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<document id="21150C28FA0E6A2A8F1E2C67D1CE8B41" ID-CLB-Dataset="31199" ID-DOI="10.11646/zootaxa.4363.2.6" ID-GBIF-Dataset="f248e89f-ca42-421b-aab5-334a2281af89" ID-ISSN="1175-5326" ID-Zenodo-Dep="1098838" ID-ZooBank="4DF8A3AC-01FA-41D1-93CE-63E23FC7728E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1512980399631" checkinUser="plazi" docAuthor="Arguez, Katherine M., Moore, Matthew R. &amp; Branham, Marc A." docDate="2017" docId="039C878AFFD2FFC00D92FBABFC5C0C3C" docLanguage="en" docName="zootaxa.4363.2.6.pdf" docOrigin="Zootaxa 4363 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phileurus truncatus" docType="treatment" docVersion="8" masterDocId="FFA5FFF2FFD6FFC60D05FFB0FFA90B43" masterDocTitle="Description of the larva and pupa of Phileurus truncatus (Palisot de Beauvois, 1806) (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) with a key to described third instars of New World Phileurini" updateTime="1698508758872" updateUser="ExternalLinkService">
<mods:mods id="22E205458898B22F481502EA9D4BBE6F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3EEA9C4777FEC59BCCAC6ACAF1EF9303">
<mods:title id="0A95528E9374E363DB97E7110112D7FD">Description of the larva and pupa of Phileurus truncatus (Palisot de Beauvois, 1806) (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) with a key to described third instars of New World Phileurini</mods:title>
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<mods:name id="D09E8F6D2263E2C8003689F438DBC649" type="personal">
<mods:role id="0E7ACC2D6F955361434C378508747E2A">
<mods:roleTerm id="BADBB72FA4769CD10E637E96C53418FF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="142CD3E26A60419EC5530595083FF7F4">Arguez, Katherine M.</mods:namePart>
</mods:name>
<mods:name id="E93F6F60492FA4D6C0C7BF57758AB876" type="personal">
<mods:role id="A9871BDFA795367523BDDA265E50BF78">
<mods:roleTerm id="0D954752A6E5B6C27D5AA17C1A529CE6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D3BCEBDFD15A550F4787594A448CA0BF">Moore, Matthew R.</mods:namePart>
</mods:name>
<mods:name id="E005CB06B703CA931A336FB04122DBCA" type="personal">
<mods:role id="09DD1EA722DF364DDA43408193AF6DD5">
<mods:roleTerm id="05312D3986D56B2542680BFC964C73F2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1FD0B8E49A8E95B9E37EB09A12E72122">Branham, Marc A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="FF4ED355EE629B89212A2DD419C78C60">text</mods:typeOfResource>
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<mods:titleInfo id="CF2D044DFA62F3D348138D51A46D5EE4">
<mods:title id="881A0B02B082BE45FF9D34014634C7B3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2A363B52F63EDE7D05AA8C8E7A924781">
<mods:date id="DFABA32CCDAFB06717A9C961B5326A61">2017</mods:date>
<mods:detail id="6A5BDF7943311F3AA1193028E8E5B190" type="pubDate">
<mods:number id="5AA552DBD84B9AB6D52E21089329AAA7">2017-12-11</mods:number>
</mods:detail>
<mods:detail id="9CFE613569710C61301477E228B0DBB0" type="volume">
<mods:number id="9FC0C625872F349B0DCD5DFAA8FB392F">4363</mods:number>
</mods:detail>
<mods:detail id="55C9FECA106F23CDFBFC686C0862AB6F" type="issue">
<mods:number id="85F5A98518CB18190B262E4BF12AC0A5">2</mods:number>
</mods:detail>
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<mods:classification id="2A7BB08BCB34E447AB2F5751157709C1">journal volume</mods:classification>
<mods:identifier id="1EA80E73066328D88E22827BED44C7EF" type="CLB-Dataset">31199</mods:identifier>
<mods:identifier id="FCA9FE15E5C51477A38C4A0344AB8133" type="DOI">10.11646/zootaxa.4363.2.6</mods:identifier>
<mods:identifier id="D83B716C46E815367DF3187C7022CA9D" type="GBIF-Dataset">f248e89f-ca42-421b-aab5-334a2281af89</mods:identifier>
<mods:identifier id="7678BD57A0A1E0515B337916558416EA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BE2782522E5A7A5A97CC0B51B9FEA0B0" type="Zenodo-Dep">1098838</mods:identifier>
<mods:identifier id="5F3EFC948755760F8C6E48906E37975B" type="ZooBank">4DF8A3AC-01FA-41D1-93CE-63E23FC7728E</mods:identifier>
</mods:mods>
<treatment id="039C878AFFD2FFC00D92FBABFC5C0C3C" ID-DOI="http://doi.org/10.5281/zenodo.6046115" ID-GBIF-Taxon="137665556" ID-Zenodo-Dep="6046115" LSID="urn:lsid:plazi:treatment:039C878AFFD2FFC00D92FBABFC5C0C3C" httpUri="http://treatment.plazi.org/id/039C878AFFD2FFC00D92FBABFC5C0C3C" lastPageId="6" pageId="4">
<subSubSection id="C32F6517FFD2FFC20D92FBABFF7F0F14" pageId="4" type="nomenclature">
<paragraph id="8B8A369CFFD2FFC20D92FBABFD740F75" blockId="4.[151,733,1051,1147]" box="[151,733,1051,1078]" pageId="4">
<heading id="D0C281F0FFD2FFC20D92FBABFD740F75" bold="true" box="[151,733,1051,1078]" fontSize="11" level="1" pageId="4" reason="1">
<taxonomicName id="4C354D1FFFD2FFC20D92FBABFD740F75" authority="(Palisot de Beauvois, 1806)" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1806" box="[151,733,1051,1078]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B941EA8EFFD2FFC20D92FBABFD740F75" bold="true" box="[151,733,1051,1078]" pageId="4">
<emphasis id="B941EA8EFFD2FFC20D92FBABFE200F76" bold="true" box="[151,393,1051,1077]" italics="true" pageId="4">Phileurus truncatus</emphasis>
(Palisot de Beauvois, 1806)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8A369CFFD2FFC20D92FB8EFF7F0F14" blockId="4.[151,733,1051,1147]" box="[151,214,1086,1111]" pageId="4">
<emphasis id="B941EA8EFFD2FFC20D92FB8EFF7F0F14" bold="true" box="[151,214,1086,1111]" pageId="4">Pupa</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32F6517FFD2FFC20D92FBD2FE8C0F38" box="[151,293,1122,1147]" pageId="4" type="description">
<paragraph id="8B8A369CFFD2FFC20D92FBD2FE8C0F38" blockId="4.[151,733,1051,1147]" box="[151,293,1122,1147]" pageId="4">
(
<figureCitation id="130E2A19FFD2FFC20D9AFBD2FEB40F38" box="[159,285,1122,1147]" captionStart="FIGURES 11–13" captionStartId="6.[151,264,1387,1410]" captionTargetBox="[305,1276,205,1360]" captionTargetId="figure@6.[297,1290,193,1366]" captionTargetPageId="6" captionText="FIGURES 11–13. Phileurus truncatus, pupa.11, Dorsalhabitus; 12, ventral habitus; 13, lateral habitus." httpUri="https://zenodo.org/record/1098846/files/figure.png" pageId="4">Figs 11–13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32F6517FFD2FFC20D92FB1AFD7E0E48" pageId="4" type="materials_examined">
<paragraph id="8B8A369CFFD2FFC20D92FB1AFD7E0E48" blockId="4.[151,1437,1194,1867]" pageId="4">
<emphasis id="B941EA8EFFD2FFC20D92FB1AFE2B0F80" bold="true" box="[151,386,1194,1219]" pageId="4">Material examined.</emphasis>
Two pupae with the following data: “
<collectingRegion id="49F1F87EFFD2FFC20E37FB1BFC1B0F80" box="[818,946,1195,1219]" country="Uruguay" name="Florida" pageId="4">FLORIDA</collectingRegion>
: Alachua Co/Gainesville; turkey oak;/larva
<date id="FF8B105CFFD2FFC20D92FB7FFEA90FA4" box="[151,256,1230,1255]" pageId="4" value="1976-02-07">7-II-1976</date>
/pupa
<date id="FF8B105CFFD2FFC20C55FB7EFE640FA4" box="[336,461,1230,1255]" pageId="4" value="1976-04-15">15-IV-1976</date>
/
<taxonomicName id="4C354D1FFFD2FFC20CD0FB7EFDC40FA4" authorityName="Latreille" authorityYear="1802" box="[469,621,1230,1255]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
<taxonomicName id="4C354D1FFFD2FFC20F78FB7EFD430FA4" authorityName="Latreille" authorityYear="1807" box="[637,746,1230,1255]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="genus">
<emphasis id="B941EA8EFFD2FFC20F78FB7EFD430FA4" box="[637,746,1230,1255]" italics="true" pageId="4">Phileurus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C354D1FFFD2FFC20FF7FB7FFCF20FA5" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1806" box="[754,859,1231,1254]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B941EA8EFFD2FFC20FF7FB7FFCF20FA5" box="[754,859,1231,1254]" italics="true" pageId="4">truncatus</emphasis>
</taxonomicName>
/P.M. Choate collr.”; “
<collectingRegion id="49F1F87EFFD2FFC20969FB7FFB450FA4" box="[1132,1260,1231,1255]" country="Uruguay" name="Florida" pageId="4">FLORIDA</collectingRegion>
: Alachua Co./ Gainesville;
<collectionCode id="ED24AE59FFD2FFC20C20FB43FEC60E48" box="[293,367,1267,1291]" pageId="4">NATL</collectionCode>
,
<date id="FF8B105CFFD2FFC20C7EFB42FE600E48" box="[379,457,1266,1291]" pageId="4" value="2016-01">
<collectionCode id="ED24AE59FFD2FFC20C7EFB42FE240E48" box="[379,397,1266,1291]" pageId="4">I-</collectionCode>
2016
</date>
./coll. Gabe Somarriba”.
</paragraph>
</subSubSection>
<subSubSection id="C32F6517FFD2FFC20DC2FAA6FD1B0C09" pageId="4" type="description">
<paragraph id="8B8A369CFFD2FFC20DC2FAA6FD1B0C09" blockId="4.[151,1437,1194,1867]" pageId="4">
<emphasis id="B941EA8EFFD2FFC20DC2FAA6FEF00E6C" bold="true" box="[199,345,1302,1327]" pageId="4">Description.</emphasis>
Length
<quantity id="4CCD9B79FFD2FFC20CBDFAA7FD9C0E6D" box="[440,565,1303,1327]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" metricValueMax="4.4" metricValueMin="4.2" pageId="4" unit="mm" value="43.0" valueMax="44.0" valueMin="42.0">42–44 mm</quantity>
; greatest width
<quantity id="4CCD9B79FFD2FFC20FECFAA7FCCD0E6D" box="[745,868,1303,1327]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="4" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">18–20 mm</quantity>
(
<figureCitation id="130E2A19FFD2FFC20E71FAA6FC5F0E6C" box="[884,1014,1302,1327]" captionStart="FIGURES 11–13" captionStartId="6.[151,264,1387,1410]" captionTargetBox="[305,1276,205,1360]" captionTargetId="figure@6.[297,1290,193,1366]" captionTargetPageId="6" captionText="FIGURES 11–13. Phileurus truncatus, pupa.11, Dorsalhabitus; 12, ventral habitus; 13, lateral habitus." httpUri="https://zenodo.org/record/1098846/files/figure.png" pageId="4">Figs 11–13</figureCitation>
). Body elongate, oval, stout, exarate. Color dark reddish brown. Entire body with very fine, small, golden setae.
<emphasis id="B941EA8EFFD2FFC20EF4FA8AFB930E10" bold="true" box="[1009,1082,1338,1363]" pageId="4">Head:</emphasis>
Surface glabrous, bent sharply beneath thorax, mouthparts directed ventrally; antennae, labrum, mandibles, maxillary, and labial palps discernible; antennal thecae expanded, stout, with apices rounded. Compound eyes sunken, visible, with 2 tubercles between eyes.
<emphasis id="B941EA8EFFD2FFC20DDFFA16FE920EFC" bold="true" box="[218,315,1446,1471]" pageId="4">Thorax:</emphasis>
Pronotal surface glabrous. Form transverse, with wide, anteromedial, rounded apex. Mesonotum and metanotum well differentiated. Elytral and posterior wing thecae closely appressed, curved ventrally around body; elytral thecae extending to end of abdominal segment I; posterior wing thecae extending to middle of abdominal segment III. Protibia with 3 distinct teeth on external edge. Mesotibia and metatibia with 3 swellings at apices; characters indicative to spines and spurs seen in adults, with the external swelling most likely the spine and the 2 inner swellings most likely the 2 apical spurs (1 spine on each tibia in adults).
<emphasis id="B941EA8EFFD2FFC20964F9EAFB760D30" bold="true" box="[1121,1247,1626,1651]" pageId="4">Abdomen:</emphasis>
Segments III–X (ventral view) well defined. Segment VII slightly long than preceding segment and segment VIII 0.25 times longer than segment VII; segments VIII and IX fused. Segments I–X (dorsal view) with well-defined, dioneiform (gin trap) organs between segments I–II, II–III, III–IV, IV–V, V–VI, and VI–VII. Pleural lobes rounded. Spiracle I ovate, covered by wing thecae, more broadly opened than spiracles II–IV; spiracles II–IV ovate, prominent, each with strongly sclerotized peritreme; spiracles V–VI less sclerotized compared to I–IV; spiracles V–VIII closed. Abdominal apex rounded, with fine, short setae.
</paragraph>
</subSubSection>
<caption id="DF4A6614FFD3FFC30D92F831FDE80C96" httpUri="https://zenodo.org/record/1098844/files/figure.png" pageId="5" targetBox="[254,1305,195,1870]" targetPageId="5">
<paragraph id="8B8A369CFFD3FFC30D92F831FDE80C96" blockId="5.[151,1435,1921,2005]" pageId="5">
<emphasis id="B941EA8EFFD3FFC30D92F831FEE50CD4" bold="true" box="[151,332,1921,1944]" pageId="5">FIGURES 7–10.</emphasis>
<taxonomicName id="4C354D1FFFD3FFC30C5DF832FD8F0CDB" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1806" box="[344,550,1922,1944]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B941EA8EFFD3FFC30C5DF832FD8F0CDB" box="[344,550,1922,1944]" italics="true" pageId="5">Phileurus truncatus</emphasis>
</taxonomicName>
, third instar.
<emphasis id="B941EA8EFFD3FFC30FB9F832FD660CD4" bold="true" box="[700,719,1922,1943]" pageId="5">7,</emphasis>
Maxillary stridulatory area. SD—stridulatory teeth;
<emphasis id="B941EA8EFFD3FFC30803F832FAB30CD4" bold="true" box="[1286,1306,1922,1943]" pageId="5">8,</emphasis>
labium and maxillae, dorsal view. HSC—hypopharyngeal sclerome; TP—truncate process;
<emphasis id="B941EA8EFFD3FFC30904F810FBB60CF4" bold="true" box="[1025,1055,1952,1975]" pageId="5">(9)</emphasis>
antennomeres II–IV;
<emphasis id="B941EA8EFFD3FFC3081DF810FA9E0CF6" bold="true" box="[1304,1335,1952,1973]" pageId="5">10,</emphasis>
terminal antennomere. DSS—dorsal sensory spots.
</paragraph>
</caption>
<caption id="DF4A6614FFD0FFC00D92FADBFB1C0EC1" box="[151,1205,1387,1410]" httpUri="https://zenodo.org/record/1098846/files/figure.png" pageId="6" targetBox="[305,1276,205,1360]" targetPageId="6">
<paragraph id="8B8A369CFFD0FFC00D92FADBFB1C0EC1" blockId="6.[151,1205,1387,1410]" box="[151,1205,1387,1410]" pageId="6">
<emphasis id="B941EA8EFFD0FFC00D92FADBFEFB0EC2" bold="true" box="[151,338,1387,1410]" pageId="6">FIGURES 11–13.</emphasis>
<taxonomicName id="4C354D1FFFD0FFC00C5DFADCFD880EC1" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1806" box="[344,545,1388,1410]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B941EA8EFFD0FFC00C5DFADCFD880EC1" box="[344,545,1388,1410]" italics="true" pageId="6">Phileurus truncatus</emphasis>
</taxonomicName>
, pupa.
<emphasis id="B941EA8EFFD0FFC00F6EFADCFD200EC2" bold="true" box="[619,649,1388,1409]" pageId="6">11,</emphasis>
Dorsal habitus;
<emphasis id="B941EA8EFFD0FFC00E35FADCFCE60EC2" bold="true" box="[816,847,1388,1409]" pageId="6">12,</emphasis>
ventral habitus;
<emphasis id="B941EA8EFFD0FFC00EFCFADCFBB10EC2" bold="true" box="[1017,1048,1388,1409]" pageId="6">13,</emphasis>
lateral habitus.
</paragraph>
</caption>
<subSubSection id="C32F6517FFD0FFC00DC2FA01FC5C0C3C" pageId="6" type="diagnosis">
<paragraph id="8B8A369CFFD0FFC00DC2FA01FB270DC0" blockId="6.[151,1436,1454,1919]" pageId="6">
<emphasis id="B941EA8EFFD0FFC00DC2FA01FE930E89" bold="true" box="[199,314,1457,1482]" pageId="6">Diagnosis</emphasis>
. Third-instar
<taxonomicName id="4C354D1FFFD0FFC00CD9FA02FDCC0E8A" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1806" box="[476,613,1458,1481]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B941EA8EFFD0FFC00CD9FA02FDCC0E8A" box="[476,613,1458,1481]" italics="true" pageId="6">P. truncatus</emphasis>
</taxonomicName>
will key out to the terminal 4th couplet in the phileurine larval key to species of
<bibRefCitation id="EFA44B6DFFD0FFC00C17FA65FDE30EAD" box="[274,586,1493,1518]" pageId="6" refString="Ibarra-Polesel, M. G., Valle N. G., Neita-Moreno J. C. &amp; Damborsky, M. P. (2017) Description of the pupa and redescription of the third instar of Phileurus valgus (Olivier) (Coleoptera: Scarabaeidae: Dynastinae: Phileurini). Zootaxa, 4290, 571 - 580. https: // doi. org / 10.11646 / zootaxa. 4290.3.10" type="journal article">
Ibarra-Polesel
<emphasis id="B941EA8EFFD0FFC00CBDFA66FE420EAD" box="[440,491,1493,1518]" italics="true" pageId="6">et al</emphasis>
. (2017)
</bibRefCitation>
. This is based on the lack of anterior frontal setae and the presence of an angulate margin on the left side of the labrum in
<taxonomicName id="4C354D1FFFD0FFC00FD8FA4DFCCE0D57" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1806" box="[733,871,1533,1556]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B941EA8EFFD0FFC00FD8FA4DFD420D57" box="[733,747,1533,1556]" italics="true" pageId="6">P</emphasis>
.
<emphasis id="B941EA8EFFD0FFC00FF8FA4DFCCE0D57" box="[765,871,1533,1556]" italics="true" pageId="6">truncatus</emphasis>
</taxonomicName>
. In the modified key below, the 4th couplet now separates
<taxonomicName id="4C354D1FFFD0FFC00C0FF990FEDE0D7A" authorityName="Latreille" authorityYear="1807" box="[266,375,1568,1593]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
<emphasis id="B941EA8EFFD0FFC00C0FF990FEDE0D7A" box="[266,375,1568,1593]" italics="true" pageId="6">Phileurus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C354D1FFFD0FFC00CB2F990FDF70D7A" authorityName="Kolbe" authorityYear="1910" box="[439,606,1568,1593]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
<emphasis id="B941EA8EFFD0FFC00CB2F990FDF70D7A" box="[439,606,1568,1593]" italics="true" pageId="6">Hemiphileurus</emphasis>
</taxonomicName>
species based on the number of dorsal sensory spots on the terminal antennomere.
<taxonomicName id="4C354D1FFFD0FFC00C32F9F6FDBF0D1C" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1806" box="[311,534,1606,1631]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B941EA8EFFD0FFC00C32F9F6FDBF0D1C" box="[311,534,1606,1631]" italics="true" pageId="6">Phileurus truncatus</emphasis>
</taxonomicName>
keys to the 5th couplet where it is separated from
<taxonomicName id="4C354D1FFFD0FFC00953F9F8FF660DC0" authority="Neita &amp; Ratcliffe, 2010" authorityName="Neita &amp; Ratcliffe" authorityYear="2010" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="elbitae">
<emphasis id="B941EA8EFFD0FFC00953F9F8FB6E0D1C" box="[1110,1223,1606,1631]" italics="true" pageId="6">H. elbitae</emphasis>
Neita &amp; Ratcliffe, 2010
</taxonomicName>
by the size of the abdominal spiracles and the number of maxillary stridulatory teeth.
</paragraph>
<paragraph id="8B8A369CFFD0FFC00DC2F93EFC5C0C3C" blockId="6.[151,1436,1454,1919]" pageId="6">
The third-instar
<taxonomicName id="4C354D1FFFD0FFC00C79F93EFDF00DE4" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1806" box="[380,601,1678,1703]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B941EA8EFFD0FFC00C79F93EFDF00DE4" box="[380,601,1678,1703]" italics="true" pageId="6">Phileurus truncatus</emphasis>
</taxonomicName>
examined for this study lack long, thin anterior frontal setae. The presence of anterior frontal setae, along with similar sized tarsal claws on all the legs, are characters shared between the other known third instars of
<taxonomicName id="4C354D1FFFD0FFC00CDBF966FDE20DAC" authorityName="Latreille" authorityYear="1807" box="[478,587,1750,1775]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
<emphasis id="B941EA8EFFD0FFC00CDBF966FDE20DAC" box="[478,587,1750,1775]" italics="true" pageId="6">Phileurus</emphasis>
</taxonomicName>
species (
<bibRefCitation id="EFA44B6DFFD0FFC00FB1F966FC750DAC" author="Ibarra-Polesel" box="[692,988,1750,1775]" pageId="6" refString="Ibarra-Polesel, M. G., Valle N. G., Neita-Moreno J. C. &amp; Damborsky, M. P. (2017) Description of the pupa and redescription of the third instar of Phileurus valgus (Olivier) (Coleoptera: Scarabaeidae: Dynastinae: Phileurini). Zootaxa, 4290, 571 - 580. https: // doi. org / 10.11646 / zootaxa. 4290.3.10" type="journal article" year="2017">
Ibarra-Polesel
<emphasis id="B941EA8EFFD0FFC00E59F968FC260DAC" box="[860,911,1750,1775]" italics="true" pageId="6">et al</emphasis>
. 2017
</bibRefCitation>
). The lack of long, thin anterior frontal setae in
<taxonomicName id="4C354D1FFFD0FFC00DF7F94CFEDE0C50" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1806" box="[242,375,1788,1811]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B941EA8EFFD0FFC00DF7F94CFEDE0C50" box="[242,375,1788,1811]" italics="true" pageId="6">P. truncatus</emphasis>
</taxonomicName>
thus broadens the diagnosis for the genus. Examined specimens of third-instar
<taxonomicName id="4C354D1FFFD0FFC009E9F94CFAD30C50" box="[1260,1402,1788,1811]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="trunctatus">
<emphasis id="B941EA8EFFD0FFC009E9F94CFAD30C50" box="[1260,1402,1788,1811]" italics="true" pageId="6">P. trunctatus</emphasis>
</taxonomicName>
do have setose punctures across the anterior margin of frons. However, these punctures are sporadically arranged and the short, stout setae in them does not reach past the margin of the punctures. We do not think that these punctures are associated with the anterior frontal setae found in other phileurine larvae.
</paragraph>
</subSubSection>
</treatment>
</document>
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