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<document id="870E7B85C75C46BE8D23A2955FC4ACEA" ID-CLB-Dataset="32751" ID-DOI="10.11646/zootaxa.4286.2.6" ID-GBIF-Dataset="1bad4e53-fbe6-4037-9f7b-dd82ca165daf" ID-ISSN="1175-5326" ID-Zenodo-Dep="828435" ID-ZooBank="B42E72F2-30F2-4228-BEFA-A8AB3239916A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500108820316" checkinUser="plazi" docAuthor="Cumberlidge, Neil" docDate="2017" docId="038F3B05FF83FFD3FF2DCA50FCE6FCB6" docLanguage="en" docName="zootaxa.4286.2.6.pdf" docOrigin="Zootaxa 4286 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Erimetopus brazzae A. Milne-Edwards 1886" docType="treatment" docVersion="7" lastPageNumber="238" masterDocId="FFB6437DFF8BFFD9FFBAC966FFD6FFA3" masterDocTitle="Potamonautes nheenae, a new species of freshwater crab from Gabon, Central Africa, with a description of the first known male of Erimetopus brazzae (A. Milne-Edwards, 1886) from the Democratic Republic of the Congo, Central Africa (Brachyura: Potamoidea: Potamonautidae)" masterLastPageNumber="240" masterPageNumber="228" pageNumber="236" updateTime="1698465160371" updateUser="ExternalLinkService">
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<mods:title id="0A54AFA5A7E2E1E21F80096BBB50F2AE">Potamonautes nheenae, a new species of freshwater crab from Gabon, Central Africa, with a description of the first known male of Erimetopus brazzae (A. Milne-Edwards, 1886) from the Democratic Republic of the Congo, Central Africa (Brachyura: Potamoidea: Potamonautidae)</mods:title>
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<mods:namePart id="1C520B2B4D53B4AB84F77AC351EBC036">Cumberlidge, Neil</mods:namePart>
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<treatment id="038F3B05FF83FFD3FF2DCA50FCE6FCB6" ID-DOI="http://doi.org/10.5281/zenodo.6015969" ID-GBIF-Taxon="132548741" ID-Zenodo-Dep="6015969" LSID="urn:lsid:plazi:treatment:038F3B05FF83FFD3FF2DCA50FCE6FCB6" httpUri="http://treatment.plazi.org/id/038F3B05FF83FFD3FF2DCA50FCE6FCB6" lastPageId="10" lastPageNumber="238" pageId="8" pageNumber="236">
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<paragraph id="8B998A13FF83FFD1FF2DCA50FD06FCF3" blockId="8.[151,720,822,882]" box="[151,720,822,848]" pageId="8" pageNumber="236">
<heading id="D0D13D7FFF83FFD1FF2DCA50FD06FCF3" bold="true" box="[151,720,822,848]" fontSize="11" level="1" pageId="8" pageNumber="236" reason="1">
<taxonomicName id="4C26F190FF83FFD1FF2DCA50FD06FCF3" ID-CoL="3B29P" authority="A. Milne-Edwards, 1886" authorityName="A. Milne-Edwards" authorityYear="1886" box="[151,720,822,848]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="236" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF83FFD1FF2DCA50FD06FCF3" bold="true" box="[151,720,822,848]" pageId="8" pageNumber="236">
<emphasis id="B9525601FF83FFD1FF2DCA50FE53FCF3" bold="true" box="[151,389,822,848]" italics="true" pageId="8" pageNumber="236">Erimetopus brazzae</emphasis>
(
<bibRefCitation id="EFB7F7E2FF83FFD1FE2FCA50FD1EFCF3" author="Milne-Edwards" box="[405,712,822,848]" pageId="8" pageNumber="236" refString="Milne-Edwards, A. (1886) La description de quelques Crustaces du genre Thelphusa recueillis par M. de Brazza dans les regions du Congo. Bulletin de la Soci e t e philosophique, 7 (10), 148 - 151. [Paris]" type="journal article" year="1886">
<collectingRegion id="49E244F1FF83FFD1FE2FCA50FE7BFCF3" box="[405,429,822,848]" country="Nigeria" name="Rivers" pageId="8" pageNumber="236">A</collectingRegion>
. Milne-Edwards, 1886
</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B998A13FF83FFD1FF2DCA3FFF3DFCD1" blockId="8.[151,720,822,882]" box="[151,235,857,882]" pageId="8" pageNumber="236">
(
<figureCitation id="131D9696FF83FFD1FF25CA3FFF35FCD1" box="[159,227,857,882]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="8" pageNumber="236">Fig. 6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33CD998FF83FFD1FF2DCAC7FAC1FC7D" pageId="8" pageNumber="236" type="materials_examined">
<paragraph id="8B998A13FF83FFD1FF2DCAC7FAC1FC7D" blockId="8.[151,1437,929,2034]" pageId="8" pageNumber="236">
<materialsCitation id="3B4E804EFF83FFD1FF2DCAC7FAC5FC7D" ID-GBIF-Occurrence="1572325059" collectionCode="MCZ" country="Democratic Republic of the Congo" latitude="-5.260628" location="Material" longLatPrecision="1" longitude="13.7678" municipality="Kinganga" pageId="8" pageNumber="236" specimenCount="1" specimenCount-male="1" stateProvince="Congo">
<emphasis id="B9525601FF83FFD1FF2DCAC7FDE8FC19" bold="true" box="[151,574,929,954]" pageId="8" pageNumber="236">
<location id="8EF9DCC8FF83FFD1FF2DCAC7FF29FC19" LSID="urn:lsid:plazi:treatment:038F3B05FF83FFD3FF2DCA50FCE6FCB6:8EF9DCC8FF83FFD1FF2DCAC7FF29FC19" box="[151,255,929,954]" country="Democratic Republic of the Congo" latitude="-5.260628" longLatPrecision="1" longitude="13.7678" municipality="Kinganga" name="Material" pageId="8" pageNumber="236" stateProvince="Congo">Material</location>
examined. D. R.
<collectingCountry id="F331CA83FF83FFD1FE71CAC7FDEFFC19" box="[459,569,929,954]" name="Democratic Republic of the Congo" pageId="8" pageNumber="236">CONGO</collectingCountry>
:
</emphasis>
several kilometers northeast of
<collectingMunicipality id="6BFD1069FF83FFD1FC19CAC7FBC5FC19" box="[931,1043,929,954]" pageId="8" pageNumber="236">Kinganga</collectingMunicipality>
, Bas
<collectingRegion id="49E244F1FF83FFD1FBF5CAC4FB4BFC19" box="[1103,1181,930,954]" country="Republic of the Congo" name="Bas-Congo" pageId="8" pageNumber="236">Congo</collectingRegion>
, on the
<collectingRegion id="49E244F1FF83FFD1FB49CAC4FA54FC19" box="[1267,1410,929,954]" isEnumeration="true" pageId="8" pageNumber="236">Congo River</collectingRegion>
(-
<geoCoordinate id="EE12ECD4FF83FFD1FF2DCAA3FECEFC7D" box="[151,280,965,990]" direction="south" orientation="latitude" pageId="8" pageNumber="236" precision="1" value="-5.260628">5.260628 S</geoCoordinate>
,
<geoCoordinate id="EE12ECD4FF83FFD1FE9ECAA3FE60FC7D" box="[292,438,965,990]" direction="east" orientation="longitude" pageId="8" pageNumber="236" precision="1" value="13.7678">13.767800 E</geoCoordinate>
), subadult
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, CW 11, CL
<quantity id="4CDE27F6FF83FFD1FD66CAA0FCCAFC7E" box="[732,796,966,990]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="236" unit="mm" value="9.0">9 mm</quantity>
(collector and date unknown) (
<collectionCode id="ED3712D6FF83FFD1FBC1CAA0FB6CFC7D" box="[1147,1210,966,990]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="8" pageNumber="236">MCZ</collectionCode>
67421)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33CD998FF83FFD1FF7DCA8FFEEBF9C5" pageId="8" pageNumber="236" type="diagnosis">
<paragraph id="8B998A13FF83FFD1FF7DCA8FFBF5F9BD" blockId="8.[151,1437,929,2034]" pageId="8" pageNumber="236">
<emphasis id="B9525601FF83FFD1FF7DCA8FFEECFBA1" bold="true" box="[199,314,1001,1026]" pageId="8" pageNumber="236">Diagnosis</emphasis>
(emended from
<bibRefCitation id="EFB7F7E2FF83FFD1FE4CCA8FFCFAFBA1" author="Cumberlidge" box="[502,812,1001,1026]" pageId="8" pageNumber="236" refString="Cumberlidge, N. &amp; Reed, S. K. (2004) Erimetopus vandenbrandeni (Balss, 1936) n. comb., with notes on the taxonomy of the genus Erimetopus Rathbun, 1894 (Brachyura: Potamoidea: Potamonautidae) from Central Africa. Zootaxa, 422 (1), 1 - 27. https: // doi. org / 10.11646 / zootaxa. 422.1.1" type="journal article" year="2004">Cumberlidge &amp; Reed 2004</bibRefCitation>
). Carapace outline subhexagonal/rounded; epibranchial tooth large, sharp, pointing forwards, positioned well behind postfrontal crest; orbit narrow (1/7 CW), upper orbital margin semi-circular; anterolateral margin between exorbital, epibranchial teeth very long, curving slightly outward, with several small pointed teeth, without visible intermediate tooth (
<figureCitation id="131D9696FF83FFD1FC44CD33FB97FBCD" box="[1022,1089,1109,1134]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="8" pageNumber="236">Fig. 6</figureCitation>
<collectingRegion id="49E244F1FF83FFD1FBFBCD33FB8EFBCD" box="[1089,1112,1109,1134]" country="Nigeria" name="Rivers" pageId="8" pageNumber="236">A</collectingRegion>
). Exopod of third maxilliped with long flagellum, ischium smooth lacking vertical sulcus. Distal tooth on inner margin of cheliped carpus large, slender, pointed, curving forwards; outer margin of cheliped carpus with series of pointed teeth (
<figureCitation id="131D9696FF83FFD1FB6FCDFBFACFFB15" box="[1237,1305,1181,1206]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="8" pageNumber="236">Fig. 6</figureCitation>
F); superior margins of meri of p2p5 with 2 large, pointed distal teeth; anterior margins of carpi of p2p4 spiny, posterior margins smooth; anterior margins of propodi of p2p4 spiny, posterior margins smooth; propodus of p5 with spines on anterior, posterior margins. S1/s2 either faint or absent; s2/s3 shallow, completely crossing thoracic sternum; s3/ s4 reduced to 2 small side notches; s4 completely smooth; s4/e4, s5/e5, s6/e6, s7/e7 each marked by visible sulcus (
<figureCitation id="131D9696FF83FFD1FF25CC37FF35FAC9" box="[159,227,1361,1386]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="8" pageNumber="236">Fig. 6</figureCitation>
B). Pleon slim, outline triangular, tapered, widest at base, telson outline forming straight-sided triangle with broad base, rounded apex; s6/s7 meeting abdomen at a5/a6; s5/s6 meeting a6 one half of somite length from a6/a5 (
<figureCitation id="131D9696FF83FFD1FF25CCFFFF33FA11" box="[159,229,1433,1458]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="8" pageNumber="236">Fig. 6</figureCitation>
B). G1 terminal article one-third as long as subterminal segment, terminal article slim throughout length, tapering to pointed tip (
<figureCitation id="131D9696FF83FFD1FE1CCCDBFE3DFA75" box="[422,491,1469,1494]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="8" pageNumber="236">Fig. 6</figureCitation>
C, D), G1 terminal article straight basally turning sharply outwards at 45° angle at halfway point (
<figureCitation id="131D9696FF83FFD1FEF2CC87FE59FA59" box="[328,399,1505,1530]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="8" pageNumber="236">Fig. 6</figureCitation>
C, D), G1 dorsal membrane narrow, rectangular, sides parallel (
<figureCitation id="131D9696FF83FFD1FBCCCC87FB6BFA59" box="[1142,1213,1505,1530]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="8" pageNumber="236">Fig. 6</figureCitation>
D), G2 subterminal segment long, slender, G2 terminal article long, slender, flagellum-like (
<figureCitation id="131D9696FF83FFD1FC05CF63FBD4F9BD" box="[959,1026,1541,1566]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="8" pageNumber="236">Fig. 6</figureCitation>
E).
</paragraph>
<paragraph id="8B998A13FF83FFD1FF7DCF4FFEEBF9C5" blockId="8.[151,1437,929,2034]" pageId="8" pageNumber="236">
<emphasis id="B9525601FF83FFD1FF7DCF4FFF28F9E1" bold="true" box="[199,254,1577,1602]" pageId="8" pageNumber="236">Size.</emphasis>
<taxonomicName id="4C26F190FF83FFD1FEBECF4CFE35F9E1" box="[260,483,1577,1602]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="236" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF83FFD1FEBECF4CFE35F9E1" box="[260,483,1577,1602]" italics="true" pageId="8" pageNumber="236">Erimetopus brazzae</emphasis>
</taxonomicName>
is a small species with an adult size range between CW
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(
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).
</paragraph>
</subSubSection>
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<paragraph id="8B998A13FF83FFD1FF7DCF17FF24F955" blockId="8.[151,1437,929,2034]" pageId="8" pageNumber="236">
<materialsCitation id="3B4E804EFF83FFD1FF7DCF17FC4FF90E" ID-GBIF-Occurrence="1572325063" country="Republic of the Congo" latitude="-3.310963" location="Kwamouth" longLatPrecision="1" longitude="16.121973" municipality="Pool District" pageId="8" pageNumber="236" specimenCount="1" stateProvince="Congo" typeStatus="holotype">
<emphasis id="B9525601FF83FFD1FF7DCF17FEB3F929" bold="true" box="[199,357,1649,1674]" pageId="8" pageNumber="236">
<typeStatus id="549D34B1FF83FFD1FF7DCF17FED4F929" box="[199,258,1649,1674]" pageId="8" pageNumber="236">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F331CA83FF83FFD1FED6CF17FDBAF929" box="[364,620,1649,1674]" name="Republic of the Congo" pageId="8" pageNumber="236">Republic of the Congo</collectingCountry>
, Ngabe (= Ngancin) (-
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,
<geoCoordinate id="EE12ECD4FF83FFD1FC42CF17FB5CF929" box="[1016,1162,1649,1674]" direction="east" orientation="longitude" pageId="8" pageNumber="236" precision="1" value="16.121973">16.121973 E</geoCoordinate>
), in the
<collectingMunicipality id="6BFD1069FF83FFD1FB5ECF17FAA3F929" box="[1252,1397,1649,1674]" pageId="8" pageNumber="236">Pool District</collectingMunicipality>
, on the right bank of the
<collectingRegion id="49E244F1FF83FFD1FE36CFF0FE01F90D" box="[396,471,1686,1710]" country="Republic of the Congo" name="Bas-Congo" pageId="8" pageNumber="236">Congo</collectingRegion>
River opposite
<location id="8EF9DCC8FF83FFD1FD35CFF3FCDAF90D" LSID="urn:lsid:plazi:treatment:038F3B05FF83FFD3FF2DCA50FCE6FCB6:8EF9DCC8FF83FFD1FD35CFF3FCDAF90D" box="[655,780,1685,1710]" country="Republic of the Congo" latitude="-3.310963" longLatPrecision="1" longitude="16.121973" municipality="Pool District" name="Kwamouth" pageId="8" pageNumber="236" stateProvince="Congo">Kwamouth</location>
in the D. R
</materialsCitation>
.
<materialsCitation id="3B4E804EFF83FFD1FC1CCFF0FD2AF971" ID-GBIF-Occurrence="1572325061" country="Democratic Republic of the Congo" location="A" pageId="8" pageNumber="236" specimenCount="1" stateProvince="A" typeStatus="lectotype">
<collectingCountry id="F331CA83FF83FFD1FC1CCFF0FC25F90D" box="[934,1011,1686,1710]" name="Democratic Republic of the Congo" pageId="8" pageNumber="236">Congo</collectingCountry>
, adult female
<typeStatus id="549D34B1FF83FFD1FB2DCFF3FB28F90D" box="[1175,1278,1685,1710]" pageId="8" pageNumber="236" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C26F190FF83FFD1FA9DCFF3FDC3F971" authority="A. Milne-Edwards, 1886" authorityName="A. Milne-Edwards" authorityYear="1886" class="Malacostraca" family="Potamonautidae" genus="Thelphusa" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="236" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF83FFD1FA9DCFF3FF38F971" italics="true" pageId="8" pageNumber="236">Thelphusa brazzae</emphasis>
<bibRefCitation id="EFB7F7E2FF83FFD1FF4DCFDCFDC3F971" author="Milne-Edwards" box="[247,533,1721,1746]" pageId="8" pageNumber="236" refString="Milne-Edwards, A. (1886) La description de quelques Crustaces du genre Thelphusa recueillis par M. de Brazza dans les regions du Congo. Bulletin de la Soci e t e philosophique, 7 (10), 148 - 151. [Paris]" type="journal article" year="1886">
<collectingRegion id="49E244F1FF83FFD1FF4DCFDCFED8F971" box="[247,270,1722,1746]" country="Nigeria" name="Rivers" pageId="8" pageNumber="236">A</collectingRegion>
. Milne-Edwards, 1886
</bibRefCitation>
</taxonomicName>
, CW 23.5, CL 18.5
</materialsCitation>
,
<materialsCitation id="3B4E804EFF83FFD1FCB0CFDCFF24F955" ID-GBIF-Occurrence="1572325062" collectingDate="1884-04" collectionCode="MNHN" collectorName="M. de Brazza" country="Switzerland" location="CH" pageId="8" pageNumber="236" specimenCount="1" typeStatus="lectotype">
<collectingCountry id="F331CA83FF83FFD1FCB0CFDCFCE7F971" box="[778,817,1722,1746]" name="Switzerland" pageId="8" pageNumber="236">CH</collectingCountry>
8.0, FW 7.0 mm,
<collectingDate id="EFDC553BFF83FFD1FBB0CFDCFBABF971" box="[1034,1149,1722,1746]" pageId="8" pageNumber="236" value="1884-04">
<date id="FF98ACD3FF83FFD1FBB0CFDCFBABF971" box="[1034,1149,1722,1746]" pageId="8" pageNumber="236" value="1884-04">Apr. 1884</date>
</collectingDate>
(
<collectorName id="26D3EFC5FF83FFD1FB35CFDCFAFAF971" box="[1167,1324,1721,1746]" pageId="8" pageNumber="236">M. de Brazza</collectorName>
) (
<collectionCode id="ED3712D6FF83FFD1FAFFCFDCFA4AF971" box="[1349,1436,1722,1746]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="236">MNHN</collectionCode>
B5069).
</materialsCitation>
</paragraph>
<paragraph id="8B998A13FF83FFD0FF7DCE67FC76FF5B" blockId="8.[151,1437,929,2034]" lastBlockId="9.[151,1436,151,464]" lastPageId="9" lastPageNumber="237" pageId="8" pageNumber="236">
<emphasis id="B9525601FF83FFD1FF7DCE67FEFCF8B9" bold="true" box="[199,298,1793,1818]" pageId="8" pageNumber="236">Habitat.</emphasis>
<taxonomicName id="4C26F190FF83FFD1FE88CE64FDC5F8B9" box="[306,531,1793,1818]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="236" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF83FFD1FE88CE64FDC5F8B9" box="[306,531,1793,1818]" italics="true" pageId="8" pageNumber="236">Erimetopus brazzae</emphasis>
</taxonomicName>
is found on both sides of the lower Congo River in the Republic of the Congo and in the D. R. Congo where the river is about
<quantity id="4CDE27F6FF83FFD1FD79CE43FCC5F89D" box="[707,787,1829,1854]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="236" unit="km" value="1.5">1.5 km</quantity>
wide and the water is deep and fast flowing because the Congo River is joined by the Ubangi, Sangha, and Kwa rivers. This species is also found in the Malebo Pool, a large, lake-like widened stretch of the Congo River (about
<quantity id="4CDE27F6FF83FFD1FC9DCE0BFCB8F825" box="[807,878,1901,1926]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="236" unit="km" value="25.0">25 km</quantity>
by
<quantity id="4CDE27F6FF83FFD1FC22CE08FC36F825" box="[920,992,1901,1926]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="236" unit="km" value="30.0">30 km</quantity>
) close to Kinshasa, D. R. Congo. All of the adult egg-bearing females of
<taxonomicName id="4C26F190FF83FFD1FDB0CEF4FD56F809" box="[522,640,1937,1962]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="236" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF83FFD1FDB0CEF4FD56F809" box="[522,640,1937,1962]" italics="true" pageId="8" pageNumber="236">E. brazzae</emphasis>
</taxonomicName>
so far have been found either in the swampy areas of the lower Congo River and its tributaries or on the banks of the Malebo Pool, rather than in the river itself (
<bibRefCitation id="EFB7F7E2FF83FFD1FB1DCED3FF02F851" author="Cumberlidge" pageId="8" pageNumber="236" refString="Cumberlidge, N. &amp; Reed, S. K. (2004) Erimetopus vandenbrandeni (Balss, 1936) n. comb., with notes on the taxonomy of the genus Erimetopus Rathbun, 1894 (Brachyura: Potamoidea: Potamonautidae) from Central Africa. Zootaxa, 422 (1), 1 - 27. https: // doi. org / 10.11646 / zootaxa. 422.1.1" type="journal article" year="2004">Cumberlidge &amp; Reed 2004</bibRefCitation>
). The breeding season for
<taxonomicName id="4C26F190FF83FFD1FE45CEBCFDA0F851" box="[511,630,2009,2034]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="236" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF83FFD1FE45CEBCFDA0F851" box="[511,630,2009,2034]" italics="true" pageId="8" pageNumber="236">E. brazzae</emphasis>
</taxonomicName>
is between April and August based on when egg-bearing females have been collected, and this species appears to favor the muddy ground near river banks where it lives either under stones, under boardwalks, or tangled in water plants. (
<bibRefCitation id="EFB7F7E2FF82FFD0FCB1C9DAFB9CFF77" author="Cumberlidge" box="[779,1098,188,213]" pageId="9" pageNumber="237" refString="Cumberlidge, N. &amp; Reed, S. K. (2004) Erimetopus vandenbrandeni (Balss, 1936) n. comb., with notes on the taxonomy of the genus Erimetopus Rathbun, 1894 (Brachyura: Potamoidea: Potamonautidae) from Central Africa. Zootaxa, 422 (1), 1 - 27. https: // doi. org / 10.11646 / zootaxa. 422.1.1" type="journal article" year="2004">Cumberlidge &amp; Reed 2004</bibRefCitation>
). At Malebo Pool
<taxonomicName id="4C26F190FF82FFD0FA99C9DBFA4AFF76" box="[1315,1436,188,213]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="237" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF82FFD0FA99C9DBFA4AFF76" box="[1315,1436,188,213]" italics="true" pageId="9" pageNumber="237">E. brazzae</emphasis>
</taxonomicName>
occurs together with
<taxonomicName id="4C26F190FF82FFD0FE38C987FE21FF5B" box="[386,503,223,248]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="237" phylum="Arthropoda" rank="species" species="paecilei">
<emphasis id="B9525601FF82FFD0FE38C987FE21FF5B" box="[386,503,223,248]" italics="true" pageId="9" pageNumber="237">P. paecilei</emphasis>
</taxonomicName>
in the mud under boards and timbers.
</paragraph>
</subSubSection>
<subSubSection id="C33CD998FF82FFD0FF7DC862FD57FE6C" pageId="9" pageNumber="237" type="distribution">
<paragraph id="8B998A13FF82FFD0FF7DC862FD57FE6C" blockId="9.[151,1436,151,464]" pageId="9" pageNumber="237">
<emphasis id="B9525601FF82FFD0FF7DC862FE8FFEBE" bold="true" box="[199,345,260,285]" pageId="9" pageNumber="237">Distribution</emphasis>
.
<taxonomicName id="4C26F190FF82FFD0FEDDC863FD91FEBE" box="[359,583,260,285]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="237" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF82FFD0FEDDC863FD91FEBE" box="[359,583,260,285]" italics="true" pageId="9" pageNumber="237">Erimetopus brazzae</emphasis>
</taxonomicName>
is a
<collectingRegion id="49E244F1FF82FFD0FDC6C862FD19FEBE" box="[636,719,260,285]" country="Togo" name="Centrale" pageId="9" pageNumber="237">Central</collectingRegion>
African species associated with the lower reaches of the
<collectingRegion id="49E244F1FF82FFD0FAEBC862FA4AFEBF" box="[1361,1436,260,284]" country="Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="237">
<collectingCountry id="F331CA83FF82FFD0FAEBC862FA4AFEBF" box="[1361,1436,260,284]" name="Democratic Republic of the Congo" pageId="9" pageNumber="237">Congo</collectingCountry>
</collectingRegion>
<collectingRegion id="49E244F1FF82FFD0FF2DC841FF03FEE3" box="[151,213,295,320]" country="Nigeria" name="Rivers" pageId="9" pageNumber="237">River</collectingRegion>
at Ngabe (= Ngancin), Pool District,
<collectingCountry id="F331CA83FF82FFD0FD3CC841FC5FFEE3" box="[646,905,295,320]" name="Republic of the Congo" pageId="9" pageNumber="237">
Republic of the
<collectingRegion id="49E244F1FF82FFD0FC84C84EFC5FFEE3" box="[830,905,296,320]" country="Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="237">Congo</collectingRegion>
</collectingCountry>
(-
<geoCoordinate id="EE12ECD4FF82FFD0FC1AC84EFBF1FEE3" box="[928,1063,296,320]" direction="south" orientation="latitude" pageId="9" pageNumber="237" precision="1" value="-4.910339">4.910339 S</geoCoordinate>
,
<geoCoordinate id="EE12ECD4FF82FFD0FB8FC841FB11FEE3" box="[1077,1223,295,320]" direction="east" orientation="longitude" pageId="9" pageNumber="237" precision="1" value="14.688858">14.688858 E</geoCoordinate>
) across the
<collectingRegion id="49E244F1FF82FFD0FAEBC84EFA4AFEE3" box="[1361,1436,296,320]" country="Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="237">
<collectingCountry id="F331CA83FF82FFD0FAEBC84EFA4AFEE3" box="[1361,1436,296,320]" name="Democratic Republic of the Congo" pageId="9" pageNumber="237">Congo</collectingCountry>
</collectingRegion>
<collectingRegion id="49E244F1FF82FFD0FF2DC82AFF03FEC6" box="[151,213,332,357]" country="Nigeria" name="Rivers" pageId="9" pageNumber="237">River</collectingRegion>
from Gombe-Matadi, D. R.
<collectingRegion id="49E244F1FF82FFD0FDA7C82AFDBDFEC7" box="[541,619,332,356]" country="Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="237">
<collectingCountry id="F331CA83FF82FFD0FDA7C82AFDBDFEC7" box="[541,619,332,356]" name="Democratic Republic of the Congo" pageId="9" pageNumber="237">Congo</collectingCountry>
</collectingRegion>
. This species is also found in the Kwa, Kwilu, and
<collectingRegion id="49E244F1FF82FFD0FB03C82BFACEFEC6" box="[1209,1304,333,357]" country="Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="237">Kwango</collectingRegion>
<collectingRegion id="49E244F1FF82FFD0FA9BC82AFABCFEC6" box="[1313,1386,332,357]" country="Nigeria" name="Rivers" pageId="9" pageNumber="237">Rivers</collectingRegion>
that are all southeastern tributaries of the
<collectingRegion id="49E244F1FF82FFD0FDFBC816FD5AFE2B" box="[577,652,368,392]" country="Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="237">
<collectingCountry id="F331CA83FF82FFD0FDFBC816FD5AFE2B" box="[577,652,368,392]" name="Democratic Republic of the Congo" pageId="9" pageNumber="237">Congo</collectingCountry>
</collectingRegion>
<collectingRegion id="49E244F1FF82FFD0FD2EC809FD04FE2B" box="[660,722,367,392]" country="Nigeria" name="Rivers" pageId="9" pageNumber="237">River</collectingRegion>
in the D. R.
<collectingRegion id="49E244F1FF82FFD0FCD1C816FC60FE2B" box="[875,950,368,392]" country="Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="237">
<collectingCountry id="F331CA83FF82FFD0FCD1C816FC60FE2B" box="[875,950,368,392]" name="Democratic Republic of the Congo" pageId="9" pageNumber="237">Congo</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="EFB7F7E2FF82FFD0FC7CC809FA8DFE2B" author="Cumberlidge" box="[966,1371,367,392]" pageId="9" pageNumber="237" refString="Cumberlidge, N. &amp; Reed, S. K. (2004) Erimetopus vandenbrandeni (Balss, 1936) n. comb., with notes on the taxonomy of the genus Erimetopus Rathbun, 1894 (Brachyura: Potamoidea: Potamonautidae) from Central Africa. Zootaxa, 422 (1), 1 - 27. https: // doi. org / 10.11646 / zootaxa. 422.1.1" type="journal article" year="2004">Cumberlidge &amp; Reed 2004: fig. 45</bibRefCitation>
). The report by
<bibRefCitation id="EFB7F7E2FF82FFD0FEBFC8F2FE60FE0F" author="Rathbun" box="[261,438,404,429]" pageId="9" pageNumber="237" refString="Rathbun, M. J. (1894) Descriptions of a new genus and two new species of African fresh-water crabs. Proceedings of the United States National Museum, 17 (180), 25 - 27." type="journal article" year="1894">Rathbun (1894)</bibRefCitation>
that this species occurs in
<collectingCountry id="F331CA83FF82FFD0FD5CC8F2FCE7FE0E" box="[742,817,404,429]" name="Gabon" pageId="9" pageNumber="237">Gabon</collectingCountry>
remains unconfirmed and it is not included here in the species list for that country for the moment.
</paragraph>
</subSubSection>
<caption id="DF59DA9BFF82FFD0FF2DCC83FBCBF99B" httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="9" pageNumber="237" targetBox="[252,1324,546,1477]" targetPageId="9">
<paragraph id="8B998A13FF82FFD0FF2DCC83FBCBF99B" blockId="9.[151,1436,1508,1592]" pageId="9" pageNumber="237">
<emphasis id="B9525601FF82FFD0FF2DCC83FEC5FA59" bold="true" box="[151,275,1509,1531]" pageId="9" pageNumber="237">FIGURE 6.</emphasis>
<taxonomicName id="4C26F190FF82FFD0FEA0CC80FD2BFA59" authority="A. Milne-Edwards, 1886" authorityName="A. Milne-Edwards" authorityYear="1886" box="[282,765,1508,1531]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="237" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF82FFD0FEA0CC80FE33FA59" box="[282,485,1508,1531]" italics="true" pageId="9" pageNumber="237">Erimetopus brazzae</emphasis>
(A. Milne-Edwards, 1886)
</taxonomicName>
. Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G1 ventral view. D. G1 dorsal view. E. G2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (CE).
</paragraph>
</caption>
<subSubSection id="C33CD998FF82FFD3FF7DCF02FCE6FCB6" lastPageId="10" lastPageNumber="238" pageId="9" pageNumber="237" type="discussion">
<paragraph id="8B998A13FF82FFD3FF7DCF02FDBCFE73" blockId="9.[151,1437,1636,2022]" lastBlockId="10.[151,1437,151,789]" lastPageId="10" lastPageNumber="238" pageId="9" pageNumber="237">
<emphasis id="B9525601FF82FFD0FF7DCF02FEEDF9DE" bold="true" box="[199,315,1636,1661]" pageId="9" pageNumber="237">Remarks.</emphasis>
<taxonomicName id="4C26F190FF82FFD0FEF9CF00FDF2F9DE" box="[323,548,1636,1661]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="237" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF82FFD0FEF9CF00FDF2F9DE" box="[323,548,1636,1661]" italics="true" pageId="9" pageNumber="237">Erimetopus brazzae</emphasis>
</taxonomicName>
was redescribed by
<bibRefCitation id="EFB7F7E2FF82FFD0FCB5CF02FB81F9DE" author="Cumberlidge" box="[783,1111,1636,1661]" pageId="9" pageNumber="237" refString="Cumberlidge, N. &amp; Reed, S. K. (2004) Erimetopus vandenbrandeni (Balss, 1936) n. comb., with notes on the taxonomy of the genus Erimetopus Rathbun, 1894 (Brachyura: Potamoidea: Potamonautidae) from Central Africa. Zootaxa, 422 (1), 1 - 27. https: // doi. org / 10.11646 / zootaxa. 422.1.1" type="journal article" year="2004">Cumberlidge &amp; Reed (2004)</bibRefCitation>
, who referred to all relevant
<typeStatus id="549D34B1FF82FFD0FF2DCFEFFF11F902" box="[151,199,1673,1697]" pageId="9" pageNumber="237">type</typeStatus>
material plus the descriptions and illustrations of the dorsal carapace of the female
<typeStatus id="549D34B1FF82FFD0FB21CFEEFAD4F902" box="[1179,1282,1672,1697]" pageId="9" pageNumber="237" type="lectotype">lectotype</typeStatus>
from Ngabé,
<collectingCountry id="F331CA83FF82FFD0FF2DCFCBFE77F966" box="[151,417,1709,1734]" name="Republic of the Congo" pageId="9" pageNumber="237">
Republic of the
<collectingRegion id="49E244F1FF82FFD0FEECCFCBFE77F966" box="[342,417,1709,1733]" country="Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="237">Congo</collectingRegion>
</collectingCountry>
by
<bibRefCitation id="EFB7F7E2FF82FFD0FE69CFCBFD56F966" author="Rathbun" box="[467,640,1709,1734]" pageId="9" pageNumber="237" refString="Rathbun, M. J. (1894) Descriptions of a new genus and two new species of African fresh-water crabs. Proceedings of the United States National Museum, 17 (180), 25 - 27." type="journal article" year="1894">Rathbun (1894</bibRefCitation>
,
<bibRefCitation id="EFB7F7E2FF82FFD0FD35CFCBFD1EF966" author="Rathbun" box="[655,712,1709,1733]" pageId="9" pageNumber="237" refString="Rathbun, M. J. (1921) Brachyuran Crabs of the Belgian Congo. Bulletin of the American Museum of Natural History, 43, 379 - 468." type="journal article" year="1921">1921</bibRefCitation>
) and
<bibRefCitation id="EFB7F7E2FF82FFD0FCB5CFCBFBC4F966" author="Capart" box="[783,1042,1709,1734]" pageId="9" pageNumber="237" refString="Capart, A. (1954) Revision des types des especes de Potamonidae de l'Afrique Tropicale conserves au Museum d'Histoire Naturelle de Paris. Folume Jubilaire de Fictor Fan Strallen, Director de l'Institut royal des Sciences naturelles de Belgique, 1925 - 1934, II, 819 - 847." type="book chapter" year="1954">Capart (1954: fig. 43)</bibRefCitation>
. The description of the species is enhanced here by the addition of characters of the sternum and pleon of the male specimen of
<taxonomicName id="4C26F190FF82FFD0FB02CFB4FAF8F94A" box="[1208,1326,1744,1769]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="237" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF82FFD0FB02CFB4FAF8F94A" box="[1208,1326,1744,1769]" italics="true" pageId="9" pageNumber="237">E. brazzae</emphasis>
</taxonomicName>
(
<figureCitation id="131D9696FF82FFD0FA87CFB6FA56F94A" box="[1341,1408,1744,1769]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="9" pageNumber="237">Fig. 6</figureCitation>
B) that was judged to be a subadult based on the size of adult females of this species that measure between CW 23.5 and
<quantity id="4CDE27F6FF82FFD0FF72CE7EFEF8F893" box="[200,302,1816,1841]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" pageId="9" pageNumber="237" unit="mm" value="33.5">33.5 mm</quantity>
(
<bibRefCitation id="EFB7F7E2FF82FFD0FE87CE7EFDAEF892" author="Cumberlidge" box="[317,632,1816,1841]" pageId="9" pageNumber="237" refString="Cumberlidge, N. &amp; Reed, S. K. (2004) Erimetopus vandenbrandeni (Balss, 1936) n. comb., with notes on the taxonomy of the genus Erimetopus Rathbun, 1894 (Brachyura: Potamoidea: Potamonautidae) from Central Africa. Zootaxa, 422 (1), 1 - 27. https: // doi. org / 10.11646 / zootaxa. 422.1.1" type="journal article" year="2004">Cumberlidge &amp; Reed 2004</bibRefCitation>
). Many of the morphological characters commonly used for taxonomy (such as those of the third maxilliped, thoracic sternum, mandible, and the merus and carpus of the cheliped) undergo isometric growth and are not likely to be significantly different in subadults and adults. Because the G1, G2 and the major cheliped of male freshwater crabs grow allometrically, however, the subadult male of
<taxonomicName id="4C26F190FF82FFD0FA9FCEE0FA4AF83D" box="[1317,1436,1925,1950]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="237" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF82FFD0FA9FCEE0FA4AF83D" box="[1317,1436,1925,1950]" italics="true" pageId="9" pageNumber="237">E. brazzae</emphasis>
</taxonomicName>
has incompletely developed gonopods and chelipeds (
<figureCitation id="131D9696FF82FFD0FCA2CECEFCB7F862" box="[792,865,1960,1985]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="9" pageNumber="237">Fig. 6</figureCitation>
<collectingRegion id="49E244F1FF82FFD0FCDBCECEFCAEF862" box="[865,888,1960,1985]" country="Nigeria" name="Rivers" pageId="9" pageNumber="237">A</collectingRegion>
). Despite this, it is still possible to make the following comparisons with the adult gonopods of
<taxonomicName id="4C26F190FF82FFD0FD53CEA8FBB2F845" authority="Balss, 1936" authorityName="Balss" authorityYear="1936" box="[745,1124,1997,2022]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="237" phylum="Arthropoda" rank="species" species="vandenbrandeni">
<emphasis id="B9525601FF82FFD0FD53CEA8FC17F845" box="[745,961,1997,2022]" italics="true" pageId="9" pageNumber="237">E. vandenbrandeni</emphasis>
(Balss, 1936)
</taxonomicName>
(see
<bibRefCitation id="EFB7F7E2FF82FFD3FB1FCEABFF02FF13" author="Cumberlidge" lastPageId="10" lastPageNumber="238" pageId="9" pageNumber="237" refString="Cumberlidge, N. &amp; Reed, S. K. (2004) Erimetopus vandenbrandeni (Balss, 1936) n. comb., with notes on the taxonomy of the genus Erimetopus Rathbun, 1894 (Brachyura: Potamoidea: Potamonautidae) from Central Africa. Zootaxa, 422 (1), 1 - 27. https: // doi. org / 10.11646 / zootaxa. 422.1.1" type="journal article" year="2004">Cumberlidge &amp; Reed 2004</bibRefCitation>
: figs. 3133, 4244). The G1 of
<taxonomicName id="4C26F190FF81FFD3FD85C9FFFD63FF13" box="[575,693,151,176]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF81FFD3FD85C9FFFD63FF13" box="[575,693,151,176]" italics="true" pageId="10" pageNumber="238">E. brazzae</emphasis>
</taxonomicName>
(
<figureCitation id="131D9696FF81FFD3FD7FC9F1FCDEFF13" box="[709,776,151,176]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="10" pageNumber="238">Fig. 6</figureCitation>
C, D) is similar in most respects to that of the adult male of
<taxonomicName id="4C26F190FF81FFD3FF2DC9DBFEBDFF76" box="[151,363,188,213]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="species" species="vandenbrandeni">
<emphasis id="B9525601FF81FFD3FF2DC9DBFEBDFF76" box="[151,363,188,213]" italics="true" pageId="10" pageNumber="238">E. vandenbrandeni</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB7F7E2FF81FFD3FEC3C9DAFC88FF77" author="Cumberlidge" box="[377,862,188,213]" pageId="10" pageNumber="238" refString="Cumberlidge, N. &amp; Reed, S. K. (2004) Erimetopus vandenbrandeni (Balss, 1936) n. comb., with notes on the taxonomy of the genus Erimetopus Rathbun, 1894 (Brachyura: Potamoidea: Potamonautidae) from Central Africa. Zootaxa, 422 (1), 1 - 27. https: // doi. org / 10.11646 / zootaxa. 422.1.1" type="journal article" year="2004">Cumberlidge &amp; Reed 2004: figs. 27, 29, 41</bibRefCitation>
), but the G1 of
<taxonomicName id="4C26F190FF81FFD3FBB6C9DBFB54FF76" box="[1036,1154,188,213]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF81FFD3FBB6C9DBFB54FF76" box="[1036,1154,188,213]" italics="true" pageId="10" pageNumber="238">E. brazzae</emphasis>
</taxonomicName>
(
<figureCitation id="131D9696FF81FFD3FB2BC9DAFB05FF76" box="[1169,1235,188,213]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="10" pageNumber="238">Fig. 6</figureCitation>
C, D) differs from that of
<taxonomicName id="4C26F190FF81FFD3FF5FC987FE6CFF5B" box="[229,442,223,248]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="species" species="vandenbrandeni">
<emphasis id="B9525601FF81FFD3FF5FC987FE6CFF5B" box="[229,442,223,248]" italics="true" pageId="10" pageNumber="238">E. vandenbrandeni</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB7F7E2FF81FFD3FE72C9B9FC00FF5B" author="Cumberlidge" box="[456,982,223,248]" pageId="10" pageNumber="238" refString="Cumberlidge, N. &amp; Reed, S. K. (2004) Erimetopus vandenbrandeni (Balss, 1936) n. comb., with notes on the taxonomy of the genus Erimetopus Rathbun, 1894 (Brachyura: Potamoidea: Potamonautidae) from Central Africa. Zootaxa, 422 (1), 1 - 27. https: // doi. org / 10.11646 / zootaxa. 422.1.1" type="journal article" year="2004">Cumberlidge &amp; Reed 2004: figs. 3133, 4244</bibRefCitation>
) as follows: the G1 terminal article of
<taxonomicName id="4C26F190FF81FFD3FA3EC987FF38FEBE" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF81FFD3FA3EC987FF38FEBE" italics="true" pageId="10" pageNumber="238">E. brazzae</emphasis>
</taxonomicName>
is slim and tapers to a pointed tip (
<figureCitation id="131D9696FF81FFD3FD3EC862FD1FFEBE" box="[644,713,260,285]" captionStart="FIGURE 6" captionStartId="9.[151,250,1509,1531]" captionTargetBox="[252,1324,546,1477]" captionTargetId="figure@9.[238,1348,517,1487]" captionTargetPageId="9" captionText="FIGURE 6. Erimetopus brazzae (A. Milne-Edwards, 1886). Subadult male, CW 11 mm, from Kinganga, D. R. Congo (MCZ 67421). A. Carapace, dorsal view. B. thoracic sternum and pleon. C. G 1 ventral view. D. G 1 dorsal view. E. G 2 ventral view. F. merus, carpus of right cheliped, dorsal view. Scale bar: 3.0 mm (A, B, F), 1.4 mm (C E)." httpUri="https://zenodo.org/record/828449/files/figure.png" pageId="10" pageNumber="238">Fig. 6</figureCitation>
C, D) (tubular with parallel sides and ends in a broad tip with a distinctly upturned triangular corner in
<taxonomicName id="4C26F190FF81FFD3FDF5C84FFCF5FEE3" box="[591,803,295,320]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="species" species="vandenbrandeni">
<emphasis id="B9525601FF81FFD3FDF5C84FFCF5FEE3" box="[591,803,295,320]" italics="true" pageId="10" pageNumber="238">E. vandenbrandeni</emphasis>
</taxonomicName>
), the G1 terminal article of
<taxonomicName id="4C26F190FF81FFD3FBE6C84FFB04FEE3" box="[1116,1234,295,320]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF81FFD3FBE6C84FFB04FEE3" box="[1116,1234,295,320]" italics="true" pageId="10" pageNumber="238">E. brazzae</emphasis>
</taxonomicName>
is straight basally and then turns sharply outward at a 45° angle to the longitudinal axis of the gonopod (straight along its entire length in
<taxonomicName id="4C26F190FF81FFD3FF0EC817FE5CFE2B" box="[180,394,367,392]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="species" species="vandenbrandeni">
<emphasis id="B9525601FF81FFD3FF0EC817FE5CFE2B" box="[180,394,367,392]" italics="true" pageId="10" pageNumber="238">E. vandenbrandeni</emphasis>
</taxonomicName>
), and the G1 dorsal membrane of
<taxonomicName id="4C26F190FF81FFD3FCB7C817FC55FE2B" box="[781,899,367,392]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="B9525601FF81FFD3FCB7C817FC55FE2B" box="[781,899,367,392]" italics="true" pageId="10" pageNumber="238">E. brazzae</emphasis>
</taxonomicName>
is a narrow rectangle with parallel sides (broad membrane with a straight superior margin and medially-indented inferior margin in
<taxonomicName id="4C26F190FF81FFD3FB09C8F3FA44FE0E" box="[1203,1426,404,429]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="species" species="vandenbrandeni">
<emphasis id="B9525601FF81FFD3FB09C8F3FA44FE0E" box="[1203,1426,404,429]" italics="true" pageId="10" pageNumber="238">E. vandenbrandeni</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFB7F7E2FF81FFD3FF24C8D1FD8BFE73" author="Cumberlidge" box="[158,605,439,464]" pageId="10" pageNumber="238" refString="Cumberlidge, N. &amp; Reed, S. K. (2004) Erimetopus vandenbrandeni (Balss, 1936) n. comb., with notes on the taxonomy of the genus Erimetopus Rathbun, 1894 (Brachyura: Potamoidea: Potamonautidae) from Central Africa. Zootaxa, 422 (1), 1 - 27. https: // doi. org / 10.11646 / zootaxa. 422.1.1" type="journal article" year="2004">Cumberlidge &amp; Reed 2004: figs. 3233</bibRefCitation>
).
</paragraph>
<paragraph id="8B998A13FF81FFD3FF7DC8BAFCE6FCB6" blockId="10.[151,1437,151,789]" pageId="10" pageNumber="238">
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The two species of
<taxonomicName id="4C26F190FF81FFD3FE24C8BBFDC9FE57" box="[414,543,477,500]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="B9525601FF81FFD3FE24C8BBFDC9FE57" box="[414,543,477,500]" italics="true" pageId="10" pageNumber="238">Erimetopus</emphasis>
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are found in the
<collectingRegion id="49E244F1FF81FFD3FD61C8BBFCA0FE57" box="[731,886,476,500]" country="Republic of the Congo" name="Bas-Congo" pageId="10" pageNumber="238">Lower Congo</collectingRegion>
River basin in the
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and the D. R
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. Congo (
<bibRefCitation id="EFB7F7E2FF81FFD3FE8DC899FDA7FDBB" author="Cumberlidge" box="[311,625,511,536]" pageId="10" pageNumber="238" refString="Cumberlidge, N. &amp; Reed, S. K. (2004) Erimetopus vandenbrandeni (Balss, 1936) n. comb., with notes on the taxonomy of the genus Erimetopus Rathbun, 1894 (Brachyura: Potamoidea: Potamonautidae) from Central Africa. Zootaxa, 422 (1), 1 - 27. https: // doi. org / 10.11646 / zootaxa. 422.1.1" type="journal article" year="2004">Cumberlidge &amp; Reed 2004</bibRefCitation>
).
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<collectionCode id="ED3712D6FF81FFD3FD3ECB66FD17FDBB" box="[644,705,512,536]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="10" pageNumber="238">DNA</collectionCode>
recently extracted from an egg of an adult female
<typeStatus id="549D34B1FF81FFD3FABFCB66FA8AFDBB" box="[1285,1372,512,536]" pageId="10" pageNumber="238" type="syntype">syntype</typeStatus>
of
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<emphasis id="B9525601FF81FFD3FA3ECB67FF38FD9E" italics="true" pageId="10" pageNumber="238">E. brazzae</emphasis>
</taxonomicName>
(
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<collectionCode id="ED3712D6FF81FFD3FF46CB42FEEDFD9F" box="[252,315,548,572]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="10" pageNumber="238">MCZ</collectionCode>
4255
</specimenCode>
) from Malebo Pool in the
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in the D. R
</materialsCitation>
.
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<collectingCountry id="F331CA83FF81FFD3FC04CB42FBDFFD9F" box="[958,1033,548,572]" name="Democratic Republic of the Congo" pageId="10" pageNumber="238">Congo</collectingCountry>
allowed this taxon to be included in a molecular phylogenetic analysis of the entire Afrotropical freshwater crab fauna (
<bibRefCitation id="EFB7F7E2FF81FFD3FB83CB21FAC4FDC3" author="Daniels" box="[1081,1298,583,608]" pageId="10" pageNumber="238" refString="Daniels, S. R., Phiri, E. E., Klaus, S., Albrecht, C. &amp; Cumberlidge, N. (2015) Multilocus phylogeny of the Afrotropical freshwater crab fauna reveals historical drainage connectivity and transoceanic dispersal since the Eocene. Systematic Biology, 64, 549 - 567." type="journal article" year="2015">
Daniels
<emphasis id="B9525601FF81FFD3FB2DCB2FFB1EFDC3" box="[1175,1224,583,608]" italics="true" pageId="10" pageNumber="238">et al</emphasis>
. 2015
</bibRefCitation>
: table 1, fig. 2), a study that positioned
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<emphasis id="B9525601FF81FFD3FE7ECB0BFD93FD27" box="[452,581,621,644]" italics="true" pageId="10" pageNumber="238">Erimetopus</emphasis>
</taxonomicName>
as a sister group to
<taxonomicName id="4C26F190FF81FFD3FC93CB0BFC45FD26" box="[809,915,620,645]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="species" species="ballayi">
<emphasis id="B9525601FF81FFD3FC93CB0BFC45FD26" box="[809,915,620,645]" italics="true" pageId="10" pageNumber="238">P. ballayi</emphasis>
</taxonomicName>
from the
<collectingRegion id="49E244F1FF81FFD3FBB9CB0BFB32FD26" box="[1027,1252,620,645]" isEnumeration="true" pageId="10" pageNumber="238">Lower Congo River</collectingRegion>
basin in a clade that also included
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<emphasis id="B9525601FF81FFD3FED5CBF7FE3CFD0B" box="[367,490,657,680]" italics="true" pageId="10" pageNumber="238">P. ecorssei</emphasis>
(
<bibRefCitation id="EFB7F7E2FF81FFD3FE46CBE9FD68FD0B" author="Marchand" box="[508,702,655,680]" pageId="10" pageNumber="238" refString="Marchand, E. (1902) Description de deux Arthropodes nouveaux provenant du Soudan francais (Trax Borgnognoi) (Coleopt.) et Pot. (Potamonautes) Ecorssei (Crust.). Bulletin de la Societe des Sciences naturelles de l'Ouest de la France (Nantes), 2, 331 - 342." type="journal article" year="1902">Marchand, 1902</bibRefCitation>
)
</taxonomicName>
, from
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</materialsCitation>
,
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<location id="8EF9DCC8FF81FFD3FCE7CBF6FC3FFD0B" LSID="urn:lsid:plazi:treatment:038F3B05FF83FFD3FF2DCA50FCE6FCB6:8EF9DCC8FF81FFD3FCE7CBF6FC3FFD0B" box="[861,1001,655,680]" country="Sao Tome and Principe" name="West Africa" pageId="10" pageNumber="238">West Africa</location>
, and another species from
<collectingCountry id="F331CA83FF81FFD3FA92CBF6FA4AFD0B" box="[1320,1436,656,680]" name="Sao Tome and Principe" pageId="10" pageNumber="238">São Tomé</collectingCountry>
(
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<emphasis id="B9525601FF81FFD3FF13CBD3FE9FFD6F" box="[169,329,693,716]" italics="true" pageId="10" pageNumber="238">Potamonautes</emphasis>
</taxonomicName>
sp. 1 in
<bibRefCitation id="EFB7F7E2FF81FFD3FE0CCBD2FD46FD6E" author="Daniels" box="[438,656,692,717]" pageId="10" pageNumber="238" refString="Daniels, S. R., Phiri, E. E., Klaus, S., Albrecht, C. &amp; Cumberlidge, N. (2015) Multilocus phylogeny of the Afrotropical freshwater crab fauna reveals historical drainage connectivity and transoceanic dispersal since the Eocene. Systematic Biology, 64, 549 - 567." type="journal article" year="2015">
Daniels
<emphasis id="B9525601FF81FFD3FDA9CBD3FD93FD6E" box="[531,581,692,717]" italics="true" pageId="10" pageNumber="238">et al</emphasis>
. 2015
</bibRefCitation>
: table 1, fig. 2). This discovery raises questions about the stability of
<taxonomicName id="4C26F190FF81FFD3FF2DCBBFFEE1FD53" box="[151,311,729,752]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="B9525601FF81FFD3FF2DCBBFFEE1FD53" box="[151,311,729,752]" italics="true" pageId="10" pageNumber="238">Potamonautes</emphasis>
</taxonomicName>
because this genus is rendered paraphyletic if
<taxonomicName id="4C26F190FF81FFD3FCF4CBBFFC19FD53" box="[846,975,729,752]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="B9525601FF81FFD3FCF4CBBFFC19FD53" box="[846,975,729,752]" italics="true" pageId="10" pageNumber="238">Erimetopus</emphasis>
</taxonomicName>
is nested within it. This is a noteworthy finding that requires further taxonomic studies to elucidate.
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</paragraph>
</subSubSection>
</treatment>
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