120 lines
15 KiB
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120 lines
15 KiB
XML
<document id="C45B726E38F02EBA5CFD1BC66222EEC4" ID-CLB-Dataset="32201" ID-DOI="10.11646/zootaxa.4317.3.4" ID-GBIF-Dataset="ca0d653d-32de-4c75-a95d-d9a4690d6fda" ID-ISSN="1175-5326" ID-Zenodo-Dep="884766" ID-ZooBank="5Fd492A4-D41D-4F37-A121-Ffa680E7E778" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504596385874" checkinUser="plazi" docAuthor="Fery, Hans" docDate="2017" docId="039C87CE0F27FF85FF1F6679CD5C57D2" docLanguage="en" docName="zootaxa.4317.3.4.pdf" docOrigin="Zootaxa 4317 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clemnius Fery 2017, n. gen." docType="treatment" docVersion="7" lastPageNumber="503" masterDocId="FFA5FFB60F23FF81FF886361C8305140" masterDocTitle="A new classification of the tribe Hygrotini Portevin, 1929 (Coleoptera: Dytiscidae: Hydroporinae)" masterLastPageNumber="529" masterPageNumber="499" pageNumber="503" updateTime="1698467863718" updateUser="ExternalLinkService">
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<mods:title id="9CCECC4CC81979C7702206B7896A88C9">A new classification of the tribe Hygrotini Portevin, 1929 (Coleoptera: Dytiscidae: Hydroporinae)</mods:title>
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<mods:namePart id="00D6A66A6945E1C188A1F13F9DBB3B34">Fery, Hans</mods:namePart>
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<mods:title id="6691FE62C2DE4A7126AA6AB5F439E5D7">Zootaxa</mods:title>
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<mods:part id="1C579088E48683D2192B007681118F43">
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<mods:date id="B6E7828C839D30B6C4FCEF653544D9C0">2017</mods:date>
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<mods:number id="9E40429153D8970485670132CF26BB57">2017-05-05</mods:number>
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<mods:number id="32FC832AAAE91F19728E5756A4FB0BC3">4317</mods:number>
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<mods:classification id="F43A265FF9CE14C606155327900978C0">journal article</mods:classification>
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<mods:identifier id="428AC874C95271874429198349013882" type="DOI">10.11646/zootaxa.4317.3.4</mods:identifier>
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<treatment id="039C87CE0F27FF85FF1F6679CD5C57D2" ID-DOI="http://doi.org/10.5281/zenodo.6026847" ID-GBIF-Taxon="133183479" ID-Zenodo-Dep="6026847" LSID="urn:lsid:plazi:treatment:039C87CE0F27FF85FF1F6679CD5C57D2" httpUri="http://treatment.plazi.org/id/039C87CE0F27FF85FF1F6679CD5C57D2" lastPageNumber="503" pageId="4" pageNumber="503">
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<subSubSection id="C32F65530F27FF85FF1F6679C98B5472" box="[151,443,1303,1330]" pageId="4" pageNumber="503" type="nomenclature">
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<paragraph id="8B8A36D80F27FF85FF1F6679C98B5472" blockId="4.[151,443,1303,1330]" box="[151,443,1303,1330]" pageId="4" pageNumber="503">
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<heading id="D0C281B40F27FF85FF1F6679C98B5472" bold="true" box="[151,443,1303,1330]" fontSize="11" level="1" pageId="4" pageNumber="503" reason="1">
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<emphasis id="B941EACA0F27FF85FF1F6679C98B5472" bold="true" box="[151,443,1303,1330]" pageId="4" pageNumber="503">
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Genus
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<taxonomicName id="4C354D5B0F27FF85FF656676C9515471" ID-CoL="8LV54" authority="Fery, 2017" authorityName="Fery" authorityYear="2017" box="[237,353,1303,1329]" class="Insecta" family="Dytiscidae" genus="Clemnius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="503" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B941EACA0F27FF85FF656676C9515471" bold="true" box="[237,353,1303,1329]" italics="true" pageId="4" pageNumber="503">Clemnius</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27257B10F27FF85FEE16679C98B5472" box="[361,443,1304,1330]" pageId="4" pageNumber="503" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F65530F27FF85FF1F663FCD59549E" pageId="4" pageNumber="503" type="materials_examined">
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<paragraph id="8B8A36D80F27FF85FF1F663FCBEF5434" blockId="4.[151,991,1374,1396]" box="[151,991,1374,1396]" pageId="4" pageNumber="503">
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<emphasis id="B941EACA0F27FF85FF1F663FC92C5434" bold="true" box="[151,284,1374,1396]" pageId="4" pageNumber="503">Type species</emphasis>
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:
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<taxonomicName id="4C354D5B0F27FF85FEA2663FCA805434" authority="Gyllenhal, 1810" authorityName="Gyllenhal" authorityYear="1810" box="[298,688,1374,1396]" class="Insecta" family="Dytiscidae" genus="Hyphydrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="503" phylum="Arthropoda" rank="species" species="decoratus">
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<emphasis id="B941EACA0F27FF85FEA2663FCA335434" box="[298,515,1374,1396]" italics="true" pageId="4" pageNumber="503">Hyphydrus decoratus</emphasis>
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<bibRefCitation id="EFA44B290F27FF85FD82663FCA805434" author="Gyllenhal" box="[522,688,1374,1396]" pageId="4" pageNumber="503" refString="Gyllenhal, L. (1810) Insecta Suecica descripta. Classis I. Coleoptera sive Eleutherata. Vol. 1. Part 2. F. J. Leverentz, Scaris, 660 pp. [pp. III - XIX (Addenda ad partem prima) + 1 + 660 pp.]" type="book" year="1810">Gyllenhal, 1810</bibRefCitation>
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</taxonomicName>
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: XVI, by present designation.
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</paragraph>
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<paragraph id="8B8A36D80F27FF85FF1F66C0CD59549E" blockId="4.[151,1437,1441,1682]" pageId="4" pageNumber="503">
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The new genus
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<taxonomicName id="4C354D5B0F27FF85FECE66C0C98054FA" authority="Fery, 2017" authorityName="Fery" authorityYear="2017" box="[326,432,1441,1466]" class="Insecta" family="Dytiscidae" genus="Clemnius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="503" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B941EACA0F27FF85FECE66C0C98054FA" box="[326,432,1441,1466]" italics="true" pageId="4" pageNumber="503">Clemnius</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27257B10F27FF85FE3E66C0CA3554F9" box="[438,517,1441,1466]" pageId="4" pageNumber="503" rank="genus">
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<emphasis id="B941EACA0F27FF85FE3E66C0C9CE54FA" bold="true" box="[438,510,1441,1466]" pageId="4" pageNumber="503">n. gen</emphasis>
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.
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</taxonomicNameLabel>
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includes eight described species distributed in the Nearctic (including the north of MeXico, sometimes treated as Neotropical) and in the Palaearctic zoogeographical region (
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<figureCitation id="130E2A5D0F27FF85FB0566A4CCE0549E" box="[1165,1232,1477,1502]" captionStart="FIGURE 2" captionStartId="12.[151,250,1941,1963]" captionTargetBox="[208,1378,197,1919]" captionTargetId="figure@12.[189,1396,193,1921]" captionTargetPageId="12" captionText="FIGURE 2. Dendrogram showing the new classification of Hygrotini. A number of species without molecular data (no voucher number) have been inserted near morphologically similar species with molecular data. Acronyms refer to biogeographical regions." httpUri="https://zenodo.org/record/884770/files/figure.png" pageId="4" pageNumber="503">Fig. 2</figureCitation>
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and
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<tableCitation id="C6B703630F27FF85FA8E66A4CD68549E" box="[1286,1368,1477,1502]" captionStart="TABLE 1" captionStartId="5.[125,200,512,532]" captionText="TABLE 1. Checklist of all 137 species of Hygrotini with original, new and former (Nilsson & Há ek 2017 a) generic and subgeneric names; last three columns indicating new combinations, distribution and methods applied to material studied. See Villastrigo et al. (in press) for details on the specimens used in the molecular phylogeny and accession numbers of the sequences." pageId="4" pageNumber="503">Table 1</tableCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F65530F27FF85FF4F6688CD5C57D2" pageId="4" pageNumber="503" type="diagnosis">
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<paragraph id="8B8A36D80F27FF85FF4F6688CD5C57D2" blockId="4.[151,1437,1441,1682]" pageId="4" pageNumber="503">
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<emphasis id="B941EACA0F27FF85FF4F6688C9715742" bold="true" box="[199,321,1513,1538]" pageId="4" pageNumber="503">Diagnosis.</emphasis>
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In the new classification
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<taxonomicName id="4C354D5B0F27FF85FDD56688CAF75742" authority="Fery, 2017" authorityName="Fery" authorityYear="2017" box="[605,711,1513,1538]" class="Insecta" family="Dytiscidae" genus="Clemnius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="503" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B941EACA0F27FF85FDD56688CAF75742" box="[605,711,1513,1538]" italics="true" pageId="4" pageNumber="503">Clemnius</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27257B10F27FF85FD456688CB2B5741" box="[717,795,1513,1538]" pageId="4" pageNumber="503" rank="genus">
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<emphasis id="B941EACA0F27FF85FD456688CB245742" bold="true" box="[717,788,1513,1538]" pageId="4" pageNumber="503">n. gen</emphasis>
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.
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</taxonomicNameLabel>
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is one of two genera of the tribe
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<taxonomicName id="4C354D5B0F27FF85FB016688CCCA5742" authorityName="Portevin" authorityYear="1929" box="[1161,1274,1513,1538]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="503" phylum="Arthropoda" rank="tribe" tribe="Hygrotini">Hygrotini</taxonomicName>
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. All species of this and the second genus
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<taxonomicName id="4C354D5B0F27FF85FE40656FCA1F5765" authorityName="sensu Stephens" authorityYear="1828" box="[456,559,1550,1573]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="503" phylum="Arthropoda" rank="genus">
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<emphasis id="B941EACA0F27FF85FE40656FCA1F5765" box="[456,559,1550,1573]" italics="true" pageId="4" pageNumber="503">Hygrotus</emphasis>
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</taxonomicName>
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have two carinae in each antennal cavity (see
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<figureCitation id="130E2A5D0F27FF85FBC5656CCC845766" box="[1101,1204,1549,1574]" captionStart-0="FIGURES 3 – 4" captionStart-1="FIGURES 5 – 6" captionStartId-0="13.[151,264,1658,1680]" captionStartId-1="14.[151,264,1930,1952]" captionTargetBox-0="[353,1132,193,1614]" captionTargetBox-1="[264,1323,403,1909]" captionTargetId-0="figure@13.[353,1233,193,1614]" captionTargetId-1="figure@14.[264,1323,403,1909]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 3 – 4. Antennal cavities of: (3) Hydroporus dorsalis (Hydroporini) and (4) Hygrotus (Leptolambus) lagari (Fery, 1992) (a: carina delimiting capsule for scape; b: fronto-lateral border of clypeus; c: second carina)." captionText-1="FIGURES 5 – 6. SEM micrographs with details of head and antennal cavities of: (5) Hygrotus (Leptolambus) impressopunctatus (frontal view) and (6) Hygrotus (s. str.) inaequalis (oblique ventral view) (a, b and c as in Figs 3 – 4; d: clypeal stripe; e: labrum)." httpUri-0="https://zenodo.org/record/884772/files/figure.png" httpUri-1="https://zenodo.org/record/884774/files/figure.png" pageId="4" pageNumber="503">Figs 4–6</figureCitation>
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), a unique character among the entire subfamily
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<taxonomicName id="4C354D5B0F27FF85FE656550CABC570A" box="[493,652,1585,1610]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="503" phylum="Arthropoda" rank="subFamily" subFamily="Hydroporinae">Hydroporinae</taxonomicName>
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. The new genus—which corresponds to clade A in
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<figureCitation id="130E2A5D0F27FF85FA836550CD7E570A" box="[1291,1358,1585,1610]" captionStart="FIGURE 1" captionStartId="11.[151,250,1975,1997]" captionTargetBox="[188,1357,201,1928]" captionTargetId="figure@11.[185,1388,195,1940]" captionTargetPageId="11" captionText="FIGURE 1. Phylogeny of Hygrotini, modified from Villastrigo et al. (in press). Numbers above nodes are bootstrap support values. Note that here the former generic and subgeneric names are still used." httpUri="https://zenodo.org/record/884768/files/figure.png" pageId="4" pageNumber="503">Fig. 1</figureCitation>
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—is a heterogeneous assembly of relatively small species with different morphological characters. It is subdivided into two sister-clades each treated as a subgenus:
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<emphasis id="B941EACA0F27FF85FD076518CACF57D2" box="[655,767,1657,1682]" italics="true" pageId="4" pageNumber="503">Cyclopius</emphasis>
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<emphasis id="B941EACA0F27FF85FC8E6518CB4F57D2" bold="true" box="[774,895,1657,1682]" pageId="4" pageNumber="503">n. subgen.</emphasis>
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and
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<taxonomicName id="4C354D5B0F27FF85FC3D6518CC2E57D2" authority="Fery, 2017" authorityName="Fery" authorityYear="2017" box="[949,1054,1657,1682]" class="Insecta" family="Dytiscidae" genus="Clemnius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="524" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B941EACA0F27FF85FC3D6518CC2E57D2" box="[949,1054,1657,1682]" italics="true" pageId="4" pageNumber="503">Clemnius</emphasis>
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</taxonomicName>
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s. str.
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<emphasis id="B941EACA0F27FF85FBEF6518CCEF57D2" bold="true" box="[1127,1247,1657,1682]" pageId="4" pageNumber="503">n. subgen.</emphasis>
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(see
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<figureCitation id="130E2A5D0F27FF85FA936518CD6F57D2" box="[1307,1375,1657,1682]" captionStart="FIGURE 2" captionStartId="12.[151,250,1941,1963]" captionTargetBox="[208,1378,197,1919]" captionTargetId="figure@12.[189,1396,193,1921]" captionTargetPageId="12" captionText="FIGURE 2. Dendrogram showing the new classification of Hygrotini. A number of species without molecular data (no voucher number) have been inserted near morphologically similar species with molecular data. Acronyms refer to biogeographical regions." httpUri="https://zenodo.org/record/884770/files/figure.png" pageId="4" pageNumber="503">Fig. 2</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |