treatments-xml/data/03/83/87/038387E74B360832946BBD5731E4FCF5.xml

<document id="4B3CD16B2950205D77891E8B8795923D" ID-CLB-Dataset="27013" ID-DOI="10.5281/zenodo.2646440" ID-GBIF-Dataset="91632c26-bc53-48bc-86ef-7ba516964f61" ID-ISSN="1175-5326" ID-Zenodo-Dep="2646440" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555588053644" checkinUser="plazi" docAuthor="Taylor, Christopher A., Soucek, David J. &amp; Organ, Edward L." docDate="2006" docId="038387E74B360832946BBD5731E4FCF5" docLanguage="en" docName="zt01200p041.pdf" docOrigin="Zootaxa 1200" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Cambarus clivosus Taylor &amp; Soucek &amp; Organ 2006, new species" docType="treatment" docVersion="9" lastPageNumber="41" masterDocId="FFBAFF9F4B34083E9563BC4B326FFFE6" masterDocTitle="A new crayfish of the genus Cambarus Erichson, 1846 (Decapoda: Cambaridae) from an under-sampled habitat type in central Ten- nessee, USA" masterLastPageNumber="41" masterPageNumber="29" pageNumber="31" updateTime="1698723041197" updateUser="ExternalLinkService">
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<mods:title id="8B7647FA19712127CA99685C2385D101">A new crayfish of the genus Cambarus Erichson, 1846 (Decapoda: Cambaridae) from an under-sampled habitat type in central Ten- nessee, USA</mods:title>
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<mods:namePart id="1856A117944F5E2CAAA381E6069C8A2C">Taylor, Christopher A.</mods:namePart>
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<mods:namePart id="5F32600BAF13CBE7CF019ABDDE8754AA">Soucek, David J.</mods:namePart>
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<mods:namePart id="CEBBA2BE06E91A27E9A9670D585336E3">Organ, Edward L.</mods:namePart>
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<subSubSection id="C330657A4B36083C946BBD5733B5FEBF" pageId="2" pageNumber="31" type="nomenclature">
<paragraph id="8B9536F14B36083C946BBD5731EBFED3" blockId="2.[264,900,283,345]" box="[264,900,283,310]" pageId="2" pageNumber="31">
<heading id="D0DD819D4B36083C946BBD5731EBFED3" bold="true" box="[264,900,283,310]" fontSize="11" level="1" pageId="2" pageNumber="31" reason="1">
<emphasis id="B95EEAE34B36083C946BBD5731EBFED3" bold="true" box="[264,900,283,310]" pageId="2" pageNumber="31">
<taxonomicName id="4C2A4D724B36083C946BBD57308AFED3" ID-CoL="7RTWX" authority="Taylor and Soucek" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[264,741,283,310]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B36083C946BBD573382FED0" bold="true" box="[264,493,284,310]" italics="true" pageId="2" pageNumber="31">Cambarus clivosus</emphasis>
, Taylor and Soucek
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<paragraph id="8B9536F14B36083C946BBD7433B5FEBF" blockId="2.[264,900,283,345]" box="[264,474,319,345]" pageId="2" pageNumber="31">
Figs. 2, 3, 
<tableCitation id="C6A8034A4B36083C94E7BD7433B5FEBF" box="[388,474,319,345]" captionStart="TABLE 1" captionStartId="4.[264,354,294,318]" captionTargetPageId="4" captionText="TABLE 1. Measurements (mm) of Cambarus clivosus, new species." httpUri="http://table.plazi.org/id/DF5566794B30083A946BBD6D3183FED9" pageId="2" pageNumber="31" tableUuid="DF5566794B30083A946BBD6D3183FED9">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C330657A4B36083D946BBDC43105F8FC" lastPageId="3" lastPageNumber="32" pageId="2" pageNumber="31" type="diagnosis">
<paragraph id="8B9536F14B36083C946BBDC43352FAA4" blockId="2.[264,1324,399,1906]" pageId="2" pageNumber="31">
<emphasis id="B95EEAE34B36083C946BBDC43310FE4E" box="[264,383,399,424]" italics="true" pageId="2" pageNumber="31">Diagnosis</emphasis>
.—Body and eyes pigmented. Rostrum triangular and moderately excavated, terminating in short, upturned acumen, weakly decurved in lateral view; median carina absent. Rostral margins thickened and bordered mesially with tufts of short setae; margins straight and converging. Areola 33.3–39.2 % (= 37.0, n = 38, 
<emphasis id="B95EEAE34B36083C9168BE4C3640FDC6" box="[1035,1071,519,544]" italics="true" pageId="2" pageNumber="31">SD</emphasis>
= 1.5) of total length of carapace (CL = distance from distal tip of acumen to dorsocaudal edge of cephalothorax) and 38.9–44.3 % (= 42.3, n = 40, 
<emphasis id="B95EEAE34B36083C96F5BE1331D6FD97" box="[918,953,600,625]" italics="true" pageId="2" pageNumber="31">SD</emphasis>
= 1.3) of postorbital carapace length (PCL = distance from anteriolateral edge of cephalothorax immediately behind eye to dorsocaudal edge of cephalothorax)). Areola sparsely punctate, narrowest at midpoint, 6.4–11.8 (= 8.6, n = 35, 
<emphasis id="B95EEAE34B36083C972EBE9B301EFD0F" box="[589,625,720,745]" italics="true" pageId="2" pageNumber="31">SD</emphasis>
= 1.2) times as long as wide with 2 to 4 (mode = 3, n = 35, 
<emphasis id="B95EEAE34B36083C946BBEB33344FCF7" box="[264,299,760,785]" italics="true" pageId="2" pageNumber="31">SD</emphasis>
= 0.2) punctations across narrowest part (Fig. 2A). Postorbital carapace length 84.4– 89.7 % (= 87.6, n = 40, 
<emphasis id="B95EEAE34B36083C9721BF6B3009FCDF" box="[578,614,800,825]" italics="true" pageId="2" pageNumber="31">SD</emphasis>
= 1.2) of CL. Cervical spines absent. Postorbital ridges low with smoothly rounded terminal ends (Fig. 2A). Suborbital angle approximately 90 degrees or slightly larger. Antennal scale broadest distal to midlength, distal margin either nearly perpendicular to lateral margin or angular to lateral margin (see Variation), thickened lateral margin terminating in short corneous spine (Figs. 2E, 2F). Incisor region of mandible deeply indented with two large cusps. Cephalomedian lobe of epistome bellshaped with small, triangular cephalomedian projection; epistomal zygoma strongly arched. Ischia of third pereiopods of males with large hooks, hooks overreaching basioischial articulation in form I males only. Coxa of fourth pereiopods of males with caudomesial boss. Chela with 1 row of tubercles (see Variation) along mesial margin of palm forming cristiform row, usually 6 or 7 tubercles (range 5–8, mode = 6, n = 40, 
<emphasis id="B95EEAE34B36083C918DB8FB377DFB2F" box="[1262,1298,1200,1225]" italics="true" pageId="2" pageNumber="31">SD</emphasis>
= 0.6) in row; small tufts of setae along opposable margins, and dorsal, lateral, and mesial surfaces of fingers; dorsal surfaces of fingers with well defined longitudinal ridges (Fig. 2H).
</paragraph>
<paragraph id="8B9536F14B36083D945BB91B3105F8FC" blockId="2.[264,1324,399,1906]" lastBlockId="3.[264,1322,1696,1818]" lastPageId="3" lastPageNumber="32" pageId="2" pageNumber="31">
First pleopods of form I male symmetrical and continuous at bases (Fig. 
<date id="FF9410314B36083C91E4B91B36CDFA8C" box="[1159,1186,1360,1386]" pageId="2" pageNumber="31">2I</date>
), extending to midpoint of bases of third pereiopods when abdomen flexed. First pleopod of form I male terminating in two elements: corneous central projection strongly recurved caudodistally at an angle of approximately 110 degrees, bladelike, with subapical notch, and reaching proximally to or just beyond distal base of mesial process (Fig. 2J); mesial process non­corneous, bulbous at base, distal end with multiple caudodistally directed small projections, distal half directed caudolaterally at 15 to 25 degrees, proximal edge usually at 90 degree angle to main axis of pleopod. First pleopod of form II male noncorneous, both elements arched caudodistally at 90 degrees to main axis of pleopod: mesial process of larger individuals directed caudolaterally at 15 to 25 degrees (Fig. 2G). Annulus ventralis immovable, subrhomboidal; cephalic half with wide median trough; caudal half strongly elevated; transverse fossa well defined with tongue extending either dextrally or sinistrally, disappearing under caudolateral wall; sinuate sinus running from center of fossa to caudal edge (Fig. 2D). view of right antennal scale; F, Dorsal view of right antennal scale; G, Lateral view of first pleopod of form II male; H, Dorsal view of right chela; I, Caudal view of first pleopods of form I male; J, Mesial view of first pleopod of form I male. A, C, F, H, I and J of 
<typeStatus id="549188534B37083D96B2BAAA3641F91E" box="[977,1070,1761,1784]" pageId="3" pageNumber="32" type="holotype">holotype</typeStatus>
(INHS 10235); 
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D, E of 
<typeStatus id="549188534B37083D946BBB483331F8FC" box="[264,350,1795,1818]" pageId="3" pageNumber="32" type="allotype">allotype</typeStatus>
(
<specimenCode id="DB8C9E8A4B37083D940FBB483396F8FC" box="[364,505,1795,1818]" collectionCode="INHS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="3" pageNumber="32" type="Other type of research institution/biorepository">INHS 10236</specimenCode>
)
</materialsCitation>
; 
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G of morphotype (
<specimenCode id="DB8C9E8A4B37083D97B2BB483131F8FC" box="[721,862,1795,1818]" collectionCode="INHS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="3" pageNumber="32" type="Other type of research institution/biorepository">INHS 10232</specimenCode>
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.
</paragraph>
</subSubSection>
<caption id="DF5566794B30083A946BBD6D3183FED9" ID-Table-UUID="DF5566794B30083A946BBD6D3183FED9" box="[264,1004,291,319]" httpUri="http://table.plazi.org/id/DF5566794B30083A946BBD6D3183FED9" pageId="4" pageNumber="33" targetBox="[278,1188,357,1129]" targetIsTable="true" targetPageId="4">
<paragraph id="8B9536F14B30083A946BBD6D3183FED9" blockId="4.[264,1004,291,319]" box="[264,1004,291,319]" pageId="4" pageNumber="33">
<emphasis id="B95EEAE34B30083A946BBD6D3312FED8" bold="true" box="[264,381,294,318]" pageId="4" pageNumber="33">TABLE 1.</emphasis>
Measurements (mm) of 
<taxonomicName id="4C2A4D724B30083A97E8BD6C3138FED8" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[651,855,295,318]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B30083A97E8BD6C3138FED8" box="[651,855,295,318]" italics="true" pageId="4" pageNumber="33">Cambarus clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B30083A9607BD6D318BFEDB" box="[868,996,294,317]" pageId="4" pageNumber="33" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="8B9536F14B30083A97B7BD2E365CFB8F" pageId="4" pageNumber="33">
<table id="F92AC4514B30F7C19475BD2E36CBFB8F" box="[278,1188,357,1129]" gridcols="4" gridrows="6" pageId="4" pageNumber="33">
<tr id="351A34B34B30F7C19475BD2E36CBFE9A" box="[278,1188,357,380]" gridrow="0" pageId="4" pageNumber="33" rowspan-0="1">
<th id="76CB5DCF4B30F7C197A4BD2E3158FE9A" box="[711,823,357,380]" gridcol="1" gridrow="0" pageId="4" pageNumber="33">Holotype</th>
<th id="76CB5DCF4B30F7C19638BD2E31B8FE9A" box="[859,983,357,380]" gridcol="2" gridrow="0" pageId="4" pageNumber="33">Allotype</th>
<th id="76CB5DCF4B30F7C1916ABD2E36CBFE9A" box="[1033,1188,357,380]" gridcol="3" gridrow="0" pageId="4" pageNumber="33">Morphotype</th>
</tr>
<tr id="351A34B34B30F7C19475BDDC36CBFDCA" box="[278,1188,407,556]" gridrow="1" pageId="4" pageNumber="33">
<th id="76CB5DCF4B30F7C19475BDDC3052FDCA" box="[278,573,407,556]" gridcol="0" gridrow="1" pageId="4" pageNumber="33">Carapace: Total length Postorbital length Width Height</th>
<td id="76CB5DCF4B30F7C197A4BDDC3158FDCA" box="[711,823,407,556]" gridcol="1" gridrow="1" pageId="4" pageNumber="33">27.3 24.1 14.2 10.5</td>
<td id="76CB5DCF4B30F7C19638BDDC31B8FDCA" box="[859,983,407,556]" gridcol="2" gridrow="1" pageId="4" pageNumber="33">32.7 29.1 16.2 12.8</td>
<td id="76CB5DCF4B30F7C1916ABDDC36CBFDCA" box="[1033,1188,407,556]" gridcol="3" gridrow="1" pageId="4" pageNumber="33">18.7 16.3 8.9 5.6</td>
</tr>
<tr id="351A34B34B30F7C19475BE1C36CBFD4D" box="[278,1188,599,683]" gridrow="2" pageId="4" pageNumber="33">
<th id="76CB5DCF4B30F7C19475BE1C3052FD4D" box="[278,573,599,683]" gridcol="0" gridrow="2" pageId="4" pageNumber="33">Areola: Width Length</th>
<td id="76CB5DCF4B30F7C197A4BE1C3158FD4D" box="[711,823,599,683]" gridcol="1" gridrow="2" pageId="4" pageNumber="33">1.2 10.5</td>
<td id="76CB5DCF4B30F7C19638BE1C31B8FD4D" box="[859,983,599,683]" gridcol="2" gridrow="2" pageId="4" pageNumber="33">1.3 12.7</td>
<td id="76CB5DCF4B30F7C1916ABE1C36CBFD4D" box="[1033,1188,599,683]" gridcol="3" gridrow="2" pageId="4" pageNumber="33">0.7 6.8</td>
</tr>
<tr id="351A34B34B30F7C19475BE9D36CBFCCD" box="[278,1188,726,811]" gridrow="3" pageId="4" pageNumber="33">
<th id="76CB5DCF4B30F7C19475BE9D3052FCCD" box="[278,573,726,811]" gridcol="0" gridrow="3" pageId="4" pageNumber="33">Rostrum: Width Length</th>
<td id="76CB5DCF4B30F7C197A4BE9D3158FCCD" box="[711,823,726,811]" gridcol="1" gridrow="3" pageId="4" pageNumber="33">3.3 4.1</td>
<td id="76CB5DCF4B30F7C19638BE9D31B8FCCD" box="[859,983,726,811]" gridcol="2" gridrow="3" pageId="4" pageNumber="33">3.9 5.0</td>
<td id="76CB5DCF4B30F7C1916ABE9D36CBFCCD" box="[1033,1188,726,811]" gridcol="3" gridrow="3" pageId="4" pageNumber="33">2.4 3.2</td>
</tr>
<tr id="351A34B34B30F7C19475BF1D36CBFC0C" box="[278,1188,854,1002]" gridrow="4" pageId="4" pageNumber="33">
<th id="76CB5DCF4B30F7C19475BF1D3052FC0C" box="[278,573,854,1002]" gridcol="0" gridrow="4" pageId="4" pageNumber="33">Chela, right: Length, palm mesial margin Palm width Length, lateral margin Dactyl length</th>
<td id="76CB5DCF4B30F7C197A4BF1D3158FC0C" box="[711,823,854,1002]" gridcol="1" gridrow="4" pageId="4" pageNumber="33">7.4 11.1 20.8 11.9</td>
<td id="76CB5DCF4B30F7C19638BF1D31B8FC0C" box="[859,983,854,1002]" gridcol="2" gridrow="4" pageId="4" pageNumber="33">8.6 11.7 22.9 13.4</td>
<td id="76CB5DCF4B30F7C1916ABF1D36CBFC0C" box="[1033,1188,854,1002]" gridcol="3" gridrow="4" pageId="4" pageNumber="33">4.6 6.5 12.1 6.5</td>
</tr>
<tr id="351A34B34B30F7C19475B85E36CBFB8F" box="[278,1188,1045,1129]" gridrow="5" pageId="4" pageNumber="33">
<th id="76CB5DCF4B30F7C19475B85E3052FB8F" box="[278,573,1045,1129]" gridcol="0" gridrow="5" pageId="4" pageNumber="33">Abdomen: Width Length</th>
<td id="76CB5DCF4B30F7C197A4B85E3158FB8F" box="[711,823,1045,1129]" gridcol="1" gridrow="5" pageId="4" pageNumber="33">12.0 28.1</td>
<td id="76CB5DCF4B30F7C19638B85E31B8FB8F" box="[859,983,1045,1129]" gridcol="2" gridrow="5" pageId="4" pageNumber="33">13.8 33.3</td>
<td id="76CB5DCF4B30F7C1916AB85E36CBFB8F" box="[1033,1188,1045,1129]" gridcol="3" gridrow="5" pageId="4" pageNumber="33">7.6 18.6</td>
</tr>
</table>
</paragraph>
<subSubSection id="C330657A4B300838945BB8F933D1FBC8" lastPageId="6" lastPageNumber="35" pageId="4" pageNumber="33" type="description">
<paragraph id="8B9536F14B30083A945BB8F931ACF94A" blockId="4.[264,1324,1202,1868]" pageId="4" pageNumber="33">
<emphasis id="B95EEAE34B30083A945BB8F93164FB2D" box="[312,779,1202,1227]" italics="true" pageId="4" pageNumber="33">Description of holotypic male, form I</emphasis>
.—Body depressed dorsoventrally, at widest points carapace wider than abdomen (14.2 and 12.0 mm, respectively). Greatest width of carapace larger than height at caudodorsal margin of cervical groove (14.2 and 
<quantity id="4CD29B144B30083A91D9B9493748FAFA" box="[1210,1319,1282,1308]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="33" unit="mm" value="10.5">10.5 mm</quantity>
, respectively). Postorbital carapace length 88.3% of total length of carapace. Areola 8.8 times longer (
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) than wide (
<quantity id="4CD29B144B30083A97A3B919314DFA8A" box="[704,802,1362,1388]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="33" unit="mm" value="1.2">1.2 mm</quantity>
) with 3 punctations across narrowest part; length of areola 38.5% of total length of carapace. Rostrum moderately excavated dorsally, floor smooth, lacking carina; margins thickened, straight and converging. Each margin bordered mesially by row of tufts of short setae. Acumen short and terminating in small corneous spine, reaching just posterior to distal margin of third antennal endopod. Postorbital ridges weakly developed, termini smoothly rounded and lacking spine or tubercle. Suborbital angles approximately 90 degrees. Cervical spines or tubercles absent. Branchiostegal areas and lateral surfaces of carapace covered with low tubercles, tubercles in branchiostegal areas larger than those on lateral surfaces.
</paragraph>
<paragraph id="8B9536F14B30083A945BBAF13744F8AA" blockId="4.[264,1324,1202,1868]" pageId="4" pageNumber="33">
Abdomen longer than carapace (28.1 and 
<quantity id="4CD29B144B30083A9629BAF131D3F932" box="[842,956,1722,1748]" metricMagnitude="-2" metricUnit="m" metricValue="2.73" pageId="4" pageNumber="33" unit="mm" value="27.3">27.3 mm</quantity>
, respectively). Anteriolateral margins of pleura smoothly rounded, posteriolateral margins of second, fourth, and fifth segments rounded, posteriolateral margin of third segment slightly angular. Cephalic section of telson bearing 3 spines in each caudolateral corner, more mesial pairs movable.
</paragraph>
<paragraph id="8B9536F14B31083B946BBD573643FE18" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="34">Proximal podomere of uropod with large spine extending over mesial ramus and small spine in caudolateral corner extending over lateral ramus. Caudal margin of cephalic section of lateral ramus with 10 fixed spines and 1 movable spine in caudolateral corner, cephalic and caudal sections of lateral ramus with median ridge. Lateral margin of mesial ramus terminating in spine; mesial ramus with prominent median ridge terminating in premarginal spine. Dorsal surfaces of telson and uropods setiferous.</paragraph>
<paragraph id="8B9536F14B31083B945BBE4733ACFD90" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="34">
Antennal scale with distal margin nearly perpendicular to lateral margin, as in Diagnosis in other respects. Right antennal scale 
<quantity id="4CD29B144B31083B9603BE7F31D4FDA8" box="[864,955,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="34" unit="mm" value="3.5">3.5 mm</quantity>
long, 
<quantity id="4CD29B144B31083B9164BE7F360DFDA8" box="[1031,1122,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="34" unit="mm" value="1.4">1.4 mm</quantity>
wide. Epistome as in Diagnosis.
</paragraph>
<paragraph id="8B9536F14B31083B945BBECF310EFBB0" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="34">Lateral margin of right chela slightly indented near base of fixed finger of propodus; mesial surface of palm with single cristiform row of 8 tubercles; dorsodistal margin of palm region at base of dactyl with two large bulbous tubercles (Fig. 2H). Mesial and lateral surfaces of chela, dorsal and ventral surfaces of fingers, and opposable margins of fingers covered with setiferous punctations. Dorsal surface of finger of propodus with strong submedian longitudinal ridge flanked by setiferous punctations; basal half of opposable margin with 3 tubercles which gradually increase in size, third tubercle from base of finger largest, 1 corneous tubercle just distal to midlength between third tubercle and distal tip of finger. Dorsal surface of dactyl with strong submedian longitudinal ridge flanked by setiferous punctations; basal half of opposable margin with 4 tubercles of equal size, tubercles smaller than largest tubercle on opposable margin of propodus. Finger of propodus and dactyl with subterminal corneous tip.</paragraph>
<paragraph id="8B9536F14B31083B945BB82F308BFAF8" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="34">Right carpus with deep oblique furrow dorsally; mesial margin with 1 large and 2 small spines, large spine distal­most; ventral surface with 1 small centrally located spine. Ventral surface of merus with 1 large corneous spine at distolateral corner and mesial row of 7 spines; row terminating in large corneous spine. Ischium with 1 weak tubercle just proximal to midlength of mesial margin.</paragraph>
<paragraph id="8B9536F14B31083B945BB96731ECF9E8" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="34">
Hook on ischium of third pereiopod only; hook long, overreaching basioischial articulation, not opposed by tubercle on basis. First pleopods of form I male symmetrical and continuous at bases (Fig. 
<date id="FF9410314B31083B9707B9373010FA70" box="[612,639,1404,1430]" pageId="5" pageNumber="34">2I</date>
), extending to midpoint of bases of third pereiopods when abdomen flexed. First pleopod of form I male as described in Diagnosis but with distal half of mesial process directed caudolaterally approximately 15 degrees, proximal edge at 90 degree angle to main axis of the pleopod. (
<figureCitation id="13112A744B31083B9642B9BF311CF9E8" box="[801,883,1524,1550]" captionStart="FIGURE 1" captionStartId="1.[264,368,1325,1349]" captionTargetBox="[449,1140,846,1305]" captionTargetId="figure@1.[449,1140,846,1305]" captionTargetPageId="1" captionText="FIGURE 1. Typical habitat for Cambarus clivosus, new species—a high gradient, seep­fed creek in Cannon Co., TN." httpUri="https://zenodo.org/record/2646442/files/figure.png" pageId="5" pageNumber="34">Fig. 1J</figureCitation>
).
</paragraph>
<paragraph id="8B9536F14B31083B945BBA573766F930" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="34">
<emphasis id="B95EEAE34B31083B945BBA5730A4F9D3" box="[312,715,1564,1589]" italics="true" pageId="5" pageNumber="34">Description of allotypic female</emphasis>
.—Differing from 
<typeStatus id="549188534B31083B96DDBA57364AF9D0" box="[958,1061,1564,1590]" pageId="5" pageNumber="34" type="holotype">holotype</typeStatus>
as follows: Areola constituting 38.8% of length of carapace and 9.8 times longer than wide. Postorbital carapace length 89.0% of length of carapace. Mesial surface of palm of right chela with single row of 7 tubercles. Caudal margin of cephalic section of lateral ramus of uropod with 9 fixed spines. Distal margin of antennal scale angular to lateral margin (Fig. 2E).
</paragraph>
<paragraph id="8B9536F14B31083B944ABAAF3190F8A8" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="34">Sternum between third and fourth pereiopods narrowly V­shaped. Annulus as described in Diagnosis but with tongue in sulcus extending sinistrally. First pleopod uniramous, barely reaching caudal margin of annulus when abdomen flexed.</paragraph>
<paragraph id="8B9536F14B320838945BBD5731EDFE18" blockId="6.[264,1323,284,1070]" pageId="6" pageNumber="35">
<emphasis id="B95EEAE34B320838945BBD57312BFED3" box="[312,836,284,309]" italics="true" pageId="6" pageNumber="35">Description of morphotypic male, form II</emphasis>
.—Differing from 
<typeStatus id="549188534B320838914FBD5736FCFED0" box="[1068,1171,284,310]" pageId="6" pageNumber="35" type="holotype">holotype</typeStatus>
as follows: Areola constituting 36.4% of length of carapace, 9.7 times longer than wide. Postorbital carapace length 87.1% of length of carapace. Mesial surface of palm of right chela with single row of 7 tubercles, 2 proximal­most tubercles fused. Ventral surface of merus with mesial row of 6 spines and 2 spines in distolateral corner. Caudal margin of cephalic section of lateral ramus of uropod with 9 fixed spines.
</paragraph>
<paragraph id="8B9536F14B320838945BBE4730B2FDA8" blockId="6.[264,1323,284,1070]" pageId="6" pageNumber="35">Hook on ischium of third pereiopod short, not overreaching basioischial articulation. First pleopod as described in Diagnosis.</paragraph>
<paragraph id="8B9536F14B320838945BBE1731A9FD20" blockId="6.[264,1323,284,1070]" pageId="6" pageNumber="35">
<emphasis id="B95EEAE34B320838945BBE173308FD93" box="[312,359,604,629]" italics="true" pageId="6" pageNumber="35">Size</emphasis>
.—The largest specimen examined is a 35.0 mm CL female. Females (
<emphasis id="B95EEAE34B32083891ACBE1736B1FD93" box="[1231,1246,604,629]" italics="true" pageId="6" pageNumber="35">n</emphasis>
= 16) range in size from 11.9 to 35.0 mm CL. Form I males (
<emphasis id="B95EEAE34B32083896E9BECF31F6FD7B" box="[906,921,644,669]" italics="true" pageId="6" pageNumber="35">n</emphasis>
= 10) range from 
<quantity id="4CD29B144B320838910FBECF3744FD78" box="[1132,1323,644,670]" metricMagnitude="-2" metricUnit="m" metricValue="2.395" metricValueMax="2.82" metricValueMin="1.97" pageId="6" pageNumber="35" unit="mm" value="23.95" valueMax="28.2" valueMin="19.7">19.7 to 28.2 mm</quantity>
CL. Form II males (
<emphasis id="B95EEAE34B3208389498BEE73065FD23" box="[507,522,684,709]" italics="true" pageId="6" pageNumber="35">n</emphasis>
= 9) range from 13.3 to 28.0 mm CL.
</paragraph>
<paragraph id="8B9536F14B320838945BBE9F33D1FBC8" blockId="6.[264,1323,284,1070]" pageId="6" pageNumber="35">
<emphasis id="B95EEAE34B320838945BBE9F3314FD0B" box="[312,379,724,749]" italics="true" pageId="6" pageNumber="35">Color</emphasis>
.— Base color of dorsal and lateral surfaces of chelae, cephalothorax, abdomen, pereiopods, and tail fan rather uniform and light to dark brown (
<figureCitation id="13112A744B3208389699BEB7362DFCF0" box="[1018,1090,764,790]" captionStart="FIGURE 3" captionStartId="6.[264,368,1857,1881]" captionTargetBox="[269,1317,1169,1845]" captionTargetId="figure@6.[269,1317,1169,1845]" captionTargetPageId="6" captionText="FIGURE 3. Cambarus clivosus, new species, Holotype." httpUri="https://zenodo.org/record/2646446/files/figure.png" pageId="6" pageNumber="35">Fig. 3</figureCitation>
). Dorsal surface of abdominal segments 1 through 5 usually darker brown than base color, giving the appearance of dark wedge running down center of dorsum. Pleurites of first abdominal segment cream colored. Tubercles on chelae and spines on carpus light orange or cream colored. Rostral margins orange or crimson red (
<figureCitation id="13112A744B3208389633BFD731F8FC50" box="[848,919,924,950]" captionStart="FIGURE 3" captionStartId="6.[264,368,1857,1881]" captionTargetBox="[269,1317,1169,1845]" captionTargetId="figure@6.[269,1317,1169,1845]" captionTargetPageId="6" captionText="FIGURE 3. Cambarus clivosus, new species, Holotype." httpUri="https://zenodo.org/record/2646446/files/figure.png" pageId="6" pageNumber="35">Fig. 3</figureCitation>
). Faint U­shaped saddle centered at caudal edge of cephalothorax and extending cephalically may be present on individuals with light brown base color. Ventral surfaces of chelae, cephalothorax, and abdomen cream to white.
</paragraph>
</subSubSection>
<caption id="DF5566794B320838946BBB0A3106F8BE" box="[264,873,1857,1881]" httpUri="https://zenodo.org/record/2646446/files/figure.png" pageId="6" pageNumber="35" startId="6.[264,368,1857,1881]" targetBox="[269,1317,1169,1845]" targetPageId="6">
<paragraph id="8B9536F14B320838946BBB0A3106F8BE" blockId="6.[264,873,1857,1881]" box="[264,873,1857,1881]" pageId="6" pageNumber="35">
<emphasis id="B95EEAE34B320838946BBB0A33E5F8BF" bold="true" box="[264,394,1857,1881]" pageId="6" pageNumber="35">FIGURE 3.</emphasis>
<taxonomicName id="4C2A4D724B32083894FBBB09300CF8BF" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[408,611,1858,1881]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B32083894FBBB09300CF8BF" box="[408,611,1858,1881]" italics="true" pageId="6" pageNumber="35">Cambarus clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B3208389712BB0A309BF8BE" box="[625,756,1857,1880]" pageId="6" pageNumber="35" rank="species">new species</taxonomicNameLabel>
, Holotype.
</paragraph>
</caption>
<subSubSection id="C330657A4B330837945BBD573677FE60" lastPageId="9" lastPageNumber="38" pageId="7" pageNumber="36" type="materials_examined">
<paragraph id="8B9536F14B330839945BBD5730AFFD90" blockId="7.[264,1324,284,1830]" pageId="7" pageNumber="36">
<emphasis id="B95EEAE34B330839945BBD5733A0FED3" box="[312,463,284,309]" italics="true" pageId="7" pageNumber="36">Type locality</emphasis>
.—Seep­fed tributary of Dry Creek at Pea Ridge Wildlife Management Area, approximately 
<quantity id="4CD29B144B3308399763BD0F3030FEB8" box="[512,607,324,350]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="7" pageNumber="36" unit="km" value="11.5">11.5 km</quantity>
WSW Smithville, De Kalb County, 
<collectingRegion id="49EEF8134B330839916ABD0F36EEFEB8" box="[1033,1153,324,350]" country="United States of America" name="Tennessee" pageId="7" pageNumber="36">Tennessee</collectingRegion>
(
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, ­ 
<geoCoordinate id="EE1E50364B330839946BBD2733E1FE60" box="[264,398,364,390]" direction="west" orientation="longitude" pageId="7" pageNumber="36" precision="5" value="-85.9271">85.9271°W</geoCoordinate>
NAD 83). The tributary lies in a densely wooded ravine with walls that sloped at approximately 60 degrees. The 
<typeStatus id="549188534B33083997FABDDF3090FE48" box="[665,767,404,430]" pageId="7" pageNumber="36" type="holotype">holotype</typeStatus>
was collected from under a piece of flat rock in mid­channel while the 
<typeStatus id="549188534B330839977ABDF73018FE30" box="[537,631,444,470]" pageId="7" pageNumber="36" type="allotype">allotype</typeStatus>
was collected from a shallow burrow dug into the stream bank. The morphotype was collected from another site. At the type locality, the seep­fed creek was approximately 
<quantity id="4CD29B144B3308399728BE4730F9FDC0" box="[587,662,524,550]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="36" unit="m" value="0.5">0.5 m</quantity>
wide with gravel and small­sized cobble and shale substrates. Rock outcroppings occurred along both banks. Water depth was 
<quantity id="4CD29B144B33083991E8BE7F3697FDA8" box="[1163,1272,564,590]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="7" pageNumber="36" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1 to 3 cm</quantity>
and flow was not constant in all stretches.
</paragraph>
<paragraph id="8B9536F14B330839945FBECF3135FC68" blockId="7.[264,1324,284,1830]" pageId="7" pageNumber="36">
<emphasis id="B95EEAE34B330839945FBECF3044FD7B" box="[316,555,644,669]" italics="true" pageId="7" pageNumber="36">Disposition of types</emphasis>
.—The 
<typeStatus id="549188534B33083997EBBECF309DFD78" box="[648,754,644,670]" pageId="7" pageNumber="36" type="holotype">holotype</typeStatus>
, 
<typeStatus id="549188534B3308399662BECF310DFD78" box="[769,866,644,670]" pageId="7" pageNumber="36" type="allotype">allotype</typeStatus>
, and morphotype are deposited in the 
<collectingRegion id="49EEF8134B330839946BBEE73334FD20" box="[264,347,684,710]" country="United States of America" name="Illinois" pageId="7" pageNumber="36">Illinois</collectingRegion>
Natural History Survey Crustacean Collection as INHS10235, INHS 10236, and INHS 10232, respectively, as are the following 
<typeStatus id="549188534B330839962CBE9F31ABFD08" box="[847,964,724,750]" pageId="7" pageNumber="36" type="paratype">paratypes</typeStatus>
: 1 MI and 1F, INHS 10237. 
<typeStatus id="549188534B330839946BBEB73315FCF0" box="[264,378,764,790]" pageId="7" pageNumber="36" type="paratype">Paratypes</typeStatus>
are deposited at the National Museum of Natural History, Smithsonian Institution, 
<collectingRegion id="49EEF8134B33083994F1BF6F304FFCD8" box="[402,544,804,830]" country="United States of America" name="Washington" pageId="7" pageNumber="36">Washington</collectingRegion>
, D. C. as follows: 1 MI, 1 MII, and 1 F, USNM 1083161. The localities and dates of collection for 
<typeStatus id="549188534B3308399783BF07313FFC80" box="[736,848,844,870]" pageId="7" pageNumber="36" type="paratype">paratypes</typeStatus>
and morphotype are provided in the following Range and specimens examined section.
</paragraph>
<paragraph id="8B9536F14B330839944FBFD730B8FAA0" blockId="7.[264,1324,284,1830]" pageId="7" pageNumber="36">
<emphasis id="B95EEAE34B330839944FBFD730CEFC53" box="[300,673,924,949]" italics="true" pageId="7" pageNumber="36">Range and specimens examined</emphasis>
.— 
<taxonomicName id="4C2A4D724B33083997A5BFD731CAFC53" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[710,933,924,949]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B33083997A5BFD731CAFC53" box="[710,933,924,949]" italics="true" pageId="7" pageNumber="36">Cambarus clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B33083996D7BFD7362BFC50" box="[948,1092,924,950]" pageId="7" pageNumber="36" rank="species">new species</taxonomicNameLabel>
, is currently known to occur in a portion of the Eastern Highland Rim in central 
<collectingRegion id="49EEF8134B330839969DBF8F3619FC38" box="[1022,1142,964,990]" country="United States of America" name="Tennessee" pageId="7" pageNumber="36">Tennessee</collectingRegion>
(
<figureCitation id="13112A744B33083991E8BF8F36B7FC38" box="[1163,1240,964,990]" captionStart="FIGURE 4" captionStartId="8.[264,368,1127,1151]" captionTargetBox="[323,1493,285,1034]" captionTargetId="figure@8.[300,1337,270,1071]" captionTargetPageId="8" captionText="FIGURE 4. Known range of Cambarus clivosus, new species, in central Tennessee, USA. Closed circles indicate sites at which C. clivosus was found; open circles indicate sites at which C. clivosus was absent." httpUri="https://zenodo.org/record/2646448/files/figure.svg" pageId="7" pageNumber="36">Fig. 4</figureCitation>
). The species seems to be restricted to a narrow (
<quantity id="4CD29B144B330839979FBFA73127FBE0" box="[764,840,1004,1030]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="36" unit="km" value="14.0">14 km</quantity>
wide) peninsula of elevated Mississipian age limestone fortified with chert and shale formations (
<bibRefCitation id="EFBB4B004B33083996A3B85F3685FBC8" author="Etnier, D. A. &amp; Starnes, W. C." box="[960,1258,1044,1070]" pageId="7" pageNumber="36" refId="ref5031" refString="Etnier, D. A. &amp; Starnes, W. C. (1993) The Fishes of Tennessee. University of Tennessee Press, Knoxville, xiv + 681 pp." type="book" year="1993">Etnier and Starnes 1993</bibRefCitation>
) that extends south­southwestward from the Highland Rim through the counties of De Kalb and Cannon. Creeks in the peninsula are characterized by chert gravel and sand substrates interspersed with areas of bedrock and drain to the north and east into the Caney Fork River drainage and to the west into the Stones River drainage. Both the Caney Fork and Stones flow north and northwest into the Cumberland River. At 
<quantity id="4CD29B144B3308399175B897360CFB10" box="[1046,1123,1244,1270]" metricMagnitude="2" metricUnit="m" metricValue="6.37" pageId="7" pageNumber="36" unit="m" value="637.0">637 m</quantity>
above sea level, Short Mountain is the highest point in the aforementioned peninsula; other hills average 
<quantity id="4CD29B144B330839946BB96733CBFAA0" box="[264,420,1324,1350]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="7" pageNumber="36" unit="m" value="325.0" valueMax="350.0" valueMin="300.0">300 to 350 m</quantity>
with slopes of 30 to 60%.
</paragraph>
<paragraph id="8B9536F14B3308399451B91F331EF8C0" blockId="7.[264,1324,284,1830]" pageId="7" pageNumber="36">
Collections made in several high gradient seep­fed creeks in nearby Eastern Highland Rim sites north, south, and east of the above described peninsula failed to record the presence of 
<taxonomicName id="4C2A4D724B33083994F4B9EF3077FA5B" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[407,536,1444,1470]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B33083994F4B9EF33C5FA5B" box="[407,426,1444,1469]" italics="true" pageId="7" pageNumber="36">C</emphasis>
. 
<emphasis id="B95EEAE34B33083994D9B9EF3077FA5B" box="[442,536,1444,1469]" italics="true" pageId="7" pageNumber="36">clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B3308399744B9EF30DAFA58" box="[551,693,1444,1470]" pageId="7" pageNumber="36" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="13112A744B33083997A5B9EF3160FA58" box="[710,783,1444,1470]" captionStart="FIGURE 4" captionStartId="8.[264,368,1127,1151]" captionTargetBox="[323,1493,285,1034]" captionTargetId="figure@8.[300,1337,270,1071]" captionTargetPageId="8" captionText="FIGURE 4. Known range of Cambarus clivosus, new species, in central Tennessee, USA. Closed circles indicate sites at which C. clivosus was found; open circles indicate sites at which C. clivosus was absent." httpUri="https://zenodo.org/record/2646448/files/figure.svg" pageId="7" pageNumber="36">Fig. 4</figureCitation>
). Additionally, a search of the over 4000 lots of 
<collectingRegion id="49EEF8134B3308399449B98733CDFA00" box="[298,418,1484,1510]" country="United States of America" name="Tennessee" pageId="7" pageNumber="36">Tennessee</collectingRegion>
crayfishes held at the National Museum of Natural History and the 
<collectingRegion id="49EEF8134B33083991BBB9873744FA00" box="[1240,1323,1484,1510]" country="United States of America" name="Illinois" pageId="7" pageNumber="36">Illinois</collectingRegion>
Natural History Survey Crustacean Collection did not produce additional specimens of 
<taxonomicName id="4C2A4D724B3308399073B9BF3309F9D3" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B3308399073B9BF374CF9EB" box="[1296,1315,1524,1549]" italics="true" pageId="7" pageNumber="36">C</emphasis>
. 
<emphasis id="B95EEAE34B330839946BBA573309F9D3" box="[264,358,1564,1589]" italics="true" pageId="7" pageNumber="36">clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B330839941BBA573065F9D0" box="[376,522,1564,1590]" pageId="7" pageNumber="36" rank="species">new species</taxonomicNameLabel>
. While it is highly likely that additional locations for 
<taxonomicName id="4C2A4D724B33083991C3BA57374CF9D3" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[1184,1315,1564,1590]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B33083991C3BA5736DCF9D3" box="[1184,1203,1564,1589]" italics="true" pageId="7" pageNumber="36">C</emphasis>
. 
<emphasis id="B95EEAE34B33083991A6BA57374CF9D3" box="[1221,1315,1564,1589]" italics="true" pageId="7" pageNumber="36">clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B330839946BBA0F33F6F9B8" box="[264,409,1604,1630]" pageId="7" pageNumber="36" rank="species">new species</taxonomicNameLabel>
will be found in the elevated peninsula in Cannon and De Kalb counties described above, it is unknown if the species will be found in other regions of central 
<collectingRegion id="49EEF8134B330839946BBADF33EFF948" box="[264,384,1684,1710]" country="United States of America" name="Tennessee" pageId="7" pageNumber="36">Tennessee</collectingRegion>
or the Eastern Highland Rim. Additional collecting efforts are needed in the Caney Fork River drainage both north and east of the currently known range of 
<taxonomicName id="4C2A4D724B3308399073BAF73309F91B" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B3308399073BAF7374CF933" box="[1296,1315,1724,1749]" italics="true" pageId="7" pageNumber="36">C</emphasis>
. 
<emphasis id="B95EEAE34B330839946BBAAF3309F91B" box="[264,358,1764,1789]" italics="true" pageId="7" pageNumber="36">clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B330839941ABAAF3065F918" box="[377,522,1764,1790]" pageId="7" pageNumber="36" rank="species">new species</taxonomicNameLabel>
to determine if this species is restricted to Cannon and De Kalb counties.
</paragraph>
<caption id="DF5566794B3C0836946BB82C33EBFB2D" httpUri="https://zenodo.org/record/2646448/files/figure.svg" pageId="8" pageNumber="37" startId="8.[264,368,1127,1151]" targetBox="[323,1493,285,1034]" targetPageId="8">
<paragraph id="8B9536F14B3C0836946BB82C33EBFB2D" blockId="8.[264,1323,1127,1227]" pageId="8" pageNumber="37">
<emphasis id="B95EEAE34B3C0836946BB82C33E5FB99" bold="true" box="[264,394,1127,1151]" pageId="8" pageNumber="37">FIGURE 4.</emphasis>
Known range of 
<taxonomicName id="4C2A4D724B3C0836972FB8233178FB99" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[588,791,1128,1151]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B3C0836972FB8233178FB99" box="[588,791,1128,1151]" italics="true" pageId="8" pageNumber="37">Cambarus clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B3C08369646B82C31C7FB98" box="[805,936,1127,1150]" pageId="8" pageNumber="37" rank="species">new species</taxonomicNameLabel>
, in central Tennessee, USA. Closed circles indicate sites at which 
<taxonomicName id="4C2A4D724B3C08369725B8C630D4FB42" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[582,699,1165,1188]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3C08369725B8C63037FB42" box="[582,600,1165,1188]" italics="true" pageId="8" pageNumber="37">C</emphasis>
. 
<emphasis id="B95EEAE34B3C08369706B8C630D4FB42" box="[613,699,1165,1188]" italics="true" pageId="8" pageNumber="37">clivosus</emphasis>
</taxonomicName>
was found; open circles indicate sites at which 
<taxonomicName id="4C2A4D724B3C083691D5B8C63745FB42" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[1206,1322,1165,1188]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3C083691D5B8C636A7FB42" box="[1206,1224,1165,1188]" italics="true" pageId="8" pageNumber="37">C</emphasis>
. 
<emphasis id="B95EEAE34B3C083691B7B8C63745FB42" box="[1236,1322,1165,1188]" italics="true" pageId="8" pageNumber="37">clivosus</emphasis>
</taxonomicName>
was absent.
</paragraph>
</caption>
<paragraph id="8B9536F14B3C08379448B9493677FE60" blockId="8.[264,1324,1282,1868]" lastBlockId="9.[264,1325,284,1870]" lastPageId="9" lastPageNumber="38" pageId="8" pageNumber="37">
A total of 
<specimenCount id="9D2CFD784B3C083694C2B9493050FAFA" box="[417,575,1282,1308]" pageId="8" pageNumber="37" type="generic">68 specimens</specimenCount>
has been examined from five locations in the Caney Fork River drainage (sites 1, 3–6) and one site in the East Fork Stones River drainage (Site 2) in Tennessee. Specific locations are as follows: 
<emphasis id="B95EEAE34B3C08369640B91931C5FA8D" box="[803,938,1362,1387]" italics="true" pageId="8" pageNumber="37">Cannon Co</emphasis>
.: 1) seep­fed tributary to 
<collectingCountry id="F33D76614B3C083691B4B9193744FA8A" box="[1239,1323,1362,1388]" name="Turkey" pageId="8" pageNumber="37">Turkey</collectingCountry>
Branch approx. 
<quantity id="4CD29B144B3C083694B4B931305DFA72" box="[471,562,1402,1428]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="37" unit="km" value="4.8">4.8 km</quantity>
NE Pleasant Ridge, 
<geoCoordinate id="EE1E50364B3C0836962EB93131BFFA72" box="[845,976,1402,1428]" direction="north" orientation="latitude" pageId="8" pageNumber="37" precision="5" value="35.8929">35.8929°N</geoCoordinate>
, ­ 
<geoCoordinate id="EE1E50364B3C0836968EB931361AFA72" box="[1005,1141,1402,1428]" direction="west" orientation="longitude" pageId="8" pageNumber="37" precision="5" value="-86.0167">86.0167°W</geoCoordinate>
NAD 83, 11 December 2004 (INHS 10234­ 1 F, 
<specimenCount id="9D2CFD784B3C083697A3B9E9316AFA5A" box="[704,773,1442,1468]" pageId="8" pageNumber="37" type="generic">3 juv.</specimenCount>
F); 2) seep­fed tributary to East Fork Stones River approx. 
<quantity id="4CD29B144B3C083694D0B9813069FA02" box="[435,518,1482,1508]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="37" unit="km" value="1.6">1.6 km</quantity>
SW town of Short Mountain, 
<geoCoordinate id="EE1E50364B3C08369612B981319BFA02" box="[881,1012,1482,1508]" direction="north" orientation="latitude" pageId="8" pageNumber="37" precision="5" value="35.8473">35.8473°N</geoCoordinate>
, ­ 
<geoCoordinate id="EE1E50364B3C0836916AB98136FEFA02" box="[1033,1169,1482,1508]" direction="west" orientation="longitude" pageId="8" pageNumber="37" precision="5" value="-85.9534">85.9534°W</geoCoordinate>
NAD 83, 28 September 2005 (INHS 10232 ­ morphotype), (USNM 1083161 – 
<typeStatus id="549188534B3C0836917FB9B936E3F9EA" box="[1052,1164,1522,1548]" pageId="8" pageNumber="37" type="paratype">paratypes</typeStatus>
1 M1, 1 M2, 1 F), (INHS 10233 – 3 MI, 3 MII, 7 F, 
<specimenCount id="9D2CFD784B3C083697B7BA51317AF9D2" box="[724,789,1562,1588]" pageId="8" pageNumber="37" type="generic">1 juv.</specimenCount>
M, 
<specimenCount id="9D2CFD784B3C08369626BA5131E8F9D2" box="[837,903,1562,1588]" pageId="8" pageNumber="37" type="generic">2 juv.</specimenCount>
F); 3) seep­fed tributary to Connell Creek at end of Tate Road, 
<quantity id="4CD29B144B3C0836972EBA0930CFF9BA" box="[589,672,1602,1628]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="37" unit="km" value="4.8">4.8 km</quantity>
NE Pleasant Ridge, 
<geoCoordinate id="EE1E50364B3C083696F5BA093676F9BA" box="[918,1049,1602,1628]" direction="north" orientation="latitude" pageId="8" pageNumber="37" precision="5" value="35.8941">35.8941°N</geoCoordinate>
, ­ 
<geoCoordinate id="EE1E50364B3C0836914DBA0936D9F9BA" box="[1070,1206,1602,1628]" direction="west" orientation="longitude" pageId="8" pageNumber="37" precision="5" value="-86.0215">86.0215°W</geoCoordinate>
NAD 83, 
<materialsCitation id="3B423CAC4B3C0836946BBA2131C7F962" ID-GBIF-Occurrence="2236573275" box="[264,936,1642,1668]" collectingDate="2005-03-06" collectionCode="INHS" pageId="8" pageNumber="37" specimenCode="INHS 10226 -2" specimenCount="3">
<collectingDate id="EFD0E9D94B3C0836946BBA2133DEF962" box="[264,433,1642,1668]" pageId="8" pageNumber="37" value="2005-03-06">
<date id="FF9410314B3C0836946BBA2133DEF962" box="[264,433,1642,1668]" pageId="8" pageNumber="37" value="2005-03-06">6 March 2005</date>
</collectingDate>
(
<specimenCode id="DB8C9E8A4B3C083694A1BA2130EEF962" box="[450,641,1642,1668]" collectionCode="INHS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="8" pageNumber="37" type="Other type of research institution/biorepository">INHS 10226 –2</specimenCode>
MII, 
<specimenCount id="9D2CFD784B3C083697AABA213163F962" box="[713,780,1642,1668]" pageId="8" pageNumber="37" type="generic">1 juv.</specimenCount>
M, 
<specimenCount id="9D2CFD784B3C08369623BA2131ECF962" box="[832,899,1642,1668]" pageId="8" pageNumber="37" type="generic">2 juv.</specimenCount>
F)
</materialsCitation>
, 
<materialsCitation id="3B423CAC4B3C083696D4BA213744F962" ID-GBIF-Occurrence="2236573270" box="[951,1323,1642,1668]" collectingDate="2005-04-07" collectionCode="INHS" pageId="8" pageNumber="37" specimenCode="INHS 10224 - 5" specimenCount="1">
<collectingDate id="EFD0E9D94B3C083696D4BA21363EF962" box="[951,1105,1642,1668]" pageId="8" pageNumber="37" value="2005-04-07">
<date id="FF9410314B3C083696D4BA21363EF962" box="[951,1105,1642,1668]" pageId="8" pageNumber="37" value="2005-04-07">7 April 2005</date>
</collectingDate>
(
<specimenCode id="DB8C9E8A4B3C08369101BA213744F962" box="[1122,1323,1642,1668]" collectionCode="INHS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="8" pageNumber="37" type="Other type of research institution/biorepository">INHS 10224 – 5</specimenCode>
</materialsCitation>
MII, 8 F, 
<specimenCount id="9D2CFD784B3C083694E0BAD933A6F94A" box="[387,457,1682,1708]" pageId="8" pageNumber="37" type="generic">2 juv.</specimenCount>
M, 
<specimenCount id="9D2CFD784B3C08369762BAD93028F94A" box="[513,583,1682,1708]" pageId="8" pageNumber="37" type="generic">2 juv.</specimenCount>
F), 
<date id="FF9410314B3C0836971CBAD9312CF94A" box="[639,835,1682,1708]" pageId="8" pageNumber="37" value="2005-08-20">20 August 2005</date>
(INHS 10227 –3 MI, 3 F); 4) seep­fed tributary to Wilmouth Creek on N side of Short Mountain, 
<geoCoordinate id="EE1E50364B3C083696A0BAF13629F932" box="[963,1094,1722,1748]" direction="north" orientation="latitude" pageId="8" pageNumber="37" precision="5" value="35.875">35.8750°N</geoCoordinate>
, ­ 
<geoCoordinate id="EE1E50364B3C08369139BAF1368DF932" box="[1114,1250,1722,1748]" direction="west" orientation="longitude" pageId="8" pageNumber="37" precision="5" value="-85.9741">85.9741°W</geoCoordinate>
NAD 83, 
<materialsCitation id="3B423CAC4B3C08369457BAA93745F91A" ID-GBIF-Occurrence="2236573269" box="[308,1322,1762,1788]" collectingDate="2005-06-12" collectionCode="INHS" county="DeKalb Co." location="Dry Creek" pageId="8" pageNumber="37" specimenCode="INHS 10229 - 2" specimenCount="1">
<collectingDate id="EFD0E9D94B3C08369457BAA933BFF91A" box="[308,464,1762,1788]" pageId="8" pageNumber="37" value="2005-06-12">
<date id="FF9410314B3C08369457BAA933BFF91A" box="[308,464,1762,1788]" pageId="8" pageNumber="37" value="2005-06-12">12 June 2005</date>
</collectingDate>
(
<specimenCode id="DB8C9E8A4B3C083694BDBAA930CFF91A" box="[478,672,1762,1788]" collectionCode="INHS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="8" pageNumber="37" type="Other type of research institution/biorepository">INHS 10229 – 2</specimenCode>
juv. F); 
<collectingCounty id="62F44E7D4B3C08369661BAA931E1F91A" box="[770,910,1762,1788]" pageId="8" pageNumber="37">
<emphasis id="B95EEAE34B3C08369661BAA931E9F91D" box="[770,902,1762,1787]" italics="true" pageId="8" pageNumber="37">DeKalb Co</emphasis>
.
</collectingCounty>
: 5) seep­fed tributary to 
<location id="8EF5602A4B3C083691D3BAA93745F91A" LSID="urn:lsid:plazi:treatment:038387E74B360832946BBD5731E4FCF5:8EF5602A4B3C083691D3BAA93745F91A" box="[1200,1322,1762,1788]" county="DeKalb Co." name="Dry Creek" pageId="8" pageNumber="37">Dry Creek</location>
</materialsCitation>
approx. 4.0 km W Snows Hill, 
<geoCoordinate id="EE1E50364B3C083697EABB413163F8C2" box="[649,780,1802,1828]" direction="north" orientation="latitude" pageId="8" pageNumber="37" precision="5" value="35.9848">35.9848°N</geoCoordinate>
, ­ 
<geoCoordinate id="EE1E50364B3C08369640BB4131C4F8C2" box="[803,939,1802,1828]" direction="west" orientation="longitude" pageId="8" pageNumber="37" precision="5" value="-85.9126">85.9126°W</geoCoordinate>
NAD 83, 
<date id="FF9410314B3C0836914CBB4136BCF8C2" box="[1071,1235,1802,1828]" pageId="8" pageNumber="37" value="2005-06-12">12 June 2005</date>
(INHS 10230 – 1 MII, 1 F, 
<specimenCount id="9D2CFD784B3C0836976CBB79303AF8AA" box="[527,597,1842,1868]" pageId="8" pageNumber="37" type="generic">1 juv.</specimenCount>
<materialsCitation id="3B423CAC4B3C08379703BB79300EFE60" ID-GBIF-Occurrence="2236573271" collectingDate="2005-09-27" collectionCode="INHS" lastPageId="9" lastPageNumber="38" latitude="35.9205" location="Dry Creek" longLatPrecision="6" longitude="-85.9271" municipality="Pea Ridge Wildlife Management Area" pageId="8" pageNumber="37" specimenCode="INHS 10235, INHS 10236, INHS 10237" specimenCount="3" typeStatus="holotype">
M); 6) Type locality, tributary of 
<location id="8EF5602A4B3C08369163BB7936EFF8AA" LSID="urn:lsid:plazi:treatment:038387E74B360832946BBD5731E4FCF5:8EF5602A4B3C08369163BB7936EFF8AA" box="[1024,1152,1842,1868]" latitude="35.9205" longLatPrecision="6" longitude="-85.9271" municipality="Pea Ridge Wildlife Management Area" name="Dry Creek" pageId="8" pageNumber="37">Dry Creek</location>
at 
<collectingMunicipality id="6BF1AC8B4B3C083791CFBB79303EFED0" lastPageId="9" lastPageNumber="38" pageId="8" pageNumber="37">Pea Ridge Wildlife Management Area</collectingMunicipality>
, approx. 
<locationDeviation id="9A272FD34B3D083797A2BD573194FED0" box="[705,1019,284,310]" location="Smithville" pageId="9" pageNumber="38">
<quantity id="4CD29B144B3D083797A2BD57314DFED0" box="[705,802,284,310]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="9" pageNumber="38" unit="km" value="11.5">11.5 km</quantity>
WSW Smithville
</locationDeviation>
, 
<geoCoordinate id="EE1E50364B3D0837916ABD5736E3FED0" box="[1033,1164,284,310]" direction="north" orientation="latitude" pageId="9" pageNumber="38" precision="5" value="35.9205">35.9205°N</geoCoordinate>
, ­ 
<geoCoordinate id="EE1E50364B3D083791C1BD573745FED0" box="[1186,1322,284,310]" direction="west" orientation="longitude" pageId="9" pageNumber="38" precision="5" value="-85.9271">85.9271°W</geoCoordinate>
NAD 83, 
<collectingDate id="EFD0E9D94B3D0837941FBD0F300DFEB8" box="[380,610,324,350]" pageId="9" pageNumber="38" value="2005-09-27">
<date id="FF9410314B3D0837941FBD0F300DFEB8" box="[380,610,324,350]" pageId="9" pageNumber="38" value="2005-09-27">27 September 2005</date>
</collectingDate>
(
<specimenCode id="DB8C9E8A4B3D08379712BD0F3167FEB8" box="[625,776,324,350]" collectionCode="INHS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="9" pageNumber="38" type="Other type of research institution/biorepository">INHS 10235</specimenCode>
– 
<typeStatus id="549188534B3D08379644BD0F31FDFEB8" box="[807,914,324,350]" pageId="9" pageNumber="38" type="holotype">holotype</typeStatus>
), (
<specimenCode id="DB8C9E8A4B3D083796CCBD0F3629FEB8" box="[943,1094,324,350]" collectionCode="INHS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="9" pageNumber="38" type="Other type of research institution/biorepository">INHS 10236</specimenCode>
– 
<typeStatus id="549188534B3D08379106BD0F36A7FEB8" box="[1125,1224,324,350]" pageId="9" pageNumber="38" type="allotype">allotype</typeStatus>
), (
<specimenCode id="DB8C9E8A4B3D08379186BD0F333EFE60" collectionCode="INHS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="9" pageNumber="38" type="Other type of research institution/biorepository">INHS 10237</specimenCode>
– 
<typeStatus id="549188534B3D0837940DBD2733B1FE60" box="[366,478,364,390]" pageId="9" pageNumber="38" type="paratype">paratypes</typeStatus>
1 MI, 1 F)
</materialsCitation>
, 
<materialsCitation id="3B423CAC4B3D0837970EBD27367BFE60" ID-GBIF-Occurrence="2236573272" box="[621,1044,364,390]" collectionCode="INHS" pageId="9" pageNumber="38" specimenCode="INHS 10238 - 1" specimenCount="3" typeStatus="holotype">
(
<specimenCode id="DB8C9E8A4B3D08379716BD273157FE60" box="[629,824,364,390]" collectionCode="INHS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="9" pageNumber="38" type="Other type of research institution/biorepository">INHS 10238 – 1</specimenCode>
MI, 2 F, 
<specimenCount id="9D2CFD784B3D083796C7BD273189FE60" box="[932,998,364,390]" pageId="9" pageNumber="38" type="generic">3 juv.</specimenCount>
M)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330657A4B3D0837945BBDDF311FFB28" pageId="9" pageNumber="38" type="etymology">
<paragraph id="8B9536F14B3D0837945BBDDF30D6FE30" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="38">
<emphasis id="B95EEAE34B3D0837945BBDDF33DCFE4B" box="[312,435,404,429]" italics="true" pageId="9" pageNumber="38">Etymology</emphasis>
.— 
<taxonomicName id="4C2A4D724B3D083794BBBDDF3059FE4B" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[472,566,404,429]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3D083794BBBDDF3059FE4B" box="[472,566,404,429]" italics="true" pageId="9" pageNumber="38">clivosus</emphasis>
</taxonomicName>
L. = hilly, steep; in reference to the high gradient, hillside habitat in which the species is found.
</paragraph>
<paragraph id="8B9536F14B3D0837945BBDAF311FFB28" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="38">
<emphasis id="B95EEAE34B3D0837945BBDAF30CDFE1B" box="[312,674,484,509]" italics="true" pageId="9" pageNumber="38">Habitat and life­history notes</emphasis>
.— 
<taxonomicName id="4C2A4D724B3D083797A4BDAF31C3FE1B" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[711,940,484,509]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B3D083797A4BDAF31C3FE1B" box="[711,940,484,509]" italics="true" pageId="9" pageNumber="38">Cambarus clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B3D083796A3BDAF3639FE18" box="[960,1110,484,510]" pageId="9" pageNumber="38" rank="species">new species</taxonomicNameLabel>
, occurs in small, high­gradient, seep­fed creeks with substrates of chert gravel, limestones, and shales (
<figureCitation id="13112A744B3D0837919BBE473379FDA8" captionStart="FIGURE 1" captionStartId="1.[264,368,1325,1349]" captionTargetBox="[449,1140,846,1305]" captionTargetId="figure@1.[449,1140,846,1305]" captionTargetPageId="1" captionText="FIGURE 1. Typical habitat for Cambarus clivosus, new species—a high gradient, seep­fed creek in Cannon Co., TN." httpUri="https://zenodo.org/record/2646442/files/figure.png" pageId="9" pageNumber="38">Fig. 1</figureCitation>
). Creek widths at sites containing the species ranged from 
<quantity id="4CD29B144B3D08379694BE7F36C7FDA8" box="[1015,1192,564,590]" metricMagnitude="-1" metricUnit="m" metricValue="4.25" metricValueMax="6.0" metricValueMin="2.5" pageId="9" pageNumber="38" unit="m" value="0.425" valueMax="0.6" valueMin="0.25">0.25 to 0.6 m</quantity>
and water depths were less than 5.0 cm. At several sites, water alternated between surface and subsurface flow regimes in given reaches. In creeks harboring the species, individuals were encountered under flat pieces of gravel or shale or in burrows. The species was found in two 
<typeStatus id="549188534B3D083794CBBE9F3389FD08" box="[424,486,724,750]" pageId="9" pageNumber="38">types</typeStatus>
of burrows, either shallow (&gt; 
<quantity id="4CD29B144B3D08379632BE9F31E2FD08" box="[849,909,724,750]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="38" unit="cm" value="5.0">5 cm</quantity>
) burrows under mid­channel shale or gravel or in deep (
<quantity id="4CD29B144B3D08379771BEB730C8FCF0" box="[530,679,764,790]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="9" pageNumber="38" unit="m" value="0.3" valueMax="0.5" valueMin="0.1">0.1 – 0.5 m</quantity>
), winding burrows dug into creek banks. The latter burrow 
<typeStatus id="549188534B3D08379406BF6F33F8FCD8" box="[357,407,804,830]" pageId="9" pageNumber="38">type</typeStatus>
usually migrated horizontally more than vertically amongst subsurface rocks and boulders. Given the small number of collections available, determining definitive lifehistory aspects is difficult. The species has been collected in the months of March, April, June, August, September, and December. Of those collections, form I males were found in August and September. No ovigerous females have been collected or observed. The presence of juveniles with carapace lengths of 
<quantity id="4CD29B144B3D08379649BFA731CEFBE0" box="[810,929,1004,1030]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="9" pageNumber="38" unit="mm" value="7.0" valueMax="9.0" valueMin="5.0">5 to 9 mm</quantity>
in September collections suggest that young 
<taxonomicName id="4C2A4D724B3D083794F0B85F3078FBCB" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[403,535,1044,1070]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3D083794F0B85F33C9FBCB" box="[403,422,1044,1069]" italics="true" pageId="9" pageNumber="38">C</emphasis>
. 
<emphasis id="B95EEAE34B3D083794DBB85F3078FBCB" box="[440,535,1044,1069]" italics="true" pageId="9" pageNumber="38">clivosus</emphasis>
</taxonomicName>
hatch in summer months. Form I males collected in September were usually found under rocks in mid­channel rather than in stream bank burrows while in spring months most males were found in burrows. Fall breeding is not uncommon in burrowing crayfish species and the observations may be a mate­seeking behavior by males attempting to locate females sequestered in burrows.
</paragraph>
</subSubSection>
<subSubSection id="C330657A4B3D0832945BB89731E4FCF5" lastPageId="12" lastPageNumber="41" pageId="9" pageNumber="38" type="description">
<paragraph id="8B9536F14B3D0837945BB89733AAFA58" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="38">
<emphasis id="B95EEAE34B3D0837945BB8973070FB13" box="[312,543,1244,1269]" italics="true" pageId="9" pageNumber="38">Crayfish associates</emphasis>
.—The following species were collected from the same seep­fed creeks containing 
<taxonomicName id="4C2A4D724B3D08379483B94F300DFAFB" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[480,610,1284,1310]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B3D08379483B94F339CFAFB" box="[480,499,1284,1309]" italics="true" pageId="9" pageNumber="38">C</emphasis>
. 
<emphasis id="B95EEAE34B3D08379767B94F300DFAFB" box="[516,610,1284,1309]" italics="true" pageId="9" pageNumber="38">clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B3D08379710B94F316AFAF8" box="[627,773,1284,1310]" pageId="9" pageNumber="38" rank="species">new species</taxonomicNameLabel>
: 
<taxonomicName id="4C2A4D724B3D08379678B94F332CFAA0" authority="Hay, 1902" authorityName="Hay" authorityYear="1902" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="tenebrosus" subGenus="Erebicambarus">
<emphasis id="B95EEAE34B3D08379678B94F31FAFAFB" box="[795,917,1284,1309]" italics="true" pageId="9" pageNumber="38">Cambarus</emphasis>
(
<emphasis id="B95EEAE34B3D083796C4B94F3634FAFB" box="[935,1115,1284,1309]" italics="true" pageId="9" pageNumber="38">Erebicambarus</emphasis>
) 
<emphasis id="B95EEAE34B3D0837910EB94F3684FAFB" box="[1133,1259,1284,1309]" italics="true" pageId="9" pageNumber="38">tenebrosus</emphasis>
<bibRefCitation id="EFBB4B004B3D08379197B94F332CFAA0" author="Hay, W. P." pageId="9" pageNumber="38" pagination="223 - 236" refId="ref5124" refString="Hay, W. P. (1902) Observations on the crustacean fauna of the region about Mammoth Cave, Kentucky. Proceedings of the United States National Museum, 25, 223 - 236." type="journal article" year="1902">Hay, 1902</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C2A4D724B3D083794E0B96731D1FAA0" authority="Hobbs, 1953" authorityName="Hobbs" authorityYear="1953" box="[387,958,1324,1350]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="friaufi" subGenus="Jugicambarus">
<emphasis id="B95EEAE34B3D083794E0B9673392FAA3" box="[387,509,1324,1349]" italics="true" pageId="9" pageNumber="38">Cambarus</emphasis>
(
<emphasis id="B95EEAE34B3D08379771B96730D7FAA3" box="[530,696,1324,1349]" italics="true" pageId="9" pageNumber="38">Jugicambarus</emphasis>
) 
<emphasis id="B95EEAE34B3D083797AEB9673178FAA3" box="[717,791,1324,1349]" italics="true" pageId="9" pageNumber="38">friaufi</emphasis>
<bibRefCitation id="EFBB4B004B3D08379642B96731D1FAA0" author="Hobbs, H. H. Jr." box="[801,958,1324,1350]" pageId="9" pageNumber="38" pagination="20 - 27" refId="ref5161" refString="Hobbs, H. H. Jr. (1953) Two new crayfishes from the Highland Rim in Tennessee (Decapoda, Astacidae). Journal of the Tennessee Academy of Science, 28, 20 - 27." type="journal article" year="1953">Hobbs, 1953</bibRefCitation>
</taxonomicName>
. At one site, small 
<taxonomicName id="4C2A4D724B3D083791D8B9673745FAA3" authorityName="Hobbs" authorityYear="1953" box="[1211,1322,1324,1350]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="friaufi">
<emphasis id="B95EEAE34B3D083791D8B96736A1FAA3" box="[1211,1230,1324,1349]" italics="true" pageId="9" pageNumber="38">C</emphasis>
. 
<emphasis id="B95EEAE34B3D08379182B9673745FAA3" box="[1249,1322,1324,1349]" italics="true" pageId="9" pageNumber="38">friaufi</emphasis>
</taxonomicName>
were found in the same burrows as 
<taxonomicName id="4C2A4D724B3D083797C5B91F3149FA8B" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[678,806,1364,1390]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3D083797C5B91F30D6FA8B" box="[678,697,1364,1389]" italics="true" pageId="9" pageNumber="38">C</emphasis>
. 
<emphasis id="B95EEAE34B3D083797ABB91F3149FA8B" box="[712,806,1364,1389]" italics="true" pageId="9" pageNumber="38">clivosus</emphasis>
</taxonomicName>
. In those cases young 
<taxonomicName id="4C2A4D724B3D08379155B91F36CEFA8B" authorityName="Hobbs" authorityYear="1953" box="[1078,1185,1364,1390]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="friaufi">
<emphasis id="B95EEAE34B3D08379155B91F3626FA8B" box="[1078,1097,1364,1389]" italics="true" pageId="9" pageNumber="38">C</emphasis>
. 
<emphasis id="B95EEAE34B3D0837913BB91F36CEFA8B" box="[1112,1185,1364,1389]" italics="true" pageId="9" pageNumber="38">friaufi</emphasis>
</taxonomicName>
individuals were found within the first 
<quantity id="4CD29B144B3D08379732B93730F4FA70" box="[593,667,1404,1430]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="38" unit="cm" value="10.0">10 cm</quantity>
of the burrow while the 
<taxonomicName id="4C2A4D724B3D083796A8B9373622FA73" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[971,1101,1404,1430]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3D083796A8B93731B1FA73" box="[971,990,1404,1429]" italics="true" pageId="9" pageNumber="38">C</emphasis>
. 
<emphasis id="B95EEAE34B3D0837968CB9373622FA73" box="[1007,1101,1404,1429]" italics="true" pageId="9" pageNumber="38">clivosus</emphasis>
</taxonomicName>
were found in the deeper portions.
</paragraph>
<paragraph id="8B9536F14B3D08349459B9873323FE60" blockId="9.[264,1325,284,1870]" lastBlockId="10.[264,1324,284,1870]" lastPageId="10" lastPageNumber="39" pageId="9" pageNumber="38">
<emphasis id="B95EEAE34B3D08379459B98733CBFA03" box="[314,420,1484,1509]" italics="true" pageId="9" pageNumber="38">Variation</emphasis>
.—Several ontogenetic variations are observed in 
<taxonomicName id="4C2A4D724B3D0837969CB98736EDFA03" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[1023,1154,1484,1510]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B3D0837969CB987367DFA03" box="[1023,1042,1484,1509]" italics="true" pageId="9" pageNumber="38">C</emphasis>
. 
<emphasis id="B95EEAE34B3D08379147B98736EDFA03" box="[1060,1154,1484,1509]" italics="true" pageId="9" pageNumber="38">clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B3D083791F7B9873748FA00" box="[1172,1319,1484,1510]" pageId="9" pageNumber="38" rank="species">new species</taxonomicNameLabel>
, one of which shows a geographic pattern of distribution. Specimens collected from the East Fork Stones River drainage have a slightly narrower areola, averaging 9.6 times as long as wide compared to areolae of individuals from the Caney Fork drainage which average 8.1 times as long as wide. Other random ontogenetic variations include the degree of excavation of the rostrum, the shape of the antennal scale, and the tubercle pattern on the mesial margin of the palm of the chela. In most individuals the rostrum is moderately excavated while a few specimens had more deeply excavated rostrums. The antennal scale shape is variable in respect to the distal edge. In some individuals that edge is nearly perpendicular to the lateral margin (Fig. 2F) while in others it is angled proximally (Fig. 2E). Finally, the number of palmar tubercles along the mesial margin is variable, ranging from five to eight. In many cases, the proximal­most two tubercles are fused to form a ridge.
</paragraph>
<paragraph id="8B9536F14B3E0834944EBDDF3010FB40" blockId="10.[264,1324,284,1870]" pageId="10" pageNumber="39">
<emphasis id="B95EEAE34B3E0834944EBDDF33A6FE4B" box="[301,457,404,429]" italics="true" pageId="10" pageNumber="39">Comparisons</emphasis>
.— 
<taxonomicName id="4C2A4D724B3E0834948DBDDF30A2FE4B" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[494,717,404,429]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B3E0834948DBDDF30A2FE4B" box="[494,717,404,429]" italics="true" pageId="10" pageNumber="39">Cambarus clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B3E083497BEBDDF3102FE48" box="[733,877,404,430]" pageId="10" pageNumber="39" rank="species">new species</taxonomicNameLabel>
, differs from all other members of the subgenus 
<taxonomicName id="4C2A4D724B3E083494E0BDF73046FE33" box="[387,553,444,469]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Jugicambarus">
<emphasis id="B95EEAE34B3E083494E0BDF73046FE33" box="[387,553,444,469]" italics="true" pageId="10" pageNumber="39">Jugicambarus</emphasis>
</taxonomicName>
in possessing a unique combination of form I male pleopod, carapace, and chelae characters. Other pigmented members of the subgenus that possess a single row of tubercles along the mesial margin of the palm, a wide areola and nonlaterally compressed carapace, and lack long setae on the dorsal and lateral surfaces of the palm and fingers of the chelae include 
<taxonomicName id="4C2A4D724B3E083497AFBE173678FD90" authority="Faxon 1914" authorityName="Faxon" authorityYear="1914" box="[716,1047,604,630]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="asperimanus">
<emphasis id="B95EEAE34B3E083497AFBE1730B0FD93" box="[716,735,604,629]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E0834978EBE1731ECFD93" box="[749,899,604,629]" italics="true" pageId="10" pageNumber="39">asperimanus</emphasis>
<bibRefCitation id="EFBB4B004B3E083496E9BE173678FD90" author="Faxon, W." box="[906,1047,604,630]" pageId="10" pageNumber="39" pagination="351 - 427" refId="ref5064" refString="Faxon, W. (1914) Notes on the crayfishes in the United States National Museum and the Museum of Comparative Zoology with descriptions of new species and subspecies to which is appended a catalogue of the known species and subspecies. Memoirs of the Museum of Comparative Zoology at Harvard College, 40, 351 - 427." type="journal article" year="1914">Faxon 1914</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C2A4D724B3E08349140BE173320FD78" authority="(Erichson 1846)" authorityName="Erichson" authorityYear="1846" baseAuthorityName="Erichson" baseAuthorityYear="1846" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="carolinus">
<emphasis id="B95EEAE34B3E08349140BE173659FD93" box="[1059,1078,604,629]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E08349127BE1736DDFD93" box="[1092,1202,604,629]" italics="true" pageId="10" pageNumber="39">carolinus</emphasis>
(
<bibRefCitation id="EFBB4B004B3E083491A3BE173329FD78" author="Erichson, W. F." pageId="10" pageNumber="39" pagination="86 - 103" refId="ref5005" refString="Erichson, W. F. (1846) Uebersicht der arten der gattung Astacus. Archir fur Naturgeschichte, 12, 86 - 103." type="journal article" year="1846">Erichson 1846</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C2A4D724B3E0834943FBECF3134FD78" authority="Hobbs and Hobbs 1962" authorityName="Hobbs and Hobbs" authorityYear="1962" box="[348,859,644,670]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="conasaugaensis">
<emphasis id="B95EEAE34B3E0834943FBECF3300FD7B" box="[348,367,644,669]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E0834941CBECF3056FD7B" box="[383,569,644,669]" italics="true" pageId="10" pageNumber="39">conasaugaensis</emphasis>
<bibRefCitation id="EFBB4B004B3E08349722BECF3134FD78" author="Hobbs, H. H. Jr. &amp; Hobbs, H. H. III." box="[577,859,644,670]" pageId="10" pageNumber="39" pagination="41 - 45" refId="ref5350" refString="Hobbs, H. H. Jr. &amp; Hobbs, H. H. III. (1962) A new crayfish of the genus Cambarus from Georgia (Decapoda, Astacidae). Proceedings of the Biological Society of Washington, 75, 41 - 45." type="journal article" year="1962">Hobbs and Hobbs 1962</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C2A4D724B3E08349604BECF36C7FD78" authority="Bouchard 1973" authorityName="Bouchard" authorityYear="1973" box="[871,1192,644,670]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="crinipes">
<emphasis id="B95EEAE34B3E08349604BECF3115FD7B" box="[871,890,644,669]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E083496E9BECF3186FD7B" box="[906,1001,644,669]" italics="true" pageId="10" pageNumber="39">crinipes</emphasis>
<bibRefCitation id="EFBB4B004B3E08349693BECF36C7FD78" author="Bouchard, R. W." box="[1008,1192,644,670]" pageId="10" pageNumber="39" pagination="103 - 111" refId="ref4908" refString="Bouchard, R. W. (1973) A new crayfish of the subgenus Jugicambarus from Tennessee with an emended definition of the subgenus (Astacidae, Decapoda). The American Midland Naturalist, 89, 103 - 111." type="journal article" year="1973">Bouchard 1973</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C2A4D724B3E083491D7BECF33D9FD20" authority="Rhoades 1944" authorityName="Rhoades" authorityYear="1944" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="distans">
<emphasis id="B95EEAE34B3E083491D7BECF36A8FD7B" box="[1204,1223,644,669]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E083491B4BECF3745FD7B" box="[1239,1322,644,669]" italics="true" pageId="10" pageNumber="39">distans</emphasis>
<bibRefCitation id="EFBB4B004B3E0834946BBEE733D9FD20" author="Rhoades, R." box="[264,438,684,710]" pageId="10" pageNumber="39" pagination="111 - 149" refId="ref5512" refString="Rhoades, R. (1944) The crayfishes of Kentucky, with notes on variation, distribution, and descriptions of new species and subspecies. American Midland Naturalist, 31, 111 - 149." type="journal article" year="1944">Rhoades 1944</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C2A4D724B3E083494A5BEE73166FD20" authority="Thoma 2000" authorityName="Thoma" authorityYear="2000" box="[454,777,684,710]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="jezerinaci">
<emphasis id="B95EEAE34B3E083494A5BEE733B6FD23" box="[454,473,684,709]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E0834948EBEE7300EFD23" box="[493,609,684,709]" italics="true" pageId="10" pageNumber="39">jezerinaci</emphasis>
<bibRefCitation id="EFBB4B004B3E0834970FBEE73166FD20" author="Thoma" box="[620,777,684,710]" pageId="10" pageNumber="39" pagination="731 - 738" refId="ref5585" refString="Thoma ,, R. F. (2000) Cambarus (Jugicambarus) jezerinaci (Crustacea: Decapoda: Cambaridae), a new species of crayfish from the Powell River drainage of Tennessee and Virginia. Proceedings of the Biological Society of Washington, 113, 731 - 738." type="journal article" year="2000">Thoma 2000</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C2A4D724B3E0834962CBEE73748FD20" authority="Hobbs and Shoup 1947" authorityName="Hobbs and Shoup" authorityYear="1947" box="[847,1319,684,710]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="parvoculus">
<emphasis id="B95EEAE34B3E0834962CBEE7310DFD23" box="[847,866,684,709]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E08349616BEE73198FD23" box="[885,1015,684,709]" italics="true" pageId="10" pageNumber="39">parvoculus</emphasis>
<bibRefCitation id="EFBB4B004B3E08349167BEE73748FD20" author="Hobbs, H. H. Jr. &amp; Shoup, C. S." box="[1028,1319,684,710]" pageId="10" pageNumber="39" pagination="138 - 145" refId="ref5399" refString="Hobbs, H. H. Jr. and Shoup, C. S. (1947) Two new crayfishes (Decapoda, Astacidae) from the Obey River drainage in Tennessee. Journal of the Tennessee Academy of Science, 22, 138 - 145." type="journal article" year="1947">Hobbs and Shoup 1947</bibRefCitation>
</taxonomicName>
. 
<taxonomicName id="4C2A4D724B3E0834946BBE9F3384FD0B" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[264,491,724,749]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3E0834946BBE9F3384FD0B" box="[264,491,724,749]" italics="true" pageId="10" pageNumber="39">Cambarus clivosus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C2A4D724B3E083497EEBE9F3127FD0B" authorityName="Faxon" authorityYear="1914" box="[653,840,724,750]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="asperimanus">
<emphasis id="B95EEAE34B3E083497EEBE9F30CFFD0B" box="[653,672,724,749]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E083497D1BE9F3127FD0B" box="[690,840,724,749]" italics="true" pageId="10" pageNumber="39">asperimanus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2A4D724B3E083496E5BE9F360AFD0B" authorityName="Hobbs and Hobbs" authorityYear="1962" box="[902,1125,724,750]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="conasaugaensis">
<emphasis id="B95EEAE34B3E083496E5BE9F31F6FD0B" box="[902,921,724,749]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E083496C8BE9F360AFD0B" box="[939,1125,724,749]" italics="true" pageId="10" pageNumber="39">conasaugaensis</emphasis>
</taxonomicName>
by possessing a suborbital angle of approximately 90 degrees (Fig. 2C), both of the latter species lack an angle. 
<taxonomicName id="4C2A4D724B3E0834940ABF6F3020FCDB" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[361,591,804,829]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3E0834940ABF6F3020FCDB" box="[361,591,804,829]" italics="true" pageId="10" pageNumber="39">Cambarus clivosus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C2A4D724B3E08349798BF6F31FEFCDB" baseAuthorityName="Erichson" baseAuthorityYear="1846" box="[763,913,804,830]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="carolinus">
<emphasis id="B95EEAE34B3E08349798BF6F3161FCDB" box="[763,782,804,829]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E08349640BF6F31FEFCDB" box="[803,913,804,829]" italics="true" pageId="10" pageNumber="39">carolinus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2A4D724B3E083496B6BF6F3634FCDB" authorityName="Bouchard" authorityYear="1973" box="[981,1115,804,830]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="crinipes">
<emphasis id="B95EEAE34B3E083496B6BF6F3187FCDB" box="[981,1000,804,829]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E0834969EBF6F3634FCDB" box="[1021,1115,804,829]" italics="true" pageId="10" pageNumber="39">crinipes</emphasis>
</taxonomicName>
by possessing a subapical notch on the central projection of the form I males (Fig. 2J) and differs from 
<taxonomicName id="4C2A4D724B3E08349073BF0733E5FC6B" authorityName="Hobbs and Shoup" authorityYear="1947" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="parvoculus">
<emphasis id="B95EEAE34B3E08349073BF07374CFC83" box="[1296,1315,844,869]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E0834946BBF3F33E5FC6B" box="[264,394,884,909]" italics="true" pageId="10" pageNumber="39">parvoculus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2A4D724B3E083494F9BF3F305EFC6B" authorityName="Thoma" authorityYear="2000" box="[410,561,884,910]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="jezerinaci">
<emphasis id="B95EEAE34B3E083494F9BF3F33C2FC6B" box="[410,429,884,909]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E083494DEBF3F305EFC6B" box="[445,561,884,909]" italics="true" pageId="10" pageNumber="39">jezerinaci</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C2A4D724B3E08349710BF3F3086FC6B" authorityName="Rhoades" authorityYear="1944" box="[627,745,884,910]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="distans">
<emphasis id="B95EEAE34B3E08349710BF3F30E9FC6B" box="[627,646,884,909]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E083497F5BF3F3086FC6B" box="[662,745,884,909]" italics="true" pageId="10" pageNumber="39">distans</emphasis>
</taxonomicName>
by possessing a triangular rostrum with margins that strongly converge (Fig. 2A). The rostral margins of 
<taxonomicName id="4C2A4D724B3E083496C4BFD73623FC53" authorityName="Hobbs and Shoup" authorityYear="1947" box="[935,1100,924,950]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="parvoculus">
<emphasis id="B95EEAE34B3E083496C4BFD731D5FC53" box="[935,954,924,949]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E083496A9BFD73623FC53" box="[970,1100,924,949]" italics="true" pageId="10" pageNumber="39">parvoculus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2A4D724B3E08349138BFD7369EFC53" authorityName="Thoma" authorityYear="2000" box="[1115,1265,924,950]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="jezerinaci">
<emphasis id="B95EEAE34B3E08349138BFD73601FC53" box="[1115,1134,924,949]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E0834911EBFD7369EFC53" box="[1149,1265,924,949]" italics="true" pageId="10" pageNumber="39">jezerinaci</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C2A4D724B3E0834946BBF8F3310FC3B" authorityName="Rhoades" authorityYear="1944" box="[264,383,964,990]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="distans">
<emphasis id="B95EEAE34B3E0834946BBF8F3374FC3B" box="[264,283,964,989]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E0834944FBF8F3310FC3B" box="[300,383,964,989]" italics="true" pageId="10" pageNumber="39">distans</emphasis>
</taxonomicName>
are sub­parallel and terminate at the base of a well­defined acumen (Fig. 2B). The triangular shape of the rostrum in 
<taxonomicName id="4C2A4D724B3E083497AABFA73126FBE3" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[713,841,1004,1030]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3E083497AABFA730B3FBE3" box="[713,732,1004,1029]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E08349788BFA73126FBE3" box="[747,841,1004,1029]" italics="true" pageId="10" pageNumber="39">clivosus</emphasis>
</taxonomicName>
also helps distinguish it from 
<taxonomicName id="4C2A4D724B3E083491C9BFA73745FBE3" authorityName="Bouchard" authorityYear="1973" box="[1194,1322,1004,1030]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="crinipes">
<emphasis id="B95EEAE34B3E083491C9BFA736D2FBE3" box="[1194,1213,1004,1029]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E083491AFBFA73745FBE3" box="[1228,1322,1004,1029]" italics="true" pageId="10" pageNumber="39">crinipes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2A4D724B3E0834945DB85F3394FBCB" authorityName="Faxon" authorityYear="1914" box="[318,507,1044,1070]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="asperimanus">
<emphasis id="B95EEAE34B3E0834945DB85F333EFBCB" box="[318,337,1044,1069]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E08349406B85F3394FBCB" box="[357,507,1044,1069]" italics="true" pageId="10" pageNumber="39">asperimanus</emphasis>
</taxonomicName>
. The rostral margins of both of the latter species are subparallel. 
<taxonomicName id="4C2A4D724B3E0834946BB8773380FBB3" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[264,495,1084,1109]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3E0834946BB8773380FBB3" box="[264,495,1084,1109]" italics="true" pageId="10" pageNumber="39">Cambarus clivosus</emphasis>
</taxonomicName>
differs from the newly described 
<emphasis id="B95EEAE34B3E083496C8B87731D1FBB3" box="[939,958,1084,1109]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<taxonomicName id="4C2A4D724B3E083496B7B8773317FB98" authority="Cooper and Schofield" authorityName="Cooper and Schofield" authorityYear="2002" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="tuckasegee" subGenus="Jugicambarus">
(
<emphasis id="B95EEAE34B3E083496BDB8773184FBB3" box="[990,1003,1084,1109]" italics="true" pageId="10" pageNumber="39">J</emphasis>
.) 
<emphasis id="B95EEAE34B3E08349168B87736E4FBB3" box="[1035,1163,1084,1109]" italics="true" pageId="10" pageNumber="39">tuckasegee</emphasis>
Cooper and Schofield
</taxonomicName>
in lacking the second row of two to four weak tubercles found along the mesial margin of the palm of the chela.
</paragraph>
<paragraph id="8B9536F14B3E0835945BB8FF31C9FEB8" blockId="10.[264,1324,284,1870]" lastBlockId="11.[264,1323,284,430]" lastPageId="11" lastPageNumber="40" pageId="10" pageNumber="39">
<emphasis id="B95EEAE34B3E0834945BB8FF33B8FB2B" box="[312,471,1204,1229]" italics="true" pageId="10" pageNumber="39">Relationships</emphasis>
.—Following Hobbs’ (1969) use of chelae morphology as the primary character for assessing evolutionary relationships within the genus 
<taxonomicName id="4C2A4D724B3E08349148B89736CAFB13" authorityName="Erichson" authorityYear="1846" box="[1067,1189,1244,1269]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAE34B3E08349148B89736CAFB13" box="[1067,1189,1244,1269]" italics="true" pageId="10" pageNumber="39">Cambarus</emphasis>
</taxonomicName>
, we assign 
<taxonomicName id="4C2A4D724B3E0834946BB94F33E1FAFB" authority="Taylor &amp; Soucek &amp; Organ, 2006" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[264,398,1284,1310]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="clivosus" status="sp. nov.">
<emphasis id="B95EEAE34B3E0834946BB94F3374FAFB" box="[264,283,1284,1309]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E08349453B94F33E1FAFB" box="[304,398,1284,1309]" italics="true" pageId="10" pageNumber="39">clivosus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57984B3E083494C7B94F3056FAF8" box="[420,569,1284,1310]" pageId="10" pageNumber="39" rank="species">new species</taxonomicNameLabel>
, to the subgenus 
<taxonomicName id="4C2A4D724B3E08349678B94F31AEFAFB" box="[795,961,1284,1309]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Jugicambarus">
<emphasis id="B95EEAE34B3E08349678B94F31AEFAFB" box="[795,961,1284,1309]" italics="true" pageId="10" pageNumber="39">Jugicambarus</emphasis>
</taxonomicName>
. Members of that subgenus possess subrectangular chelae with a cristiform or serrate row of turbercles along the mesial margins of the palm (
<bibRefCitation id="EFBB4B004B3E08349706B91F3093FA88" author="Hobbs, H. H. Jr." box="[613,764,1364,1390]" pageId="10" pageNumber="39" pagination="93 - 178" refId="ref5201" refString="Hobbs, H. H. Jr. (1969) On the distribution and phylogeny of the crayfish genus Cambarus. In Holt, P. C., Hoffman, R. L, &amp; Hart, C. W. Jr. (Eds.), The distributional history of the biota of the southern Appalachians, Part I: Invertebrates. Virginia Polytechnic Institute, Research Division Monograph, 1, 93 - 178." type="journal article" year="1969">Hobbs 1969</bibRefCitation>
; 
<bibRefCitation id="EFBB4B004B3E08349668B91F31A9FA88" author="Bouchard, R. W." box="[779,966,1364,1390]" pageId="10" pageNumber="39" pagination="103 - 111" refId="ref4908" refString="Bouchard, R. W. (1973) A new crayfish of the subgenus Jugicambarus from Tennessee with an emended definition of the subgenus (Astacidae, Decapoda). The American Midland Naturalist, 89, 103 - 111." type="journal article" year="1973">Bouchard 1973</bibRefCitation>
), both characters found in 
<taxonomicName id="4C2A4D724B3E08349073B91F3309FA73" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3E08349073B91F374CFA8B" box="[1296,1315,1364,1389]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E0834946BB9373309FA73" box="[264,358,1404,1429]" italics="true" pageId="10" pageNumber="39">clivosus</emphasis>
</taxonomicName>
. While their rostral shapes differ, 
<taxonomicName id="4C2A4D724B3E0834966AB9373185FA73" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[777,1002,1404,1429]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3E0834966AB9373185FA73" box="[777,1002,1404,1429]" italics="true" pageId="10" pageNumber="39">Cambarus clivosus</emphasis>
</taxonomicName>
shares a central projection with a subapical notch and a suborbital angle with 
<taxonomicName id="4C2A4D724B3E08349614B9EF3671FA5B" authorityName="Hobbs and Shoup" authorityYear="1947" box="[887,1054,1444,1470]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="parvoculus">
<emphasis id="B95EEAE34B3E08349614B9EF31E5FA5B" box="[887,906,1444,1469]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E083496FFB9EF3671FA5B" box="[924,1054,1444,1469]" italics="true" pageId="10" pageNumber="39">parvoculus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2A4D724B3E08349153B9EF36A6FA5B" authorityName="Thoma" authorityYear="2000" box="[1072,1225,1444,1470]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="jezerinaci">
<emphasis id="B95EEAE34B3E08349153B9EF362CFA5B" box="[1072,1091,1444,1469]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E08349136B9EF36A6FA5B" box="[1109,1225,1444,1469]" italics="true" pageId="10" pageNumber="39">jezerinaci</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C2A4D724B3E08349073B9EF3330FA03" authorityName="Rhoades" authorityYear="1944" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="distans">
<emphasis id="B95EEAE34B3E08349073B9EF374CFA5B" box="[1296,1315,1444,1469]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E0834946BB9873330FA03" box="[264,351,1484,1509]" italics="true" pageId="10" pageNumber="39">distans</emphasis>
</taxonomicName>
<emphasis id="B95EEAE34B3E0834943CB987330CFA03" box="[351,355,1484,1509]" italics="true" pageId="10" pageNumber="39">.</emphasis>
However, given that 
<taxonomicName id="4C2A4D724B3E0834971FB9873149FA03" authorityName="Hobbs and Shoup" authorityYear="1947" box="[636,806,1484,1510]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="parvoculus">
<emphasis id="B95EEAE34B3E0834971FB98730E0FA03" box="[636,655,1484,1509]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E083497C0B9873149FA03" box="[675,806,1484,1509]" italics="true" pageId="10" pageNumber="39">parvoculus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2A4D724B3E0834960BB987366EFA03" authorityName="Thoma" authorityYear="2000" box="[872,1025,1484,1510]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="jezerinaci">
<emphasis id="B95EEAE34B3E0834960BB9873114FA03" box="[872,891,1484,1509]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E083496EDB987366EFA03" box="[910,1025,1484,1509]" italics="true" pageId="10" pageNumber="39">jezerinaci</emphasis>
</taxonomicName>
both possess a strongly recurved central projection and occur in the same 
<typeStatus id="549188534B3E08349603B9BF31FCF9E8" box="[864,915,1524,1550]" pageId="10" pageNumber="39">type</typeStatus>
of habitat as 
<taxonomicName id="4C2A4D724B3E08349154B9BF36D6F9EB" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[1079,1209,1524,1550]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3E08349154B9BF3625F9EB" box="[1079,1098,1524,1549]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E08349138B9BF36D6F9EB" box="[1115,1209,1524,1549]" italics="true" pageId="10" pageNumber="39">clivosus</emphasis>
</taxonomicName>
, it seems logical to conclude that these three species most likely share a recent common ancestor. 
<taxonomicName id="4C2A4D724B3E0834946BBA0F307BF9BB" authorityName="Hobbs and Shoup" authorityYear="1947" box="[264,532,1604,1629]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="parvoculus">
<emphasis id="B95EEAE34B3E0834946BBA0F307BF9BB" box="[264,532,1604,1629]" italics="true" pageId="10" pageNumber="39">Cambarus parvoculus</emphasis>
</taxonomicName>
occurs in the Appalachian Uplands of eastern 
<collectingRegion id="49EEF8134B3E0834911DBA0F369FF9B8" box="[1150,1264,1604,1630]" country="United States of America" name="Kentucky" pageId="10" pageNumber="39">Kentucky</collectingRegion>
and 
<collectingRegion id="49EEF8134B3E0834946BBA2733EFF960" box="[264,384,1644,1670]" country="United States of America" name="Tennessee" pageId="10" pageNumber="39">Tennessee</collectingRegion>
and extreme northwestern Geogia (
<bibRefCitation id="EFBB4B004B3E08349636BA2736CCF960" author="Taylor, C. A. &amp; Schuster, G. A." box="[853,1187,1644,1670]" pageId="10" pageNumber="39" refId="ref5549" refString="Taylor, C. A. &amp; Schuster, G. A. (2004) The Crayfishes of Kentucky. Illinois Natural History Survey Special Publication No. 28, viii + 219 pp." type="book" year="2004">Taylor and Schuster 2004</bibRefCitation>
) while 
<taxonomicName id="4C2A4D724B3E08349073BA273313F94B" authorityName="Thoma" authorityYear="2000" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="jezerinaci">
<emphasis id="B95EEAE34B3E08349073BA27374CF963" box="[1296,1315,1644,1669]" italics="true" pageId="10" pageNumber="39">C</emphasis>
. 
<emphasis id="B95EEAE34B3E0834946BBADF3313F94B" box="[264,380,1684,1709]" italics="true" pageId="10" pageNumber="39">jezerinaci</emphasis>
</taxonomicName>
is currently known from a restricted portion of the Appalachian Uplands in the 
<collectingRegion id="49EEF8134B3E0834946BBAF73310F930" box="[264,383,1724,1750]" country="United States of America" name="Tennessee" pageId="10" pageNumber="39">Tennessee</collectingRegion>
/ 
<collectingRegion id="49EEF8134B3E083494EBBAF73387F930" box="[392,488,1724,1750]" country="United States of America" name="Virginia" pageId="10" pageNumber="39">Virginia</collectingRegion>
border area (
<bibRefCitation id="EFBB4B004B3E083497C7BAF73123F930" author="Thoma" box="[676,844,1724,1750]" pageId="10" pageNumber="39" pagination="731 - 738" refId="ref5585" refString="Thoma ,, R. F. (2000) Cambarus (Jugicambarus) jezerinaci (Crustacea: Decapoda: Cambaridae), a new species of crayfish from the Powell River drainage of Tennessee and Virginia. Proceedings of the Biological Society of Washington, 113, 731 - 738." type="journal article" year="2000">Thoma 2000</bibRefCitation>
). Given the paucity of informative morphological characters and molecular datasets, the monophyly of the subgenus 
<taxonomicName id="4C2A4D724B3E0834946BBB4733C1F8C3" box="[264,430,1804,1829]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Jugicambarus">
<emphasis id="B95EEAE34B3E0834946BBB4733C1F8C3" box="[264,430,1804,1829]" italics="true" pageId="10" pageNumber="39">Jugicambarus</emphasis>
</taxonomicName>
and interrelationships of its members have never been rigorously analyzed using cladistic techniques. Future analyses using a combined data set of both molecular and morphological characters should help elucidate the relationships within the subgenus and help resolve the phylogenetic position of 
<taxonomicName id="4C2A4D724B3F0835967CBD0F31F0FEBB" authorityName="Taylor &amp; Soucek &amp; Organ" authorityYear="2006" box="[799,927,324,350]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="clivosus">
<emphasis id="B95EEAE34B3F0835967CBD0F315DFEBB" box="[799,818,324,349]" italics="true" pageId="11" pageNumber="40">C</emphasis>
. 
<emphasis id="B95EEAE34B3F08359622BD0F31F0FEBB" box="[833,927,324,349]" italics="true" pageId="11" pageNumber="40">clivosus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9536F14B3F0835945BBD27310EFE48" blockId="11.[264,1323,284,430]" pageId="11" pageNumber="40">
<emphasis id="B95EEAE34B3F0835945BBD2730F2FE63" box="[312,669,364,389]" italics="true" pageId="11" pageNumber="40">Suggested vernacular name</emphasis>
.—Short Mountain Crayfish, in reference to the geographic feature near which the species is found.
</paragraph>
<paragraph id="8B9536F14B3F0835946BBE403382FDC3" blockId="11.[264,493,523,549]" box="[264,493,523,549]" pageId="11" pageNumber="40">
<emphasis id="B95EEAE34B3F0835946BBE403382FDC3" bold="true" box="[264,493,523,549]" pageId="11" pageNumber="40">Acknowledgments</emphasis>
</paragraph>
<paragraph id="8B9536F14B3F0835946BBE1C304DFCF7" blockId="11.[264,1323,599,785]" pageId="11" pageNumber="40">We are grateful to M. Retzer for reviewing an earlier version of this manuscript. We are also grateful to D. Withers, R. McCoy, and H. Garland for assisting with sampling locality access and permitting and to R. Lemaitre and K. Reed for graciously providing records from and access to the crustacean collections at the National Museum of Natural History, Smithsonian Institution.</paragraph>
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</paragraph>
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&amp; 
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<emphasis id="B95EEAE34B3F083597A4B8113159FB97" box="[711,822,1114,1137]" italics="true" pageId="11" pageNumber="40">Cambarus</emphasis>
(
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<emphasis id="B95EEAE34B3F08359693B811360AFB97" box="[1008,1125,1114,1137]" italics="true" pageId="11" pageNumber="40">tuckasegee</emphasis>
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L, &amp; 
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</subSubSection>
</treatment>
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