299 lines
51 KiB
XML
299 lines
51 KiB
XML
<document id="7BF76C71796AE0C0DE09B8EB9CA62C91" ID-CLB-Dataset="4220" ID-DOI="10.12782/sd.22_29" ID-GBIF-Dataset="111f6060-421e-4f53-bfb2-63a7d605f1a4" ID-ISSN="2189-7301" ID-Zenodo-Dep="4583717" ID-ZooBank="EFF0CA4B-AD0A-44B4-99BA-79446785ED0A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614892972232" checkinUser="felipe" docAuthor="Okumura, Ken-ichi" docDate="2017" docId="039C1A521C5CA558FC5C7B90CF244670" docLanguage="en" docName="SpeciesDiversity.22.1.29-36.pdf" docOrigin="Species Diversity 22 (1)" docSource="http://dx.doi.org/10.12782/sd.22_29" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Dichodactylus Okumura 2017, gen. nov." docType="treatment" docUuid="10EBEA2B-FFD8-41A9-A0D5-211AC2129A46" docUuidSource="ZooBank" docVersion="5" lastPageNumber="31" masterDocId="FFA5622A1C5CA55AFF947C13CD204006" masterDocTitle="Dichodactylus gen. nov. (Araneae: Agelenidae: Coelotinae) from Japan" masterLastPageNumber="36" masterPageNumber="29" pageNumber="29" updateTime="1698932642149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="659CD3A6ABEEE48D956980C6B73A7E22">
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<mods:title id="203D572DFA18F4426C82F52E7057D8CA">Dichodactylus gen. nov. (Araneae: Agelenidae: Coelotinae) from Japan</mods:title>
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<mods:namePart id="317A0C47FC9304604F4E63510A72E650">Okumura, Ken-ichi</mods:namePart>
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<mods:titleInfo id="541032047B75491FBE49796E95796DE6">
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<mods:title id="A9B5857DD4C7BCCD8FCE51FB1AB011EF">Species Diversity</mods:title>
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</mods:titleInfo>
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<mods:part id="BF77B85CC052A6934809F5EF8DD73858">
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<mods:date id="3EB8632A1739C9FCBC4B73BFF7CB7F70">2017</mods:date>
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<mods:number id="5A0005525382152ADB8DA64C97EB4235">2017-05-25</mods:number>
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<treatment id="039C1A521C5CA558FC5C7B90CF244670" ID-DOI="http://doi.org/10.5281/zenodo.5526965" ID-GBIF-Taxon="188493169" ID-Zenodo-Dep="5526965" LSID="urn:lsid:zoobank.org:act:10EBEA2B-FFD8-41A9-A0D5-211AC2129A46" httpUri="http://treatment.plazi.org/id/039C1A521C5CA558FC5C7B90CF244670" lastPageId="2" lastPageNumber="31" pageId="0" pageNumber="29">
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<subSubSection id="C32FF8CF1C5CA55AFC5C7B90C825479C" box="[968,1285,1922,1947]" pageId="0" pageNumber="29" type="nomenclature">
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<paragraph id="8B8AAB441C5CA55AFC5C7B90C825479C" blockId="0.[892,1360,1890,1947]" box="[968,1285,1922,1947]" pageId="0" pageNumber="29">
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<heading id="D0C21C281C5CA55AFC5C7B90C825479C" box="[968,1285,1922,1947]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="29" reason="2">
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Genus
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<taxonomicName id="4C35D0C71C5CA55AFB9B7B91C984479C" ID-CoL="62PFY" ID-ENA="2018510" authority="Okumura, 2017" authorityName="Okumura" authorityYear="2017" box="[1039,1188,1922,1946]" class="Arachnida" family="Agelenidae" genus="Dichodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="29" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B94177561C5CA55AFB9B7B91C984479C" bold="true" box="[1039,1188,1922,1946]" italics="true" pageId="0" pageNumber="29">Dichodactylus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A272CA2D1C5CA55AFB3E7B90C825479C" box="[1194,1285,1923,1947]" pageId="0" pageNumber="29" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32FF8CF1C5CA55AFCDA7BD7C82A47DD" box="[846,1290,1987,2012]" pageId="0" pageNumber="29" type="type_taxon">
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<paragraph id="8B8AAB441C5CA55AFCDA7BD7C82A47DD" blockId="0.[821,1432,1987,2077]" box="[846,1290,1987,2012]" pageId="0" pageNumber="29">
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<emphasis id="B94177561C5CA55AFCDA7BD7CEF747DD" bold="true" box="[846,983,1987,2012]" pageId="0" pageNumber="29">
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<typeStatus id="548E15E61C5CA55AFCDA7BD7CEA247DA" box="[846,898,1988,2012]" pageId="0" pageNumber="29">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C35D0C71C5CA55AFC657BD7C82A47DD" authority="Arita, 1976" authorityName="Arita" authorityYear="1976" box="[1009,1290,1987,2012]" class="Arachnida" family="Amaurobiidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="tarumii">
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<emphasis id="B94177561C5CA55AFC657BD7C9B347DD" box="[1009,1171,1988,2012]" italics="true" pageId="0" pageNumber="29">Coelotes tarumii</emphasis>
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<bibRefCitation id="EFA4D6B51C5CA55AFB0D7BD7C82A47DD" author="Arita, T." box="[1177,1290,1987,2012]" pageId="0" pageNumber="29" pagination="197 - 204" refId="ref4504" refString="Arita, T. 1976. Three new Coelotes (Araneae, Agelenidae) from Tottori Prefecture, west Japan. Annotationes Zoologicae Japonenses 49: 197 - 204." type="journal article" year="1976">Arita, 1976</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32FF8CF1C5CA558FCDA7BF7CCB04263" lastPageId="2" lastPageNumber="31" pageId="0" pageNumber="29" type="diagnosis">
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<paragraph id="8B8AAB441C5CA558FCDA7BF7CCB04263" blockId="0.[821,1432,1987,2077]" lastBlockId="2.[156,768,198,1654]" lastPageId="2" lastPageNumber="31" pageId="0" pageNumber="29">
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<emphasis id="B94177561C5CA55AFCDA7BF7CE9E47FA" bold="true" box="[846,958,2020,2044]" pageId="0" pageNumber="29">Diagnosis.</emphasis>
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Males of
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<taxonomicName id="4C35D0C71C5CA55AFBA97BF7C9EA47FA" authorityName="Okumura" authorityYear="2017" box="[1085,1226,2020,2044]" class="Arachnida" family="Agelenidae" genus="Dichodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B94177561C5CA55AFBA97BF7C9EA47FA" box="[1085,1226,2020,2044]" italics="true" pageId="0" pageNumber="29">Dichodactylus</emphasis>
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</taxonomicName>
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can be clearly distinguished from males of all other coelotine genera except
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<taxonomicName id="4C35D0C71C5DA55BFF087CD5CC2A40D8" authorityName="Kocak and Kemal" authorityYear="2008" box="[156,266,198,222]" class="Arachnida" family="Amaurobiidae" genus="Orumcekia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="B94177561C5DA55BFF087CD5CC2A40D8" box="[156,266,198,222]" italics="true" pageId="1" pageNumber="30">Orumcekia</emphasis>
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</taxonomicName>
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, some species of
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<taxonomicName id="4C35D0C71C5DA55BFE407CD5CF7240D8" authorityName=": Wang" authorityYear="2002" box="[468,594,198,222]" class="Arachnida" family="Amaurobiidae" genus="Platocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="B94177561C5DA55BFE407CD5CF7240D8" box="[468,594,198,222]" italics="true" pageId="1" pageNumber="30">Platocoelotes</emphasis>
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</taxonomicName>
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and some species of
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<taxonomicName id="4C35D0C71C5DA55BFF7F7CF4CC4140F9" authorityName="Ovtchinnikov" authorityYear="1999" box="[235,353,231,255]" class="Arachnida" family="Amaurobiidae" genus="Tegecoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="B94177561C5DA55BFF7F7CF4CC4140F9" box="[235,353,231,255]" italics="true" pageId="1" pageNumber="30">Tegecoelotes</emphasis>
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</taxonomicName>
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by having two patellar apophyses and reduced LTA in male palp (
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<figureCitation id="130EB7C11C5DA55BFE5C7D14CF3B4126" box="[456,539,263,288]" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="1" pageNumber="30">Figs 1B</figureCitation>
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,
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<figureCitation id="130EB7C11C5DA55BFDBE7D14CF154119" box="[554,565,263,287]" captionStart="Fig" captionStartId="5.[156,187,1156,1178]" captionTargetBox="[258,1330,205,1117]" captionTargetId="figure-449@5.[258,1330,205,1117]" captionTargetPageId="5" captionText="Fig. 5. Comparison of genitalic characters of three species of Dichodactylus gen. nov. from Japan: A, D, D. shinshuensis sp. nov., male (A: NSMT-Ar 13339), and female (D: NSMT-Ar 13341); B, E, D. tarumii (Arita, 1976), male (B: NSMT-Ar 14732), and female (E: NSMT-Ar 14733); C, F, D. satoi (Nishikawa, 2003), male (C: NSMT-Ar 14735), and female (F: NSMT-Ar 14734). A–C, retrolateral views of male palp; D–F, ventral views of epigynum. Abbreviations: CO, conductor; ET, epigynal teeth; LAM, lamellas; PA, patellar apophyses; SW, swollen portion of tegulum. Black circle on tibia shows the vestigial LTA." figureDoi="http://doi.org/10.5281/zenodo.4583728" httpUri="https://zenodo.org/record/4583728/files/figure.png" pageId="1" pageNumber="30">5</figureCitation>
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A–C). Females can be distinguished from all other coelotine genera except for some species of
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<taxonomicName id="4C35D0C71C5DA55BFED87D5BCCBD4166" authorityName="Blackwall" authorityYear="1841" box="[332,413,328,352]" class="Arachnida" family="Amaurobiidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="B94177561C5DA55BFED87D5BCCBD4166" box="[332,413,328,352]" italics="true" pageId="1" pageNumber="30">Coelotes</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C35D0C71C5DA55BFE397D5ACFDF4167" authority="Ovtchinnikov, 1999" authorityName="Ovtchinnikov" authorityYear="1999" box="[429,767,328,353]" class="Arachnida" family="Amaurobiidae" genus="Draconarius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="B94177561C5DA55BFE397D5ACF094166" box="[429,553,329,352]" italics="true" pageId="1" pageNumber="30">Draconarius</emphasis>
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Ovtchinnikov, 1999
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</taxonomicName>
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and
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<taxonomicName id="4C35D0C71C5DA55BFF597D79CCF44187" authority="Kishida, 1955" authorityName="Kishida" authorityYear="1955" box="[205,468,361,385]" class="Arachnida" family="Amaurobiidae" genus="Pireneitega" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="B94177561C5DA55BFF597D79CC1B4187" box="[205,315,362,385]" italics="true" pageId="1" pageNumber="30">Pireneitega</emphasis>
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Kishida, 1955
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</taxonomicName>
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by having anteroposteriorly elongated and convoluted spermathecae in an internal genitalia (
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<figureCitation id="130EB7C11C5DA55BFF407DB9CC0541C4" box="[212,293,426,450]" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="1" pageNumber="30">Figs 1D</figureCitation>
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,
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<figureCitation id="130EB7C11C5DA55BFEBB7DB9CC6F41C4" box="[303,335,426,450]" captionStart="Fig" captionStartId="3.[156,187,1637,1659]" captionTargetBox="[278,1319,206,1597]" captionTargetId="figure-161@3.[265,1323,205,1598]" captionTargetPageId="3" captionText="Fig. 2. Dichodactylus shinshuensis sp. nov., male (A, B: holotype, NSMT-Ar 13339), female (C, D: paratype, NSMT-Ar 13341). A, ventral view of male palp; B, retrolateral view of male palp; C, ventral view of epigynum; D, dorsal view of internal female genitalia. Abbreviations: CD, copulatory duct; CO, conductor; CY, cymbial furrow; EM, embolus; LAM, lamellas; MA, median apophysis; PA, patellar apophyses; RTA, retrolateral tibial apophysis; SP, spermatheca. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4583722" httpUri="https://zenodo.org/record/4583722/files/figure.png" pageId="1" pageNumber="30">2D</figureCitation>
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,
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<figureCitation id="130EB7C11C5DA55BFECC7DB9CC5841C4" box="[344,376,426,450]" captionStart="Fig" captionStartId="4.[156,187,1051,1073]" captionTargetBox="[305,1283,207,982]" captionTargetId="figure-7@4.[302,1285,205,1012]" captionTargetPageId="4" captionText="Fig. 3. Dichodactylus tarumii (Arita, 1976). male (A, B: NSMT-Ar 14732), female (C, D: NSMT-Ar 14733). A, ventral view of male palp; B, retrolateral view of male palp; C, ventral view of epigynum; D, dorsal view of internal female genitalia. Abbreviations: CO, conductor; CY, cymbial furrow; EM, embolus; ET, epigynal teeth; MA, median apophysis; PA, patellar apophyses; RTA, retrolateral tibial apophysis; SP, spermatheca; SH, spermathecal head. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4583724" httpUri="https://zenodo.org/record/4583724/files/figure.png" pageId="1" pageNumber="30">3D</figureCitation>
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,
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<figureCitation id="130EB7C11C5DA55BFE167DB9CC8441C4" box="[386,420,426,450]" captionStart="Fig" captionStartId="4.[156,187,1972,1994]" captionTargetBox="[264,1322,1212,1934]" captionTargetId="figure-80@4.[264,1322,1212,1934]" captionTargetPageId="4" captionText="Fig. 4. Dichodactylus satoi (Nishikawa, 2003). male (A, B: NSMT-Ar 14735), female (C, D: NSMT-Ar 14734). A, ventral view of male palp; B, retrolateral view of male palp; C, ventral view of epigynum; D, dorsal view of internal female genitalia. Abbreviations: CO, conductor; CY, cymbial furrow; EM, embolus; MA, median apophysis; PA, patellar apophyses; RTA, retrolateral tibial apophysis; SP, spermatheca; SH, spermathecal head. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4583726" httpUri="https://zenodo.org/record/4583726/files/figure.png" pageId="1" pageNumber="30">4D</figureCitation>
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). Males can be distinguished from males of
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<taxonomicName id="4C35D0C71C5DA55BFF697DD8CC4B41E5" authorityName="Kocak and Kemal" authorityYear="2008" box="[253,363,459,483]" class="Arachnida" family="Amaurobiidae" genus="Orumcekia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="B94177561C5DA55BFF697DD8CC4B41E5" box="[253,363,459,483]" italics="true" pageId="1" pageNumber="30">Orumcekia</emphasis>
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</taxonomicName>
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by the absence of a ventral apophysis of the conductor and the semi-circular or crescent-shaped median apophysis (
|
||
<figureCitation id="130EB7C11C5DA55BFEFD7E1FCCFA4222" box="[361,474,524,548]" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="1" pageNumber="30">Fig. 1A, B</figureCitation>
|
||
) (
|
||
<emphasis id="B94177561C5DA55BFE617E1FCF2F4225" box="[501,527,524,547]" italics="true" pageId="1" pageNumber="30">vs.</emphasis>
|
||
present, and spatulate or hook-shaped, respectively, in
|
||
<taxonomicName id="4C35D0C71C5DA55BFB187CD5C9DA40D8" authorityName="Kocak and Kemal" authorityYear="2008" box="[1164,1274,198,222]" class="Arachnida" family="Amaurobiidae" genus="Orumcekia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94177561C5DA55BFB187CD5C9DA40D8" box="[1164,1274,198,222]" italics="true" pageId="1" pageNumber="30">Orumcekia</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<figureCitation id="130EB7C11C5DA55BFA9F7CD5C85440D8" box="[1291,1396,198,222]" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="1" pageNumber="30">Fig. 1E, F</figureCitation>
|
||
; in an apparent simple labeling error in
|
||
<bibRefCitation id="EFA4D6B51C5DA55BFB227CF4C81E40F9" author="Wang, X. P." box="[1206,1342,231,255]" pageId="1" pageNumber="30" pagination="1 - 150" refId="ref4764" refString="Wang, X. P. 2002. A generic-level revision of the spider subfamily Coelotinae (Araneae, Amaurobiidae). Bulletin of the American Museum of Natural History 269: 1 - 150." type="journal article" year="2002">Wang (2002)</bibRefCitation>
|
||
, median apophysis was marked as an embolus in figs 161, 162, however, it was correctly labeled elsewhere in the same paper,
|
||
<emphasis id="B94177561C5DA55BFCA07D5ACE6F4166" box="[820,847,329,352]" italics="true" pageId="1" pageNumber="30">e.g</emphasis>
|
||
., figs 166, 167); females are distinguished by the absence of a posterolaterally expanded margin in the epigynum (
|
||
<figureCitation id="130EB7C11C5DA55BFAE47D7ACE7241A4" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="1" pageNumber="30">Fig. 1C</figureCitation>
|
||
) (
|
||
<emphasis id="B94177561C5DA55BFCFF7D99CEA641A7" box="[875,902,394,417]" italics="true" pageId="1" pageNumber="30">vs.</emphasis>
|
||
present in
|
||
<taxonomicName id="4C35D0C71C5DA55BFC687D9AC94A41A7" authorityName="Kocak and Kemal" authorityYear="2008" box="[1020,1130,393,417]" class="Arachnida" family="Amaurobiidae" genus="Orumcekia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94177561C5DA55BFC687D9AC94A41A7" box="[1020,1130,393,417]" italics="true" pageId="1" pageNumber="30">Orumcekia</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<figureCitation id="130EB7C11C5DA55BFBED7D9AC9E741A4" box="[1145,1223,393,418]" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="1" pageNumber="30">Fig. 1G</figureCitation>
|
||
). Moreover, females of
|
||
<taxonomicName id="4C35D0C71C5DA55BFCC47DB9CEFD41C4" authorityName="Okumura" authorityYear="2017" box="[848,989,426,450]" class="Arachnida" family="Agelenidae" genus="Dichodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94177561C5DA55BFCC47DB9CEFD41C4" box="[848,989,426,450]" italics="true" pageId="1" pageNumber="30">Dichodactylus</emphasis>
|
||
</taxonomicName>
|
||
have anteroposteriorly elongated and convoluted spermathecae (
|
||
<figureCitation id="130EB7C11C5DA55BFBB77DD9C95341E5" box="[1059,1139,458,483]" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="1" pageNumber="30">Fig. 1D</figureCitation>
|
||
), while the spermathecae of
|
||
<taxonomicName id="4C35D0C71C5DA55BFCA07DF8CE824205" authorityName="Kocak and Kemal" authorityYear="2008" box="[820,930,491,515]" class="Arachnida" family="Amaurobiidae" genus="Orumcekia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94177561C5DA55BFCA07DF8CE824205" box="[820,930,491,515]" italics="true" pageId="1" pageNumber="30">Orumcekia</emphasis>
|
||
</taxonomicName>
|
||
are more compact and transversely oriented in the posterior portion of the vulva (
|
||
<figureCitation id="130EB7C11C5DA55BFB357E1FC9D34222" box="[1185,1267,524,548]" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="1" pageNumber="30">Fig. 1H</figureCitation>
|
||
). Species of
|
||
<emphasis id="B94177561C5DA558FAE27E1FCC3040D8" italics="true" lastPageId="2" lastPageNumber="31" pageId="1" pageNumber="30">Dichodactylus</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="4C35D0C71C5EA558FDD17CD5CFE340D8" authorityName=": Wang" authorityYear="2002" box="[581,707,198,222]" class="Arachnida" family="Amaurobiidae" genus="Platocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94177561C5EA558FDD17CD5CFE340D8" box="[581,707,198,222]" italics="true" pageId="2" pageNumber="31">Platocoelotes</emphasis>
|
||
</taxonomicName>
|
||
by a spatulate conductor, short cymbial furrow, short embolus, and no epigynal hood (
|
||
<figureCitation id="130EB7C11C5EA558FE037D14CCF14126" box="[407,465,263,288]" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="2" pageNumber="31">Fig. 1</figureCitation>
|
||
A–C) (
|
||
<emphasis id="B94177561C5EA558FD8B7D1BCF1A4119" box="[543,570,264,287]" italics="true" pageId="2" pageNumber="31">vs.</emphasis>
|
||
a conductor elongated to the base of cymbium obliquely, elongated cymbial furrow, long filiform embolus, and the presence of epigynal hood in
|
||
<taxonomicName id="4C35D0C71C5EA558FF637D7ACF1A4187" authority=": Wang 2002" authorityName=": Wang" authorityYear="2002" box="[247,570,361,386]" class="Arachnida" family="Amaurobiidae" genus="Platocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94177561C5EA558FF637D7ACC554187" box="[247,373,361,385]" italics="true" pageId="2" pageNumber="31">Platocoelotes</emphasis>
|
||
:
|
||
<bibRefCitation id="EFA4D6B51C5EA558FE177D79CCD94187" author="Wang, X. P." box="[387,505,361,386]" pageId="2" pageNumber="31" pagination="1 - 150" refId="ref4764" refString="Wang, X. P. 2002. A generic-level revision of the spider subfamily Coelotinae (Araneae, Amaurobiidae). Bulletin of the American Museum of Natural History 269: 1 - 150." type="journal article" year="2002">Wang 2002</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA4D6B51C5EA558FD917D7ACF1A4187" author="Wang, X. P." box="[517,570,361,385]" pageId="2" pageNumber="31" pagination="499 - 662" refId="ref4800" refString="Wang, X. P. 2003. Species revision of the coelotine spider genera Bifi- docoelotes, Coronilla, Draconarius, Femoracoelotes, Leptocoelotes, Longicoelotes, Platocoelotes, Spiricoelotes, Tegecoelotes, and Tonsilla (Araneae: Amaurobiidae). Proceedings of the California Academy of Sciences 54: 499 - 662." type="journal article" year="2003">2003</bibRefCitation>
|
||
</taxonomicName>
|
||
). Specimens of
|
||
<taxonomicName id="4C35D0C71C5EA558FD497D79CC3041A7" authorityName="Okumura" authorityYear="2017" class="Arachnida" family="Agelenidae" genus="Dichodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94177561C5EA558FD497D79CC3041A7" italics="true" pageId="2" pageNumber="31">Dichodactylus</emphasis>
|
||
</taxonomicName>
|
||
also can be distinguished from
|
||
<taxonomicName id="4C35D0C71C5EA558FDF37D9ACFFD41A7" authorityName="Ovtchinnikov" authorityYear="1999" box="[615,733,393,417]" class="Arachnida" family="Amaurobiidae" genus="Tegecoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94177561C5EA558FDF37D9ACFFD41A7" box="[615,733,393,417]" italics="true" pageId="2" pageNumber="31">Tegecoelotes</emphasis>
|
||
</taxonomicName>
|
||
by a semi-circular or crescent-shaped median apophysis (
|
||
<figureCitation id="130EB7C11C5EA558FD437DB9CDF841E5" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="2" pageNumber="31">Fig. 1A, B</figureCitation>
|
||
), no or nipple-shaped epigynal teeth (
|
||
<figureCitation id="130EB7C11C5EA558FDF57DD8CFBA41E4" box="[609,666,458,483]" captionStart="Fig" captionStartId="5.[156,187,1156,1178]" captionTargetBox="[258,1330,205,1117]" captionTargetId="figure-449@5.[258,1330,205,1117]" captionTargetPageId="5" captionText="Fig. 5. Comparison of genitalic characters of three species of Dichodactylus gen. nov. from Japan: A, D, D. shinshuensis sp. nov., male (A: NSMT-Ar 13339), and female (D: NSMT-Ar 13341); B, E, D. tarumii (Arita, 1976), male (B: NSMT-Ar 14732), and female (E: NSMT-Ar 14733); C, F, D. satoi (Nishikawa, 2003), male (C: NSMT-Ar 14735), and female (F: NSMT-Ar 14734). A–C, retrolateral views of male palp; D–F, ventral views of epigynum. Abbreviations: CO, conductor; ET, epigynal teeth; LAM, lamellas; PA, patellar apophyses; SW, swollen portion of tegulum. Black circle on tibia shows the vestigial LTA." figureDoi="http://doi.org/10.5281/zenodo.4583728" httpUri="https://zenodo.org/record/4583728/files/figure.png" pageId="2" pageNumber="31">Fig. 5</figureCitation>
|
||
D–F) and anteroposteriorly elongated and convoluted spermathecae (
|
||
<figureCitation id="130EB7C11C5EA558FF377E1FCDD14222" box="[163,241,524,548]" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="2" pageNumber="31">Fig. 1D</figureCitation>
|
||
) (
|
||
<emphasis id="B94177561C5EA558FE9E7E1FCC054225" box="[266,293,524,547]" italics="true" pageId="2" pageNumber="31">vs.</emphasis>
|
||
hook-shaped median apophysis, broad epigynal teeth and posteriorly situated spermathecae in
|
||
<taxonomicName id="4C35D0C71C5EA558FD387E3ECCA54263" authority=": Wang 2002" authorityName=": Wang" authorityYear="2002" class="Arachnida" family="Amaurobiidae" genus="Tegecoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94177561C5EA558FD387E3ECDE84263" italics="true" pageId="2" pageNumber="31">Tegecoelotes</emphasis>
|
||
:
|
||
<bibRefCitation id="EFA4D6B51C5EA558FF477E5ECC674263" author="Wang, X. P." box="[211,327,589,613]" pageId="2" pageNumber="31" pagination="1 - 150" refId="ref4764" refString="Wang, X. P. 2002. A generic-level revision of the spider subfamily Coelotinae (Araneae, Amaurobiidae). Bulletin of the American Museum of Natural History 269: 1 - 150." type="journal article" year="2002">Wang 2002</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA4D6B51C5EA558FEC47E5ECCA54263" author="Wang, X. P." box="[336,389,589,613]" pageId="2" pageNumber="31" pagination="499 - 662" refId="ref4800" refString="Wang, X. P. 2003. Species revision of the coelotine spider genera Bifi- docoelotes, Coronilla, Draconarius, Femoracoelotes, Leptocoelotes, Longicoelotes, Platocoelotes, Spiricoelotes, Tegecoelotes, and Tonsilla (Araneae: Amaurobiidae). Proceedings of the California Academy of Sciences 54: 499 - 662." type="journal article" year="2003">2003</bibRefCitation>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4AFBCC1C5DA55BFF087B8BC9614818" ID-DOI="http://doi.org/10.5281/zenodo.4583719" ID-Zenodo-Dep="4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="1" pageNumber="30" startId="1.[156,187,1944,1966]" targetBox="[265,1322,597,1906]" targetPageId="1">
|
||
<paragraph id="8B8AAB441C5DA55BFF087B8BC9614818" blockId="1.[156,1432,1944,2078]" pageId="1" pageNumber="30">
|
||
Fig. 1. Comparison between the genera
|
||
<taxonomicName id="4C35D0C71C5DA55BFDB77B8BCF8147AB" authority="Okumura, 2017" authorityName="Okumura" authorityYear="2017" box="[547,673,1944,1965]" class="Arachnida" family="Agelenidae" genus="Dichodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94177561C5DA55BFDB77B8BCF8147AB" box="[547,673,1944,1965]" italics="true" pageId="1" pageNumber="30">Dichodactylus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A272CA2D1C5DA55BFD3C7B8BCFDC47AB" box="[680,764,1944,1965]" pageId="1" pageNumber="30" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C35D0C71C5DA55BFCBF7B8BC95447AB" authority="Kocak and Kemal, 2008" authorityName="Kocak and Kemal" authorityYear="2008" box="[811,1140,1944,1966]" class="Arachnida" family="Amaurobiidae" genus="Orumcekia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94177561C5DA55BFCBF7B8BCEAE47AB" box="[811,910,1944,1965]" italics="true" pageId="1" pageNumber="30">Orumcekia</emphasis>
|
||
Koçak and Kemal, 2008
|
||
</taxonomicName>
|
||
: A–D,
|
||
<taxonomicName id="4C35D0C71C5DA55BFB217B8BC8B447A8" authority="(Arita, 1974)" baseAuthorityName="Arita" baseAuthorityYear="1974" box="[1205,1428,1944,1966]" class="Arachnida" family="Amaurobiidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="tarumii">
|
||
<emphasis id="B94177561C5DA55BFB217B8BC83447AB" box="[1205,1300,1944,1965]" italics="true" pageId="1" pageNumber="30">D. tarumii</emphasis>
|
||
(Arita, 1974)
|
||
</taxonomicName>
|
||
, male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H,
|
||
<taxonomicName id="4C35D0C71C5DA55BFCAE7BA7C93747CC" authority="(Wang, 1994)" baseAuthorityName="Wang" baseAuthorityYear="1994" box="[826,1047,1972,1994]" class="Arachnida" family="Amaurobiidae" genus="Orumcekia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="gemata">
|
||
<emphasis id="B94177561C5DA55BFCAE7BA7CEB647CF" box="[826,918,1972,1993]" italics="true" pageId="1" pageNumber="30">O. gemata</emphasis>
|
||
(Wang, 1994)
|
||
</taxonomicName>
|
||
, male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32FF8CF1C5EA558FF217E7ECFD64536" pageId="2" pageNumber="31" type="description">
|
||
<paragraph id="8B8AAB441C5EA558FF217E7ECDF043CC" blockId="2.[156,768,198,1654]" pageId="2" pageNumber="31">
|
||
<emphasis id="B94177561C5EA558FF217E7ECC194283" bold="true" box="[181,313,621,645]" pageId="2" pageNumber="31">Description.</emphasis>
|
||
Small- to medium-sized coelotine spiders, 5.0–10.0 mm in length. Cheliceral promargin with three teeth, retromargin with four. Leg formula IV-I-II-III. Carapace brown or yellowish brown with grayish brown radial flecks; dorsum of abdomen grayish brown with yellowish brown chevrons or irregular spots, and venter yellowish brown with grayish brown irregular flecks; sternum yellowish brown and its marginal region brown; chelicerae and maxillae reddish brown; labium brown; legs yellowish brown with grayish brown ring flecks in femur, patella and tibia.
|
||
</paragraph>
|
||
<paragraph id="8B8AAB441C5EA558FF217FC0CFBC446B" blockId="2.[156,768,198,1654]" pageId="2" pageNumber="31">
|
||
<emphasis id="B94177561C5EA558FF217FC0CDD043ED" bold="true" box="[181,240,979,1003]" pageId="2" pageNumber="31">Male.</emphasis>
|
||
Palp (
|
||
<figureCitation id="130EB7C11C5EA558FEDC7FC0CC9E43ED" box="[328,446,979,1003]" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="2" pageNumber="31">Figs 1A, B</figureCitation>
|
||
,
|
||
<figureCitation id="130EB7C11C5EA558FE5A7FC0CCFB43ED" box="[462,475,979,1003]" captionStart="Fig" captionStartId="4.[156,187,1972,1994]" captionTargetBox="[264,1322,1212,1934]" captionTargetId="figure-80@4.[264,1322,1212,1934]" captionTargetPageId="4" captionText="Fig. 4. Dichodactylus satoi (Nishikawa, 2003). male (A, B: NSMT-Ar 14735), female (C, D: NSMT-Ar 14734). A, ventral view of male palp; B, retrolateral view of male palp; C, ventral view of epigynum; D, dorsal view of internal female genitalia. Abbreviations: CO, conductor; CY, cymbial furrow; EM, embolus; MA, median apophysis; PA, patellar apophyses; RTA, retrolateral tibial apophysis; SP, spermatheca; SH, spermathecal head. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4583726" httpUri="https://zenodo.org/record/4583726/files/figure.png" pageId="2" pageNumber="31">4</figureCitation>
|
||
A–C): two patellar apophyses present; LTA reduced; RTA large and elongated; cymbial furrow short, one-sixth or one-seventh of the cymbial length; median apophysis crescent-shaped or semicircular; conductor spatulate, resembles that of
|
||
<taxonomicName id="4C35D0C71C5EA558FDB47846CFB6446B" authorityName=": Wang" authorityYear="2002" box="[544,662,1109,1133]" class="Arachnida" family="Amaurobiidae" genus="Tegecoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94177561C5EA558FDB47846CFB6446B" box="[544,662,1109,1133]" italics="true" pageId="2" pageNumber="31">Tegecoelotes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8AAB441C5EA558FF217865CFD64536" blockId="2.[156,768,198,1654]" pageId="2" pageNumber="31">
|
||
<emphasis id="B94177561C5EA558FF217865CC274488" bold="true" box="[181,263,1142,1166]" pageId="2" pageNumber="31">Female.</emphasis>
|
||
Epigynum and internal genitalia (
|
||
<figureCitation id="130EB7C11C5EA558FD117865CFDB448B" box="[645,763,1142,1166]" captionStart="Fig" captionStartId="1.[156,187,1944,1966]" captionTargetBox="[265,1322,597,1906]" captionTargetId="figure-213@1.[265,1323,596,1906]" captionTargetPageId="1" captionText="Fig. 1. Comparison between the genera Dichodactylus gen. nov. and Orumcekia Koçak and Kemal, 2008: A–D, D. tarumii (Arita, 1974), male (A, B: NSMT-Ar 14732), and female (C, D: NSMT-Ar 14733); E–H, O. gemata (Wang, 1994), male (E, F: NSMT-Ar 14737), and female (G, H: NSMT-Ar 14736). A, E, ventral views of male palp; B, F, retrolateral views of male palp; C, G, ventral views of epigynum; D, H, dorsal views of internal female genitalia. Abbreviations: CON, conductor; CVA, conductor ventral apophysis; MA, median apophysis; PA, patellar apophyses; PEM, posterolaterally expanded margin; RTA, retrolateral tibial apophysis; SP, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4583719" httpUri="https://zenodo.org/record/4583719/files/figure.png" pageId="2" pageNumber="31">Figs 1C, D</figureCitation>
|
||
,
|
||
<figureCitation id="130EB7C11C5EA558FF087885CD8944A8" box="[156,169,1174,1198]" captionStart="Fig" captionStartId="4.[156,187,1972,1994]" captionTargetBox="[264,1322,1212,1934]" captionTargetId="figure-80@4.[264,1322,1212,1934]" captionTargetPageId="4" captionText="Fig. 4. Dichodactylus satoi (Nishikawa, 2003). male (A, B: NSMT-Ar 14735), female (C, D: NSMT-Ar 14734). A, ventral view of male palp; B, retrolateral view of male palp; C, ventral view of epigynum; D, dorsal view of internal female genitalia. Abbreviations: CO, conductor; CY, cymbial furrow; EM, embolus; MA, median apophysis; PA, patellar apophyses; RTA, retrolateral tibial apophysis; SP, spermatheca; SH, spermathecal head. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4583726" httpUri="https://zenodo.org/record/4583726/files/figure.png" pageId="2" pageNumber="31">4</figureCitation>
|
||
D–F): epigynum hexagonal or hourglass-shaped; epigynal teeth absent, or nipple-shaped and situated in posterior portion; epigynal hood absent; spermathecae elongated anteroposteriorly, convoluted and close to each other; spermathecal heads situated on anterolateral portion of spermathecae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32FF8CF1C5EA558FF21792ACF3445B4" pageId="2" pageNumber="31" type="discussion">
|
||
<paragraph id="8B8AAB441C5EA558FF21792ACCCE4557" blockId="2.[156,768,198,1654]" box="[181,494,1337,1361]" pageId="2" pageNumber="31">
|
||
<emphasis id="B94177561C5EA558FF21792ACC674557" bold="true" box="[181,327,1337,1361]" pageId="2" pageNumber="31">Composition.</emphasis>
|
||
Three species:
|
||
</paragraph>
|
||
<paragraph id="8B8AAB441C5EA558FF087949CFAF4577" blockId="2.[156,768,198,1654]" box="[156,655,1369,1394]" pageId="2" pageNumber="31">
|
||
1.
|
||
<taxonomicName id="4C35D0C71C5EA558FF21794ACF3B4577" authority="(Arita, 1976)" authorityName="Okumura" authorityYear="2017" baseAuthorityName="Arita" baseAuthorityYear="1976" box="[181,539,1369,1393]" class="Arachnida" family="Agelenidae" genus="Dichodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="tarumii" status="comb. nov.">
|
||
<emphasis id="B94177561C5EA558FF21794ACCB34577" box="[181,403,1369,1393]" italics="true" pageId="2" pageNumber="31">Dichodactylus tarumii</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA4D6B51C5EA558FE34794ACF334577" author="Arita, T." box="[416,531,1369,1393]" pageId="2" pageNumber="31" pagination="197 - 204" refId="ref4504" refString="Arita, T. 1976. Three new Coelotes (Araneae, Agelenidae) from Tottori Prefecture, west Japan. Annotationes Zoologicae Japonenses 49: 197 - 204." type="journal article" year="1976">Arita, 1976</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A272CA2D1C5EA558FDB5794ACFAF4577" box="[545,655,1369,1393]" pageId="2" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8B8AAB441C5EA558FF087969CF884597" blockId="2.[156,768,198,1654]" box="[156,680,1402,1426]" pageId="2" pageNumber="31">
|
||
2.
|
||
<taxonomicName id="4C35D0C71C5EA558FF217969CF144594" authority="(Nishikawa, 2003)" authorityName="Okumura" authorityYear="2017" baseAuthorityName="Nishikawa" baseAuthorityYear="2003" box="[181,564,1402,1426]" class="Arachnida" family="Agelenidae" genus="Dichodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="satoi" status="comb. nov.">
|
||
<emphasis id="B94177561C5EA558FF217969CC574594" box="[181,375,1402,1426]" italics="true" pageId="2" pageNumber="31">Dichodactylus satoi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA4D6B51C5EA558FE177969CF0E4594" author="Nishikawa, Y." box="[387,558,1402,1426]" pageId="2" pageNumber="31" pagination="37 - 40" refId="ref4624" refString="Nishikawa, Y. 2003. A new spider of the genus Coelotes (Araneae, Agel- enidae) from Gifu Prefecture, Central Japan. Special Bulletin of the Japanese Society of Coleopterology 6: 37 - 40." type="journal article" year="2003">Nishikawa, 2003</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A272CA2D1C5EA558FDAE7969CF884597" box="[570,680,1402,1426]" pageId="2" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8B8AAB441C5EA558FF087988CF3445B4" blockId="2.[156,768,198,1654]" box="[156,532,1434,1459]" pageId="2" pageNumber="31">
|
||
3.
|
||
<taxonomicName id="4C35D0C71C5EA558FF217989CCE245B4" authority="Okumura, 2017" authorityName="Okumura" authorityYear="2017" box="[181,450,1434,1458]" class="Arachnida" family="Agelenidae" genus="Dichodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="shinshuensis" status="sp. nov.">
|
||
<emphasis id="B94177561C5EA558FF217989CCE245B4" box="[181,450,1434,1458]" italics="true" pageId="2" pageNumber="31">Dichodactylus shinshuensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A272CA2D1C5EA558FE5C7988CF3445B4" box="[456,532,1435,1458]" pageId="2" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32FF8CF1C5EA558FF2179A8CF0145F5" pageId="2" pageNumber="31" type="distribution">
|
||
<paragraph id="8B8AAB441C5EA558FF2179A8CF0145F5" blockId="2.[156,768,198,1654]" pageId="2" pageNumber="31">
|
||
<emphasis id="B94177561C5EA558FF2179A8CC6145D5" bold="true" box="[181,321,1467,1491]" pageId="2" pageNumber="31">Distribution.</emphasis>
|
||
<collectingCountry id="F322EBD41C5EA558FECE79A8CCB245D4" box="[346,402,1467,1490]" name="Japan" pageId="2" pageNumber="31">Japan</collectingCountry>
|
||
(Chubu, Kinki and Chugoku Districts of Honshu, and Kyushu) (
|
||
<figureCitation id="130EB7C11C5EA558FE4D79C8CF3445F5" box="[473,532,1499,1523]" captionStart="Fig" captionStartId="6.[156,187,941,963]" captionTargetBox="[340,1247,205,902]" captionTargetId="figure-443@6.[340,1247,205,902]" captionTargetPageId="6" captionText="Fig. 6. Distribution of three species of the genus Dichodactylus gen. nov. from Japan. Closed circles denote D. shinshuensis sp. nov.; closed inverted triangles, D. tarumii (Arita, 1976); crosses, D. satoi (Nishikawa, 2003)." figureDoi="http://doi.org/10.5281/zenodo.4583730" httpUri="https://zenodo.org/record/4583730/files/figure.png" pageId="2" pageNumber="31">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32FF8CF1C5EA558FF2179EFCF244670" pageId="2" pageNumber="31" type="etymology">
|
||
<paragraph id="8B8AAB441C5EA558FF2179EFCF244670" blockId="2.[156,768,198,1654]" pageId="2" pageNumber="31">
|
||
<emphasis id="B94177561C5EA558FF2179EFCC0D4612" bold="true" box="[181,301,1532,1556]" pageId="2" pageNumber="31">Etymology.</emphasis>
|
||
The generic name is a compound noun derived from the Greek prefix dicho-"paired", and dactylus- "finger" referring to having two patellar apophyses in the male palp. The gender is masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |