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<mods:title id="E74608E8F9C37101E7E27919E46D18AF">A taxonomic synopsis of Chrysomydas Wilcox, Papavero &amp; Pimentel (Diptera: Mydidae: Mydinae), with first bionomic records for the genus and description of a new species from Brazil</mods:title>
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<emphasis id="B95EEAFBF22AFFDC2BF1D46009B2802D" bold="true" box="[151,419,1365,1392]" italics="true" pageId="4" pageNumber="107">Chrysomydas phoenix</emphasis>
Calhau &amp; Lamas
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(
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A–F, 3H–K, 4F–J, 6)
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<emphasis id="B95EEAFBF22AFFDC2BF1D4EB090380A4" bold="true" box="[151,274,1502,1529]" pageId="4" pageNumber="107">Diagnosis.</emphasis>
Labellum about 1.7x longer than wide. T2–T5 with discrete green metallic reflection, with long black setulae, directed outwards. Scutum without distinct pruinose spots. Male cerci exposed, uncovered by epandrial lamellae; distal process of the gonocoxite-hypandrial complex apically wide, truncate, with roundish dorsal tip and acute ventral tip.
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Description. Male 
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.
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Greatest length, excluding antennae 
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. 
<emphasis id="B95EEAFBF22AFFDC2F54D75A0C6383D4" bold="true" box="[1074,1138,1647,1673]" pageId="4" pageNumber="107">Head</emphasis>
(
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). 4.0 mm high in lateral view, 
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wide in anterior view. Black, facial gibbosity dark brown, about as high as wide. Setulae dark brown to black; mystax dark brown, centrally sparse. Pruinescence scarce, gray. Mouthparts: proboscis dark brown. Labellum about 1.7x longer than wide. Antenna: dark brown. Setulae of scape and pedicel all black. Scape 
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long. Pedicel 
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long; proximal postpedicel 
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long; distal postpedicel not distinctly inflated, 
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long, 
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wide, broader at about three-quarters of its length; sensory area rounded, restricted to apex on inner surface of antenna. 
<emphasis id="B95EEAFBF22AFFDC2AD1D6720A01823C" bold="true" box="[439,528,1863,1889]" pageId="4" pageNumber="107">Thorax</emphasis>
(
<figureCitation id="13112A6CF22AFFDC2946D6720A88823C" box="[544,665,1863,1889]" captionStart="FIGURE 2" captionStartId="5.[151,250,1977,2002]" captionTargetBox="[151,1435,841,1957]" captionTargetId="figure@5.[151,1435,841,1957]" captionTargetPageId="5" captionText="FIGURE 2. Chrysomydas phoenix sp. nov., male holotype (MNRJ), before dissection of terminalia. A. Dorsal habitus; B. Lateral habitus; C. Labels; D. Antenna, inner surface; E. Head, frontal view; F. Wing." httpUri="https://zenodo.org/record/3383566/files/figure.png" pageId="4" pageNumber="107">Figs 2A,B</figureCitation>
). Dark brown. Setulae of scutum black, present anteriorly on dorsocentral area, lateral margins, postalar callus and posteriorly close to scutellum; pruinescence grayish, scarce. Scutum without distinct pruinose spots. Scutellum black setulose, lightly pruinose. Mediotergite lightly pruinose. Pleura dark brown; lightly gray-pruinose. Anepisternum black setulose on dorsal and posterior margins. Anepimeron long black setulose. Katepimeron black setulose. Prosternum and proepisternum long black and white setulose. Meso- sternum long black-setulose. Metakatepisternum ventrally long black setulose. Metepimeron black setulose. 
<emphasis id="B95EEAFBF22BFFDD2E02D1AC0D8D85EE" bold="true" box="[1380,1436,153,179]" pageId="5" pageNumber="108">Legs</emphasis>
(
<figureCitation id="13112A6CF22BFFDD2BC6D18808E6858A" box="[160,247,189,215]" captionStart="FIGURE 2" captionStartId="5.[151,250,1977,2002]" captionTargetBox="[151,1435,841,1957]" captionTargetId="figure@5.[151,1435,841,1957]" captionTargetPageId="5" captionText="FIGURE 2. Chrysomydas phoenix sp. nov., male holotype (MNRJ), before dissection of terminalia. A. Dorsal habitus; B. Lateral habitus; C. Labels; D. Antenna, inner surface; E. Head, frontal view; F. Wing." httpUri="https://zenodo.org/record/3383566/files/figure.png" pageId="5" pageNumber="108">Fig. 2B</figureCitation>
). Dark brown; setulae black, fore and mid tibiae densely setulose. Hind femur 
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long, 
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wide; black spiniform macrosetae distributed in anteroventral and anterodorsal rows. Hind femur with long, erect setulae dorsally. Hind tibia 
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long, 
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wide; ventral keel developed; apical spur acute, 
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long. Hind tibia with strong anteroventral setulae at base of spine. Hind tarsomere 1 
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long; 
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wide; hind tarsomere 2 
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long; hind tarsomere 3 
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long; hind tarsomere 4 
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long; hind tarsomere 5 
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long. Pulvillus pale brown, shorter than fore, mid and hind tarsomere 5. Claw reddish brown, tip black. 
<emphasis id="B95EEAFBF22BFFDD2E79D0450D7184D6" bold="true" box="[1311,1376,368,395]" pageId="5" pageNumber="108">Wing</emphasis>
(
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). 
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long, 
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high. Alula 
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long, 
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high. Membrane hyaline, very light brown; microtrichiae absent. Veins orange to brown. C with short black setulae on anterior wing margin and very short brown microtrichia on posterior wing margin. Sc ending in C, reaching about two thirds of length of cell r 
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. R 
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(“stump vein”) emerging from R 
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, a little after its base; cell r 
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closed (R 
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terminating in R 
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); M 
<subScript id="17AE34ACF22BFFDD2F17D3270C9A877D" attach="left" box="[1137,1163,530,544]" fontSize="6" pageId="5" pageNumber="108">3+4</subScript>
present. Upper calypter pale brown with margin yellow; broader medially; marginal squamous setulae yellowish-white. Halter dark brown. 
<emphasis id="B95EEAFBF22BFFDD2BF1D37D091D873E" bold="true" box="[151,268,584,611]" pageId="5" pageNumber="108">Abdomen</emphasis>
(
<figureCitation id="13112A6CF22BFFDD2A7CD37C09B6873E" box="[282,423,585,611]" captionStart="FIGURE 2" captionStartId="5.[151,250,1977,2002]" captionTargetBox="[151,1435,841,1957]" captionTargetId="figure@5.[151,1435,841,1957]" captionTargetPageId="5" captionText="FIGURE 2. Chrysomydas phoenix sp. nov., male holotype (MNRJ), before dissection of terminalia. A. Dorsal habitus; B. Lateral habitus; C. Labels; D. Antenna, inner surface; E. Head, frontal view; F. Wing." httpUri="https://zenodo.org/record/3383566/files/figure.png" pageId="5" pageNumber="108">Figs 2A,B,F</figureCitation>
). Tergites and sternites mostly with dark brown to black setulae; T1 and T2 with few golden setulae; T1 black; T2–5 with discrete green metallic reflection, with long black setulae, directed outwards. Sternites black setulose. Tergites and sternites very lightly pruinose. Anterior portion of T1 convex, slightly pointed, asetulose. Bulla reddish brown, about 
<date id="FF941029F22BFFDD2961D3800A328792" box="[519,547,693,719]" pageId="5" pageNumber="108">4x</date>
as long as wide. 
<emphasis id="B95EEAFBF22BFFDD2987D3800BB18792" bold="true" box="[737,928,692,719]" pageId="5" pageNumber="108">Male terminalia</emphasis>
(
<figureCitation id="13112A6CF22BFFDD28C9D3800BE28792" box="[943,1011,693,719]" captionStart="FIGURE 3" captionStartId="6.[151,250,1496,1521]" captionTargetBox="[151,1435,174,1457]" captionTargetId="figure@6.[151,1435,174,1457]" captionTargetPageId="6" captionText="FIGURE 3. Male and female terminalia of Chrysomydas. A–D. Chrysomydas nitidulus (male, UFMG); A. Epandrium and proctiger, dorsal view; B. Gonocoxite-hypandrial complex and associate structures, dorsal view; C. same, ventral view; D. same, lateral view; E–G. Chrysomydas nitidulus (female, INPA); E. Sternite 8; F. Female terminalia, dorsal view; G. same, ventral view; H–K. Chrysomydas phoenix sp. nov. (male holotype, MNRJ); H. Epandrium and proctiger, dorsal view; I. Gonocoxite-hypandrial complex and associate structures, dorsal view; J. same, ventral view; K. same, lateral view. Abbreviations: cerc, cercus; ph, phallus; phc, phallic crest; sps, sclerite of spermathecal ducts; S, sternite; S9, genital fork; T, tergite." httpUri="https://zenodo.org/record/3383568/files/figure.png" pageId="5" pageNumber="108">Figs 3</figureCitation>
A–D). Black, setulae black. Epandrial lamella with distinct posterior spur, tip slightly curved. Cercus uncovered by epandrial lamellae. Phallus with bifid dorsal crest. Distal process of gonocoxite-hypandrial complex apically wide, truncate, with roundish dorsal tip and acute ventral tip.
</paragraph>
<caption id="DF556661F22BFFDD2BF1D68C0BEC82AB" httpUri="https://zenodo.org/record/3383566/files/figure.png" pageId="5" pageNumber="108" startId="5.[151,250,1977,2002]" targetBox="[151,1435,841,1957]" targetPageId="5">
<paragraph id="8B9536E9F22BFFDD2BF1D68C0BEC82AB" blockId="5.[151,1436,1977,2038]" pageId="5" pageNumber="108">
<emphasis id="B95EEAFBF22BFFDD2BF1D68C0907828F" bold="true" box="[151,278,1977,2002]" pageId="5" pageNumber="108">FIGURE 2.</emphasis>
<taxonomicName id="4C2A4D6AF22BFFDD2A79D68F0A13828F" authority="Calhau &amp; Lima &amp; Rafael &amp; Lamas, 2019" authorityName="Calhau &amp; Lima &amp; Rafael &amp; Lamas" authorityYear="2019" box="[287,514,1978,2002]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="phoenix" status="sp. nov.">
<emphasis id="B95EEAFBF22BFFDD2A79D68F0A13828F" box="[287,514,1978,2002]" italics="true" pageId="5" pageNumber="108">Chrysomydas phoenix</emphasis>
</taxonomicName>
<emphasis id="B95EEAFBF22BFFDD296AD68F0A4E828F" bold="true" box="[524,607,1978,2002]" pageId="5" pageNumber="108">
<taxonomicNameLabel id="A26D5780F22BFFDD296AD68F0A4E828F" box="[524,607,1978,2002]" pageId="5" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (MNRJ), before dissection of terminalia. 
<emphasis id="B95EEAFBF22BFFDD2FDBD68C0CC7828F" bold="true" box="[1213,1238,1977,2002]" pageId="5" pageNumber="108">A.</emphasis>
Dorsal habitus; 
<emphasis id="B95EEAFBF22BFFDD2EE3D68F0D8D828F" bold="true" box="[1413,1436,1978,2002]" pageId="5" pageNumber="108">B.</emphasis>
Lateral habitus; 
<emphasis id="B95EEAFBF22BFFDD2A5AD6E8094582AB" bold="true" box="[316,340,2013,2038]" pageId="5" pageNumber="108">C.</emphasis>
Labels; 
<emphasis id="B95EEAFBF22BFFDD2ACAD6EB09D582AB" bold="true" box="[428,452,2014,2038]" pageId="5" pageNumber="108">D.</emphasis>
Antenna, inner surface; 
<emphasis id="B95EEAFBF22BFFDD29DBD6EB0AC282AB" bold="true" box="[701,723,2014,2038]" pageId="5" pageNumber="108">E.</emphasis>
Head, frontal view; 
<emphasis id="B95EEAFBF22BFFDD28C3D6EB0BA982AB" bold="true" box="[933,952,2014,2038]" pageId="5" pageNumber="108">F.</emphasis>
Wing.
</paragraph>
</caption>
<caption id="DF556661F228FFDE2BF1D4ED0D2983F8" httpUri="https://zenodo.org/record/3383568/files/figure.png" pageId="6" pageNumber="109" startId="6.[151,250,1496,1521]" targetBox="[151,1435,174,1457]" targetPageId="6">
<paragraph id="8B9536E9F228FFDE2BF1D4ED0D2983F8" blockId="6.[151,1437,1496,1701]" pageId="6" pageNumber="109">
<emphasis id="B95EEAFBF228FFDE2BF1D4ED090480AC" bold="true" box="[151,277,1496,1521]" pageId="6" pageNumber="109">FIGURE 3.</emphasis>
Male and female terminalia of 
<taxonomicName id="4C2A4D6AF228FFDE2902D4EC0AFF80AC" authorityName="Wilcox, Papavero &amp; Pimentel" authorityYear="1989" box="[612,750,1497,1521]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAFBF228FFDE2902D4EC0AFF80AC" box="[612,750,1497,1521]" italics="true" pageId="6" pageNumber="109">Chrysomydas</emphasis>
</taxonomicName>
. 
<emphasis id="B95EEAFBF228FFDE299BD4ED0B3F80AC" bold="true" box="[765,814,1496,1521]" pageId="6" pageNumber="109">A–D</emphasis>
. 
<taxonomicName id="4C2A4D6AF228FFDE2858D4EC0C3B80AC" box="[830,1066,1497,1521]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B95EEAFBF228FFDE2858D4EC0C3B80AC" box="[830,1066,1497,1521]" italics="true" pageId="6" pageNumber="109">Chrysomydas nitidulus</emphasis>
</taxonomicName>
(male, UFMG); 
<emphasis id="B95EEAFBF228FFDE2FBBD4ED0CE480AC" bold="true" box="[1245,1269,1496,1521]" pageId="6" pageNumber="109">A.</emphasis>
Epandrium and proctiger, dorsal view; 
<emphasis id="B95EEAFBF228FFDE2AECD4C809B08348" bold="true" box="[394,417,1533,1557]" pageId="6" pageNumber="109">B.</emphasis>
Gonocoxite-hypandrial complex and associate structures, dorsal view; 
<emphasis id="B95EEAFBF228FFDE2FEBD4C90CB48348" bold="true" box="[1165,1189,1532,1557]" pageId="6" pageNumber="109">C.</emphasis>
same, ventral view; 
<emphasis id="B95EEAFBF228FFDE2EE2D4C80D8D8348" bold="true" box="[1412,1436,1533,1557]" pageId="6" pageNumber="109">D.</emphasis>
same, lateral view; 
<emphasis id="B95EEAFBF228FFDE2A04D715098B8364" bold="true" box="[354,410,1568,1593]" pageId="6" pageNumber="109">E–G.</emphasis>
<taxonomicName id="4C2A4D6AF228FFDE2AC4D7140A9C8364" box="[418,653,1569,1593]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B95EEAFBF228FFDE2AC4D7140A9C8364" box="[418,653,1569,1593]" italics="true" pageId="6" pageNumber="109">Chrysomydas nitidulus</emphasis>
</taxonomicName>
(female, INPA); 
<emphasis id="B95EEAFBF228FFDE2824D7140B488364" bold="true" box="[834,857,1569,1593]" pageId="6" pageNumber="109">E.</emphasis>
Sternite 8; 
<emphasis id="B95EEAFBF228FFDE28B3D7140BF98364" bold="true" box="[981,1000,1569,1593]" pageId="6" pageNumber="109">F.</emphasis>
Female terminalia, dorsal view; 
<emphasis id="B95EEAFBF228FFDE2E59D7150D4B8364" bold="true" box="[1343,1370,1568,1593]" pageId="6" pageNumber="109">G.</emphasis>
same, ventral view; 
<emphasis id="B95EEAFBF228FFDE2A44D770094C8300" bold="true" box="[290,349,1605,1629]" pageId="6" pageNumber="109">H–K.</emphasis>
<taxonomicName id="4C2A4D6AF228FFDE2A04D7700A538300" authority="Calhau &amp; Lima &amp; Rafael &amp; Lamas, 2019" authorityName="Calhau &amp; Lima &amp; Rafael &amp; Lamas" authorityYear="2019" box="[354,578,1605,1629]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="phoenix" status="sp. nov.">
<emphasis id="B95EEAFBF228FFDE2A04D7700A538300" box="[354,578,1605,1629]" italics="true" pageId="6" pageNumber="109">Chrysomydas phoenix</emphasis>
</taxonomicName>
<emphasis id="B95EEAFBF228FFDE292ED7700A898300" bold="true" box="[584,664,1605,1629]" pageId="6" pageNumber="109">
<taxonomicNameLabel id="A26D5780F228FFDE292ED7700A898300" box="[584,664,1605,1629]" pageId="6" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(male holotype, MNRJ); 
<emphasis id="B95EEAFBF228FFDE28FAD7700BA78300" bold="true" box="[924,950,1605,1629]" pageId="6" pageNumber="109">H.</emphasis>
Epandrium and proctiger, dorsal view; 
<emphasis id="B95EEAFBF228FFDE2E23D7700D448300" bold="true" box="[1349,1365,1605,1629]" pageId="6" pageNumber="109">I.</emphasis>
Gonocoxite-hypandrial complex and associate structures, dorsal view; 
<emphasis id="B95EEAFBF228FFDE2853D75C0B5983DC" bold="true" box="[821,840,1641,1665]" pageId="6" pageNumber="109">J.</emphasis>
same, ventral view; 
<emphasis id="B95EEAFBF228FFDE2F79D75C0C2883DC" bold="true" box="[1055,1081,1641,1665]" pageId="6" pageNumber="109">K.</emphasis>
same, lateral view. Abbreviations: cerc, cercus; ph, phallus; phc, phallic crest; sps, sclerite of spermathecal ducts; S, sternite; S9, genital fork; T, tergite.
</paragraph>
</caption>
<paragraph id="8B9536E9F228FFDF2BA1D7E609DE8762" blockId="6.[151,1437,1747,2026]" lastBlockId="7.[151,1437,153,683]" lastPageId="7" lastPageNumber="110" pageId="6" pageNumber="109">
<emphasis id="B95EEAFBF228FFDE2BA1D7E6091483B0" bold="true" box="[199,261,1747,1773]" pageId="6" pageNumber="109">Pupa</emphasis>
(
<figureCitation id="13112A6CF228FFDE2A72D7E6094B83B0" box="[276,346,1747,1773]" captionStart="FIGURE 4" captionStartId="7.[151,250,1854,1879]" captionTargetBox="[151,1435,697,1804]" captionTargetId="figure@7.[151,1435,697,1804]" captionTargetPageId="7" captionText="FIGURE 4. Pupal exuviae of Chrysomydas. A–E. Chrysomydas nitidulus (female, INPA). F–J. Chrysomydas phoenix sp. nov. (male, MNRJ). Abbreviations: aap, anterior antennal process; abs, abdominal spiracles; amsp, anterior mesothoracic spines; dpp, dorsal posterolateral process; ls, labral sheath; ls1, leg sheath 1; ls2, leg sheath 2; ls3, leg sheath 3; ms, maxillary sheath; pap, posterior antennal processes; pmc, posterior mesothoracic callosity; prs, proboscidal sheath; ps; palpal sheath; pths, prothoracic spiracle; vc, ventral callosity; vpp, ventral posterolateral process; ws, wing sheath." httpUri="https://zenodo.org/record/3383570/files/figure.png" pageId="6" pageNumber="109">Figs 4</figureCitation>
A–E). Greatest length, not including anterior antennal or dorsal posterolateral processes 
<quantity id="4CD29B0CF228FFDE2E54D7E60D8883B3" box="[1330,1433,1747,1774]" metricMagnitude="-2" metricUnit="m" metricValue="2.77" pageId="6" pageNumber="109" unit="mm" value="27.7">27.7 mm</quantity>
. Greatest width of thorax 
<quantity id="4CD29B0CF228FFDE2ADED7C30A02824F" box="[440,531,1782,1810]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="6" pageNumber="109" unit="mm" value="5.8">5.8 mm</quantity>
. Greatest width of abdomen 
<quantity id="4CD29B0CF228FFDE2806D7C20BAD824F" box="[864,956,1783,1810]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="6" pageNumber="109" unit="mm" value="6.4">6.4 mm</quantity>
. Body pale brown. Spines reddish brown, distally darker. Anterior antennal processes spiniform, reddish on proximal third, black on distal two thirds, rugose, curved outwards. Maxillary sheath rugose. Proboscidal sheath short, rugose, truncate. Prothoracic spiracle laterally at anterior margin of thorax on highly rugose, raised area. Anterior lateral mesothoracic callosity reddish, flat, rugose. Wing sheath with small basal reclinate spine. Thoracic area above wing sheath smoothly transversely rugose. Leg sheaths rugose. Leg sheath 3 reaching to middle of abdominal segment 2. Spiracles of abdominal segments 1–7 situated along midline laterally, raised, shining, reddish. Spiracle of segment 8 not as conspicuous as the remain- ing abdominal spiracles. Abdominal segment 1 anteriorly with 14 flattened proclinate spines, posterolaterally with 10–12 reclinate spines, some of them bifid. Abdominal segments 2–7 each with 39–57 posterior flattened erect to reclinate spines. Abdominal segment 2 with 57 posterior flattened erect to reclinate spines, with 18 dorsal, 8–10 on each lateral and 21 ventral. Abdominal segment 3 with 49 posterior flattened erect to reclinate spines, with 18 dorsal, 9 on each lateral and 22 ventral. Abdominal segment 4 with 54 posterior flattened erect to reclinate spines, with 17 dorsal, 8–9 on each lateral and 20 ventral. Abdominal segment 5 with 56 posterior flattened erect to reclinate spines, with 18 dorsal, 7–8 on each lateral and 23 ventral. Abdominal segment 6 with 48 posterior flattened erect to reclinate spines, with 15 dorsal, 8 on each lateral, and 17 ventral. Abdominal segment 7 with 39 posterior flattened erect to reclinate spines, with 15 dorsal, 6 on each lateral and 12 ventral. Abdominal segment 8 dorsally with 4 reclinate short spines, laterally with 1 long and 1–2 small reclinate spine on each of two raised tubercular areas on each side, with 8 erect ventral spines. Abdominal segment 9 with pair of divergent long posterior spiniform processes, ventrally curved.
</paragraph>
<paragraph id="8B9536E9F229FFDF2BA1D37C0B9987F6" blockId="7.[151,1437,153,683]" pageId="7" pageNumber="110">
<emphasis id="B95EEAFBF229FFDF2BA1D37C095F873E" bold="true" box="[199,334,585,611]" pageId="7" pageNumber="110">Etymology.</emphasis>
The specific epithet is an allusion to the Greek mythological bird Phoenix, which obtains new life by arising from its own ashes. This name represents the resurgence of the 
<collectionCode id="ED3BAE2CF229FFDF28AED3580C0287DA" box="[968,1043,621,647]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="7" pageNumber="110">MNRJ</collectionCode>
after the terrible fire that destroyed most of its collections, including nearly the entire insect collection.
</paragraph>
<caption id="DF556661F229FFDF2BF1D60B0C3A82BA" httpUri="https://zenodo.org/record/3383570/files/figure.png" pageId="7" pageNumber="110" startId="7.[151,250,1854,1879]" targetBox="[151,1435,697,1804]" targetPageId="7">
<paragraph id="8B9536E9F229FFDF2BF1D60B0C3A82BA" blockId="7.[151,1436,1854,2023]" pageId="7" pageNumber="110">
<emphasis id="B95EEAFBF229FFDF2BF1D60B0902820A" bold="true" box="[151,275,1854,1879]" pageId="7" pageNumber="110">FIGURE 4.</emphasis>
Pupal exuviae of 
<taxonomicName id="4C2A4D6AF229FFDF2AAED60B0A43820B" authorityName="Wilcox, Papavero &amp; Pimentel" authorityYear="1989" box="[456,594,1854,1878]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAFBF229FFDF2AAED60B0A43820B" box="[456,594,1854,1878]" italics="true" pageId="7" pageNumber="110">Chrysomydas</emphasis>
</taxonomicName>
. 
<emphasis id="B95EEAFBF229FFDF2938D60B0A85820A" bold="true" box="[606,660,1854,1879]" pageId="7" pageNumber="110">A–E.</emphasis>
<taxonomicName id="4C2A4D6AF229FFDF29FDD60B0B95820B" box="[667,900,1854,1878]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="110" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B95EEAFBF229FFDF29FDD60B0B95820B" box="[667,900,1854,1878]" italics="true" pageId="7" pageNumber="110">Chrysomydas nitidulus</emphasis>
</taxonomicName>
(female, INPA). 
<emphasis id="B95EEAFBF229FFDF2F56D60A0C71820A" bold="true" box="[1072,1120,1855,1879]" pageId="7" pageNumber="110">F–J.</emphasis>
<taxonomicName id="4C2A4D6AF229FFDF2F00D60B0D54820B" authority="Calhau &amp; Lima &amp; Rafael &amp; Lamas, 2019" authorityName="Calhau &amp; Lima &amp; Rafael &amp; Lamas" authorityYear="2019" box="[1126,1349,1854,1878]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="110" phylum="Arthropoda" rank="species" species="phoenix" status="sp. nov.">
<emphasis id="B95EEAFBF229FFDF2F00D60B0D54820B" box="[1126,1349,1854,1878]" italics="true" pageId="7" pageNumber="110">Chrysomydas phoenix</emphasis>
</taxonomicName>
<emphasis id="B95EEAFBF229FFDF2E2AD60A0D8D820A" bold="true" box="[1356,1436,1855,1879]" pageId="7" pageNumber="110">
<taxonomicNameLabel id="A26D5780F229FFDF2E2AD60A0D8D820A" box="[1356,1436,1855,1879]" pageId="7" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(male, MNRJ). Abbreviations: aap, anterior antennal process; abs, abdominal spiracles; amsp, anterior mesothoracic spines; dpp, dorsal posterolateral process; ls, labral sheath; ls1, leg sheath 1; ls2, leg sheath 2; ls3, leg sheath 3; ms, maxillary sheath; pap, posterior antennal processes; pmc, posterior mesothoracic callosity; prs, proboscidal sheath; ps; palpal sheath; pths, prothoracic spiracle; vc, ventral callosity; vpp, ventral posterolateral process; ws, wing sheath.
</paragraph>
</caption>
<paragraph id="8B9536E9F226FFD02BA1D1AC0B1F841F" blockId="8.[151,1436,152,323]" pageId="8" pageNumber="111">
<emphasis id="B95EEAFBF226FFD02BA1D1AC096485EE" bold="true" box="[199,373,152,179]" pageId="8" pageNumber="111">Type material.</emphasis>
<materialsCitation id="3B423CB4F226FFD02A1BD1AD0B1F841F" ID-GBIF-Occurrence="2382327817" collectingDate="1964-11" collectingDateMax="2018-09" collectingDateMin="1964-11" collectionCode="MNRJ, MZUSP" collectorName="P. Buhrnheim &amp; It" country="Brazil" location="Male" pageId="8" pageNumber="111" specimenCount="1" specimenCount-male="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<emphasis id="B95EEAFBF226FFD02A1BD1AD0A0D85EE" bold="true" box="[381,540,152,179]" pageId="8" pageNumber="111">
<typeStatus id="5491884BF226FFD02A1BD1AD0A0D85EE" box="[381,540,152,179]" pageId="8" pageNumber="111" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="9D2CFD60F226FFD02945D1AC0A2A85EE" box="[547,571,153,179]" pageId="8" pageNumber="111" type="male">♂</specimenCount>
: [
<collectingCountry id="F33D7679F226FFD0292BD1AD0A8685EF" box="[589,663,152,178]" name="Brazil" pageId="8" pageNumber="111">Brazil</collectingCountry>
, 
<collectingRegion id="49EEF80BF226FFD029C5D1AD0B9985EE" box="[675,904,152,179]" country="Brazil" name="Rio de Janeiro" pageId="8" pageNumber="111">Rio de Janeiro State</collectingRegion>
] 
<collectingRegion id="49EEF80BF226FFD028F3D1AD0C2A85EF" box="[917,1083,152,178]" country="Brazil" name="Rio de Janeiro" pageId="8" pageNumber="111">Rio de Janeiro</collectingRegion>
, ii. 
<date id="FF941029F226FFD02F3DD1AD0CA585EF" box="[1115,1204,152,178]" pageId="8" pageNumber="111" value="1964-11">
<collectingDate id="EFD0E9C1F226FFD02F3DD1AD0CA585EF" box="[1115,1204,152,178]" pageId="8" pageNumber="111" value="1964-11">xi.1964</collectingDate>
</date>
, 
<collectorName id="26DF533FF226FFD02FA6D1AC0D4885EF" box="[1216,1369,152,179]" pageId="8" pageNumber="111">P. Buhrnheim</collectorName>
[col.] [
<location id="8EF56032F226FFD02BF8D18908CB858B" LSID="urn:lsid:plazi:treatment:038387FFF22AFFD02BF1D4600B1F841F:8EF56032F226FFD02BF8D18908CB858B" box="[158,218,188,214]" country="Brazil" name="Male" pageId="8" pageNumber="111" stateProvince="Rio de Janeiro">Male</location>
imago with associated pupal exuviae on two separate pins]. (
<collectionCode id="ED3BAE2CF226FFD028E3D1880BC7858A" box="[901,982,189,215]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="8" pageNumber="111">MNRJ</collectionCode>
)* * It should be noted that this specimen was borrowed from the Museo Nacional several years ago. It was not impacted by the loss of the collection due to the fire in 
<date id="FF941029F226FFD02A6AD03109D28442" box="[268,451,260,287]" pageId="8" pageNumber="111" value="2018-09">
<collectingDate id="EFD0E9C1F226FFD02A6AD03109D28442" box="[268,451,260,287]" pageId="8" pageNumber="111" value="2018-09">September 2018</collectingDate>
</date>
as the specimen remained on loan to the senior author at 
<collectionCode id="ED3BAE2CF226FFD02F2AD0300CBB8442" box="[1100,1194,261,287]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="111">MZUSP</collectionCode>
. 
<collectorName id="26DF533FF226FFD02FD3D0300CD78442" box="[1205,1222,261,287]" pageId="8" pageNumber="111">It</collectorName>
will be returned to 
<collectionCode id="ED3BAE2CF226FFD02BF1D01C08F3841E" box="[151,226,297,323]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="8" pageNumber="111">MNRJ</collectionCode>
when appropriate collection spaces are available.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>