treatments-xml/data/03/8F/3B/038F3B1FFFE0FF9EFE8BFC07982FFA1D.xml

<document id="30B8C376E1AD477E643212F46F692526" ID-CLB-Dataset="7097" ID-DOI="10.1051/acarologia/20122047" ID-GBIF-Dataset="2a27e96d-26db-479a-a450-396b8089a867" ID-ISSN="2107-7207" ID-Zenodo-Dep="4667079" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617736673151" checkinUser="carolina" docAuthor="Nazari, A., Khanjani, M. &amp; Kamali, K." docDate="2012" docId="038F3B1FFFE0FF9EFE8BFC07982FFA1D" docLanguage="en" docName="Acarologia.52.2.173-181.pdf" docOrigin="Acarologia 52 (2)" docSource="http://dx.doi.org/10.1051/acarologia/20122047" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Stigmaeus LADANAE N., n. sp." docType="treatment" docVersion="7" lastPageNumber="177" masterDocId="FFB64367FFE3FF9BFFB3FFC29D6CFFA5" masterDocTitle="Two New Eyeless Mite Species From The Western Provinces Of Iran: Stigmaeus Ladanae N. Sp. And Stigmaeus Nasrinae N. Sp. (Acari: Stigmaeidae)" masterLastPageNumber="181" masterPageNumber="173" pageNumber="175" updateTime="1698935987680" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:title id="2B4A14C8D182E2943E69FD58AE9848C7">Two New Eyeless Mite Species From The Western Provinces Of Iran: Stigmaeus Ladanae N. Sp. And Stigmaeus Nasrinae N. Sp. (Acari: Stigmaeidae)</mods:title>
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<mods:namePart id="3A179E209BE4BF83F9E9BDB264685463">Khanjani, M.</mods:namePart>
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<mods:namePart id="3F9986AE1BD7D296AD55E3F92BD49F6A">Kamali, K.</mods:namePart>
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<mods:title id="3E0F668820898210F294A85CA0A9D986">Acarologia</mods:title>
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<treatment id="038F3B1FFFE0FF9EFE8BFC07982FFA1D" ID-DOI="http://doi.org/10.5281/zenodo.4737648" ID-GBIF-Taxon="181872028" ID-Zenodo-Dep="4737648" LSID="urn:lsid:plazi:treatment:038F3B1FFFE0FF9EFE8BFC07982FFA1D" httpUri="http://treatment.plazi.org/id/038F3B1FFFE0FF9EFE8BFC07982FFA1D" lastPageId="5" lastPageNumber="177" pageId="3" pageNumber="175">
<subSubSection id="C33CD982FFE0FF98FE8BFC079F0DFC78" box="[312,609,964,989]" pageId="3" pageNumber="175" type="nomenclature">
<paragraph id="8B998A09FFE0FF98FE8BFC079F0DFC78" blockId="3.[312,609,964,1024]" box="[312,609,964,989]" pageId="3" pageNumber="175">
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<emphasis id="B952561BFFE0FF98FE8BFC079F0DFC78" bold="true" box="[312,609,964,989]" pageId="3" pageNumber="175">
<taxonomicName id="4C26F18AFFE0FF98FE8BFC079F74FC79" authority="ladanae n." authorityName="LADANAE N." box="[312,536,964,989]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="173" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B952561BFFE0FF98FE8BFC079F74FC79" bold="true" box="[312,536,964,989]" italics="true" pageId="3" pageNumber="175">Stigmaeus ladanae</emphasis>
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<taxonomicNameLabel id="A261EB60FFE0FF98FDADFC079F0DFC78" box="[542,609,965,989]" pageId="3" pageNumber="175" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8B998A09FFE0FF98FE22FC2A9F64FBA5" blockId="3.[312,609,964,1024]" box="[401,520,1000,1024]" pageId="3" pageNumber="175">
<emphasis id="B952561BFFE0FF98FE22FC2A9F64FBA5" bold="true" box="[401,520,1000,1024]" pageId="3" pageNumber="175">
(
<figureCitation id="131D968CFFE0FF98FE2AFC2A9C92FBA5" box="[409,510,1000,1024]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStartId-0="2.[392,404,1883,1902]" captionStartId-1="4.[397,409,1695,1714]" captionTargetBox-0="[334,1266,340,1852]" captionTargetBox-1="[242,1345,302,1662]" captionTargetId-0="figure-4@2.[322,1266,340,1852]" captionTargetId-1="figure-4@4.[242,1345,302,1663]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 1: Stigmaeus ladanae n. sp. (female): A – Dorsum; B – Venter; C – gnathosoma." captionText-1="FIGURE 2: Stigmaeus ladanae n. sp. (female): A – leg I; B – leg II; C – leg III; D – leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.4667082" figureDoi-1="http://doi.org/10.5281/zenodo.4667084" httpUri-0="https://zenodo.org/record/4667082/files/figure.png" httpUri-1="https://zenodo.org/record/4667084/files/figure.png" pageId="3" pageNumber="175">Figs. 1-2</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B998A09FFE0FF98FF76FBE69DABFADD" blockId="3.[159,762,1060,1949]" pageId="3" pageNumber="175">
Diagnosis — All dorsal, ventral and suranal shields reticulated, dorsum with 14 pairs of setae (
<emphasis id="B952561BFFE0FF98FF1BFBA89DD2FB21" box="[168,190,1130,1156]" italics="true" pageId="3" pageNumber="175">
h 
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present), eyes and post ocular bodies absent, median hysterosomal shield with two setae (
<emphasis id="B952561BFFE0FF98FD1CFB4C9FAEFB02" box="[687,706,1166,1191]" italics="true" pageId="3" pageNumber="175">
c 
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</emphasis>
, 
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d 
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</emphasis>
), genital setae two pairs, aggenital setae four pairs, genua 5(+1 
<emphasis id="B952561BFFE0FF98FE97FB119C5FFB4E" box="[292,307,1235,1259]" italics="true" pageId="3" pageNumber="175">κ</emphasis>
) - 5 - 2 - 2, propodosomal setae 
<emphasis id="B952561BFFE0FF98FD76FB169F96FB4E" box="[709,762,1236,1259]" italics="true" pageId="3" pageNumber="175">ve/vi</emphasis>
1.26 (1.29 – 1.35), ratio 
<emphasis id="B952561BFFE0FF98FE7AFB359F7FFAB5" box="[457,531,1271,1296]" italics="true" pageId="3" pageNumber="175">
<subScript id="17A2884CFFE0FF98FE7AFB359CB7FAB5" attach="left" box="[457,475,1271,1296]" fontSize="7" pageId="3" pageNumber="175">vi</subScript>
/ 
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</emphasis>
0.85 (0.54 – 0.80), 
<emphasis id="B952561BFFE0FF98FF2CFAD89DDFFA91" box="[159,179,1306,1332]" italics="true" pageId="3" pageNumber="175">
c 
<subScript id="17A2884CFFE0FF98FF19FAE19DDFFA91" attach="left" box="[170,179,1315,1332]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
/ 
<emphasis id="B952561BFFE0FF98FF77FAD89D9AFA91" box="[196,246,1306,1332]" italics="true" pageId="3" pageNumber="175">
c 
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-c 
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</emphasis>
0.52 (0.46 – 0.51); 
<emphasis id="B952561BFFE0FF98FE6FFAD89F63FA91" box="[476,527,1306,1332]" italics="true" pageId="3" pageNumber="175">
c 
<subScript id="17A2884CFFE0FF98FE54FAE19C9CFA91" attach="left" box="[487,496,1315,1332]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-c 
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</emphasis>
: 
<emphasis id="B952561BFFE0FF98FD95FADB9F31FA91" box="[550,605,1305,1332]" italics="true" pageId="3" pageNumber="175">
d 
<subScript id="17A2884CFFE0FF98FD80FAE19F50FA91" attach="left" box="[563,572,1315,1332]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-d 
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</emphasis>
: 
<emphasis id="B952561BFFE0FF98FDC7FAD89FC9FA91" box="[628,677,1306,1332]" italics="true" pageId="3" pageNumber="175">
e 
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-e 
<subScript id="17A2884CFFE0FF98FD2FFAE19FC9FA91" attach="left" box="[668,677,1315,1332]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
: 
<emphasis id="B952561BFFE0FF98FD0FFADB9F8BFA91" box="[700,743,1305,1332]" italics="true" pageId="3" pageNumber="175">
f 
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-f 
<subScript id="17A2884CFFE0FF98FD6DFAE19F8BFA91" attach="left" box="[734,743,1315,1332]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
: 0.87 (1 – 1.86): 0.89 (0.81 – 1.60): 0.79 (0.81 – 1.09): 1.0.
</paragraph>
<paragraph id="8B998A09FFE0FF98FF76FA4C9FB2F88D" blockId="3.[159,762,1060,1949]" pageId="3" pageNumber="175">
Material examined — 
<typeStatus id="549D34ABFFE0FF98FE54FA4C9F3AFA03" box="[487,598,1422,1446]" pageId="3" pageNumber="175" type="holotype">Holotype</typeStatus>
female, collected from soil beneath apple trees, 
<taxonomicName id="4C26F18AFFE0FF98FDECFA739CDBFA49" authority="Borkh." authorityName="Borkh." class="Magnoliopsida" family="Rosaceae" genus="Malus" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="175" phylum="Tracheophyta" rank="species" species="domestica">
<emphasis id="B952561BFFE0FF98FDECFA739DAAFA49" italics="true" pageId="3" pageNumber="175">Malus domestica</emphasis>
Borkh. (Rosaceae)
</taxonomicName>
, 
<collectingRegion id="49E244EBFFE0FF98FE7AFA169F52FA49" box="[457,574,1492,1516]" country="Iran" name="Hamadan" pageId="3" pageNumber="175">Hamedan</collectingRegion>
(34º46’48.13&quot;N, 48º28’11.46&quot;E and altitude 
<quantity id="4CDE27ECFFE0FF98FE6DFA359F58F9AA" box="[478,564,1527,1551]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="3" pageNumber="175" unit="m" value="1930.0">1930 m</quantity>
a.s.l), 
<collectingRegion id="49E244EBFFE0FF98FD36FA359C65F997" country="Iran" name="Hamadan" pageId="3" pageNumber="175">Hamedan Province</collectingRegion>
, 
<collectingCountry id="F331CA99FFE0FF98FEAAF9D99C27F997" box="[281,331,1563,1586]" name="Iran" pageId="3" pageNumber="175">Iran</collectingCountry>
, 
<date id="FF98ACC9FFE0FF98FEEFF9D89C81F997" box="[348,493,1562,1586]" pageId="3" pageNumber="175" value="2011-06-08">8 June 2011</date>
, Alireza Nazari; 
<specimenCount id="9D204180FFE0FF98FD70F9D89D8FF9F0" pageId="3" pageNumber="175" type="female">6 females</specimenCount>
<typeStatus id="549D34ABFFE0FF98FF58F9FC9C33F9F3" box="[235,351,1598,1622]" pageId="3" pageNumber="175" type="paratype">paratypes</typeStatus>
with the same data. The 
<typeStatus id="549D34ABFFE0FF98FD22F9FF9F95F9F0" box="[657,761,1597,1621]" pageId="3" pageNumber="175" type="holotype">holotype</typeStatus>
female and 
<specimenCount id="9D204180FFE0FF98FE81F9A29CD3F9DC" box="[306,447,1632,1657]" pageId="3" pageNumber="175" type="generic" typeStatus="paratypes">5 paratypes</specimenCount>
females are deposited as slide-mounted specimens in the Collection of the Acarology Laboratory, University of Bu-Ali 
<taxonomicName id="4C26F18AFFE0FF98FD73F9659F99F91A" box="[704,757,1703,1727]" class="Insecta" genus="Sina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="173" phylum="Arthropoda" rank="genus">Sina</taxonomicName>
, 
<collectingRegion id="49E244EBFFE0FF98FF2CF9089C7AF947" box="[159,278,1738,1762]" country="Iran" name="Hamadan" pageId="3" pageNumber="175">Hamadan</collectingRegion>
, 
<collectingCountry id="F331CA99FFE0FF98FE9BF9089C34F944" box="[296,344,1738,1761]" name="Iran" pageId="3" pageNumber="175">Iran</collectingCountry>
and one 
<typeStatus id="549D34ABFFE0FF98FE67F9089F51F947" box="[468,573,1738,1762]" pageId="3" pageNumber="175" type="paratype">paratype</typeStatus>
female will be deposited in the National Collection of 
<taxonomicName id="4C26F18AFFE0FF98FDC5F92F9C37F88D" authority=", Plant Protection" authorityName="Plant Protection" class="Arachnida" kingdom="Animalia" pageId="3" pageNumber="175" phylum="Arthropoda" rank="class">Arachnida, Plant Protection</taxonomicName>
Research, Pretoria, 
<collectingCountry id="F331CA99FFE0FF98FDF1F8D29FB6F88D" box="[578,730,1808,1832]" name="South Africa" pageId="3" pageNumber="175">South Africa</collectingCountry>
.
</paragraph>
<paragraph id="8B998A09FFE0FF98FF76F8FC9857FE1D" blockId="3.[159,762,1060,1949]" lastBlockId="3.[826,1429,311,440]" pageId="3" pageNumber="175">
Female (n = 7) — Color in life red. Idiosoma oval. Measurements of 
<typeStatus id="549D34ABFFE0FF98FE67F8A39F50F8DC" box="[468,572,1889,1913]" pageId="3" pageNumber="175" type="holotype">holotype</typeStatus>
with measurements of 
<typeStatus id="549D34ABFFE0FF98FEBEF8479CEDF838" box="[269,385,1925,1949]" pageId="3" pageNumber="175" type="paratype">paratypes</typeStatus>
in parentheses: Length of body (excluding gnathosoma) 420 (419 – 441), (including gnathosoma) 523 (510 – 548); width 206 (206 – 245); length of leg I 240 (218 – 242); leg II 188 (175 – 193); leg III 185 (153 – 190), leg IV 223 (208 – 246).
</paragraph>
<paragraph id="8B998A09FFE0FF98FCD3FE1798E2FA66" blockId="3.[826,1429,469,1475]" pageId="3" pageNumber="175">
Dorsum (
<figureCitation id="131D968CFFE0FF98FC67FE179955FE48" box="[980,1081,469,493]" captionStart="FIGURE" captionStartId="2.[392,404,1883,1902]" captionTargetBox="[334,1266,340,1852]" captionTargetId="figure-4@2.[322,1266,340,1852]" captionTargetPageId="2" captionText="FIGURE 1: Stigmaeus ladanae n. sp. (female): A – Dorsum; B – Venter; C – gnathosoma." figureDoi="http://doi.org/10.5281/zenodo.4667082" httpUri="https://zenodo.org/record/4667082/files/figure.png" pageId="3" pageNumber="175">Fig. 1A</figureCitation>
) — Prodorsal shield oblong and reticulated; bearing three pairs of setae (
<emphasis id="B952561BFFE0FF98FAE1FE3B9E3BFD96" italics="true" pageId="3" pageNumber="175">vi, ve, sci</emphasis>
), eyes and post ocular bodies absent; dorsal hysterosoma with 10 pairs of setae, almost smooth, four pairs of paired and three unpaired shields and surrounded with entire and reticulated (
<figureCitation id="131D968CFFE0FF98FB58FD46982FFD39" box="[1259,1347,644,668]" captionStart="FIGURE" captionStartId="2.[392,404,1883,1902]" captionTargetBox="[334,1266,340,1852]" captionTargetId="figure-4@2.[322,1266,340,1852]" captionTargetPageId="2" captionText="FIGURE 1: Stigmaeus ladanae n. sp. (female): A – Dorsum; B – Venter; C – gnathosoma." figureDoi="http://doi.org/10.5281/zenodo.4667082" httpUri="https://zenodo.org/record/4667082/files/figure.png" pageId="3" pageNumber="175">Fig. 1A</figureCitation>
). Setae 
<emphasis id="B952561BFFE0FF98FC89FD6A9E22FD67" box="[826,846,680,706]" italics="true" pageId="3" pageNumber="175">
c 
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</emphasis>
situated laterally between coxae II-III. Setae 
<emphasis id="B952561BFFE0FF98FAEAFD6A981DFD1A" box="[1369,1393,680,703]" italics="true" pageId="3" pageNumber="175">ve</emphasis>
almost as long as or slightly longer than setae 
<emphasis id="B952561BFFE0FF98FAE3FD099809FD47" box="[1360,1381,715,738]" italics="true" pageId="3" pageNumber="175">vi</emphasis>
, setae 
<emphasis id="B952561BFFE0FF98FCD0FD2C9EEEFCA0" box="[867,898,750,773]" italics="true" pageId="3" pageNumber="175">sce</emphasis>
on lateral propodosomal shields, reticulated; lengths of dorsal setae: 
<emphasis id="B952561BFFE0FF98FBE8FCD3991DFC8E" box="[1115,1137,785,811]" italics="true" pageId="3" pageNumber="175">
v 
<subScript id="17A2884CFFE0FF98FBDBFCD8991DFC8E" attach="left" box="[1128,1137,794,811]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
23 (20 – 24), 
<emphasis id="B952561BFFE0FF98FAA5FCD39842FC8D" box="[1302,1326,785,808]" italics="true" pageId="3" pageNumber="175">ve</emphasis>
29 (27 – 31), 
<emphasis id="B952561BFFE0FF98FCDFFCF69EE7FCEB" box="[876,907,820,846]" italics="true" pageId="3" pageNumber="175">
sc 
<subScript id="17A2884CFFE0FF98FC31FCFF9EE7FCEB" attach="left" box="[898,907,829,846]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
28 (22 – 29), 
<emphasis id="B952561BFFE0FF98FB98FCF69925FCEB" box="[1067,1097,820,846]" italics="true" pageId="3" pageNumber="175">
sc 
<subScript id="17A2884CFFE0FF98FBF3FCFF9925FCEB" attach="left" box="[1088,1097,829,846]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
30 (28 – 32), 
<emphasis id="B952561BFFE0FF98FB5AFCF69991FCEB" box="[1257,1277,820,846]" italics="true" pageId="3" pageNumber="175">
c 
<subScript id="17A2884CFFE0FF98FB47FCFF9991FCEB" attach="left" box="[1268,1277,829,846]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
25 (20 – 27), 
<emphasis id="B952561BFFE0FF98FC89FC959E22FCD4" box="[826,846,855,881]" italics="true" pageId="3" pageNumber="175">
c 
<subScript id="17A2884CFFE0FF98FCF6FCA29E22FCD4" attach="left" box="[837,846,864,881]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
41 (32 – 43), 
<emphasis id="B952561BFFE0FF98FC5CFC959969FCD4" box="[1007,1029,855,881]" italics="true" pageId="3" pageNumber="175">
d 
<subScript id="17A2884CFFE0FF98FC4FFCA29969FCD4" attach="left" box="[1020,1029,864,881]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
19 (19 – 24), 
<emphasis id="B952561BFFE0FF98FB15FC9599D0FCD4" box="[1190,1212,855,881]" italics="true" pageId="3" pageNumber="175">
d 
<subScript id="17A2884CFFE0FF98FB00FCA299D0FCD4" attach="left" box="[1203,1212,864,881]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
21 (20 – 24), 
<geoCoordinate id="EE12ECCEFFE0FF98FAEEFC9598F9FCCA" box="[1373,1429,855,881]" degrees="1" direction="east" minutes="22" orientation="longitude" pageId="3" pageNumber="175" precision="925" value="1.3666667">
<emphasis id="B952561BFFE0FF98FAEEFC95981CFCD4" box="[1373,1392,855,881]" italics="true" pageId="3" pageNumber="175">
e 
<subScript id="17A2884CFFE0FF98FAD4FCA2981CFCD4" attach="left" box="[1383,1392,864,881]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
22
</geoCoordinate>
(20 – 24), 
<geoCoordinate id="EE12ECCEFFE0FF98FC1CFCB89E84FC37" box="[943,1000,890,916]" degrees="2" direction="east" minutes="21" orientation="longitude" pageId="3" pageNumber="175" precision="925" value="2.35">
<emphasis id="B952561BFFE0FF98FC1CFCB89EAEFC31" box="[943,962,890,916]" italics="true" pageId="3" pageNumber="175">
e 
<subScript id="17A2884CFFE0FF98FC0AFC419EAEFC31" attach="left" box="[953,962,899,916]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
21
</geoCoordinate>
(21 – 24), 
<emphasis id="B952561BFFE0FF98FBD6FCB89919FC31" box="[1125,1141,890,916]" italics="true" pageId="3" pageNumber="175">
f 
<subScript id="17A2884CFFE0FF98FBDFFC419919FC31" attach="left" box="[1132,1141,899,916]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
30 (28 – 33), 
<emphasis id="B952561BFFE0FF98FAABFCB89842FC31" box="[1304,1326,890,916]" italics="true" pageId="3" pageNumber="175">
h 
<subScript id="17A2884CFFE0FF98FA96FC419842FC31" attach="left" box="[1317,1326,899,916]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
32 (28 – 37), 
<emphasis id="B952561BFFE0FF98FCDFFC5F9EEFFC12" box="[876,899,925,951]" italics="true" pageId="3" pageNumber="175">
h 
<subScript id="17A2884CFFE0FF98FCC9FC649EEFFC12" attach="left" box="[890,899,934,951]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
40 (37 – 44); distances between dorsal setae: 
<emphasis id="B952561BFFE0FF98FC89FC039E00FC7D" box="[826,876,961,984]" italics="true" pageId="3" pageNumber="175">vi-vi</emphasis>
26 (25 – 44), 
<emphasis id="B952561BFFE0FF98FBB1FC039956FC7D" box="[1026,1082,961,984]" italics="true" pageId="3" pageNumber="175">ve-ve</emphasis>
50 (46 – 55), 
<emphasis id="B952561BFFE0FF98FB63FC039869FC7D" box="[1232,1285,961,984]" italics="true" pageId="3" pageNumber="175">vi-ve</emphasis>
32 (25 – 34), 
<emphasis id="B952561BFFE0FF98FC89FC269E10FC5E" box="[826,892,995,1019]" italics="true" pageId="3" pageNumber="175">sci -sci</emphasis>
85 (73 – 88), 
<emphasis id="B952561BFFE0FF98FB93FC269904FC5E" box="[1056,1128,996,1019]" italics="true" pageId="3" pageNumber="175">sce-sce</emphasis>
135 (131 – 152), 
<emphasis id="B952561BFFE0FF98FA87FC26981DFC5E" box="[1332,1393,996,1019]" italics="true" pageId="3" pageNumber="175">ve-sci</emphasis>
47 (41 – 50), 
<emphasis id="B952561BFFE0FF98FC1EFBC59E9EFBBB" box="[941,1010,1031,1054]" italics="true" pageId="3" pageNumber="175">sci-sce</emphasis>
25 (18 – 34), 
<emphasis id="B952561BFFE0FF98FB23FBC599AEFB85" box="[1168,1218,1031,1056]" italics="true" pageId="3" pageNumber="175">
c 
<subScript id="17A2884CFFE0FF98FB29FBCD99CFFB85" attach="left" box="[1178,1187,1039,1056]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-c 
<subScript id="17A2884CFFE0FF98FB0AFBCD99AEFB85" attach="left" box="[1209,1218,1039,1056]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
46 (43 – 53), 
<emphasis id="B952561BFFE0FF98FAD2FBC598FFFB85" box="[1377,1427,1031,1056]" italics="true" pageId="3" pageNumber="175">
c 
<subScript id="17A2884CFFE0FF98FADFFBCD9819FB85" attach="left" box="[1388,1397,1039,1056]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-c 
<subScript id="17A2884CFFE0FF98FA39FBCD98FFFB85" attach="left" box="[1418,1427,1039,1056]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
73 (57 – 89), 
<emphasis id="B952561BFFE0FF98FC63FBE8996EFBE1" box="[976,1026,1066,1092]" italics="true" pageId="3" pageNumber="175">
c 
<subScript id="17A2884CFFE0FF98FC69FBF19E8FFBE1" attach="left" box="[986,995,1075,1092]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
-c 
<subScript id="17A2884CFFE0FF98FC4AFBF1996EFBE1" attach="left" box="[1017,1026,1075,1092]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
191 (170 – 206), 
<emphasis id="B952561BFFE0FF98FB7AFBE89992FBE1" box="[1225,1278,1065,1092]" italics="true" pageId="3" pageNumber="175">
c 
<subScript id="17A2884CFFE0FF98FB67FBF199B1FBE1" attach="left" box="[1236,1245,1075,1092]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-d 
<subScript id="17A2884CFFE0FF98FB46FBF19992FBE1" attach="left" box="[1269,1278,1075,1092]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
59 (48 – 64), 
<emphasis id="B952561BFFE0FF98FC89FB8E9E1DFBC2" box="[826,881,1100,1127]" italics="true" pageId="3" pageNumber="175">
d 
<subScript id="17A2884CFFE0FF98FCF4FB949E3CFBC2" attach="left" box="[839,848,1110,1127]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-d 
<subScript id="17A2884CFFE0FF98FCDBFB949E1DFBC2" attach="left" box="[872,881,1110,1127]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
47 (37 – 43), 
<emphasis id="B952561BFFE0FF98FBB5FB8E9951FBC2" box="[1030,1085,1100,1127]" italics="true" pageId="3" pageNumber="175">
d 
<subScript id="17A2884CFFE0FF98FBA0FB949970FBC2" attach="left" box="[1043,1052,1110,1127]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-d 
<subScript id="17A2884CFFE0FF98FB87FB949951FBC2" attach="left" box="[1076,1085,1110,1127]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
49 (44 – 56), 
<emphasis id="B952561BFFE0FF98FB61FB8E986AFBC2" box="[1234,1286,1100,1127]" italics="true" pageId="3" pageNumber="175">
d 
<subScript id="17A2884CFFE0FF98FB6CFB949984FBC2" attach="left" box="[1247,1256,1110,1127]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-e 
<subScript id="17A2884CFFE0FF98FB4EFB94986AFBC2" attach="left" box="[1277,1286,1110,1127]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
68 (62 – 71), 
<emphasis id="B952561BFFE0FF98FC89FBAD9E02FB2F" box="[826,878,1135,1162]" italics="true" pageId="3" pageNumber="175">
d 
<subScript id="17A2884CFFE0FF98FCF4FBBB9E3CFB2F" attach="left" box="[839,848,1145,1162]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
- 
<geoCoordinate id="EE12ECCEFFE0FF98FCE8FBB29E09FB22" box="[859,869,1136,1159]" degrees="2" direction="east" minutes="51" orientation="longitude" pageId="3" pageNumber="175" precision="925" value="2.85">e</geoCoordinate>
<subScript id="17A2884CFFE0FF98FCD6FBBB9E02FB2F" attach="left" box="[869,878,1145,1162]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
51 (48 – 69), 
<emphasis id="B952561BFFE0FF98FBBFFBAD992CFB2F" box="[1036,1088,1135,1162]" italics="true" pageId="3" pageNumber="175">
d 
<subScript id="17A2884CFFE0FF98FBA9FBBB994FFB2F" attach="left" box="[1050,1059,1145,1162]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
-e 
<subScript id="17A2884CFFE0FF98FB84FBBB992CFB2F" attach="left" box="[1079,1088,1145,1162]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
73 (71 – 77), 
<emphasis id="B952561BFFE0FF98FB6CFBAD987AFB2F" box="[1247,1302,1135,1162]" italics="true" pageId="3" pageNumber="175">
d 
<subScript id="17A2884CFFE0FF98FB5FFBBB9999FB2F" attach="left" box="[1260,1269,1145,1162]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
-d 
<subScript id="17A2884CFFE0FF98FABEFBBB987AFB2F" attach="left" box="[1293,1302,1145,1162]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
128 (118 – 151), 
<emphasis id="B952561BFFE0FF98FCCBFB519EC5FB08" box="[888,937,1171,1197]" italics="true" pageId="3" pageNumber="175">
e 
<subScript id="17A2884CFFE0FF98FC31FB5E9EE7FB08" attach="left" box="[898,907,1180,1197]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
- 
<geoCoordinate id="EE12ECCEFFE0FF98FC25FB519ECCFB0F" box="[918,928,1171,1194]" degrees="1" direction="east" minutes="42" orientation="longitude" pageId="3" pageNumber="175" precision="925" value="1.7">e</geoCoordinate>
<subScript id="17A2884CFFE0FF98FC13FB5E9EC5FB08" attach="left" box="[928,937,1180,1197]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
42 (25 – 43), 
<emphasis id="B952561BFFE0FF98FBF2FB51991EFB08" box="[1089,1138,1171,1197]" italics="true" pageId="3" pageNumber="175">
e 
<subScript id="17A2884CFFE0FF98FBF8FB5E9938FB08" attach="left" box="[1099,1108,1180,1197]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
-e 
<subScript id="17A2884CFFE0FF98FBDAFB5E991EFB08" attach="left" box="[1129,1138,1180,1197]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
94 (87 – 121), 
<emphasis id="B952561BFFE0FF98FAA4FB519824FB08" box="[1303,1352,1171,1197]" italics="true" pageId="3" pageNumber="175">
e 
<subScript id="17A2884CFFE0FF98FA92FB5E9846FB08" attach="left" box="[1313,1322,1180,1197]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
- 
<geoCoordinate id="EE12ECCEFFE0FF98FA86FB519853FB0F" box="[1333,1343,1171,1194]" degrees="2" direction="east" minutes="33" orientation="longitude" pageId="3" pageNumber="175" precision="925" value="2.55">e</geoCoordinate>
<subScript id="17A2884CFFE0FF98FA8CFB5E9824FB08" attach="left" box="[1343,1352,1180,1197]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
33 (29 – 40), 
<emphasis id="B952561BFFE0FF98FC32FB749EC3FB75" box="[897,943,1205,1232]" italics="true" pageId="3" pageNumber="175">
e 
<subScript id="17A2884CFFE0FF98FC38FB7D9EF8FB75" attach="left" box="[907,916,1215,1232]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-f 
<subScript id="17A2884CFFE0FF98FC15FB7D9EC3FB75" attach="left" box="[934,943,1215,1232]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
40 (30 – 42), 
<emphasis id="B952561BFFE0FF98FBFFFB77991BFB75" box="[1100,1143,1205,1232]" italics="true" pageId="3" pageNumber="175">
f 
<subScript id="17A2884CFFE0FF98FBE7FB7D9931FB75" attach="left" box="[1108,1117,1215,1232]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-f 
<subScript id="17A2884CFFE0FF98FBDDFB7D991BFB75" attach="left" box="[1134,1143,1215,1232]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
53 (23 – 53), 
<emphasis id="B952561BFFE0FF98FAA6FB77982AFB75" box="[1301,1350,1205,1232]" italics="true" pageId="3" pageNumber="175">
f 
<subScript id="17A2884CFFE0FF98FAAFFB7D9849FB75" attach="left" box="[1308,1317,1215,1232]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-h 
<subScript id="17A2884CFFE0FF98FA8EFB7D982AFB75" attach="left" box="[1341,1350,1215,1232]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
36 (32 – 45), 
<emphasis id="B952561BFFE0FF98FC37FB1A9ED7FB56" box="[900,955,1240,1267]" italics="true" pageId="3" pageNumber="175">
h 
<subScript id="17A2884CFFE0FF98FC21FB209EF7FB56" attach="left" box="[914,923,1250,1267]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-h 
<subScript id="17A2884CFFE0FF98FC01FB209ED7FB56" attach="left" box="[946,955,1250,1267]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
33 (31 – 38), 
<emphasis id="B952561BFFE0FF98FBD2FB1A99FEFB56" box="[1121,1170,1240,1267]" italics="true" pageId="3" pageNumber="175">
f 
<subScript id="17A2884CFFE0FF98FBDAFB20991EFB56" attach="left" box="[1129,1138,1250,1267]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-h 
<subScript id="17A2884CFFE0FF98FB3AFB2099FEFB56" attach="left" box="[1161,1170,1250,1267]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
39 (36 – 43), 
<emphasis id="B952561BFFE0FF98FA8BFB1A9803FB56" box="[1336,1391,1240,1267]" italics="true" pageId="3" pageNumber="175">
h 
<subScript id="17A2884CFFE0FF98FAF6FB209822FB56" attach="left" box="[1349,1358,1250,1267]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
-h 
<subScript id="17A2884CFFE0FF98FAD5FB209803FB56" attach="left" box="[1382,1391,1250,1267]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
71 (64 – 72), 
<emphasis id="B952561BFFE0FF98FC1DFB399E89FAB3" box="[942,997,1275,1302]" italics="true" pageId="3" pageNumber="175">
h 
<subScript id="17A2884CFFE0FF98FC08FAC79EA8FAB3" attach="left" box="[955,964,1285,1302]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-h 
<subScript id="17A2884CFFE0FF98FC6FFAC79E89FAB3" attach="left" box="[988,997,1285,1302]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
17 (15 – 21); ratio: 
<emphasis id="B952561BFFE0FF98FB7CFB3E9873FAB6" box="[1231,1311,1276,1299]" italics="true" pageId="3" pageNumber="175">vi/vi-vi</emphasis>
0.88 (0.54 – 0.80), 
<emphasis id="B952561BFFE0FF98FC20FADD9ECBFA9C" box="[915,935,1311,1337]" italics="true" pageId="3" pageNumber="175">
c 
<subScript id="17A2884CFFE0FF98FC2DFAEA9ECBFA9C" attach="left" box="[926,935,1320,1337]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
/ 
<emphasis id="B952561BFFE0FF98FC0AFADD9E87FA9C" box="[953,1003,1311,1337]" italics="true" pageId="3" pageNumber="175">
c 
<subScript id="17A2884CFFE0FF98FC70FAEA9EA0FA9C" attach="left" box="[963,972,1320,1337]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-c 
<subScript id="17A2884CFFE0FF98FC51FAEA9E87FA9C" attach="left" box="[994,1003,1320,1337]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
0.52 (0.46 – 0.51), 
<emphasis id="B952561BFFE0FF98FB73FADC99BBFA9C" box="[1216,1239,1310,1337]" italics="true" pageId="3" pageNumber="175">
d 
<subScript id="17A2884CFFE0FF98FB7DFAEA99BBFA9C" attach="left" box="[1230,1239,1320,1337]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
/ 
<emphasis id="B952561BFFE0FF98FB5BFADC9873FA9C" box="[1256,1311,1310,1337]" italics="true" pageId="3" pageNumber="175">
d 
<subScript id="17A2884CFFE0FF98FB45FAEA9993FA9C" attach="left" box="[1270,1279,1320,1337]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-d 
<subScript id="17A2884CFFE0FF98FAA5FAEA9873FA9C" attach="left" box="[1302,1311,1320,1337]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
0.40 (0.51 – 0.55), 
<emphasis id="B952561BFFE0FF98FC2AFA809EC0FAF9" box="[921,940,1346,1372]" italics="true" pageId="3" pageNumber="175">
e 
<subScript id="17A2884CFFE0FF98FC10FA899EC0FAF9" attach="left" box="[931,940,1355,1372]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
/ 
<emphasis id="B952561BFFE0FF98FC0DFA809E83FAF9" box="[958,1007,1346,1372]" italics="true" pageId="3" pageNumber="175">
e 
<subScript id="17A2884CFFE0FF98FC7BFA899EBDFAF9" attach="left" box="[968,977,1355,1372]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
- 
<geoCoordinate id="EE12ECCEFFE0FF98FC68FA809E89FAFC" box="[987,997,1346,1369]" degrees="1" direction="east" minutes="0.5" orientation="longitude" pageId="3" pageNumber="175" precision="92" value="1.0083333">e</geoCoordinate>
<subScript id="17A2884CFFE0FF98FC55FA899E83FAF9" attach="left" box="[998,1007,1355,1372]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
0.5 (0.55 – 0.8), 
<emphasis id="B952561BFFE0FF98FB05FA8099ABFAF9" box="[1206,1223,1346,1372]" italics="true" pageId="3" pageNumber="175">
f 
<subScript id="17A2884CFFE0FF98FB0DFA8999ABFAF9" attach="left" box="[1214,1223,1355,1372]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
/ 
<emphasis id="B952561BFFE0FF98FB6AFA809868FAF9" box="[1241,1284,1346,1372]" italics="true" pageId="3" pageNumber="175">
f 
<subScript id="17A2884CFFE0FF98FB53FA899985FAF9" attach="left" box="[1248,1257,1355,1372]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-f 
<subScript id="17A2884CFFE0FF98FB48FA899868FAF9" attach="left" box="[1275,1284,1355,1372]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
0.52 (0.62 – 1.21), 
<emphasis id="B952561BFFE0FF98FCCDFAA79EF8FADA" box="[894,916,1381,1407]" italics="true" pageId="3" pageNumber="175">
h 
<subScript id="17A2884CFFE0FF98FC38FAAC9EF8FADA" attach="left" box="[907,916,1390,1407]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
/ 
<emphasis id="B952561BFFE0FF98FC15FAA79EB1FADA" box="[934,989,1381,1407]" italics="true" pageId="3" pageNumber="175">
h 
<subScript id="17A2884CFFE0FF98FC00FAAC9ED0FADA" attach="left" box="[947,956,1390,1407]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-h 
<subScript id="17A2884CFFE0FF98FC67FAAC9EB1FADA" attach="left" box="[980,989,1390,1407]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
0.96 (0.90 – 0.97), 
<emphasis id="B952561BFFE0FF98FB1CFAA799A9FADA" box="[1199,1221,1381,1407]" italics="true" pageId="3" pageNumber="175">
h 
<subScript id="17A2884CFFE0FF98FB0FFAAC99A9FADA" attach="left" box="[1212,1221,1390,1407]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
/ 
<emphasis id="B952561BFFE0FF98FB64FAA79862FADA" box="[1239,1294,1381,1407]" italics="true" pageId="3" pageNumber="175">
h 
<subScript id="17A2884CFFE0FF98FB57FAAC9981FADA" attach="left" box="[1252,1261,1390,1407]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
-h 
<subScript id="17A2884CFFE0FF98FAB6FAAC9862FADA" attach="left" box="[1285,1294,1390,1407]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
0.56 (0.57 – 0.61), 
<emphasis id="B952561BFFE0FF98FC33FA4A9EDBFA07" box="[896,951,1416,1442]" italics="true" pageId="3" pageNumber="175">
h 
<subScript id="17A2884CFFE0FF98FC3DFA539EFBFA07" attach="left" box="[910,919,1425,1442]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
/h 
<subScript id="17A2884CFFE0FF98FC1DFA539EDBFA07" attach="left" box="[942,951,1425,1442]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
0.80 (057 – 0.84), 
<emphasis id="B952561BFFE0FF98FB3FFA4A98E0FA07" box="[1164,1420,1416,1442]" italics="true" pageId="3" pageNumber="175">
c 
<subScript id="17A2884CFFE0FF98FB24FA5399CCFA07" attach="left" box="[1175,1184,1425,1442]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-c 
<subScript id="17A2884CFFE0FF98FB06FA5399D2FA07" attach="left" box="[1205,1214,1425,1442]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
: d 
<subScript id="17A2884CFFE0FF98FB6CFA539984FA07" attach="left" box="[1247,1256,1425,1442]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-d 
<subScript id="17A2884CFFE0FF98FAB3FA539865FA07" attach="left" box="[1280,1289,1425,1442]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
: e 
<subScript id="17A2884CFFE0FF98FA94FA53985CFA07" attach="left" box="[1319,1328,1425,1442]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-e 
<subScript id="17A2884CFFE0FF98FAF6FA539822FA07" attach="left" box="[1349,1358,1425,1442]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
: f 
<subScript id="17A2884CFFE0FF98FADAFA53981EFA07" attach="left" box="[1385,1394,1425,1442]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
-f 
<subScript id="17A2884CFFE0FF98FA30FA5398E0FA07" attach="left" box="[1411,1420,1425,1442]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
: 0.87(1 – 1.86): 0.89 (0.81 – 1.60): 0.79 (0.81 – 1.09): 1.
</paragraph>
<paragraph id="8B998A09FFE0FF9FFCD3FA229CDCF8D1" blockId="3.[826,1429,1504,1951]" lastBlockId="4.[159,762,1779,1911]" lastPageId="4" lastPageNumber="176" pageId="3" pageNumber="175">
Venter (
<figureCitation id="131D968CFFE0FF98FC0EFA22997AFA5D" box="[957,1046,1504,1528]" captionStart="FIGURE" captionStartId="2.[392,404,1883,1902]" captionTargetBox="[334,1266,340,1852]" captionTargetId="figure-4@2.[322,1266,340,1852]" captionTargetPageId="2" captionText="FIGURE 1: Stigmaeus ladanae n. sp. (female): A – Dorsum; B – Venter; C – gnathosoma." figureDoi="http://doi.org/10.5281/zenodo.4667082" httpUri="https://zenodo.org/record/4667082/files/figure.png" pageId="3" pageNumber="175">Fig. 1B</figureCitation>
) — Coxae I-IV and surrounded shields reticulated, coxisternal shields I-II and III- IV not fused in mid-line, longitudinal striae; cuticle transversely striate between coxisternal II-III; coxisternal shields I-II and III-IV surrounded by longitudinal striae (
<figureCitation id="131D968CFFE0FF98FC5BF94D992EF902" box="[1000,1090,1679,1703]" captionStart="FIGURE" captionStartId="2.[392,404,1883,1902]" captionTargetBox="[334,1266,340,1852]" captionTargetId="figure-4@2.[322,1266,340,1852]" captionTargetPageId="2" captionText="FIGURE 1: Stigmaeus ladanae n. sp. (female): A – Dorsum; B – Venter; C – gnathosoma." figureDoi="http://doi.org/10.5281/zenodo.4667082" httpUri="https://zenodo.org/record/4667082/files/figure.png" pageId="3" pageNumber="175">Fig. 1B</figureCitation>
). Length of setae 
<emphasis id="B952561BFFE0FF98FAA4F94D985CF902" box="[1303,1328,1679,1703]" italics="true" pageId="3" pageNumber="175">1a</emphasis>
24 (18 – 25), 
<emphasis id="B952561BFFE0FF98FCDDF9709EE4F96F" box="[878,904,1714,1738]" italics="true" pageId="3" pageNumber="175">1b</emphasis>
22 (8 – 24), 
<emphasis id="B952561BFFE0FF98FB92F9709955F96F" box="[1057,1081,1714,1738]" italics="true" pageId="3" pageNumber="175">1c</emphasis>
29 (21 – 32), 
<emphasis id="B952561BFFE0FF98FB6CF9709995F96F" box="[1247,1273,1714,1738]" italics="true" pageId="3" pageNumber="175">2b</emphasis>
34 (25 – 35), 
<emphasis id="B952561BFFE0FF98FC89F9179E3EF948" box="[826,850,1749,1773]" italics="true" pageId="3" pageNumber="175">2c</emphasis>
37 (28 – 37), 
<emphasis id="B952561BFFE0FF98FC5DF9179964F948" box="[1006,1032,1749,1773]" italics="true" pageId="3" pageNumber="175">3a</emphasis>
23 (18 – 25), 
<emphasis id="B952561BFFE0FF98FB17F91799D2F948" box="[1188,1214,1749,1773]" italics="true" pageId="3" pageNumber="175">3b</emphasis>
23 (18 – 24), 
<emphasis id="B952561BFFE0FF98FAE9F917981EF948" box="[1370,1394,1749,1773]" italics="true" pageId="3" pageNumber="175">3c</emphasis>
21 (17 – 24), 
<emphasis id="B952561BFFE0FF98FC02F93A9EA5F8B5" box="[945,969,1784,1808]" italics="true" pageId="3" pageNumber="175">4a</emphasis>
21 (18 – 24), 
<emphasis id="B952561BFFE0FF98FBDEF93A99EAF8B5" box="[1133,1158,1784,1808]" italics="true" pageId="3" pageNumber="175">4b</emphasis>
18 (14 – 19) and 
<emphasis id="B952561BFFE0FF98FAEAF93A981DF8B5" box="[1369,1393,1784,1808]" italics="true" pageId="3" pageNumber="175">4c</emphasis>
19 (14 – 20). Aggenital (
<emphasis id="B952561BFFE0FF98FB80F8DE9906F893" box="[1075,1130,1820,1846]" italics="true" pageId="3" pageNumber="175">
ag 
<subScript id="17A2884CFFE0FF98FBFEF8E79906F893" attach="left" box="[1101,1130,1829,1846]" fontSize="7" pageId="3" pageNumber="175">1,-4</subScript>
</emphasis>
) setae 
<emphasis id="B952561BFFE0FF98FB0DF8DE998CF893" box="[1214,1248,1820,1846]" italics="true" pageId="3" pageNumber="175">
ag 
<subScript id="17A2884CFFE0FF98FB64F8E7998CF893" attach="left" box="[1239,1248,1829,1846]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
almost as long as 
<emphasis id="B952561BFFE0FF98FCEAF8FD9EFCF8FC" box="[857,912,1855,1881]" italics="true" pageId="3" pageNumber="175">
ag 
<subScript id="17A2884CFFE0FF98FCC0F88A9EFCF8FC" attach="left" box="[883,912,1864,1881]" fontSize="7" pageId="3" pageNumber="175">2,-3</subScript>
</emphasis>
; and pseudanal seta 
<emphasis id="B952561BFFE0FF98FB3AF8FD99C6F8FC" box="[1161,1194,1855,1881]" italics="true" pageId="3" pageNumber="175">
ps 
<subScript id="17A2884CFFE0FF98FB12F88A99C6F8FC" attach="left" box="[1185,1194,1864,1881]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
less than two times longer than seta 
<emphasis id="B952561BFFE0FF98FBB7F8A09948F8D9" box="[1028,1060,1890,1916]" italics="true" pageId="3" pageNumber="175">
ps 
<subScript id="17A2884CFFE0FF98FBA8F8A99948F8D9" attach="left" box="[1051,1060,1899,1916]" fontSize="7" pageId="3" pageNumber="175">3</subScript>
</emphasis>
; measurements of setae: setae 
<emphasis id="B952561BFFE0FF98FC89F8479E30F83A" box="[826,860,1925,1951]" italics="true" pageId="3" pageNumber="175">
ag 
<subScript id="17A2884CFFE0FF98FCE0F84C9E30F83A" attach="left" box="[851,860,1934,1951]" fontSize="7" pageId="3" pageNumber="175">1</subScript>
</emphasis>
18 (14 – 21), 
<emphasis id="B952561BFFE0FF98FC41F8479978F83A" box="[1010,1044,1925,1951]" italics="true" pageId="3" pageNumber="175">
ag 
<subScript id="17A2884CFFE0FF98FBB8F84C9978F83A" attach="left" box="[1035,1044,1934,1951]" fontSize="7" pageId="3" pageNumber="175">2</subScript>
</emphasis>
18 (16 – 20), 
<emphasis id="B952561BFFE0FF98FB02F84799BFF83A" box="[1201,1235,1925,1951]" italics="true" pageId="3" pageNumber="175">
ag 
<subScript id="17A2884CFFE0FF98FB79F84C99BFF83A" attach="left" box="[1226,1235,1934,1951]" fontSize="7" pageId="3" pageNumber="175">3</subScript>
</emphasis>
22 (17 – 23), 
<emphasis id="B952561BFFE0FF98FAC2F84798FFF83A" box="[1393,1427,1925,1951]" italics="true" pageId="3" pageNumber="175">
ag 
<subScript id="17A2884CFFE0FF98FA39F84C98FFF83A" attach="left" box="[1418,1427,1934,1951]" fontSize="7" pageId="3" pageNumber="175">4</subScript>
</emphasis>
22 (19 – 26), 
<emphasis id="B952561BFFE7FF9FFE80F9369C25F8A8" box="[307,329,1780,1805]" italics="true" pageId="4" pageNumber="176">
g 
<subScript id="17A2884CFFE7FF9FFEF3F93E9C25F8A8" attach="left" box="[320,329,1788,1805]" fontSize="7" pageId="4" pageNumber="176">1</subScript>
</emphasis>
19 (16 – 20), 
<emphasis id="B952561BFFE7FF9FFE6DF9369C99F8A8" box="[478,501,1780,1805]" italics="true" pageId="4" pageNumber="176">
g 
<subScript id="17A2884CFFE7FF9FFE5FF93E9C99F8A8" attach="left" box="[492,501,1788,1805]" fontSize="7" pageId="4" pageNumber="176">2</subScript>
</emphasis>
29 (19 – 31), 
<emphasis id="B952561BFFE7FF9FFD39F9369FC7F8A8" box="[650,683,1780,1805]" italics="true" pageId="4" pageNumber="176">
ps 
<subScript id="17A2884CFFE7FF9FFD11F93E9FC7F8A8" attach="left" box="[674,683,1788,1805]" fontSize="7" pageId="4" pageNumber="176">1</subScript>
</emphasis>
52 (36 – 53), 
<emphasis id="B952561BFFE7FF9FFF50F8D59C68F895" box="[227,260,1815,1840]" italics="true" pageId="4" pageNumber="176">
ps 
<subScript id="17A2884CFFE7FF9FFF48F8DD9C68F895" attach="left" box="[251,260,1823,1840]" fontSize="7" pageId="4" pageNumber="176">2</subScript>
</emphasis>
28 (19 – 28), 
<emphasis id="B952561BFFE7FF9FFE28F8D59CD7F895" box="[411,443,1815,1840]" italics="true" pageId="4" pageNumber="176">
ps 
<subScript id="17A2884CFFE7FF9FFE01F8DD9CD7F895" attach="left" box="[434,443,1823,1840]" fontSize="7" pageId="4" pageNumber="176">3</subScript>
</emphasis>
44 (38 – 46). Distances: 
<emphasis id="B952561BFFE7FF9FFD7EF8D59DADF8F1" italics="true" pageId="4" pageNumber="176">
ag 
<subScript id="17A2884CFFE7FF9FFD55F8DD9F83F895" attach="left" box="[742,751,1823,1840]" fontSize="7" pageId="4" pageNumber="176">1</subScript>
- ag 
<subScript id="17A2884CFFE7FF9FFF0BF8819DADF8F1" attach="left" box="[184,193,1859,1876]" fontSize="7" pageId="4" pageNumber="176">1</subScript>
</emphasis>
33 (25 – 33), 
<emphasis id="B952561BFFE7FF9FFEEEF8F89CC0F8F1" box="[349,428,1850,1876]" italics="true" pageId="4" pageNumber="176">
ag 
<subScript id="17A2884CFFE7FF9FFEC5F8819C13F8F1" attach="left" box="[374,383,1859,1876]" fontSize="7" pageId="4" pageNumber="176">2</subScript>
-ag 
<subScript id="17A2884CFFE7FF9FFE10F8819CC0F8F1" attach="left" box="[419,428,1859,1876]" fontSize="7" pageId="4" pageNumber="176">2</subScript>
</emphasis>
35 (33 – 38), 
<emphasis id="B952561BFFE7FF9FFDFBF8F89FFAF8F1" box="[584,662,1850,1876]" italics="true" pageId="4" pageNumber="176">
ag 
<subScript id="17A2884CFFE7FF9FFDD2F8819F06F8F1" attach="left" box="[609,618,1859,1876]" fontSize="7" pageId="4" pageNumber="176">3</subScript>
-ag 
<subScript id="17A2884CFFE7FF9FFD3EF8819FFAF8F1" attach="left" box="[653,662,1859,1876]" fontSize="7" pageId="4" pageNumber="176">3</subScript>
</emphasis>
49 (47 – 53), 
<emphasis id="B952561BFFE7FF9FFF7CF89F9C72F8D2" box="[207,286,1885,1911]" italics="true" pageId="4" pageNumber="176">
ag 
<subScript id="17A2884CFFE7FF9FFF5AF8A49D9EF8D2" attach="left" box="[233,242,1894,1911]" fontSize="7" pageId="4" pageNumber="176">4</subScript>
-ag 
<subScript id="17A2884CFFE7FF9FFEA6F8A49C72F8D2" attach="left" box="[277,286,1894,1911]" fontSize="7" pageId="4" pageNumber="176">4</subScript>
</emphasis>
46 (46 – 61).
</paragraph>
<caption id="DF59DA81FFE7FF9FFE3EF95D99C5F917" ID-DOI="http://doi.org/10.5281/zenodo.4667084" ID-Zenodo-Dep="4667084" box="[397,1193,1694,1714]" httpUri="https://zenodo.org/record/4667084/files/figure.png" pageId="4" pageNumber="176" startId="4.[397,409,1695,1714]" targetBox="[242,1345,302,1662]" targetPageId="4">
<paragraph id="8B998A09FFE7FF9FFE3EF95D99C5F917" blockId="4.[397,1193,1694,1714]" box="[397,1193,1694,1714]" pageId="4" pageNumber="176">
FIGURE 2: 
<taxonomicName id="4C26F18AFFE7FF9FFE41F95D9FFEF914" authority="ladanae n." authorityName="LADANAE N." box="[498,658,1694,1714]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="173" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B952561BFFE7FF9FFE41F95D9FFEF914" box="[498,658,1694,1714]" italics="true" pageId="4" pageNumber="176">Stigmaeus ladanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261EB60FFE7FF9FFD24F95D9FA7F917" box="[663,715,1695,1714]" pageId="4" pageNumber="176" rank="species">
<emphasis id="B952561BFFE7FF9FFD24F95D9FA7F917" bold="true" box="[663,715,1695,1714]" pageId="4" pageNumber="176">n. sp.</emphasis>
</taxonomicNameLabel>
(female): A – leg I; B – leg II; C – leg III; D – leg IV.
</paragraph>
</caption>
<paragraph id="8B998A09FFE7FF9EFCD3F9319C85FDE1" blockId="4.[826,1429,1779,1946]" lastBlockId="5.[159,762,311,1003]" lastPageId="5" lastPageNumber="177" pageId="4" pageNumber="176">
Gnathosoma (
<figureCitation id="131D968CFFE7FF9FFBB4F9319908F8AE" box="[1031,1124,1779,1803]" captionStart="FIGURE" captionStartId="2.[392,404,1883,1902]" captionTargetBox="[334,1266,340,1852]" captionTargetId="figure-4@2.[322,1266,340,1852]" captionTargetPageId="2" captionText="FIGURE 1: Stigmaeus ladanae n. sp. (female): A – Dorsum; B – Venter; C – gnathosoma." figureDoi="http://doi.org/10.5281/zenodo.4667082" httpUri="https://zenodo.org/record/4667082/files/figure.png" pageId="4" pageNumber="176">Fig. 1C</figureCitation>
) — Subcapitulum reticulated and with two pairs of subcapitular setae, 
<emphasis id="B952561BFFE7FF9FFAECF8D59818F88B" box="[1375,1396,1815,1838]" italics="true" pageId="4" pageNumber="176">m</emphasis>
23 (19 – 25) and 
<emphasis id="B952561BFFE7FF9FFC68F8F89E86F8F4" box="[987,1002,1850,1873]" italics="true" pageId="4" pageNumber="176">n</emphasis>
18 (14 – 21), two pairs of adoral setae, 
<emphasis id="B952561BFFE7FF9FFCDEF89F9EE0F8D2" box="[877,908,1885,1911]" italics="true" pageId="4" pageNumber="176">
or 
<subScript id="17A2884CFFE7FF9FFC30F8A49EE0F8D2" attach="left" box="[899,908,1894,1911]" fontSize="7" pageId="4" pageNumber="176">1</subScript>
</emphasis>
7 (4 – 8), 
<emphasis id="B952561BFFE7FF9FFBBAF89F9944F8D2" box="[1033,1064,1885,1911]" italics="true" pageId="4" pageNumber="176">
or 
<subScript id="17A2884CFFE7FF9FFBACF8A49944F8D2" attach="left" box="[1055,1064,1894,1911]" fontSize="7" pageId="4" pageNumber="176">2</subScript>
</emphasis>
9 (7 – 10); distances: 
<emphasis id="B952561BFFE7FF9FFA87F89F9811F8D2" box="[1332,1405,1885,1911]" italics="true" pageId="4" pageNumber="176">
or 
<subScript id="17A2884CFFE7FF9FFAF9F8A4983FF8D2" attach="left" box="[1354,1363,1894,1911]" fontSize="7" pageId="4" pageNumber="176">1</subScript>
-or 
<subScript id="17A2884CFFE7FF9FFAC7F8A49811F8D2" attach="left" box="[1396,1405,1894,1911]" fontSize="7" pageId="4" pageNumber="176">1</subScript>
</emphasis>
7 (5 – 18), 
<emphasis id="B952561BFFE7FF9FFC2EF8429E8AF83F" box="[925,998,1920,1946]" italics="true" pageId="4" pageNumber="176">
or 
<subScript id="17A2884CFFE7FF9FFC00F84B9ED0F83F" attach="left" box="[947,956,1929,1946]" fontSize="7" pageId="4" pageNumber="176">2</subScript>
-or 
<subScript id="17A2884CFFE7FF9FFC6EF84B9E8AF83F" attach="left" box="[989,998,1929,1946]" fontSize="7" pageId="4" pageNumber="176">2</subScript>
</emphasis>
14 (11 – 16), 
<emphasis id="B952561BFFE7FF9FFB32F84299DFF832" box="[1153,1203,1920,1943]" italics="true" pageId="4" pageNumber="176">m-m</emphasis>
31 (29 – 36), 
<emphasis id="B952561BFFE7FF9FFAFEF842981FF832" box="[1357,1395,1920,1943]" italics="true" pageId="4" pageNumber="176">n-n</emphasis>
23 (23 – 25), 
<emphasis id="B952561BFFE6FF9EFEA1FEFA9C3CFEEA" box="[274,336,312,337]" italics="true" pageId="5" pageNumber="177">
or 
<subScript id="17A2884CFFE6FF9EFE9BFE829C5DFEF4" attach="left" box="[296,305,320,337]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
-m
</emphasis>
25 (24 – 27), 
<emphasis id="B952561BFFE6FF9EFE5DFEFA9F76FEEA" box="[494,538,312,335]" italics="true" pageId="5" pageNumber="177">m-n</emphasis>
6 (5 – 7) (
<figureCitation id="131D968CFFE6FF9EFD22FEF59F80FEEA" box="[657,748,311,335]" captionStart="FIGURE" captionStartId="2.[392,404,1883,1902]" captionTargetBox="[334,1266,340,1852]" captionTargetId="figure-4@2.[322,1266,340,1852]" captionTargetPageId="2" captionText="FIGURE 1: Stigmaeus ladanae n. sp. (female): A – Dorsum; B – Venter; C – gnathosoma." figureDoi="http://doi.org/10.5281/zenodo.4667082" httpUri="https://zenodo.org/record/4667082/files/figure.png" pageId="5" pageNumber="177">Fig. 1C</figureCitation>
). Chelicerae free 55 (50 – 63), movable digit 25(22 – 27) (
<figureCitation id="131D968CFFE6FF9EFF63FEBF9C4BFE30" box="[208,295,381,405]" captionStart="FIGURE" captionStartId="2.[392,404,1883,1902]" captionTargetBox="[334,1266,340,1852]" captionTargetId="figure-4@2.[322,1266,340,1852]" captionTargetPageId="2" captionText="FIGURE 1: Stigmaeus ladanae n. sp. (female): A – Dorsum; B – Venter; C – gnathosoma." figureDoi="http://doi.org/10.5281/zenodo.4667082" httpUri="https://zenodo.org/record/4667082/files/figure.png" pageId="5" pageNumber="177">Fig. 1C</figureCitation>
). Palpi five segmented, palp tarsus with four simple setae + one simple eupathidium + one solenidion (ω) + one tridentate eupathidium, palp tibia with two setae + one well developed claw + one accessory claw, palp genu with one setae and palp femur with three setae.
</paragraph>
<paragraph id="8B998A09FFE6FF9EFF76FD9E9C12FCE3" blockId="5.[159,762,311,1003]" pageId="5" pageNumber="177">
Legs (
<figureCitation id="131D968CFFE6FF9EFEBFFD9E9C38FDD1" box="[268,340,604,628]" captionStart="FIGURE" captionStartId="4.[397,409,1695,1714]" captionTargetBox="[242,1345,302,1662]" captionTargetId="figure-4@4.[242,1345,302,1663]" captionTargetPageId="4" captionText="FIGURE 2: Stigmaeus ladanae n. sp. (female): A – leg I; B – leg II; C – leg III; D – leg IV." figureDoi="http://doi.org/10.5281/zenodo.4667084" httpUri="https://zenodo.org/record/4667084/files/figure.png" pageId="5" pageNumber="177">Fig. 2</figureCitation>
) — Legs about half length of body. Leg segments setal formulae as follows: coxae 2 - 2 - 2 - 2; trochanters 1 - 1 - 2 - 1; femora 4 - 4 - 3 - 2, genua 5+1Κ - 5 - 2 - 2; tibiae 5+1’ρ+1’ - 5+1’ρ - 5+1’ρ - 5+1’ρ; tarsi 13 +1ω - 9 +1ω - 7 +1ω - 7 +1ω. Length of solenidia: ωI 19 (16-20), ωII 17 (14-18), ωIII 10 (7 – 12), ωIV 8 (6 – 9).
</paragraph>
<paragraph id="8B998A09FFE6FF9EFF76FC9F9CC9FCD0" blockId="5.[159,762,311,1003]" box="[197,421,861,885]" pageId="5" pageNumber="177">Male — Unknown.</paragraph>
<paragraph id="8B998A09FFE6FF9EFF76FC4F9CE1FC4E" blockId="5.[159,762,311,1003]" pageId="5" pageNumber="177">
Remarks — 
<taxonomicName id="4C26F18AFFE6FF9EFEE9FC4F9F49FC00" authority="ladanae n." authorityName="LADANAE N." box="[346,549,909,933]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="173" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B952561BFFE6FF9EFEE9FC4F9F49FC00" box="[346,549,909,933]" italics="true" pageId="5" pageNumber="177">Stigmaeus ladanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261EB60FFE6FF9EFD9CFC4F9F17FC00" box="[559,635,909,933]" pageId="5" pageNumber="177" rank="species">
<emphasis id="B952561BFFE6FF9EFD9CFC4F9F17FC00" bold="true" box="[559,635,909,933]" pageId="5" pageNumber="177">n. sp.</emphasis>
</taxonomicNameLabel>
is closely similar 
<taxonomicName id="4C26F18AFFE6FF9EFF45FC729C63FC4E" authority="Haddad, Lotfollahi and Akbari, 2010" authorityName="Haddad, Lotfollahi and Akbari" authorityYear="2010" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="shabestariensis">
<emphasis id="B952561BFFE6FF9EFF45FC729CDCFC6D" box="[246,432,944,968]" italics="true" pageId="5" pageNumber="177">S. shabestariensis</emphasis>
<bibRefCitation id="EFB7F7F8FFE6FF9EFE05FC729C63FC4E" author="Haddad Irani-Nejad, P. &amp; Lotfollahi K. &amp; Akbari A. &amp; Bagheri M. &amp; Ueckermann E. A." pageId="5" pageNumber="177" pagination="118 - 122" refId="ref5158" refString="Haddad Irani-Nejad, P., Lotfollahi K., Akbari A., Bagheri M., Ueckermann E. A. 2010 a - Stigmaeus shendabadiensis (Acari: Prostigmata: Stigmaeidae) from Northwest of Iran - Syst. Appl. Acarol., 15: 118 - 122." type="journal article" year="2010">
Haddad, Lotfollahi and Akbari, 
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in having:
</paragraph>
<paragraph id="8B998A09FFE6FF9EFF74FBD89C2DFB94" blockId="5.[199,321,1050,1076]" box="[199,321,1050,1076]" pageId="5" pageNumber="177">
– setae 
<emphasis id="B952561BFFE6FF9EFE91FBD89C55FB91" box="[290,313,1050,1076]" italics="true" pageId="5" pageNumber="177">
h 
<subScript id="17A2884CFFE6FF9EFE83FBE19C55FB91" attach="left" box="[304,313,1059,1076]" fontSize="7" pageId="5" pageNumber="177">3</subScript>
</emphasis>
,
</paragraph>
<paragraph id="8B998A09FFE6FF9EFF74FB959CBBFBCA" blockId="5.[199,471,1111,1135]" box="[199,471,1111,1135]" pageId="5" pageNumber="177">– entire suranal shield,</paragraph>
<paragraph id="8B998A09FFE6FF9EFF74FB569F62FB09" blockId="5.[199,526,1172,1196]" box="[199,526,1172,1196]" pageId="5" pageNumber="177">– reticulated dorsal shields,</paragraph>
<paragraph id="8B998A09FFE6FF9EFF74FB139F64FB4C" blockId="5.[199,520,1233,1257]" box="[199,520,1233,1257]" pageId="5" pageNumber="177">– the same legs chaetotaxy.</paragraph>
<paragraph id="8B998A09FFE6FF9EFF2CFADA9F2BFA8A" blockId="5.[159,583,1303,1327]" box="[159,583,1303,1327]" pageId="5" pageNumber="177">However, the new species differs in:</paragraph>
<paragraph id="8B998A09FFE6FF9EFF74FA9D9FF6FA3C" blockId="5.[199,761,1374,1433]" pageId="5" pageNumber="177">
– lateral prodorsal shield reticulate in 
<taxonomicName id="4C26F18AFFE6FF9EFD3FFA9D9F95FAD3" authorityName="Nazari &amp; Khanjani &amp; Kamali" authorityYear="2012" box="[652,761,1374,1398]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="ladanae">
<emphasis id="B952561BFFE6FF9EFD3FFA9D9F95FAD3" box="[652,761,1374,1398]" italics="true" pageId="5" pageNumber="177">S. ladanae</emphasis>
</taxonomicName>
instead of smooth in 
<taxonomicName id="4C26F18AFFE6FF9EFE64FA409FF8FA3C" authorityName="Haddad, Lotfollahi and Akbari" authorityYear="2010" box="[471,660,1409,1433]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="shabestariensis">
<emphasis id="B952561BFFE6FF9EFE64FA409FF8FA3C" box="[471,660,1409,1433]" italics="true" pageId="5" pageNumber="177">S. shabestariensis</emphasis>
</taxonomicName>
,
</paragraph>
<paragraph id="8B998A09FFE6FF9EFF74FA7D9CCDFA5C" blockId="5.[199,761,1470,1529]" pageId="5" pageNumber="177">– subcapiltulum faintly reticulated instead of faintly punctate,</paragraph>
<paragraph id="8B998A09FFE6FF9EFF74F9DD9FDDF993" blockId="5.[199,689,1566,1590]" box="[199,689,1566,1590]" pageId="5" pageNumber="177">– dorsal setae smooth instead of serrated,</paragraph>
<paragraph id="8B998A09FFE6FF9EFF74F99E9CAEF933" blockId="5.[199,762,1627,1686]" pageId="5" pageNumber="177">
– lateral zonal shields unique instead of divided in 
<taxonomicName id="4C26F18AFFE6FF9EFF4CF9BD9CD7F933" authorityName="Haddad, Lotfollahi and Akbari" authorityYear="2010" box="[255,443,1662,1686]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="shabestariensis">
<emphasis id="B952561BFFE6FF9EFF4CF9BD9CD7F933" box="[255,443,1662,1686]" italics="true" pageId="5" pageNumber="177">S. shabestariensis</emphasis>
</taxonomicName>
,
</paragraph>
<paragraph id="8B998A09FFE6FF9EFF74F97E9CAEF953" blockId="5.[199,762,1723,1782]" pageId="5" pageNumber="177">
– base of setae 
<emphasis id="B952561BFFE6FF9EFEC9F9799CF3F976" box="[378,415,1723,1747]" italics="true" pageId="5" pageNumber="177">ps1</emphasis>
reticulated instead of smooth in 
<taxonomicName id="4C26F18AFFE6FF9EFF4CF91D9CD7F953" authorityName="Haddad, Lotfollahi and Akbari" authorityYear="2010" box="[255,443,1758,1782]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="shabestariensis">
<emphasis id="B952561BFFE6FF9EFF4CF91D9CD7F953" box="[255,443,1758,1782]" italics="true" pageId="5" pageNumber="177">S. shabestariensis</emphasis>
</taxonomicName>
,
</paragraph>
<paragraph id="8B998A09FFE6FF9EFF74F8DE9CC8F839" blockId="5.[199,761,1819,1948]" pageId="5" pageNumber="177">
– ratio 
<emphasis id="B952561BFFE6FF9EFE96F8DE9C54F893" box="[293,312,1820,1846]" italics="true" pageId="5" pageNumber="177">
c 
<subScript id="17A2884CFFE6FF9EFE9CF8E79C54F893" attach="left" box="[303,312,1829,1846]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
</emphasis>
/ 
<emphasis id="B952561BFFE6FF9EFEF9F8DE9C10F893" box="[330,380,1820,1846]" italics="true" pageId="5" pageNumber="177">
c 
<subScript id="17A2884CFFE6FF9EFEE6F8E79C32F893" attach="left" box="[341,350,1829,1846]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
-c 
<subScript id="17A2884CFFE6FF9EFEC0F8E79C10F893" attach="left" box="[371,380,1829,1846]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
</emphasis>
0.52 (0.46 – 0.51) and 
<emphasis id="B952561BFFE6FF9EFD11F8DE9FD9F893" box="[674,693,1820,1846]" italics="true" pageId="5" pageNumber="177">
e 
<subScript id="17A2884CFFE6FF9EFD1FF8E79FD9F893" attach="left" box="[684,693,1829,1846]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
</emphasis>
/ 
<emphasis id="B952561BFFE6FF9EFD74F8DE9F94F893" box="[711,760,1820,1846]" italics="true" pageId="5" pageNumber="177">
e 
<subScript id="17A2884CFFE6FF9EFD62F8E79FB6F893" attach="left" box="[721,730,1829,1846]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
- 
<geoCoordinate id="EE12ECCEFFE6FF9EFD56F8DE9F83F896" box="[741,751,1820,1843]" degrees="1" direction="east" minutes="0.5" orientation="longitude" pageId="5" pageNumber="177" precision="92" value="1.0083333">e</geoCoordinate>
<subScript id="17A2884CFFE6FF9EFD5CF8E79F94F893" attach="left" box="[751,760,1829,1846]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
</emphasis>
0.5 (0.55 – 0.8) in 
<taxonomicName id="4C26F18AFFE6FF9EFE0DF8FD9F43F8F3" authorityName="Nazari &amp; Khanjani &amp; Kamali" authorityYear="2012" box="[446,559,1854,1878]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="ladanae">
<emphasis id="B952561BFFE6FF9EFE0DF8FD9F43F8F3" box="[446,559,1854,1878]" italics="true" pageId="5" pageNumber="177">S. ladanae</emphasis>
</taxonomicName>
opposed to 0.40 (0.20 – 0.40) and 
<emphasis id="B952561BFFE6FF9EFE03F8A09CA8F8D9" box="[432,452,1890,1916]" italics="true" pageId="5" pageNumber="177">
e 
<subScript id="17A2884CFFE6FF9EFE08F8A99CA8F8D9" attach="left" box="[443,452,1899,1916]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
</emphasis>
/ 
<emphasis id="B952561BFFE6FF9EFE66F8A09F6AF8D9" box="[469,518,1890,1916]" italics="true" pageId="5" pageNumber="177">
e 
<subScript id="17A2884CFFE6FF9EFE53F8A99C85F8D9" attach="left" box="[480,489,1899,1916]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
- 
<geoCoordinate id="EE12ECCEFFE6FF9EFE40F8A09C91F8DC" box="[499,509,1890,1913]" degrees="1" direction="east" minutes="0.34" orientation="longitude" pageId="5" pageNumber="177" precision="9" value="1.0056666">e</geoCoordinate>
<subScript id="17A2884CFFE6FF9EFE4EF8A99F6AF8D9" attach="left" box="[509,518,1899,1916]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
</emphasis>
0.34 (0.33 – 0.34) in 
<taxonomicName id="4C26F18AFFE6FF9EFF52F8479CF1F839" authorityName="Haddad, Lotfollahi and Akbari" authorityYear="2010" box="[225,413,1924,1948]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="shabestariensis">
<emphasis id="B952561BFFE6FF9EFF52F8479CF1F839" box="[225,413,1924,1948]" italics="true" pageId="5" pageNumber="177">S. shabestariensis</emphasis>
</taxonomicName>
,
</paragraph>
<paragraph id="8B998A09FFE6FF9EFCD1FEF5994FFE31" blockId="5.[866,1429,311,405]" pageId="5" pageNumber="177">
– coxal plates I-IV and legs segments reticulated in the new species versus smooth in 
<taxonomicName id="4C26F18AFFE6FF9EFACCFE989970FE30" authorityName="Haddad, Lotfollahi and Akbari" authorityYear="2010" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="shabestariensis">
<emphasis id="B952561BFFE6FF9EFACCFE989970FE30" italics="true" pageId="5" pageNumber="177">S. shabestariensis</emphasis>
</taxonomicName>
,
</paragraph>
<paragraph id="8B998A09FFE6FF9EFCD1FE7499B6FE55" blockId="5.[866,1428,438,497]" pageId="5" pageNumber="177">
– aggenital and anal shields reticulated instead of smooth in 
<taxonomicName id="4C26F18AFFE6FF9EFBA4FE1B99BFFE54" authorityName="Haddad, Lotfollahi and Akbari" authorityYear="2010" box="[1047,1235,473,497]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="shabestariensis">
<emphasis id="B952561BFFE6FF9EFBA4FE1B99BFFE54" box="[1047,1235,473,497]" italics="true" pageId="5" pageNumber="177">S. shabestariensis</emphasis>
</taxonomicName>
,
</paragraph>
<paragraph id="8B998A09FFE6FF9EFCD1FDD098F8FD59" blockId="5.[866,1429,529,764]" pageId="5" pageNumber="177">
– setae 
<emphasis id="B952561BFFE6FF9EFC70FDD09EB7FD8C" box="[963,987,530,553]" italics="true" pageId="5" pageNumber="177">ps</emphasis>
<subScript id="17A2884CFFE6FF9EFC68FDD99E88FD89" attach="left" box="[987,996,539,556]" fontSize="7" pageId="5" pageNumber="177">2</subScript>
two times longer than 
<emphasis id="B952561BFFE6FF9EFAA2FDD0985DFD89" box="[1297,1329,530,556]" italics="true" pageId="5" pageNumber="177">
ps 
<subScript id="17A2884CFFE6FF9EFA9BFDD9985DFD89" attach="left" box="[1320,1329,539,556]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
</emphasis>
instead of as long as in 
<emphasis id="B952561BFFE6FF9EFBDBFDF7985AFDE8" box="[1128,1334,565,589]" italics="true" pageId="5" pageNumber="177">
<taxonomicName id="4C26F18AFFE6FF9EFBDBFDF7985EFDE8" authorityName="Haddad, Lotfollahi and Akbari" authorityYear="2010" box="[1128,1330,565,589]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="shabestariensis">S. shabestariensis</taxonomicName>
.
</emphasis>
Also this species resembles 
<taxonomicName id="4C26F18AFFE6FF9EFB16FD9A9957FD36" authority="Kuznetsov, 1987" authorityName="Kuznetsov" authorityYear="1987" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="B952561BFFE6FF9EFB16FD9A98F9FDD5" box="[1189,1429,600,624]" italics="true" pageId="5" pageNumber="177">Stigmaeus pulchellus</emphasis>
<bibRefCitation id="EFB7F7F8FFE6FF9EFCCFFDB99957FD36" author="Kuznetsov N. N." box="[892,1083,635,659]" pageId="5" pageNumber="177" pagination="635 - 648" refId="ref5473" refString="Kuznetsov N. N. 1987 - New species of the family Stigmaeidae - Zool. Zhurnal., 54 (4): 635 - 648 [In Russian]." type="journal article" year="1987">Kuznetsov, 1987</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C26F18AFFE6FF9EFBC4FDB99930FD13" authority="Khanjani and Ueckermann, 2002" authorityName="Khanjani and Ueckermann" authorityYear="2002" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="alvandis">
<emphasis id="B952561BFFE6FF9EFBC4FDB99982FD36" box="[1143,1262,635,659]" italics="true" pageId="5" pageNumber="177">S. alvandis</emphasis>
<bibRefCitation id="EFB7F7F8FFE6FF9EFB45FDB99930FD13" author="Khanjani M. &amp; Ueckermann E. A." pageId="5" pageNumber="177" pagination="317 - 339" refId="ref5267" refString="Khanjani M., Ueckermann E. A. 2002 - The stigmaeid mites of Iran (Acari: Stigmaeidae) - Internat. J. Acarol., 28 (4): 317 - 339. doi: 10.1080 / 01647950208684309" type="journal article" year="2002">
Khanjani and Ueckermann, 
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</bibRefCitation>
</taxonomicName>
in having all dorsal and ventral shields, and leg segments reticulated, eyes absent, two pairs of median zonal shield;
</paragraph>
<paragraph id="8B998A09FFE6FF9EFC89FCE59800FC0D" blockId="5.[826,1429,807,936]" pageId="5" pageNumber="177">
however it differs from that: aggenital shields reticulated, setae 
<emphasis id="B952561BFFE6FF9EFC65FC899E99FCC7" box="[982,1013,843,866]" italics="true" pageId="5" pageNumber="177">sce</emphasis>
shorter than 
<emphasis id="B952561BFFE6FF9EFB25FC889844FCC7" box="[1174,1320,842,866]" italics="true" pageId="5" pageNumber="177">
<taxonomicName id="4C26F18AFFE6FF9EFB25FC88984FFCC7" authorityName="Kuznetsov" authorityYear="1987" box="[1174,1315,842,866]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="pulchellus">S. pulchellus</taxonomicName>
,
</emphasis>
setae 
<emphasis id="B952561BFFE6FF9EFAC1FC8998FFFCC0" box="[1394,1427,843,869]" italics="true" pageId="5" pageNumber="177">
ps 
<subScript id="17A2884CFFE6FF9EFA39FC9698FFFCC0" attach="left" box="[1418,1427,852,869]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
</emphasis>
longer than 
<emphasis id="B952561BFFE6FF9EFC7AFCAF9940FC2D" box="[969,1068,877,904]" italics="true" pageId="5" pageNumber="177">
h 
<subScript id="17A2884CFFE6FF9EFC64FCB59E8CFC2D" attach="left" box="[983,992,887,904]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
, h 
<subScript id="17A2884CFFE6FF9EFC4EFCB5996AFC2D" attach="left" box="[1021,1030,887,904]" fontSize="7" pageId="5" pageNumber="177">2</subScript>
, h 
<subScript id="17A2884CFFE6FF9EFB90FCB59940FC2D" attach="left" box="[1059,1068,887,904]" fontSize="7" pageId="5" pageNumber="177">3</subScript>
</emphasis>
instead of as long as in 
<taxonomicName id="4C26F18AFFE6FF9EFAF8FCAC9EE8FC0D" authorityName="Kuznetsov" authorityYear="1987" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="B952561BFFE6FF9EFAF8FCAC9EE8FC0D" italics="true" pageId="5" pageNumber="177">S. pulchellus</emphasis>
</taxonomicName>
and also it differs from 
<taxonomicName id="4C26F18AFFE6FF9EFB2EFC53987FFC0D" authorityName="Khanjani and Ueckermann" authorityYear="2002" box="[1181,1299,912,936]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="alvandis">
<emphasis id="B952561BFFE6FF9EFB2EFC53987FFC0D" box="[1181,1299,912,936]" italics="true" pageId="5" pageNumber="177">S. alvandis</emphasis>
</taxonomicName>
in that:
</paragraph>
<paragraph id="8B998A09FFE6FF9EFCD1FC169EB1FBAB" blockId="5.[866,1429,979,1038]" pageId="5" pageNumber="177">
– the suranal shield entire whereas divided in 
<emphasis id="B952561BFFE6FF9EFACCFC169EB1FBAB" italics="true" pageId="5" pageNumber="177">
<taxonomicName id="4C26F18AFFE6FF9EFACCFC169EB5FBAB" authorityName="Khanjani and Ueckermann" authorityYear="2002" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="alvandis">S. alvandis</taxonomicName>
,
</emphasis>
</paragraph>
<paragraph id="8B998A09FFE6FF9EFCD1FBF2997BFBCF" blockId="5.[866,1429,1071,1130]" pageId="5" pageNumber="177">
– femur IV with two setae opposed to one seta in 
<taxonomicName id="4C26F18AFFE6FF9EFC29FB91997CFBCF" authorityName="Khanjani and Ueckermann" authorityYear="2002" box="[922,1040,1106,1130]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="alvandis">
<emphasis id="B952561BFFE6FF9EFC29FB91997CFBCF" box="[922,1040,1106,1130]" italics="true" pageId="5" pageNumber="177">S. alvandis</emphasis>
</taxonomicName>
,
</paragraph>
<paragraph id="8B998A09FFE6FF9EFCD1FB49991EFB63" blockId="5.[866,1429,1163,1222]" pageId="5" pageNumber="177">
– setae 
<emphasis id="B952561BFFE6FF9EFC72FB4E9E8EFB00" box="[961,994,1164,1189]" italics="true" pageId="5" pageNumber="177">
ps 
<subScript id="17A2884CFFE6FF9EFC6AFB569E8EFB00" attach="left" box="[985,994,1172,1189]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
</emphasis>
longer than 
<emphasis id="B952561BFFE6FF9EFB37FB499982FB00" box="[1156,1262,1163,1189]" italics="true" pageId="5" pageNumber="177">
h 
<subScript id="17A2884CFFE6FF9EFB22FB5699F6FB00" attach="left" box="[1169,1178,1172,1189]" fontSize="7" pageId="5" pageNumber="177">1</subScript>
, h 
<subScript id="17A2884CFFE6FF9EFB08FB5699A8FB00" attach="left" box="[1211,1220,1172,1189]" fontSize="7" pageId="5" pageNumber="177">2</subScript>
, h 
<subScript id="17A2884CFFE6FF9EFB56FB569982FB00" attach="left" box="[1253,1262,1172,1189]" fontSize="7" pageId="5" pageNumber="177">3</subScript>
</emphasis>
instead of as long as in 
<taxonomicName id="4C26F18AFFE6FF9EFC47FB6C9906FB63" authorityName="Khanjani and Ueckermann" authorityYear="2002" box="[1012,1130,1198,1222]" class="Arachnida" family="Stigmaeidae" genus="Stigmaeus" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="177" phylum="Arthropoda" rank="species" species="alvandis">
<emphasis id="B952561BFFE6FF9EFC47FB6C9906FB63" box="[1012,1130,1198,1222]" italics="true" pageId="5" pageNumber="177">S. alvandis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B998A09FFE6FF9EFC89FB33982FFA1D" blockId="5.[826,1429,1265,1464]" pageId="5" pageNumber="177">
Etymology — The new species is named in honor of Mrs. Ladan Mohammadi, wife of senior author, who kindly helped us in mite collection and who is already post student (Agricultural Entomology), Department of Plant Protection, College of Agriculture, Bu-Ali 
<taxonomicName id="4C26F18AFFE6FF9EFC79FA629E90FA1D" box="[970,1020,1440,1464]" class="Insecta" genus="Sina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="genus">Sina</taxonomicName>
University, 
<collectingRegion id="49E244EBFFE6FF9EFB39FA62986DFA1D" box="[1162,1281,1440,1464]" country="Iran" name="Hamadan" pageId="5" pageNumber="177">Hamedan</collectingRegion>
, 
<collectingCountry id="F331CA99FFE6FF9EFABFFA639853FA1D" box="[1292,1343,1441,1464]" name="Iran" pageId="5" pageNumber="177">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>