158 lines
17 KiB
XML
158 lines
17 KiB
XML
<document id="3299BAA96DF890814452EE594C52DC42" ID-DOI="10.5252/g2011n2a10" ID-ISSN="1638-9395" ID-Zenodo-Dep="5372482" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630272317791" checkinUser="marcus" docAuthor="Delfino, Massimo, Bailon, Salvador & Pitruzzella, Gaetano" docDate="2011" docId="038387F3FFCDDE003BD9FF53B1D4FE15" docLanguage="en" docName="Geodiversitas.2011.33.2.357-382.pdf" docOrigin="Geodiversitas 33 (2)" docSource="http://dx.doi.org/10.5252/g2011n2a10" docStyle="DocumentStyle:DF8BFA78045876A77BB343B212438C9A.3:Geodiversitas.2011-2012.journal_article.type1" docStyleId="DF8BFA78045876A77BB343B212438C9A" docStyleName="Geodiversitas.2011-2012.journal_article.type1" docStyleVersion="3" docTitle="Colubridae Oppel 1811" docType="treatment" docVersion="3" lastPageNumber="375" masterDocId="FFBAFF8BFFDCDE133920FF8AB10BFFE5" masterDocTitle="The Late Pliocene amphibians and reptiles from “ Capo Mannu D 1 Local Fauna ” (Mandriola, Sardinia, Italy)" masterLastPageNumber="382" masterPageNumber="357" pageNumber="374" updateTime="1699207784534" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="383E104E855BCCC51BA3B941C4E56653">
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<mods:title id="BC1ADF3A27937FD180B0A3980B5BB0CC">The Late Pliocene amphibians and reptiles from “ Capo Mannu D 1 Local Fauna ” (Mandriola, Sardinia, Italy)</mods:title>
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<mods:name id="198800C8CFEA9A832CB8A210469C3320" type="personal">
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<mods:roleTerm id="0346372963FBB554A14B60E2BDFB04E8">Author</mods:roleTerm>
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<mods:namePart id="719D7404FCE935E960A98FCF123CA94C">Delfino, Massimo</mods:namePart>
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</mods:name>
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<mods:name id="06813AFD46AAA4AB09D503594C1521DD" type="personal">
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<mods:roleTerm id="C0F8902F331DB3C5C3B83D309BBFDB69">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DD0D50B3C167050F8F2C5648E6C54EF9">Bailon, Salvador</mods:namePart>
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</mods:name>
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<mods:name id="5327D6EDFB7F13487238350437C5B55E" type="personal">
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<mods:roleTerm id="30EDF49858A7D57CE741666F0704EDFE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0AA99F37894C84526E1EAAE7B69C18B3">Pitruzzella, Gaetano</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="99BB8EB13D80B6331910725D538567F7">text</mods:typeOfResource>
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<mods:title id="0ABED31E3C0C54FB532D0C92120F5A88">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="1253368A025D1AD68504C5FFDB6C518C">
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<mods:date id="6143B664AB17095353358FC350C4DD13">2011</mods:date>
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<mods:detail id="C2CB929270BC23F1BB65190017CC2487" type="pubDate">
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<mods:number id="8EA61C0EBACF8748BA6DACC4B143F499">2011-06-30</mods:number>
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<mods:detail id="151A4B92942A072A3C1091A87E3710B6" type="volume">
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<mods:number id="7BF116456EF214DBC1B5DC56A3836D6D">33</mods:number>
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<mods:number id="8ABFF93014028832CDEE881E16571AFD">2</mods:number>
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<mods:start id="679CC25BDE96D185FFDCD52357905A6D">357</mods:start>
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<mods:end id="27D740F60533D481681768EFD6791BA0">382</mods:end>
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<mods:url id="CA879CF73A91AAA6DA0CB18FFA2AB169">http://dx.doi.org/10.5252/g2011n2a10</mods:url>
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<mods:classification id="96D0A0BEC0677BDB93E993C8C886CE8D">journal article</mods:classification>
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<mods:identifier id="D7B49DC36FA7F42B3C17BA24654A3F30" type="DOI">10.5252/g2011n2a10</mods:identifier>
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<mods:identifier id="FAF1560101F8CAED888BC4FB04A0DF20" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="67E2B2051B5C99A5BA38B14EC7C8C1D0" type="Zenodo-Dep">5372482</mods:identifier>
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</mods:mods>
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<treatment id="038387F3FFCDDE003BD9FF53B1D4FE15" LSID="urn:lsid:plazi:treatment:038387F3FFCDDE003BD9FF53B1D4FE15" httpUri="http://treatment.plazi.org/id/038387F3FFCDDE003BD9FF53B1D4FE15" lastPageId="19" lastPageNumber="375" pageId="17" pageNumber="374">
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<subSubSection id="C330656EFFCDDE023BD9FF53B2E4FE97" pageId="17" pageNumber="374" type="nomenclature">
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<paragraph id="8B9536E5FFCDDE023BD9FF53B56EFF16" blockId="17.[761,1125,216,243]" box="[761,1125,216,243]" pageId="17" pageNumber="374">
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<heading id="D0DD8189FFCDDE023BD9FF53B56EFF16" box="[761,1125,216,243]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="374" reason="3">
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Family
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<taxonomicName id="4C2A4D66FFCDDE023A69FF53B56EFF16" authority="Oppel, 1811" authorityName="Oppel" authorityYear="1811" box="[841,1125,216,243]" class="Reptilia" family="Colubridae" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="374" phylum="Chordata" rank="family">COLUBRIDAE Oppel, 1811</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B9536E5FFCDDE023A0CFE9DB2E4FE97" blockId="17.[767,1119,279,370]" pageId="17" pageNumber="374">
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<heading id="D0DD8189FFCDDE023A0CFE9DB2E4FE97" centered="true" fontSize="11" level="2" pageId="17" pageNumber="374" reason="2">
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“Colubrine”
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<typeStatus id="54918847FFCDDE023A97FE9DB2EEFED7" box="[951,997,279,306]" pageId="17" pageNumber="374">type</typeStatus>
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(trunk vertebrae without hypapophysis) (
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<figureCitation id="13112A60FFCDDE023A57FEDDB2BEFE97" box="[887,949,343,370]" captionStart="FIG" captionStartId="20.[113,124,1635,1652]" captionTargetBox="[113,1208,233,1580]" captionTargetId="figure-19@20.[113,663,698,1167]" captionTargetPageId="20" captionText="FIG. 7. — Fossil remains of reptiles from Capo Mannu D1 Local Fauna (Sardinia): A-D, “Colubrines” indet: A, B, trunk vertebra (DSTC 6024), dorsal and posterior views; C, D, right maxilla (DSTC 6025), medial and ventral views; E, F, Vipera sp. trunk vertebra (DSTC 6026), dorsal and right lateral views. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5372498" httpUri="https://zenodo.org/record/5372498/files/figure.png" pageId="17" pageNumber="374">Fig. 7</figureCitation>
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A-D)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C330656EFFCDDE023B85FE1AB229FE27" pageId="17" pageNumber="374" type="materials_examined">
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<paragraph id="8B9536E5FFCDDE023B85FE1AB229FE27" blockId="17.[677,1209,400,450]" pageId="17" pageNumber="374">
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<collectionCode id="ED3BAE20FFCDDE023B85FE1AB202FE43" box="[677,777,400,423]" pageId="17" pageNumber="374">MATERIAL</collectionCode>
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EXAMINED. — Maxilla: 1 fragment; trunk vertebra: 21.
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</paragraph>
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</subSubSection>
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<subSubSection id="C330656EFFCDDE023B85FE7FB598F9C0" pageId="17" pageNumber="374" type="description">
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<paragraph id="8B9536E5FFCDDE023B85FE7FB23EFDE8" blockId="17.[674,1212,501,1573]" box="[677,821,501,527]" pageId="17" pageNumber="374">DESCRIPTION</paragraph>
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<paragraph id="8B9536E5FFCDDE023B85FD9FB2ADFC2E" blockId="17.[674,1212,501,1573]" pageId="17" pageNumber="374">The fragmentary right maxilla is represented by a posterior section of this element extending from an undetermined point following the prefrontal process (not preserved) and the tooth position following the ectopterygoid process (preserved); the ectopterygoid process corresponds to two tooth sockets, hosting the only preserved tooth, and part of the preceding socket; two more tooth sockets are placed anteriorly to the ectopterygoid process; the toothed region is preceded by a long diastema corresponding to at least two tooth positions; in ventrolateral view, an evident change of slope corresponds to the boundary between the diastema and the toothed region.</paragraph>
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<paragraph id="8B9536E5FFCDDE023B9BFC5AB2A7FA42" blockId="17.[674,1212,501,1573]" pageId="17" pageNumber="374">
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The most common vertebral morphotype is characterized by an elongated centrum (whose length can reach
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<quantity id="4CD29B00FFCDDE023A29FB9AB255FBCE" box="[777,862,1040,1067]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="17" pageNumber="374" unit="mm" value="2.4">2.4 mm</quantity>
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); subcircular cotyles and condyles; parapophysis and diapophysis separated by a shallow depression; parapophysis anteroposteriorly longer than diapophysis; parapophyseal processes absent; haemal keel well defined and usually modestly widened posteriorly; pre- and postzygapophyses weakly tilted dorsally; prezygapophyseal processes slender, pointed and long up to half the width of the prezygapophyseal facet; prezygapophyseal facets drop-shaped; zygosphene moderately convex in dorsal view; neural arch depressed and with dorsal edge modestly convex in posterior view; neural spine long and very low.
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</paragraph>
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<paragraph id="8B9536E5FFCDDE023B9BFA21B598F9C0" blockId="17.[674,1212,501,1573]" pageId="17" pageNumber="374">Variations of the morphology described above are mostly related to minor differences in terms of haemal keel shape (not widened posteriorly) and shape of the posterior edge of the neural arch.</paragraph>
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</subSubSection>
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<subSubSection id="C330656EFFCDDE003B85F9C0B1D4FE15" lastPageId="19" lastPageNumber="376" pageId="17" pageNumber="374" type="discussion">
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<paragraph id="8B9536E5FFCDDE023B85F9C0B22DF987" blockId="17.[677,1210,1610,1700]" box="[677,806,1610,1636]" pageId="17" pageNumber="374">DISCUSSION</paragraph>
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<paragraph id="8B9536E5FFCDDE013B85F9E3B53DF980" blockId="17.[677,1210,1610,1700]" lastBlockId="18.[676,1211,1515,1700]" lastPageId="18" lastPageNumber="375" pageId="17" pageNumber="374">
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The presence of a diastema is a character typical of the opisthoglyphous colubrines at our disposal for comparison:
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<taxonomicName id="4C2A4D66FFCEDE0139DFFA61B318F9E3" authority="Fitzinger, 1826" authorityName="Fitzinger" authorityYear="1826" box="[255,531,1515,1542]" class="Reptilia" family="Lamprophiidae" genus="Malpolon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="375" phylum="Chordata" rank="genus">
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<emphasis id="B95EEAF7FFCEDE0139DFFA61B068F9E0" box="[255,355,1515,1541]" italics="true" pageId="18" pageNumber="375">Malpolon</emphasis>
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Fitzinger, 1826
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</taxonomicName>
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,
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<taxonomicName id="4C2A4D66FFCEDE013B00FA61B1F4F9C0" authority="Wagler, 1830" authorityName="Wagler" authorityYear="1830" class="Reptilia" family="Colubridae" genus="Telescopus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="375" phylum="Chordata" rank="genus">
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<emphasis id="B95EEAF7FFCEDE013B00FA61B38DF9E0" box="[544,646,1515,1541]" italics="true" pageId="18" pageNumber="375">Telescopus</emphasis>
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Wagler, 1830
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</taxonomicName>
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, and
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<taxonomicName id="4C2A4D66FFCEDE013815F980B38BF9C3" authority="Guichenot, 1850" authorityName="Guichenot" authorityYear="1850" box="[309,640,1546,1574]" class="Reptilia" family="Colubridae" genus="Macroprotodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="375" phylum="Chordata" rank="genus">
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<emphasis id="B95EEAF7FFCEDE013815F980B0C7F9C1" box="[309,460,1546,1572]" italics="true" pageId="18" pageNumber="375">Macroprotodon</emphasis>
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Guichenot, 1850
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</taxonomicName>
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. Even if not totally congruent in morphology, the fragmentary maxilla of Capo Mannu is more similar to that of
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<taxonomicName id="4C2A4D66FFCEDE0139F7F9E3B07FF966" authorityName="Guichenot" authorityYear="1850" box="[215,372,1641,1667]" class="Reptilia" family="Colubridae" genus="Macroprotodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="375" phylum="Chordata" rank="genus">
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<emphasis id="B95EEAF7FFCEDE0139F7F9E3B07FF966" box="[215,372,1641,1667]" italics="true" pageId="18" pageNumber="375">Macroprotodon</emphasis>
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</taxonomicName>
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than to those of
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<taxonomicName id="4C2A4D66FFCEDE013B03F9E3B38DF966" authorityName="Fitzinger" authorityYear="1826" box="[547,646,1641,1667]" class="Reptilia" family="Lamprophiidae" genus="Malpolon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="375" phylum="Chordata" rank="genus">
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<emphasis id="B95EEAF7FFCEDE013B03F9E3B38DF966" box="[547,646,1641,1667]" italics="true" pageId="18" pageNumber="375">Malpolon</emphasis>
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</taxonomicName>
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or
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<taxonomicName id="4C2A4D66FFCEDE0139AFF903B1FEF946" authorityName="Wagler" authorityYear="1830" box="[143,245,1673,1699]" class="Reptilia" family="Colubridae" genus="Telescopus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="375" phylum="Chordata" rank="genus">
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<emphasis id="B95EEAF7FFCEDE0139AFF903B1FEF946" box="[143,245,1673,1699]" italics="true" pageId="18" pageNumber="375">Telescopus</emphasis>
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</taxonomicName>
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. The morphology of the fragmentary maxilla does not match with that of any known taxon so that is not demonstrated if it belongs to a new taxon or, more likely, if it is an unusual morphology of a taxon already described.
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</paragraph>
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<caption id="DF55666DFFCEDE013951FAEEB39AFA46" ID-DOI="http://doi.org/10.5281/zenodo.5372496" ID-Zenodo-Dep="5372496" httpUri="https://zenodo.org/record/5372496/files/figure.png" pageId="18" pageNumber="375" startId="18.[113,124,1380,1397]" targetBox="[150,1173,221,1337]" targetPageId="18">
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<paragraph id="8B9536E5FFCEDE013951FAEEB39AFA46" blockId="18.[113,1211,1380,1443]" pageId="18" pageNumber="375">
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FIG. 6. — Fossil remains of reptiles from Capo Mannu D1 Local Fauna (Sardinia):
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<emphasis id="B95EEAF7FFCEDE013A02FAEEB24FFA90" bold="true" box="[802,836,1380,1397]" pageId="18" pageNumber="375">A -C</emphasis>
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,
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<taxonomicName id="4C2A4D66FFCEDE013A70FAEEB2CCFA90" authorityName="Gray" authorityYear="1844" box="[848,967,1380,1397]" order="Amphisbaenia" pageId="18" pageNumber="373" rank="order">Amphisbaenia</taxonomicName>
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indet., trunk vertebra (DSTC 6021), dorsal, left lateral and ventral views;
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<emphasis id="B95EEAF7FFCEDE0138C1FAF1B30AFA69" bold="true" box="[481,513,1403,1420]" pageId="18" pageNumber="375">D -F</emphasis>
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,
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<taxonomicName id="4C2A4D66FFCEDE013B2DFAF1B33AFA69" authorityName="Daudin" authorityYear="1803" box="[525,561,1403,1420]" class="Reptilia" family="Boidae" genus="Eryx" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="375" phylum="Chordata" rank="genus">
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<emphasis id="B95EEAF7FFCEDE013B2DFAF1B33AFA69" box="[525,561,1403,1420]" italics="true" pageId="18" pageNumber="375">Eryx</emphasis>
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</taxonomicName>
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cf.
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<emphasis id="B95EEAF7FFCEDE013B71FAF1B3A3FA69" box="[593,680,1403,1420]" italics="true" pageId="18" pageNumber="375">
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<taxonomicName id="4C2A4D66FFCEDE013B71FAF1B3AFFA69" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[593,676,1403,1420]" class="Reptilia" family="Boidae" genus="Eryx" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="375" phylum="Chordata" rank="species" species="jaculus">E. jaculus</taxonomicName>
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,
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</emphasis>
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trunk vertebra (DST 6022), dorsal, ventral and posterior views;
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<emphasis id="B95EEAF7FFCEDE013951FA18B174FA46" bold="true" box="[113,127,1426,1443]" pageId="18" pageNumber="375">G</emphasis>
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, caudal vertebra (DSTC 6023), anterior view. Scale bars: 2 mm.
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</paragraph>
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</caption>
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<paragraph id="8B9536E5FFCEDE003B9BF9E3B1D4FE15" blockId="18.[676,1211,1515,1700]" lastBlockId="19.[113,647,216,496]" lastPageId="19" lastPageNumber="376" pageId="18" pageNumber="375">
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The colubrid trunk vertebrae devoid of hypapophysis are referred to an undetermined member of the working taxon “Colubrines” (sensu
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<bibRefCitation id="EFBB4B14FFCFDE003B08FF52B1B2FEF6" author="SZYNDLAR Z." pageId="19" pageNumber="376" pagination="103 - 126" refId="ref16620" refString="SZYNDLAR Z. 1991 a. - A review of Neogene and Quaternary snakes of Central and Eastern Europe. Part I: Scolecophidia, Boidae, Colubrinae. Estudios geologicos 47: 103 - 126." type="journal article" year="1991">Szyndlar 1991a</bibRefCitation>
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). The morphological uniformity of these vertebrae suggests that most of them probably belong to a single taxon, but it is not possible to exclude that some of the fragments belong to other taxa. Noteworthy is that the depressed neural arch and the long and low neural spine are two characters typical of taxa like
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<taxonomicName id="4C2A4D66FFCFDE003864FE3FB35EFE34" authority="Laurenti, 1768" authorityName="Laurenti" authorityYear="1768" box="[324,597,437,465]" class="Reptilia" family="Colubridae" genus="Coronella" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="376" phylum="Chordata" rank="genus">
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<emphasis id="B95EEAF7FFCFDE003864FE3FB0A3FE2A" box="[324,424,437,463]" italics="true" pageId="19" pageNumber="376">Coronella</emphasis>
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Laurenti, 1768
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</taxonomicName>
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and
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<taxonomicName id="4C2A4D66FFCFDE003953FE5FB1D3FE0A" authorityName="Wagler" authorityYear="1830" box="[115,216,469,495]" class="Reptilia" family="Colubridae" genus="Telescopus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="376" phylum="Chordata" rank="genus">
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<emphasis id="B95EEAF7FFCFDE003953FE5FB1D3FE0A" box="[115,216,469,495]" italics="true" pageId="19" pageNumber="376">Telescopus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |