
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C354D420650FFF4FE692AC0FCC8EAD6" authority="Calvert, 1905" authorityName="Calvert" authorityYear="1905" box="[408,829,152,179]" class="Insecta" family="Aeshnidae" genus="Aeshna" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="williamsoniana">
|
||
<emphasis id="B941EAD30650FFF4FE692AC0FD68EAD6" box="[408,669,153,179]" italics="true" pageId="8">Aeshna williamsoniana</emphasis>
|
||
<bibRefCitation id="EFA44B300650FFF4FD522AC0FCC8EAD6" author="Calvert, P. P." box="[675,829,152,179]" pageId="8" pagination="145 - 212" refId="ref6408" refString="Calvert, P. P. (1905) Insecta Neuroptera. Odonata. Biologia Centrali-Americana, 1905, 145 - 212." type="journal article" year="1905">Calvert, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C354D420650FFF4FCB62AC0FAF0EAD6" authority="(Brauer, 1865)" authorityName="Brauer" authorityYear="1865" baseAuthorityName="Brauer" baseAuthorityYear="1865" box="[839,1285,152,179]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="cornigera">
|
||
<emphasis id="B941EAD30650FFF4FCB62AC0FBACEAD6" box="[839,1113,153,179]" italics="true" pageId="8">Rhionaeschna cornigera</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA44B300650FFF4FB962AC0FB09EAD6" author="Brauer, F." box="[1127,1276,152,179]" pageId="8" pagination="903 - 908" refId="ref6373" refString="Brauer, F. (1865) Vierter Bericht uber die auf der Weltfahrt der kais. Fregatte Novara gesammelten Neuropteren. Verhandlungen der zoologisch-botanischen Gesellschaft in Wien, 15, 903 - 908." type="journal article" year="1865">Brauer, 1865</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C354D420650FFF4FACC2AC3FEB0EAB2" authority="(Calvert, 1947)" authorityName="Calvert" authorityYear="1947" baseAuthorityName="Calvert" baseAuthorityYear="1947" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="psilus">
|
||
<emphasis id="B941EAD30650FFF4FACC2AC3FA69EAD6" box="[1341,1436,153,179]" italics="true" pageId="8">R. psilus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA44B300650FFF4FF6E2AE4FEC8EAB2" author="Calvert, P. P." box="[159,317,189,215]" pageId="8" pagination="1 - 11" refId="ref6431" refString="Calvert, P. P. (1947) Aeshna psilus, a new species of the group of Ae. cornigera Brauer (Odonata: Aeshnidae). Proceedings of the Academy of Natural Sciences of Philadelphia, 194, 1 - 11." type="journal article" year="1947">Calvert, 1947</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, all of which may overlap in distribution and habitat with
|
||
<taxonomicName id="4C354D420650FFF4FC222AE4FBBDEAB2" authorityName="Haber" authorityYear="2019" box="[979,1096,189,215]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="vargasi">
|
||
<emphasis id="B941EAD30650FFF4FC222AE4FBBDEAB2" box="[979,1096,189,215]" italics="true" pageId="8">G. vargasi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130E2A440650FFF4FBA92AE4FB68EAB2" box="[1112,1181,189,215]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1969]" captionTargetBox="[196,1391,201,1883]" captionTargetId="figure@3.[196,1391,201,1885]" captionTargetPageId="3" captionText="FIGURE 2. Images of head and thorax of Gynacantha vargasi and similarly patterned species from Costa Rica. G. vargasi: a) male, b) female; G. caudata: c) male, d) female; Aeshna williamsoniana: e) male, f) female; Rhionaeschna cornigera: g) male, h) female." httpUri="https://zenodo.org/record/3233616/files/figure.png" pageId="8">Fig. 2</figureCitation>
|
||
).
|
||
<taxonomicName id="4C354D420650FFF4FB432AE4FA69EAB2" authorityName="Karsch" authorityYear="1891" box="[1202,1436,189,215]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="caudata">
|
||
<emphasis id="B941EAD30650FFF4FB432AE4FA69EAB2" box="[1202,1436,189,215]" italics="true" pageId="8">Gynacantha caudata</emphasis>
|
||
</taxonomicName>
|
||
is distinct in having entirely pale mid and hindlegs. The top of the frons of
|
||
<taxonomicName id="4C354D420650FFF4FC2E2AB8FBA1EA9E" authorityName="Haber" authorityYear="2019" box="[991,1108,225,251]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="vargasi">
|
||
<emphasis id="B941EAD30650FFF4FC2E2AB8FBA1EA9E" box="[991,1108,225,251]" italics="true" pageId="8">G. vargasi</emphasis>
|
||
</taxonomicName>
|
||
has a black T-mark lacking a stem, while the frons of
|
||
<taxonomicName id="4C354D420650FFF4FE562B5CFD86EB7A" authorityName="Calvert" authorityYear="1905" box="[423,627,261,287]" class="Insecta" family="Aeshnidae" genus="Aeshna" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="williamsoniana">
|
||
<emphasis id="B941EAD30650FFF4FE562B5CFD86EB7A" box="[423,627,261,287]" italics="true" pageId="8">A. williamsoniana</emphasis>
|
||
</taxonomicName>
|
||
is all black, and
|
||
<taxonomicName id="4C354D420650FFF4FCC02B5FFC4BEB7A" baseAuthorityName="Brauer" baseAuthorityYear="1865" box="[817,958,262,287]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="cornigera">
|
||
<emphasis id="B941EAD30650FFF4FCC02B5FFC4BEB7A" box="[817,958,262,287]" italics="true" pageId="8">R. cornigera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C354D420650FFF4FC042B5FFBA0EB7A" baseAuthorityName="Calvert" baseAuthorityYear="1947" box="[1013,1109,261,287]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="psilus">
|
||
<emphasis id="B941EAD30650FFF4FC042B5FFBA0EB7A" box="[1013,1109,261,287]" italics="true" pageId="8">R. psilus</emphasis>
|
||
</taxonomicName>
|
||
have a T-mark with a broad stem.
|
||
<emphasis id="B941EAD30652FFF6FF2B2AC0FDB0EAD6" box="[218,581,153,179]" italics="true" pageId="10">
|
||
<taxonomicName id="4C354D420652FFF6FF2B2AC0FE4AEAD6" authorityName="Haber" authorityYear="2019" box="[218,447,153,179]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="vargasi">Gynacantha vargasi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C354D420652FFF6FE382AC0FDB0EAD6" authorityName="Karsch" authorityYear="1891" box="[457,581,153,179]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="caudata">G. caudata</taxonomicName>
|
||
</emphasis>
|
||
, and the South American
|
||
<taxonomicName id="4C354D420652FFF6FC972AC0FB80EAD6" authority="(Martin, 1909)" authorityName="Martin" authorityYear="1909" baseAuthorityName="Martin" baseAuthorityYear="1909" box="[870,1141,153,179]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="tenuis">
|
||
<emphasis id="B941EAD30652FFF6FC972AC0FC3FEAD6" box="[870,970,153,179]" italics="true" pageId="10">G. tenuis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA44B300652FFF6FC292AC0FB98EAD6" author="Martin, R." box="[984,1133,153,179]" pageId="10" pagination="157 - 223" refId="ref6686" refString="Martin, R. (1909) Aeschnines. Collections Zoologiques du Baron Edmund de Selys-Longchamps, Catalogue Systematique et Descriptif, 20, 157 - 223. [Gynacantha auricularis, G. tenuis]" type="journal article" year="1909">Martin, 1909</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C354D420652FFF6FB5A2AC0FF2CEAB2" authority="(Martin, 1909)" authorityName="Martin" authorityYear="1909" baseAuthorityName="Martin" baseAuthorityYear="1909" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="francesca">
|
||
<emphasis id="B941EAD30652FFF6FB5A2AC0FACCEAD6" box="[1195,1337,153,179]" italics="true" pageId="10">G. francesca</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA44B300652FFF6FAB62AC0FF25EAB2" author="Martin, R." pageId="10" pagination="157 - 223" refId="ref6686" refString="Martin, R. (1909) Aeschnines. Collections Zoologiques du Baron Edmund de Selys-Longchamps, Catalogue Systematique et Descriptif, 20, 157 - 223. [Gynacantha auricularis, G. tenuis]" type="journal article" year="1909">Martin, 1909</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
are the only species of New World
|
||
<taxonomicName id="4C354D420652FFF6FD982AE4FD07EAB2" authorityName="Rambur" authorityYear="1842" box="[617,754,189,215]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAD30652FFF6FD982AE4FD07EAB2" box="[617,754,189,215]" italics="true" pageId="10">Gynacantha</emphasis>
|
||
</taxonomicName>
|
||
with bright-green thoracic stripes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4A66490650FFF4FF662D27FE6DEDBA" httpUri="https://zenodo.org/record/3233624/files/figure.png" pageId="8" startId="8.[151,250,1918,1943]" targetBox="[187,1421,287,1872]" targetPageId="8">
|
||
<paragraph id="8B8A36C10650FFF4FF662D27FE6DEDBA" blockId="8.[151,1437,1918,2015]" pageId="8">
|
||
<emphasis id="B941EAD30650FFF4FF662D27FEE1EDF2" bold="true" box="[151,276,1918,1943]" pageId="8">FIGURE 6.</emphasis>
|
||
Abdominal segments 1−2 (ventral view) of male
|
||
<taxonomicName id="4C354D420650FFF4FCEB2D26FC63EDF2" authorityName="Rambur" authorityYear="1842" box="[794,918,1919,1943]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAD30650FFF4FCEB2D26FC63EDF2" box="[794,918,1919,1943]" italics="true" pageId="8">Gynacantha</emphasis>
|
||
</taxonomicName>
|
||
species showing secondary genitalia and auricles: a)
|
||
<taxonomicName id="4C354D420650FFF4FF432DFAFEE9EDDF" authorityName="Haber" authorityYear="2019" box="[178,284,1955,1978]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="vargasi">
|
||
<emphasis id="B941EAD30650FFF4FF432DFAFEE9EDDF" box="[178,284,1955,1978]" italics="true" pageId="8">G. vargasi</emphasis>
|
||
</taxonomicName>
|
||
, male holotype; b)
|
||
<taxonomicName id="4C354D420650FFF4FE152DFAFD6DEDDF" authority=", Prov." authorityName="Prov." box="[484,664,1955,1979]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="caudata">
|
||
<emphasis id="B941EAD30650FFF4FE152DFAFDA3EDDE" box="[484,598,1955,1979]" italics="true" pageId="8">G. caudata</emphasis>
|
||
, Prov.
|
||
</taxonomicName>
|
||
Guanacaste, Costa Rica; c)
|
||
<taxonomicName id="4C354D420650FFF4FC492DFAFBAFEDDF" authority=", Dept." authorityName="Dept." box="[952,1114,1955,1978]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="tenuis">
|
||
<emphasis id="B941EAD30650FFF4FC492DFAFBE1EDDF" box="[952,1044,1955,1978]" italics="true" pageId="8">G. tenuis</emphasis>
|
||
, Dept.
|
||
</taxonomicName>
|
||
Loreto, Peru; and d)
|
||
<taxonomicName id="4C354D420650FFF4FAC52DFAFF38EDBB" authority=", Prov." authorityName="Prov." class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="8" phylum="Arthropoda" rank="species" species="tibiata">
|
||
<emphasis id="B941EAD30650FFF4FAC52DFAFA63EDDE" box="[1332,1430,1955,1979]" italics="true" pageId="8">G. tibiata</emphasis>
|
||
, Prov.
|
||
</taxonomicName>
|
||
Limón, Costa Rica.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4A66490651FFF5FF662DE8FA99EDAF" box="[151,1388,1969,1994]" httpUri="https://zenodo.org/record/3233626/files/figure.png" pageId="9" startId="9.[151,250,1969,1994]" targetBox="[185,1331,171,1883]" targetPageId="9">
|
||
<paragraph id="8B8A36C10651FFF5FF662DE8FA99EDAF" blockId="9.[151,1388,1969,1994]" box="[151,1388,1969,1994]" pageId="9">
|
||
<emphasis id="B941EAD30651FFF5FF662DE8FEE6EDAF" bold="true" box="[151,275,1969,1994]" pageId="9">FIGURE 7.</emphasis>
|
||
Abdominal appendages of
|
||
<taxonomicName id="4C354D420651FFF5FDDB2DE8FD53EDAC" authorityName="Rambur" authorityYear="1842" box="[554,678,1969,1993]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EAD30651FFF5FDDB2DE8FD53EDAC" box="[554,678,1969,1993]" italics="true" pageId="9">Gynacantha</emphasis>
|
||
</taxonomicName>
|
||
species, arranged from top down by increasing length of the epiproct.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32F654A0652FFF6FF362AB8FD15E896" pageId="10" type="reference_group">
|
||
<paragraph id="8B8A36C10652FFF6FF362AB8FAD2EB92" blockId="10.[151,1437,153,755]" pageId="10">
|
||
<taxonomicName id="4C354D420652FFF6FF362AB8FE5FEA9E" ID-CoL="9X2VK" authorityName="Haber" authorityYear="2019" box="[199,426,225,251]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="vargasi">
|
||
<emphasis id="B941EAD30652FFF6FF362AB8FE5FEA9E" box="[199,426,225,251]" italics="true" pageId="10">Gynacantha vargasi</emphasis>
|
||
</taxonomicName>
|
||
belongs to the group of species lacking crossveins basal to the first thickened Ax vein and Pt less than
|
||
<quantity id="4CCD9B240652FFF6FEEF2B5CFE95EB7A" box="[286,352,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="10" unit="mm" value="4.0">4 mm</quantity>
|
||
long.
|
||
<taxonomicName id="4C354D420652FFF6FE562B5CFBF2EB7A" ID-CoL="3HVTP" authority="Williamson & Williamson, 1930" authorityName="Williamson & Williamson" authorityYear="1930" box="[423,1031,261,287]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="helenga">
|
||
<emphasis id="B941EAD30652FFF6FE562B5CFD65EB7A" box="[423,656,261,287]" italics="true" pageId="10">Gynacantha helenga</emphasis>
|
||
<bibRefCitation id="EFA44B300652FFF6FD662B5CFBF2EB7A" author="Williamson, E. B. & Williamson, J. H." box="[663,1031,261,287]" pageId="10" pagination="1 - 5" refId="ref7249" refString="Williamson, E. B. & Williamson, J. H. (1930) Two new Neotropical Aeshnines (Odonata). Occasional Papers of the Museum of Zoology, University of Michigan, 218, 1 - 5. [Gynacantha helenga]" type="journal article" year="1930">Williamson & Williamson, 1930</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C354D420652FFF6FBE22B5CFAB0EB7A" ID-CoL="3HVTX" authority="Williamson, 1923" authorityName="Williamson" authorityYear="1923" box="[1043,1349,261,287]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="jessei">
|
||
<emphasis id="B941EAD30652FFF6FBE22B5CFB86EB7A" box="[1043,1139,261,287]" italics="true" pageId="10">G. jessei</emphasis>
|
||
<bibRefCitation id="EFA44B300652FFF6FB8A2B5CFAB0EB7A" author="Williamson, E. B." box="[1147,1349,261,287]" pageId="10" pagination="1 - 80" refId="ref7215" refString="Williamson, E. B. (1923) Notes on American species of Triacanthagyna and Gynacantha. Miscellaneous Publications of the Museum Zoology University of Michigan, 9, 1 - 80." type="journal article" year="1923">Williamson, 1923</bibRefCitation>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C354D420652FFF6FA702B5CFF15EB26" ID-CoL="3HVVP" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="tibiata">
|
||
<emphasis id="B941EAD30652FFF6FA702B5CFF15EB26" italics="true" pageId="10">G. tibiata</emphasis>
|
||
</taxonomicName>
|
||
differ in having the epiproct only about 1/3 the length of the cercus. Also, these species lack the distinctive thoracic stripes of the new species. The femora of the mid and hind legs are black apically, differing from
|
||
<taxonomicName id="4C354D420652FFF6FAB92B14FF3CEBEE" ID-CoL="3HVT4" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="caudata">
|
||
<emphasis id="B941EAD30652FFF6FAB92B14FF3CEBEE" italics="true" pageId="10">G. caudata</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C354D420652FFF6FF262B28FDE2EBEE" ID-CoL="6KVM2" authority="Williamson, 1923" authorityName="Williamson" authorityYear="1923" box="[215,535,369,395]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="klagesi">
|
||
<emphasis id="B941EAD30652FFF6FF262B28FEBDEBEE" box="[215,328,369,395]" italics="true" pageId="10">G. klagesi</emphasis>
|
||
<bibRefCitation id="EFA44B300652FFF6FEBF2B28FDE2EBEE" author="Williamson, E. B." box="[334,535,369,395]" pageId="10" pagination="1 - 80" refId="ref7215" refString="Williamson, E. B. (1923) Notes on American species of Triacanthagyna and Gynacantha. Miscellaneous Publications of the Museum Zoology University of Michigan, 9, 1 - 80." type="journal article" year="1923">Williamson, 1923</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C354D420652FFF6FDD32B28FD6EEBEE" ID-CoL="6L7LQ" authorityName="Williamson" authorityYear="1923" box="[546,667,369,395]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="laticeps">
|
||
<emphasis id="B941EAD30652FFF6FDD32B28FD6EEBEE" box="[546,667,369,395]" italics="true" pageId="10">G. laticeps</emphasis>
|
||
</taxonomicName>
|
||
; and
|
||
<emphasis id="B941EAD30652FFF6FD292B28FC88EBEE" box="[728,893,369,395]" italics="true" pageId="10">
|
||
<taxonomicName id="4C354D420652FFF6FD292B28FC8DEBEE" ID-CoL="3HVSQ" authorityName="Martin" authorityYear="1909" box="[728,888,369,395]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="auricularis">G. auricularis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but similar to
|
||
<taxonomicName id="4C354D420652FFF6FBEC2B28FF3AEBCA" ID-CoL="9WWBB" authority="Neiss & De Marmels, 2017" authorityName="Neiss & De Marmels" authorityYear="2017" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="dryadula">
|
||
<emphasis id="B941EAD30652FFF6FBEC2B28FB51EBEE" box="[1053,1188,369,395]" italics="true" pageId="10">G. dryadula</emphasis>
|
||
Neiss & De Marmels, 2017
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C354D420652FFF6FEF42BCCFE93EBCA" ID-CoL="3HVTX" authorityName="Williamson" authorityYear="1923" box="[261,358,405,431]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="jessei">
|
||
<emphasis id="B941EAD30652FFF6FEF42BCCFE93EBCA" box="[261,358,405,431]" italics="true" pageId="10">G. jessei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C354D420652FFF6FE852BCCFD9DEBCA" ID-CoL="9WWBB" authorityName="Neiss & De Marmels" authorityYear="2017" box="[372,616,405,431]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="dryadula">
|
||
<emphasis id="B941EAD30652FFF6FE852BCCFD9DEBCA" box="[372,616,405,431]" italics="true" pageId="10">Gynacantha dryadula</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C354D420652FFF6FD872BCCFD23EBCA" ID-CoL="3HVTX" authorityName="Williamson" authorityYear="1923" box="[630,726,405,431]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="jessei">
|
||
<emphasis id="B941EAD30652FFF6FD872BCCFD23EBCA" box="[630,726,405,431]" italics="true" pageId="10">G. jessei</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C354D420652FFF6FCE52BCCFC78EBCA" ID-CoL="6L7LQ" authorityName="Williamson" authorityYear="1923" box="[788,909,405,431]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="laticeps">
|
||
<emphasis id="B941EAD30652FFF6FCE52BCCFC78EBCA" box="[788,909,405,431]" italics="true" pageId="10">G. laticeps</emphasis>
|
||
</taxonomicName>
|
||
have a uniformly colored thorax. Also,
|
||
<taxonomicName id="4C354D420652FFF6FABA2BCCFF22EBB6" ID-CoL="9WWBB" authorityName="Neiss & De Marmels" authorityYear="2017" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="dryadula">
|
||
<emphasis id="B941EAD30652FFF6FABA2BCCFF22EBB6" italics="true" pageId="10">G. dryadula</emphasis>
|
||
</taxonomicName>
|
||
has a unique truncate cercus tip.
|
||
<taxonomicName id="4C354D420652FFF6FDB82BE0FCC7EBB6" ID-CoL="3HVT4" box="[585,818,441,467]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="caudata">
|
||
<emphasis id="B941EAD30652FFF6FDB82BE0FCC7EBB6" box="[585,818,441,467]" italics="true" pageId="10">Gynacantha caudata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C354D420652FFF6FCB12BE0FC3AEBB6" ID-CoL="3HVTJ" baseAuthorityName="Martin" baseAuthorityYear="1909" box="[832,975,441,467]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="francesca">
|
||
<emphasis id="B941EAD30652FFF6FCB12BE0FC3AEBB6" box="[832,975,441,467]" italics="true" pageId="10">G. francesca</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C354D420652FFF6FBF52BE0FB9CEBB6" ID-CoL="3HVVN" box="[1028,1129,441,467]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="tenuis">
|
||
<emphasis id="B941EAD30652FFF6FBF52BE0FB9CEBB6" box="[1028,1129,441,467]" italics="true" pageId="10">G. tenuis</emphasis>
|
||
</taxonomicName>
|
||
share green lateral thoracic stripes with
|
||
<taxonomicName id="4C354D420652FFF6FED12B84FE60EB92" ID-CoL="9X2VK" authorityName="Haber" authorityYear="2019" box="[288,405,477,503]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="vargasi">
|
||
<emphasis id="B941EAD30652FFF6FED12B84FE60EB92" box="[288,405,477,503]" italics="true" pageId="10">G. vargasi</emphasis>
|
||
</taxonomicName>
|
||
, but are smaller species (HW 39 to 41), and the femora are not tipped with black.
|
||
</paragraph>
|
||
<paragraph id="8B8A36C10652FFF6FF362859FD15E896" blockId="10.[151,1437,153,755]" pageId="10">
|
||
<emphasis id="B941EAD30652FFF6FF362859FE36E87E" bold="true" box="[199,451,512,539]" pageId="10">Habitat and Ecology.</emphasis>
|
||
Adults of
|
||
<taxonomicName id="4C354D420652FFF6FDC82858FD5AE87E" ID-CoL="9X2VK" authorityName="Haber" authorityYear="2019" box="[569,687,513,539]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="vargasi">
|
||
<emphasis id="B941EAD30652FFF6FDC82858FD5AE87E" box="[569,687,513,539]" italics="true" pageId="10">G. vargasi</emphasis>
|
||
</taxonomicName>
|
||
were all collected on the Caribbean slope of
|
||
<collectingCountry id="F32276510652FFF6FB582858FAD6E87E" box="[1193,1315,513,539]" name="Costa Rica" pageId="10">Costa Rica</collectingCountry>
|
||
during the wet season in April, June, August, and November. While the larval habitat and oviposition sites are unknown, the
|
||
<typeStatus id="548E88630652FFF6FF662810FF0CE806" box="[151,249,585,611]" pageId="10" type="holotype">holotype</typeStatus>
|
||
male was collected while patrolling over a residual pool in a seasonal stream at a time when that section of the stream had no running water. The Las Brisas female was collected in August at the edge of the same stream, then with flowing water, while perched during the morning on an understory palm in secondary forest. The Río Peñas Blancas female was collected in November when it flew into a building while foraging with other aeshnids during mid-day over a grassy yard at a height of about
|
||
<specimenCount id="9D33FD480652FFF6FD5F2880FD15E896" box="[686,736,729,755]" pageId="10" type="generic">
|
||
<quantity id="4CCD9B240652FFF6FD5F2880FD29E896" box="[686,732,729,755]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="10" unit="m" value="3.0">3 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |