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<mods:title id="80F8BC887906046755A44E16614B520F">Description of a new species of the genus Didrepanephorus Wood-Mason, 1878 (Coleoptera: Scarabaeidae: Rutelinae) from China with notes on allied species</mods:title>
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<heading id="D0DD81CDFF95F700FF2A21C63706FA61" bold="true" box="[151,868,1482,1509]" fontSize="11" level="1" pageId="1" pageNumber="494" reason="1">
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<taxonomicName id="4C2A4D22FF95F700FF2A21C636C7FA66" authority="Arrow, 1921" authorityName="Arrow" authorityYear="1921" box="[151,677,1482,1509]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="494" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B95EEAB3FF95F700FF2A21C63661FA67" bold="true" box="[151,515,1482,1509]" italics="true" pageId="1" pageNumber="494">Didrepanephorus mucronatus</emphasis>
<bibRefCitation id="EFBB4B50FF95F700FDB621C636C7FA66" author="Arrow, G. J." box="[523,677,1482,1508]" pageId="1" pageNumber="494" pagination="378 - 381" refId="ref9871" refString="Arrow, G. J. (1921) Two remarkable new Ruteline beetles from Indo-china. The Annals and Magazine of Natural History, includ- ing Zoology, Botany and Geology, Series 9, 7 (40), 378 - 381. https: // doi. org / 10.1080 / 00222932108632533" type="journal article" year="1921">Arrow, 1921</bibRefCitation>
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'腹镰'•龟
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<paragraph id="8B9536A1FF95F700FF2A21E33624F98B" blockId="1.[151,868,1482,1545]" box="[151,582,1518,1545]" pageId="1" pageNumber="494">
(
<figureCitation id="13112A24FF95F700FF2221E3357AF98B" box="[159,280,1519,1545]" captionStart-0="FIGURES 14" captionStart-1="FIGURES 510" captionStartId-0="2.[151,264,2001,2025]" captionStartId-1="3.[151,264,1215,1239]" captionTargetBox-0="[213,1380,320,1964]" captionTargetBox-1="[155,1430,193,1182]" captionTargetId-0="figure-84@2.[196,1390,306,1975]" captionTargetId-1="figure-17@3.[151,1435,181,1192]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 14. Habitus of Didrepanephorus mucronatus Arrow, 1921 (dorsal and ventral view). 12, male; 34, female." captionText-1="FIGURES 510. Habitus of Didrepanephorus mucronatus Arrow, 1921 (oblique view). 58, males in different body sizes; 910, females in different body sizes." figureDoi-0="http://doi.org/10.5281/zenodo.4556622" figureDoi-1="http://doi.org/10.5281/zenodo.4556624" httpUri-0="https://zenodo.org/record/4556622/files/figure.png" httpUri-1="https://zenodo.org/record/4556624/files/figure.png" pageId="1" pageNumber="494">Figs. 110</figureCitation>
,
<figureCitation id="13112A24FF95F700FE9821E3350FF98B" box="[293,365,1519,1545]" captionStart="FIGURES 3340" captionStartId="13.[151,264,1635,1659]" captionTargetBox="[156,1393,201,1596]" captionTargetId="figure-17@13.[151,1435,181,1611]" captionTargetPageId="13" captionText="FIGURES 3340. Male genitalia of Didrepanephorus Wood-Mason, 1878 species (showing parameres in dorsal and ventral views). 3334, D. mucronatus Arrow, 1921; 3536, D. heterocolor new species (holotype); 3738, D. nishiyamai Muramoto, 2006; 3940, D. subvittatus Benderitter, 1922." figureDoi="http://doi.org/10.5281/zenodo.4556636" httpUri="https://zenodo.org/record/4556636/files/figure.png" pageId="1" pageNumber="494">3334</figureCitation>
,
<figureCitation id="13112A24FF95F700FEC421E235A0F98B" box="[377,450,1518,1545]" captionStart="FIGURES 4156" captionStartId="15.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1882]" captionTargetId="figure-17@15.[164,1423,181,1885]" captionTargetPageId="15" captionText="FIGURES 4156. Type specimens of Didrepanephorus Wood-Mason, 1878 species. 4146, syntype, labels, and male genitalia of D. bifalcifer Wood-Mason, 1878 (male, BMNH, photograph by Chuan-Bu Gao); 4752, holotype, labels, and male genitalia of D. mucronatus Arrow, 1921 (male, BMNH, photograph by Chuan-Bu Gao); 5356, holotype and labels of D. subvittatus Benderitter, 1922 (female, IRSNB, photograph by Matthias Seidel)." figureDoi="http://doi.org/10.5281/zenodo.4556638" httpUri="https://zenodo.org/record/4556638/files/figure.png" pageId="1" pageNumber="494">4752</figureCitation>
,
<figureCitation id="13112A24FF95F700FE7021E23674F98B" box="[461,534,1518,1545]" captionStart="FIGURES 6569" captionStartId="17.[151,264,1841,1865]" captionTargetBox="[189,1398,181,1818]" captionTargetId="figure-17@17.[189,1398,181,1818]" captionTargetPageId="17" captionText="FIGURES 6569. Habitat of Didrepanephorus Wood-Mason, 1878 species. 6567, habitat D. mucronatus Arrow, 1921: 65, forest in Mount Jinuoshan, Jinghong (Yunnan, China); 6667, male attracted by light trap and stayed on a nearby bush in Dadu- gang (Yunnan, China); 68, male of D. nishiyamai Muramoto, 2006 on the ground (Guangxi, China; photograph by Chao Li); 69, male of D. subvittatus Benderitter, 1922 attracted by light trap and stayed on a nearby leaf in Mount Shiwandashan (Guangxi, China; photograph by Tao Zhang)." figureDoi="http://doi.org/10.5281/zenodo.4556642" httpUri="https://zenodo.org/record/4556642/files/figure.png" pageId="1" pageNumber="494">6567</figureCitation>
,
<figureCitation id="13112A24FF95F700FD9C21E3365CF98B" box="[545,574,1519,1545]" captionStart="FIGURE 70" captionStartId="18.[151,250,1036,1060]" captionTargetBox="[151,1435,181,1012]" captionTargetId="figure-23@18.[151,1435,181,1012]" captionTargetPageId="18" captionText="FIGURE 70. Known distribution of four Didrepanephorus Wood-Mason, 1878 species. Type locality of D. mucronatus Arrow, 1921 is missing because the precise location is unknown." figureDoi="http://doi.org/10.5281/zenodo.4556644" httpUri="https://zenodo.org/record/4556644/files/figure.png" pageId="1" pageNumber="494">70</figureCitation>
)
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<taxonomicName id="4C2A4D22FF95F700FF2A223F36E5F9C9" authority="Arrow, 1921: 379" authorityName="Arrow" authorityPageNumber="379" authorityYear="1921" box="[151,647,1587,1611]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="494" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B95EEAB3FF95F700FF2A223F35ABF9C9" box="[151,457,1587,1611]" italics="true" pageId="1" pageNumber="494">Didrepanephorus mucronatus</emphasis>
<bibRefCitation id="EFBB4B50FF95F700FE6D223F36E5F9C9" author="Arrow, G. J." box="[464,647,1587,1611]" pageId="1" pageNumber="494" pagination="378 - 381" refId="ref9871" refString="Arrow, G. J. (1921) Two remarkable new Ruteline beetles from Indo-china. The Annals and Magazine of Natural History, includ- ing Zoology, Botany and Geology, Series 9, 7 (40), 378 - 381. https: // doi. org / 10.1080 / 00222932108632533" type="journal article" year="1921">Arrow, 1921: 379</bibRefCitation>
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(type locality: “
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”), plate X, fig. 2 (habitus of
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;
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<bibRefCitation id="EFBB4B50FF95F700FF53225E35FEF9E8" author="Paulian, R." box="[238,412,1618,1642]" pageId="1" pageNumber="494" pagination="73 - 105" refId="ref10639" refString="Paulian, R. (1958) Coleopteres Scarabeides de l'Indochine (Rutelines et Cetonines). Annales de la Societe Entomologique de France, 127, 73 - 105." type="journal article" year="1958">Paulian 1958: 78</bibRefCitation>
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;
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<bibRefCitation id="EFBB4B50FF95F700FE1A225E36E0F9E8" author="Machatschke, J. W." box="[423,642,1618,1642]" pageId="1" pageNumber="494" refId="ref10112" refString="Machatschke, J. W. (1972) Coleopterorum catalogus supplementa pars 66, fascicle 1, Superfamilie Scarabaeoidea, Familie Melolonthidae, Subfamilie Rutelinae. Editio Secunda. Dr. W. Junk, s' - Gravenhage, Berlin, 361 pp." type="book" year="1972">Machatschke 1972: 8</bibRefCitation>
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;
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, fig. 2 (lateral habitus of male);
<bibRefCitation id="EFBB4B50FF95F700FB39225E31FAF9E8" author="Muramoto, R. &amp; Fujioka M." box="[1156,1432,1618,1642]" pageId="1" pageNumber="494" pagination="5 - 8" refId="ref10382" refString="Muramoto, R. &amp; Fujioka M. (2000) Notes on genus Didrepanephorus Wood-Mason, 1878 (Coleoptera: Scarabaeidae: Rutelinae). Saikaku Tsushin, 1, 5 - 8." type="journal article" year="2000">Muramoto &amp; Fujioka 2000</bibRefCitation>
, figs. 79 (habitus and labels of
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, figs. 46 (habitus, genitalia and labels of
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<bibRefCitation id="EFBB4B50FF95F700FAED227C3541F925" author="Krajcik, M." pageId="1" pageNumber="494" pagination="1 - 139" refId="ref10049" refString="Krajcik, M. (2007) Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. x, Supplementum, 4, 1 - 139." type="journal article" year="2007">Krajčík 2007: 67</bibRefCitation>
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;
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<emphasis id="B95EEAB3FF95F700FF2A22D93525F96D" bold="true" box="[151,327,1749,1775]" pageId="1" pageNumber="494">Type material.</emphasis>
<materialsCitation id="3B423CFCFF95F700FEF222D9312BF896" ID-GBIF-Occurrence="3050816332" collectingDate="1919-05-15" collectionCode="BMNH" collectorName="R. Vitalis de Salvaza" country="Laos" location="LAOS" pageId="1" pageNumber="494" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54918803FF95F700FEF222D935DEF96D" box="[335,444,1749,1775]" pageId="1" pageNumber="494" type="holotype">Holotype</typeStatus>
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(♂, by monotypy,
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,
<figureCitation id="13112A24FF95F700FD4622D937E5F972" box="[763,903,1749,1776]" captionStart="FIGURES 4156" captionStartId="15.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1882]" captionTargetId="figure-17@15.[164,1423,181,1885]" captionTargetPageId="15" captionText="FIGURES 4156. Type specimens of Didrepanephorus Wood-Mason, 1878 species. 4146, syntype, labels, and male genitalia of D. bifalcifer Wood-Mason, 1878 (male, BMNH, photograph by Chuan-Bu Gao); 4752, holotype, labels, and male genitalia of D. mucronatus Arrow, 1921 (male, BMNH, photograph by Chuan-Bu Gao); 5356, holotype and labels of D. subvittatus Benderitter, 1922 (female, IRSNB, photograph by Matthias Seidel)." figureDoi="http://doi.org/10.5281/zenodo.4556638" httpUri="https://zenodo.org/record/4556638/files/figure.png" pageId="1" pageNumber="494">Figs. 4752</figureCitation>
):
<taxonomicName id="4C2A4D22FF95F700FC2622D9308EF96D" authorityName="Arrow" authorityYear="1921" box="[923,1260,1749,1775]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="494" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B95EEAB3FF95F700FC2622D9308EF96D" box="[923,1260,1749,1775]" italics="true" pageId="1" pageNumber="494">Didrepanephorus mucronatus</emphasis>
</taxonomicName>
, Type, arrow // Indo-China., R. V. de Salvaza., 1920-47 //
<collectingCountry id="F33D7631FF95F700FDCE22F536DDF891" box="[627,703,1785,1811]" name="Laos" pageId="1" pageNumber="494">LAOS</collectingCountry>
, m. yan. le
<date id="FF941061FF95F700FC8222F537D8F896" box="[831,954,1785,1812]" pageId="1" pageNumber="494" value="1919-05-15">
<collectingDate id="EFD0E989FF95F700FC8222F537D8F896" box="[831,954,1785,1812]" pageId="1" pageNumber="494" value="1919-05-15">15-V-1919</collectingDate>
</date>
,
<collectorName id="26DF5377FF95F700FC7B22F630CCF891" box="[966,1198,1785,1812]" pageId="1" pageNumber="494">R. Vitalis de Salvaza</collectorName>
// Type, H. T.
</materialsCitation>
</paragraph>
<paragraph id="8B9536A1FF95F703FF7A23113082FE9F" blockId="1.[151,1437,1749,2030]" lastBlockId="2.[151,1437,151,285]" lastPageId="2" lastPageNumber="495" pageId="1" pageNumber="494">
<emphasis id="B95EEAB3FF95F700FF7A23113649F8B5" bold="true" box="[199,555,1821,1847]" pageId="1" pageNumber="494">Additional material examined</emphasis>
(11♂♂,
<specimenCount id="9D2CFD28FF95F700FD30231136A7F8B5" box="[653,709,1821,1847]" pageId="1" pageNumber="494" type="female">8♀♀</specimenCount>
).
<materialsCitation id="3B423CFCFF95F700FD6623113013F8B5" ID-GBIF-Occurrence="3050816338" box="[731,1137,1821,1850]" collectionCode="CZMZ" country="China" location="Yunnan" pageId="1" pageNumber="494" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<emphasis id="B95EEAB3FF95F700FD66231137CCF8B5" bold="true" box="[731,942,1821,1847]" pageId="1" pageNumber="494">
<collectingCountry id="F33D7631FF95F700FD662311375FF8B5" box="[731,829,1821,1847]" name="China" pageId="1" pageNumber="494">CHINA</collectingCountry>
:
<collectingRegion id="49EEF843FF95F700FCFA231137CBF8B5" box="[839,937,1821,1847]" country="China" name="Yunnan" pageId="1" pageNumber="494">Yunnan</collectingRegion>
:
</emphasis>
1♂,
<specimenCount id="9D2CFD28FF95F700FC5923113065F8B5" box="[996,1031,1821,1847]" pageId="1" pageNumber="494" type="female">1♀</specimenCount>
(
<collectionCode id="ED3BAE64FF95F700FBA82311300AF8B5" box="[1045,1128,1821,1847]" pageId="1" pageNumber="494">CZMZ</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B423CFCFF95F700FBC62311374EF8D9" ID-GBIF-Occurrence="3050816326" collectingDate="2014-05-15" collectionCode="MYNU" collectorName="Chao-Tai WEI" country="China" county="Mengla County" location="Xishuangbanna" pageId="1" pageNumber="494" specimenCount="1" specimenCount-female="1">
<date id="FF941061FF95F700FBC623113092F8BA" box="[1147,1264,1821,1848]" pageId="1" pageNumber="494" value="2014-05-15">
<collectingDate id="EFD0E989FF95F700FBC623113092F8BA" box="[1147,1264,1821,1848]" pageId="1" pageNumber="494" value="2014-05-15">2014.V.15</collectingDate>
</date>
,
<collectingCounty id="62F44E2DFF95F700FB46231134CCF8DE" pageId="1" pageNumber="494">Mengla County</collectingCounty>
,
<location id="8EF5607AFF95F700FF06234D350EF8D9" LSID="urn:lsid:plazi:treatment:038387B7FF95F704FF2A21C635A2FDBF:8EF5607AFF95F700FF06234D350EF8D9" box="[187,364,1857,1883]" country="China" county="Mengla County" name="Xishuangbanna" pageId="1" pageNumber="494">Xishuangbanna</location>
,
<collectorName id="26DF5377FF95F700FEC4234D367DF8DE" box="[377,543,1857,1884]" pageId="1" pageNumber="494">Chao-Tai WEI</collectorName>
leg.; 1♂,
<specimenCount id="9D2CFD28FF95F700FD28234D36DAF8D9" box="[661,696,1857,1883]" pageId="1" pageNumber="494" type="female">1♀</specimenCount>
(
<collectionCode id="ED3BAE64FF95F700FD74234D3746F8D9" box="[713,804,1857,1883]" pageId="1" pageNumber="494">MYNU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B423CFCFF95F700FC84234D3132F8D9" ID-GBIF-Occurrence="3050816337" box="[825,1360,1857,1886]" collectionCode="CLYJ" collectorName="Jinghong &amp; Xishuangbanna" country="China" pageId="1" pageNumber="494" specimenCount="1">
2015.V,
<collectorName id="26DF5377FF95F700FC24234D3061F8D9" box="[921,1027,1857,1883]" pageId="1" pageNumber="494">Jinghong</collectorName>
,
<collectorName id="26DF5377FF95F700FBB2234D30A3F8D9" box="[1039,1217,1857,1883]" pageId="1" pageNumber="494">Xishuangbanna</collectorName>
; 1♂ (
<collectionCode id="ED3BAE64FF95F700FABF234D312AF8D9" box="[1282,1352,1857,1883]" pageId="1" pageNumber="494">CLYJ</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B423CFCFF95F700FAE0234D307EF8FD" ID-GBIF-Occurrence="3050816321" collectingDate="2017-04-27" collectionCode="MYNU" collectorName="Dadugang &amp; Jinghong &amp; Xishuangbanna &amp; Chao-Tai" country="China" pageId="1" pageNumber="494" specimenCount="1">
<date id="FF941061FF95F700FAE0234D34B5F8FD" pageId="1" pageNumber="494" value="2017-04-27">
<collectingDate id="EFD0E989FF95F700FAE0234D34B5F8FD" pageId="1" pageNumber="494" value="2017-04-27">2017. IV.27</collectingDate>
</date>
,
<collectorName id="26DF5377FF95F700FF5923693538F8FD" box="[228,346,1893,1919]" pageId="1" pageNumber="494">Dadugang</collectorName>
,
<collectorName id="26DF5377FF95F700FEDA236935B3F8FD" box="[359,465,1893,1919]" pageId="1" pageNumber="494">Jinghong</collectorName>
,
<collectorName id="26DF5377FF95F700FE63236936EDF8FD" box="[478,655,1893,1919]" pageId="1" pageNumber="494">Xishuangbanna</collectorName>
,
<collectorName id="26DF5377FF95F700FD2123693766F8FD" box="[668,772,1893,1919]" pageId="1" pageNumber="494">Chao-Tai</collectorName>
WEI leg.; 1♂ (
<collectionCode id="ED3BAE64FF95F700FC0523693071F8FD" box="[952,1043,1893,1919]" pageId="1" pageNumber="494">MYNU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B423CFCFF95F700FB9423693722F821" ID-GBIF-Occurrence="3050816322" collectingDate="2018-05-20" collectionCode="MYNU" collectorName="Jian-Yue &amp; Hao XU" country="China" elevation="1" pageId="1" pageNumber="494" specimenCount="1">
<date id="FF941061FF95F700FB94236930FCF8FD" box="[1065,1182,1893,1919]" pageId="1" pageNumber="494" value="2018-05-20">
<collectingDate id="EFD0E989FF95F700FB94236930FCF8FD" box="[1065,1182,1893,1919]" pageId="1" pageNumber="494" value="2018-05-20">2018.V.20</collectingDate>
</date>
,
<collectorName id="26DF5377FF95F700FB1623693143F8FD" box="[1195,1313,1893,1919]" pageId="1" pageNumber="494">Dadugang</collectorName>
,
<collectorName id="26DF5377FF95F700FA92236931FBF8FD" box="[1327,1433,1893,1919]" pageId="1" pageNumber="494">Jinghong</collectorName>
, alt.
<quantity id="4CD29B44FF95F700FF7E23853540F826" box="[195,290,1929,1956]" metricMagnitude="0" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="494" unit="m" value="1.4">
<elevation id="0007D192FF95F700FF7E23853540F826" box="[195,290,1929,1956]" metricMagnitude="0" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="494" unit="m" value="1.4">1,400 m</elevation>
</quantity>
,
<collectorName id="26DF5377FF95F700FE8D238535F0F821" box="[304,402,1929,1955]" pageId="1" pageNumber="494">Jian-Yue</collectorName>
QIU &amp;
<collectorName id="26DF5377FF95F700FE4E23863630F826" box="[499,594,1930,1956]" pageId="1" pageNumber="494">Hao XU</collectorName>
leg.; 2♂♂ (
<collectionCode id="ED3BAE64FF95F700FD602385375AF821" box="[733,824,1929,1955]" pageId="1" pageNumber="494">MYNU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B423CFCFF95F700FCF1238536C2F845" ID-GBIF-Occurrence="3050816305" collectingDate="2018-04-04" collectionCode="MYNU" collectorName="Yu-Tang Wang" country="China" county="Mengla County" elevation="850" location="Mengla County" pageId="1" pageNumber="494" specimenCount="1">
<date id="FF941061FF95F700FCF1238537DFF821" box="[844,957,1929,1955]" pageId="1" pageNumber="494" value="2018-04-04">
<collectingDate id="EFD0E989FF95F700FCF1238537DFF821" box="[844,957,1929,1955]" pageId="1" pageNumber="494" value="2018-04-04">2018.IV.4</collectingDate>
</date>
,
<collectorName id="26DF5377FF95F700FC772385307EF821" box="[970,1052,1929,1955]" pageId="1" pageNumber="494">Mohan</collectorName>
,
<collectingCounty id="62F44E2DFF95F700FB94238530B9F821" box="[1065,1243,1929,1955]" pageId="1" pageNumber="494">Mengla County</collectingCounty>
,
<collectorName id="26DF5377FF95F700FB5A238531FAF821" box="[1255,1432,1929,1955]" pageId="1" pageNumber="494">Xishuangbanna</collectorName>
, alt.
<quantity id="4CD29B44FF95F700FF7C23A1356BF84A" box="[193,265,1965,1992]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="494" unit="m" value="850.0">
<elevation id="0007D192FF95F700FF7C23A1356BF84A" box="[193,265,1965,1992]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="494" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<collectorName id="26DF5377FF95F700FEAE23A23511F84A" box="[275,371,1966,1992]" pageId="1" pageNumber="494">Yu-Tang</collectorName>
WANG leg.; 1♂ (
<collectionCode id="ED3BAE64FF95F700FD8123A136F5F845" box="[572,663,1965,1991]" pageId="1" pageNumber="494">MYNU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B423CFCFF95F700FD1723A23721F845" ID-GBIF-Occurrence="3050816345" box="[682,835,1965,1992]" collectingDate="2018-04-04" collectionCode="MYNU" collectorName="Yu-Tang Wang" country="China" county="Mengla County" elevation="850" location="Mengla County" pageId="1" pageNumber="494" specimenCount="1">same data but</materialsCitation>
<materialsCitation id="3B423CFCFF95F700FCF723A1300DF845" ID-GBIF-Occurrence="3050816344" box="[842,1135,1965,1991]" collectingDate="2018-04-07" collectionCode="MYNU" country="China" pageId="1" pageNumber="494" specimenCount="2" specimenCount-female="2">
<date id="FF941061FF95F700FCF723A137D9F845" box="[842,955,1965,1991]" pageId="1" pageNumber="494" value="2018-04-07">
<collectingDate id="EFD0E989FF95F700FCF723A137D9F845" box="[842,955,1965,1991]" pageId="1" pageNumber="494" value="2018-04-07">2018.IV.7</collectingDate>
</date>
;
<specimenCount id="9D2CFD28FF95F700FC7B23A1379CF845" box="[966,1022,1965,1991]" pageId="1" pageNumber="494" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED3BAE64FF95F700FBB123A13005F845" box="[1036,1127,1965,1991]" pageId="1" pageNumber="494">MYNU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B423CFCFF95F700FBC723A23171F845" ID-GBIF-Occurrence="3050816330" box="[1146,1299,1965,1992]" collectingDate="2018-04-07" collectionCode="MYNU" country="China" pageId="1" pageNumber="494" specimenCount="1">same data but</materialsCitation>
<materialsCitation id="3B423CFCFF95F700FAA423A135BCF869" ID-GBIF-Occurrence="3050816328" collectingDate="2018-04-10" collectionCode="MYNU" country="China" elevation="1" pageId="1" pageNumber="494" specimenCount="2" specimenCount-female="2">
<date id="FF941061FF95F700FAA423A131F5F845" box="[1305,1431,1965,1991]" pageId="1" pageNumber="494" value="2018-04-10">
<collectingDate id="EFD0E989FF95F700FAA423A131F5F845" box="[1305,1431,1965,1991]" pageId="1" pageNumber="494" value="2018-04-10">2018.IV.10</collectingDate>
</date>
, alt.
<quantity id="4CD29B44FF95F700FF7E23DD3546F86E" box="[195,292,2001,2028]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="494" unit="m" value="1.2">
<elevation id="0007D192FF95F700FF7E23DD3546F86E" box="[195,292,2001,2028]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="494" unit="m" value="1.2">1,200 m</elevation>
</quantity>
;
<specimenCount id="9D2CFD28FF95F700FE8F23DD3508F869" box="[306,362,2001,2027]" pageId="1" pageNumber="494" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED3BAE64FF95F700FEC723DD35B7F869" box="[378,469,2001,2027]" pageId="1" pageNumber="494">MYNU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B423CFCFF95F700FE5623DE36EBF869" ID-GBIF-Occurrence="3050816301" box="[491,649,2001,2028]" collectingDate="2018-04-10" collectionCode="MYNU" country="China" elevation="1" pageId="1" pageNumber="494" specimenCount="1">same data but</materialsCitation>
<materialsCitation id="3B423CFCFF95F700FD2C23DD37C8F869" ID-GBIF-Occurrence="3050816341" box="[657,938,2001,2030]" collectingDate="2018-04-24" collectionCode="CCCC" country="China" pageId="1" pageNumber="494" specimenCount="1">
<date id="FF941061FF95F700FD2C23DD3772F869" box="[657,784,2001,2027]" pageId="1" pageNumber="494" value="2018-04-24">
<collectingDate id="EFD0E989FF95F700FD2C23DD3772F869" box="[657,784,2001,2027]" pageId="1" pageNumber="494" value="2018-04-24">2018.IV.24</collectingDate>
</date>
; 1♂ (
<collectionCode id="ED3BAE64FF95F700FCEF23DD37C0F869" box="[850,930,2001,2027]" country="USA" httpUri="http://grbio.org/cool/3dng-x47a" name="Carthage College" pageId="1" pageNumber="494">CCCC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B423CFCFF95F700FC0A23DD302DF869" ID-GBIF-Occurrence="3050816302" box="[951,1103,2001,2027]" collectionCode="MYNU" country="China" pageId="1" pageNumber="494" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D2CFD28FF95F700FC0A23DD37B8F869" box="[951,986,2001,2027]" pageId="1" pageNumber="494" type="female">1♀</specimenCount>
(
<collectionCode id="ED3BAE64FF95F700FC5623DD3024F869" box="[1003,1094,2001,2027]" pageId="1" pageNumber="494">MYNU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B423CFCFF95F703FBE123DD378DFF33" ID-GBIF-Occurrence="3050816311" collectingDate="2018-04-15" collectionCode="MYNU" collectorName="Yu-Tang Wang" country="China" county="Mengla County" elevation="1" lastPageId="2" lastPageNumber="495" location="Mengla County" pageId="1" pageNumber="494" specimenCount="1">
<date id="FF941061FF95F700FBE123DD30B8F86E" box="[1116,1242,2001,2028]" pageId="1" pageNumber="494" value="2018-04-15">
<collectingDate id="EFD0E989FF95F700FBE123DD30B8F86E" box="[1116,1242,2001,2028]" pageId="1" pageNumber="494" value="2018-04-15">2018.IV.15</collectingDate>
</date>
,
<collectorName id="26DF5377FF95F700FB5A23DD315BF869" box="[1255,1337,2001,2027]" pageId="1" pageNumber="494">Mohan</collectorName>
,
<collectingCounty id="62F44E2DFF95F703FAFA23DD3489FF33" lastPageId="2" lastPageNumber="495" pageId="1" pageNumber="494">Mengla County</collectingCounty>
,
<collectorName id="26DF5377FF96F703FF44249B35C8FF33" box="[249,426,151,177]" pageId="2" pageNumber="495">Xishuangbanna</collectorName>
, alt.
<quantity id="4CD29B44FF96F703FE58249B3624FF33" box="[485,582,151,177]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="495" unit="m" value="1.2">
<elevation id="0007D192FF96F703FE58249B3624FF33" box="[485,582,151,177]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="495" unit="m" value="1.2">1,200 m</elevation>
</quantity>
,
<collectorName id="26DF5377FF96F703FDE9249B36D6FF33" box="[596,692,151,177]" pageId="2" pageNumber="495">Yu-Tang</collectorName>
WANG leg.; 1♂ (
<collectionCode id="ED3BAE64FF96F703FC36249B3784FF33" box="[907,998,151,177]" pageId="2" pageNumber="495">MYNU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B423CFCFF96F703FC40249B36FAFF57" ID-GBIF-Occurrence="3050816323" collectingDate="2018-05-31" collectionCode="MYNU" collectorName="Yu-Tang Wang" country="China" elevation="1" pageId="2" pageNumber="495" specimenCount="1">
<date id="FF941061FF96F703FC40249B3010FF33" box="[1021,1138,151,177]" pageId="2" pageNumber="495" value="2018-05-31">
<collectingDate id="EFD0E989FF96F703FC40249B3010FF33" box="[1021,1138,151,177]" pageId="2" pageNumber="495" value="2018-05-31">2018.V.31</collectingDate>
</date>
,
<collectorName id="26DF5377FF96F703FB3D249B3091FF33" box="[1152,1267,151,177]" pageId="2" pageNumber="495">Jinuoshan</collectorName>
,
<collectorName id="26DF5377FF96F703FABF249B310EFF33" box="[1282,1388,151,177]" pageId="2" pageNumber="495">Jinghong</collectorName>
, alt.
<quantity id="4CD29B44FF96F703FF2A24B73495FF57" box="[151,247,187,213]" metricMagnitude="0" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="495" unit="m" value="1.1">
<elevation id="0007D192FF96F703FF2A24B73495FF57" box="[151,247,187,213]" metricMagnitude="0" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="495" unit="m" value="1.1">1,100 m</elevation>
</quantity>
,
<collectorName id="26DF5377FF96F703FEBE24B73501FF57" box="[259,355,187,213]" pageId="2" pageNumber="495">Yu-Tang</collectorName>
WANG leg.; 1♂ (
<collectionCode id="ED3BAE64FF96F703FD8824B736F2FF57" box="[565,656,187,213]" pageId="2" pageNumber="495">MYNU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B423CFCFF96F703FD1824B7354EFF7B" ID-GBIF-Occurrence="3050816306" collectingDate="2019-05-06" collectionCode="MYNU" collectorName="Yu-Tang Wang" country="China" elevation="1" pageId="2" pageNumber="495" specimenCount="1" specimenCount-female="1">
<date id="FF941061FF96F703FD1824B7376EFF57" box="[677,780,187,213]" pageId="2" pageNumber="495" value="2019-05-06">
<collectingDate id="EFD0E989FF96F703FD1824B7376EFF57" box="[677,780,187,213]" pageId="2" pageNumber="495" value="2019-05-06">2019.V.6</collectingDate>
</date>
,
<collectorName id="26DF5377FF96F703FCA424B737EEFF57" box="[793,908,187,213]" pageId="2" pageNumber="495">Jinuoshan</collectorName>
,
<collectorName id="26DF5377FF96F703FC2424B73061FF57" box="[921,1027,187,213]" pageId="2" pageNumber="495">Jinghong</collectorName>
, alt.
<quantity id="4CD29B44FF96F703FB8724B730F8FF57" box="[1082,1178,187,213]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="495" unit="m" value="1.2">
<elevation id="0007D192FF96F703FB8724B730F8FF57" box="[1082,1178,187,213]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="495" unit="m" value="1.2">1,200 m</elevation>
</quantity>
,
<collectorName id="26DF5377FF96F703FB1B24B73164FF57" box="[1190,1286,187,213]" pageId="2" pageNumber="495">Yu-Tang</collectorName>
WANG leg.;
<specimenCount id="9D2CFD28FF96F703FF2A24D334D8FF7B" box="[151,186,223,249]" pageId="2" pageNumber="495" type="female">1♀</specimenCount>
(
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)
</materialsCitation>
,
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<date id="FF941061FF96F703FE8524D335FDFF7B" box="[312,415,223,249]" pageId="2" pageNumber="495" value="2019-05-07">
<collectingDate id="EFD0E989FF96F703FE8524D335FDFF7B" box="[312,415,223,249]" pageId="2" pageNumber="495" value="2019-05-07">2019.V.7</collectingDate>
</date>
,
<collectingMunicipality id="6BF1ACDBFF96F703FE1724D3367FFF7B" box="[426,541,223,249]" pageId="2" pageNumber="495">Jinuoshan</collectingMunicipality>
,
<location id="8EF5607AFF96F703FD9524D336F0FF7B" LSID="urn:lsid:plazi:treatment:038387B7FF95F704FF2A21C635A2FDBF:8EF5607AFF96F703FD9524D336F0FF7B" box="[552,658,223,249]" country="China" municipality="Jinuoshan" name="Jinghong" pageId="2" pageNumber="495">Jinghong</location>
, alt.
<quantity id="4CD29B44FF96F703FD7A24D33747FF7B" box="[711,805,223,249]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="495" unit="m" value="1.2">
<elevation id="0007D192FF96F703FD7A24D33747FF7B" box="[711,805,223,249]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="495" unit="m" value="1.2">1,200 m</elevation>
</quantity>
,
<collectorName id="26DF5377FF96F703FC8D24D337F2FF7B" box="[816,912,223,249]" pageId="2" pageNumber="495">Yu-Tang</collectorName>
WANG leg
</materialsCitation>
.;
<materialsCitation id="3B423CFCFF96F703FB9724D230BEFE9F" ID-GBIF-Occurrence="3050816339" collectingDate="2005-05-05" collectingDateMax="2005-05-20" collectingDateMin="2005-05-05" collectionCode="CCZ" collectorName="Ban Van Hua &amp; M. Pejcha" country="Laos" elevation="1" location="Municipal" municipality="Phou Khao Khouay" pageId="2" pageNumber="495" specimenCount="1" stateProvince="Viangchan">
<emphasis id="B95EEAB3FF96F703FB9724D23014FF7A" bold="true" box="[1066,1142,222,248]" pageId="2" pageNumber="495">
<collectingCountry id="F33D7631FF96F703FB9724D23014FF7A" box="[1066,1142,222,248]" name="Laos" pageId="2" pageNumber="495">LAOS</collectingCountry>
</emphasis>
: 1♂ (
<collectionCode id="ED3BAE64FF96F703FB0A24D33091FF7B" box="[1207,1267,223,249]" pageId="2" pageNumber="495">CCZ</collectionCode>
),
<date id="FF941061FF96F703FABA24D231F5FF7B" box="[1287,1431,222,249]" pageId="2" pageNumber="495" value="2005-05-05" valueMax="2005-05-20" valueMin="2005-05-05">
<collectingDate id="EFD0E989FF96F703FABA24D231F5FF7B" box="[1287,1431,222,249]" pageId="2" pageNumber="495" value="2005-05-05" valueMax="2005-05-20" valueMin="2005-05-05">2005.V.520</collectingDate>
</date>
,
<collectingMunicipality id="6BF1ACDBFF96F703FF2A250F3516FE9F" box="[151,372,259,285]" pageId="2" pageNumber="495">Phou Khao Khouay</collectingMunicipality>
NBCA,
<collectorName id="26DF5377FF96F703FE6A250F360EFE9F" box="[471,620,259,285]" pageId="2" pageNumber="495">Ban Van Hua</collectorName>
env.,
<collectingRegion id="49EEF843FF96F703FD13250F377BFE9F" box="[686,793,259,285]" country="Laos" name="Viangchan" pageId="2" pageNumber="495">Vientiane</collectingRegion>
<location id="8EF5607AFF96F703FC9D250F37F4FE9F" LSID="urn:lsid:plazi:treatment:038387B7FF95F704FF2A21C635A2FDBF:8EF5607AFF96F703FC9D250F37F4FE9F" box="[800,918,259,285]" country="Laos" municipality="Phou Khao Khouay" name="Municipal" pageId="2" pageNumber="495" stateProvince="Viangchan">Municipal</location>
, 800
<quantity id="4CD29B44FF96F703FC67250F3055FE9F" box="[986,1079,259,285]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="495" unit="m" value="1.0">
<elevation id="0007D192FF96F703FC67250F3055FE9F" box="[986,1079,259,285]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="495" unit="m" value="1.0">1,000 m</elevation>
</quantity>
,
<collectorName id="26DF5377FF96F703FBFE250F30D7FE9F" box="[1091,1205,259,285]" pageId="2" pageNumber="495">M. Pejcha</collectorName>
lgt
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF556629FF96F703FF2A23DD310AF868" ID-DOI="http://doi.org/10.5281/zenodo.4556622" ID-Zenodo-Dep="4556622" box="[151,1384,2001,2026]" httpUri="https://zenodo.org/record/4556622/files/figure.png" pageId="2" pageNumber="495" startId="2.[151,264,2001,2025]" targetBox="[213,1380,320,1964]" targetPageId="2">
<paragraph id="8B9536A1FF96F703FF2A23DD310AF868" blockId="2.[151,1384,2001,2026]" box="[151,1384,2001,2026]" pageId="2" pageNumber="495">
<emphasis id="B95EEAB3FF96F703FF2A23DD3558F868" bold="true" box="[151,314,2001,2026]" pageId="2" pageNumber="495">FIGURES 14.</emphasis>
Habitus of
<taxonomicName id="4C2A4D22FF96F703FE0C23DE3705F868" authority="Arrow, 1921" authorityName="Arrow" authorityYear="1921" box="[433,871,2002,2026]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B95EEAB3FF96F703FE0C23DE3682F868" box="[433,736,2002,2026]" italics="true" pageId="2" pageNumber="495">Didrepanephorus mucronatus</emphasis>
<bibRefCitation id="EFBB4B50FF96F703FD5A23DE3705F868" author="Arrow, G. J." box="[743,871,2002,2026]" pageId="2" pageNumber="495" pagination="378 - 381" refId="ref9871" refString="Arrow, G. J. (1921) Two remarkable new Ruteline beetles from Indo-china. The Annals and Magazine of Natural History, includ- ing Zoology, Botany and Geology, Series 9, 7 (40), 378 - 381. https: // doi. org / 10.1080 / 00222932108632533" type="journal article" year="1921">Arrow, 1921</bibRefCitation>
</taxonomicName>
(dorsal and ventral view). 12, male; 34, female.
</paragraph>
</caption>
<caption id="DF556629FF97F702FF2A20B33676FB7E" ID-DOI="http://doi.org/10.5281/zenodo.4556624" ID-Zenodo-Dep="4556624" httpUri="https://zenodo.org/record/4556624/files/figure.png" pageId="3" pageNumber="496" startId="3.[151,264,1215,1239]" targetBox="[155,1430,193,1182]" targetPageId="3">
<paragraph id="8B9536A1FF97F702FF2A20B33676FB7E" blockId="3.[151,1436,1215,1276]" pageId="3" pageNumber="496">
<emphasis id="B95EEAB3FF97F702FF2A20B33528FB55" bold="true" box="[151,330,1215,1239]" pageId="3" pageNumber="496">FIGURES 510.</emphasis>
Habitus of
<taxonomicName id="4C2A4D22FF97F702FE7520CC37E7FB5A" authority="Arrow, 1921" authorityName="Arrow" authorityYear="1921" box="[456,901,1216,1240]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B95EEAB3FF97F702FE7520CC3698FB55" box="[456,762,1216,1240]" italics="true" pageId="3" pageNumber="496">Didrepanephorus mucronatus</emphasis>
<bibRefCitation id="EFBB4B50FF97F702FCBF20CC37E7FB5A" author="Arrow, G. J." box="[770,901,1216,1240]" pageId="3" pageNumber="496" pagination="378 - 381" refId="ref9871" refString="Arrow, G. J. (1921) Two remarkable new Ruteline beetles from Indo-china. The Annals and Magazine of Natural History, includ- ing Zoology, Botany and Geology, Series 9, 7 (40), 378 - 381. https: // doi. org / 10.1080 / 00222932108632533" type="journal article" year="1921">Arrow, 1921</bibRefCitation>
</taxonomicName>
(oblique view). 58, males in different body sizes; 910, females in different body sizes.
</paragraph>
</caption>
<subSubSection id="C330652AFF97F705FF7A212535F7FA1F" lastPageId="4" lastPageNumber="497" pageId="3" pageNumber="496" type="description">
<paragraph id="8B9536A1FF97F705FF7A2125307AFC6F" blockId="3.[151,1437,1321,2032]" lastBlockId="4.[151,1437,150,2013]" lastPageId="4" lastPageNumber="497" pageId="3" pageNumber="496">
<emphasis id="B95EEAB3FF97F702FF7A21253655FAC1" bold="true" box="[199,567,1321,1347]" pageId="3" pageNumber="496">Redescription (male). General:</emphasis>
Body oval and convex, hairs and setae yellowish brown (
<figureCitation id="13112A24FF97F702FB0521263167FAC6" box="[1208,1285,1322,1348]" captionStart="FIGURES 14" captionStartId="2.[151,264,2001,2025]" captionTargetBox="[213,1380,320,1964]" captionTargetId="figure-84@2.[196,1390,306,1975]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Didrepanephorus mucronatus Arrow, 1921 (dorsal and ventral view). 12, male; 34, female." figureDoi="http://doi.org/10.5281/zenodo.4556622" httpUri="https://zenodo.org/record/4556622/files/figure.png" pageId="3" pageNumber="496">Figs. 1</figureCitation>
,
<figureCitation id="13112A24FF97F702FAAF21263122FAC7" box="[1298,1344,1322,1349]" captionStart="FIGURES 510" captionStartId="3.[151,264,1215,1239]" captionTargetBox="[155,1430,193,1182]" captionTargetId="figure-17@3.[151,1435,181,1192]" captionTargetPageId="3" captionText="FIGURES 510. Habitus of Didrepanephorus mucronatus Arrow, 1921 (oblique view). 58, males in different body sizes; 910, females in different body sizes." figureDoi="http://doi.org/10.5281/zenodo.4556624" httpUri="https://zenodo.org/record/4556624/files/figure.png" pageId="3" pageNumber="496">58</figureCitation>
).
<emphasis id="B95EEAB3FF97F702FAEE212631FEFAC6" bold="true" box="[1363,1436,1322,1348]" pageId="3" pageNumber="496">Head:</emphasis>
Dark brown. Clypeus flat, trapezoidal, anterior margin feebly arcuate, anterior angle almost right angle, sides strongly convergent anteriad and slightly swollen in basal half, roundly curved in middle, then almost subparallel in apical third; with irregular and moderately dense, small punctures, somewhat less punctured near anterior margin; surface with moderately dense minute setae, margins with erect short setae. Frontal-clypeal suture broadly interrupted medially, represented by a short carina at each side. Frons and vertex densely rugopunctate with dense, semierect long setae, the setae being rather long posteriad. Eye-canthus wide and truncate at outer margin, slightly extends beyond outermost point of eye, anterior angle broadly rounded. Antennomeres 27 dark brown, antennal club and antennomere 1 yellowish brown, length of antennal club distinctly longer than antennomeres 27 combined. Labrum exposed, anterior margin slightly raised at middle, dorsal surface rather sparsely microsculptured, with sparse, erect, short and long setae. Mandible dark brown, broadest at base, upper margin with an acute and distinct protrusion near base, strongly bended upward. Larger body size with longer mandibles (
<figureCitation id="13112A24FF97F702FC0022BA304BF953" box="[957,1065,1718,1745]" captionStart="FIGURES 510" captionStartId="3.[151,264,1215,1239]" captionTargetBox="[155,1430,193,1182]" captionTargetId="figure-17@3.[151,1435,181,1192]" captionTargetPageId="3" captionText="FIGURES 510. Habitus of Didrepanephorus mucronatus Arrow, 1921 (oblique view). 58, males in different body sizes; 910, females in different body sizes." figureDoi="http://doi.org/10.5281/zenodo.4556624" httpUri="https://zenodo.org/record/4556624/files/figure.png" pageId="3" pageNumber="496">Figs. 58</figureCitation>
). Maxillary palpi and labial palpi dark reddish brown, simple.
<collectingRegion id="49EEF843FF97F702FE6422D6365AF976" box="[473,568,1754,1780]" country="Myanmar" name="Chin" pageId="3" pageNumber="496">Mentum</collectingRegion>
and maxilla with dense punctures and long setae, anterior margin of mentum concave at middle. Gula dark brown, glabrous and matt.
<emphasis id="B95EEAB3FF97F702FCB122F237EDF89A" bold="true" box="[780,911,1790,1816]" pageId="3" pageNumber="496">Pronotum:</emphasis>
Yellowish-brown. Strongly convex, about 1.35 times as wide as long, widest near middle. Anterior margin distinctly bisinuate; anterior marginal membrane absent. Sides concave and not strongly convergent posteriad in basal half, roundly and broadly curved at middle, feebly concave and strongly convergent anteriad in apical half. Posterior margin broadly protruding in middle. Marginal lines completed, basal marginal line slightly vague at middle. Anterior and posterior angles both distinctly protruding and blunt at apices. Surface with rather dense small punctures, rugopunctate along lateral margin, disc with a longitudinal medial line. Most areas of surface with semierect, minute to short setae; setae on both sides of disc long and tufted.
<emphasis id="B95EEAB3FF90F705FEF0249A35ACFF32" bold="true" box="[333,462,150,176]" pageId="4" pageNumber="497">Scutellum:</emphasis>
Dark brown. Nearly triangular to short triangular, lateral margin curved, apex rounded. Surface densely rugopunctate, with short setae on basomedial area.
<emphasis id="B95EEAB3FF90F705FC2224B73797FF57" bold="true" box="[927,1013,187,213]" pageId="4" pageNumber="497">Elytra:</emphasis>
Yellowish brown to brown. Convex, almost as wide as long, widest at middle. Surface with moderately dense, irregular, shallow, round to elliptical large punctures. Four innermost intervals recognizable but unclear, subsutural interstice widest. Surface with dense semierect setae; setae along base of elytra long and yellowish; on disc short and brownish; on distomarginal area and humeral umbone short, color gradually changed to dark brown; on distal declivity short, yellowish. Epipleura with dense minute setae.
<emphasis id="B95EEAB3FF90F705FE0825633682FE0A" bold="true" box="[437,736,366,393]" pageId="4" pageNumber="497">Ventral thoracic surface:</emphasis>
Black. Anterior margin and lateral portions of ventral prothoracic surface with dense long hairs (
<figureCitation id="13112A24FF90F705FD92259F3616FE2F" box="[559,628,403,429]" captionStart="FIGURES 14" captionStartId="2.[151,264,2001,2025]" captionTargetBox="[213,1380,320,1964]" captionTargetId="figure-84@2.[196,1390,306,1975]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Didrepanephorus mucronatus Arrow, 1921 (dorsal and ventral view). 12, male; 34, female." figureDoi="http://doi.org/10.5281/zenodo.4556622" httpUri="https://zenodo.org/record/4556622/files/figure.png" pageId="4" pageNumber="497">Fig. 2</figureCitation>
); ventral mesothoracic surface with sparse short setae; ventral metathoracic surface with rather dense long hairs. Prosternal process small, horizontally broad, shield-liked with an medial carina. Metasternum with a large, glabrous, longitudinal medial protrusion.
<emphasis id="B95EEAB3FF90F705FC4325D630C6FE76" bold="true" box="[1022,1188,474,500]" pageId="4" pageNumber="497">Propygidium:</emphasis>
Dark brown. Surface coarse, with rather dense, semierect short setae.
<emphasis id="B95EEAB3FF90F705FD7C25F2375FFD9A" bold="true" box="[705,829,510,536]" pageId="4" pageNumber="497">Pygidium:</emphasis>
Convex, apex concave. Dark brown. Surface coarse, with rather dense, semierect short setae.
<emphasis id="B95EEAB3FF90F705FDEA262E36B7FDBE" bold="true" box="[599,725,546,572]" pageId="4" pageNumber="497">Abdomen:</emphasis>
Reddish brown, with dense long hairs. Abdominal ventrites distinctly curved in lateral view; abdominal ventrites 14 strongly abbreviated; posterior margin of abdominal ventrite 6 broadly and deeply concave medially.
<emphasis id="B95EEAB3FF90F705FDE9266736F7FD07" bold="true" box="[596,661,619,645]" pageId="4" pageNumber="497">Legs:</emphasis>
Yellowish-brown. Femora with dense punctures and long hairs; tibiae with dense punctures and short setae; tarsi with dense, hard, short setae. Inner protarsal claw simple and thickened, outer protarsal claw smaller and sharp at apex. Outer meso- and metatarsal claws widely and deeply spilt into two branches respectively, the upper branches thinner and sharper, equal to the lower branches in length. Inner meso- and metatarsal claws more curved than the outer ones. Protibia with three distinct and acute teeth, the middle one is adjacent to the apical one. Protibial spur situates at the level of middle protibial tooth, vertically orient forwards. Meso- and metatibiae bidentate at apical edges, the upper tooth distinctly protruding. Protarsus strongly thickened; protarsomeres 24 abbreviated;protarsomere 4 ventrally with a short protrusion; meso- and metatarsomeres 4 each with an acute ventroapical protrusion, each protrusion with a pair of spines at base. Tarsomeres 5 of all legs with a ventral protrusion in basal third, which is distinct in protarsomere but indistinct in meso- and metatarsomere; protarsomere 5 with a distinct ventromedial protrusion.
<emphasis id="B95EEAB3FF90F705FD5E27DE3702FC6E" bold="true" box="[739,864,978,1004]" pageId="4" pageNumber="497">Aedeagus:</emphasis>
As
<figureCitation id="13112A24FF90F705FC3027DF3076FC6F" box="[909,1044,979,1005]" captionStart="FIGURES 3340" captionStartId="13.[151,264,1635,1659]" captionTargetBox="[156,1393,201,1596]" captionTargetId="figure-17@13.[151,1435,181,1611]" captionTargetPageId="13" captionText="FIGURES 3340. Male genitalia of Didrepanephorus Wood-Mason, 1878 species (showing parameres in dorsal and ventral views). 3334, D. mucronatus Arrow, 1921; 3536, D. heterocolor new species (holotype); 3738, D. nishiyamai Muramoto, 2006; 3940, D. subvittatus Benderitter, 1922." figureDoi="http://doi.org/10.5281/zenodo.4556636" httpUri="https://zenodo.org/record/4556636/files/figure.png" pageId="4" pageNumber="497">Figs. 3334</figureCitation>
.
</paragraph>
<paragraph id="8B9536A1FF90F705FF7A27FB3662FAD7" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="497">
<emphasis id="B95EEAB3FF90F705FF7A27FB354AFB93" bold="true" box="[199,296,1015,1041]" pageId="4" pageNumber="497">Female:</emphasis>
Body color darker; head, propygidium, pygidium and ventral surface black (
<figureCitation id="13112A24FF90F705FB3B27FB30B6FB93" box="[1158,1236,1015,1041]" captionStart="FIGURES 14" captionStartId="2.[151,264,2001,2025]" captionTargetBox="[213,1380,320,1964]" captionTargetId="figure-84@2.[196,1390,306,1975]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Didrepanephorus mucronatus Arrow, 1921 (dorsal and ventral view). 12, male; 34, female." figureDoi="http://doi.org/10.5281/zenodo.4556622" httpUri="https://zenodo.org/record/4556622/files/figure.png" pageId="4" pageNumber="497">Figs. 3</figureCitation>
,
<figureCitation id="13112A24FF90F705FB5C27FB317CFB93" box="[1249,1310,1015,1041]" captionStart="FIGURES 510" captionStartId="3.[151,264,1215,1239]" captionTargetBox="[155,1430,193,1182]" captionTargetId="figure-17@3.[151,1435,181,1192]" captionTargetPageId="3" captionText="FIGURES 510. Habitus of Didrepanephorus mucronatus Arrow, 1921 (oblique view). 58, males in different body sizes; 910, females in different body sizes." figureDoi="http://doi.org/10.5281/zenodo.4556624" httpUri="https://zenodo.org/record/4556624/files/figure.png" pageId="4" pageNumber="497">910</figureCitation>
). Mandible small, nearly triangular; outer edge bidentate, both teeth acute and bend upwards. Clypeus longer than in male, anterior margin truncate and distinctly convex. Antennal club slightly shorter. Punctures on dorsal surface of head and pronotum denser and coarser; punctures on elytra denser. Setae and hairs on the whole body surface sparser and shorter. Elytra distinctly and broadly bulging at middle of lateral portion. Medial portion of metasternum flat. Abdominal ventrites evenly convex in lateral view; abdominal ventrites 14 almost equal; posterior margin of abdominal ventrite 6 straight medially (
<figureCitation id="13112A24FF90F705FD8E20C33615FB6B" box="[563,631,1231,1257]" captionStart="FIGURES 14" captionStartId="2.[151,264,2001,2025]" captionTargetBox="[213,1380,320,1964]" captionTargetId="figure-84@2.[196,1390,306,1975]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Didrepanephorus mucronatus Arrow, 1921 (dorsal and ventral view). 12, male; 34, female." figureDoi="http://doi.org/10.5281/zenodo.4556622" httpUri="https://zenodo.org/record/4556622/files/figure.png" pageId="4" pageNumber="497">Fig. 4</figureCitation>
). Protibia and mesotibia shorter and thinner; metatibia shorter but wider; protibial teeth acuter; tarsi of all legs shorter and thinner than in male; tarsomeres 4 of all legs each with an acute ventroapical protrusion, each protrusion with a pair of spines at base; tarsomeres 5 of all legs with a small and blunt ventral protrusion in basal third.
</paragraph>
<paragraph id="8B9536A1FF90F705FF7A215335F7FA1F" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="497">
<emphasis id="B95EEAB3FF90F705FF7A2153351CFAFB" bold="true" box="[199,382,1375,1401]" pageId="4" pageNumber="497">Measurements.</emphasis>
Body length: male 18.322.4 mm, female 19.222.4 mm; body width: male 9.311.9 mm, female 10.012.4 mm.
</paragraph>
</subSubSection>
<subSubSection id="C330652AFF90F705FF7A21AA36DEFA43" box="[199,700,1446,1473]" pageId="4" pageNumber="497" type="diagnosis">
<paragraph id="8B9536A1FF90F705FF7A21AA36DEFA43" blockId="4.[151,1437,150,2013]" box="[199,700,1446,1473]" pageId="4" pageNumber="497">
<emphasis id="B95EEAB3FF90F705FF7A21AA35ACFA42" bold="true" box="[199,462,1446,1472]" pageId="4" pageNumber="497">Differential diagnosis.</emphasis>
See remarks section.
</paragraph>
</subSubSection>
<subSubSection id="C330652AFF90F705FF7A21C6351EF9D3" pageId="4" pageNumber="497" type="distribution">
<paragraph id="8B9536A1FF90F705FF7A21C63698FA67" blockId="4.[151,1437,150,2013]" box="[199,762,1482,1509]" pageId="4" pageNumber="497">
<emphasis id="B95EEAB3FF90F705FF7A21C63503FA66" bold="true" box="[199,353,1482,1508]" pageId="4" pageNumber="497">Distribution.</emphasis>
<collectingCountry id="F33D7631FF90F705FEDA21C735CCFA67" box="[359,430,1483,1509]" name="China" pageId="4" pageNumber="497">China</collectingCountry>
:
<collectingRegion id="49EEF843FF90F705FE0521C7366CFA67" box="[440,526,1483,1509]" country="China" name="Yunnan" pageId="4" pageNumber="497">Yunnan</collectingRegion>
(
<emphasis id="B95EEAB3FF90F705FDA321C736C6FA67" bold="true" box="[542,676,1483,1509]" pageId="4" pageNumber="497">new record</emphasis>
);
<collectingCountry id="F33D7631FF90F705FD0121C73694FA67" box="[700,758,1483,1509]" name="Laos" pageId="4" pageNumber="497">Laos</collectingCountry>
.
</paragraph>
<paragraph id="8B9536A1FF90F705FF7A21E3351EF9D3" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="497">
<emphasis id="B95EEAB3FF90F705FF7A21E335E4F98A" bold="true" box="[199,390,1518,1545]" pageId="4" pageNumber="497">Natural history.</emphasis>
This species inhabits the tropical rainforests at elevations of 850
<quantity id="4CD29B44FF90F705FBCD21E230AEF98B" box="[1136,1228,1518,1545]" metricMagnitude="0" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="497" unit="m" value="1.4">1,400 m</quantity>
around the border of
<collectingCountry id="F33D7631FF90F705FF0B221F349BF9AF" box="[182,249,1555,1581]" name="China" pageId="4" pageNumber="497">China</collectingCountry>
and
<collectingCountry id="F33D7631FF90F705FE8D221F3508F9AF" box="[304,362,1555,1581]" name="Laos" pageId="4" pageNumber="497">Laos</collectingCountry>
. Adults can be collected by light trap from early April to later May (
<figureCitation id="13112A24FF90F705FBD4221F3090F9AF" box="[1129,1266,1554,1581]" captionStart="FIGURES 6569" captionStartId="17.[151,264,1841,1865]" captionTargetBox="[189,1398,181,1818]" captionTargetId="figure-17@17.[189,1398,181,1818]" captionTargetPageId="17" captionText="FIGURES 6569. Habitat of Didrepanephorus Wood-Mason, 1878 species. 6567, habitat D. mucronatus Arrow, 1921: 65, forest in Mount Jinuoshan, Jinghong (Yunnan, China); 6667, male attracted by light trap and stayed on a nearby bush in Dadu- gang (Yunnan, China); 68, male of D. nishiyamai Muramoto, 2006 on the ground (Guangxi, China; photograph by Chao Li); 69, male of D. subvittatus Benderitter, 1922 attracted by light trap and stayed on a nearby leaf in Mount Shiwandashan (Guangxi, China; photograph by Tao Zhang)." figureDoi="http://doi.org/10.5281/zenodo.4556642" httpUri="https://zenodo.org/record/4556642/files/figure.png" pageId="4" pageNumber="497">Figs. 6567</figureCitation>
), but immature stages are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C330652AFF90F704FF7A225735A2FDBF" lastPageId="5" lastPageNumber="498" pageId="4" pageNumber="497" type="discussion">
<paragraph id="8B9536A1FF90F705FF7A225730E6F83B" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="497">
<emphasis id="B95EEAB3FF90F705FF7A22573559F9F7" bold="true" box="[199,315,1627,1653]" pageId="4" pageNumber="497">Remarks.</emphasis>
Although
<taxonomicName id="4C2A4D22FF90F705FE04225737C4F9F7" authority="Arrow, 1921" authorityName="Arrow" authorityYear="1921" box="[441,934,1627,1653]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B95EEAB3FF90F705FE0422573769F9F6" box="[441,779,1627,1653]" italics="true" pageId="4" pageNumber="497">Didrepanephorus mucronatus</emphasis>
<bibRefCitation id="EFBB4B50FF90F705FCA8225737C4F9F7" author="Arrow, G. J." box="[789,934,1627,1653]" pageId="4" pageNumber="497" pagination="378 - 381" refId="ref9871" refString="Arrow, G. J. (1921) Two remarkable new Ruteline beetles from Indo-china. The Annals and Magazine of Natural History, includ- ing Zoology, Botany and Geology, Series 9, 7 (40), 378 - 381. https: // doi. org / 10.1080 / 00222932108632533" type="journal article" year="1921">Arrow, 1921</bibRefCitation>
</taxonomicName>
has been mentioned several times (
<bibRefCitation id="EFBB4B50FF90F705FAFB225734B0F91B" author="Paulian, R." pageId="4" pageNumber="497" pagination="73 - 105" refId="ref10639" refString="Paulian, R. (1958) Coleopteres Scarabeides de l'Indochine (Rutelines et Cetonines). Annales de la Societe Entomologique de France, 127, 73 - 105." type="journal article" year="1958">Paulian 1958</bibRefCitation>
;
<bibRefCitation id="EFBB4B50FF90F705FF602273350BF91B" author="Young, R. M." box="[221,361,1663,1689]" pageId="4" pageNumber="497" pagination="355 - 358" refId="ref10868" refString="Young, R. M. (1999) A new Fruhstorferia Kolbe from the Dawna Ra Mountains, Myanmar. The Coleopterists Bulletin, 53 (4), 355 - 358." type="journal article" year="1999">Young 1999</bibRefCitation>
;
<bibRefCitation id="EFBB4B50FF90F705FECB227336C8F91B" author="Muramoto, R. &amp; Fujioka M." box="[374,682,1663,1689]" pageId="4" pageNumber="497" pagination="5 - 8" refId="ref10382" refString="Muramoto, R. &amp; Fujioka M. (2000) Notes on genus Didrepanephorus Wood-Mason, 1878 (Coleoptera: Scarabaeidae: Rutelinae). Saikaku Tsushin, 1, 5 - 8." type="journal article" year="2000">Muramoto &amp; Fujioka 2000</bibRefCitation>
;
<bibRefCitation id="EFBB4B50FF90F705FD082273370DF91B" author="Muramoto, R." box="[693,879,1663,1689]" pageId="4" pageNumber="497" pagination="30 - 32" refId="ref10203" refString="Muramoto, R. (2003) A new species of the genus Didrepanephorus from southern Vietnam (Coleoptera, Scarabaeidae, Rutelinae). Kogane, 4, 30 - 32." type="journal article" year="2003">Muramoto 2003</bibRefCitation>
), no further specimens were reported since it was described. The new specimens collected in the past two decades from
<collectingCountry id="F33D7631FF90F705FC0B22AF379BF93F" box="[950,1017,1699,1725]" name="China" pageId="4" pageNumber="497">China</collectingCountry>
do not only represent a new country record, but also allowed us to observe detailed morphological characters of this rare species and its variability.
<taxonomicName id="4C2A4D22FF90F705FF2A22E7358AF886" authorityName="Arrow" authorityYear="1921" box="[151,488,1771,1797]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B95EEAB3FF90F705FF2A22E7358AF886" box="[151,488,1771,1797]" italics="true" pageId="4" pageNumber="497">Didrepanephorus mucronatus</emphasis>
</taxonomicName>
, together with
<taxonomicName id="4C2A4D22FF90F705FD2522E73069F887" authority="Wood-Mason, 1878" authorityName="Wood-Mason" authorityYear="1878" box="[664,1035,1771,1797]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="bifalcifer">
<emphasis id="B95EEAB3FF90F705FD2522E73741F887" box="[664,803,1771,1797]" italics="true" pageId="4" pageNumber="497">D. bifalcifer</emphasis>
<bibRefCitation id="EFBB4B50FF90F705FC9622E73069F887" author="Wood-Mason, J." box="[811,1035,1771,1797]" pageId="4" pageNumber="497" pagination="422 - 426" refId="ref10798" refString="Wood-Mason, J. (1878) Description of Didrepanephorus bifalcifer, the type of a new genus and species of Rutelidae, remarkable for the huge sickle-shaped mandibular horns of the males. Annals and Magazine of Natural History, including Zoology, Botany and Geology, Series 5, 2 (11), 422 - 426. https: // doi. org / 10.1080 / 00222937808682447" type="journal article" year="1878">Wood-Mason, 1878</bibRefCitation>
</taxonomicName>
(Northeast
<collectingCountry id="F33D7631FF90F705FB2E22E730B3F884" box="[1171,1233,1771,1798]" name="India" pageId="4" pageNumber="497">India</collectingCountry>
),
<taxonomicName id="4C2A4D22FF90F705FB5B22E7354DF8AB" authority="Muramoto, 2003" authorityName="Muramoto" authorityYear="2003" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="arnaudi">
<emphasis id="B95EEAB3FF90F705FB5B22E73101F887" box="[1254,1379,1771,1797]" italics="true" pageId="4" pageNumber="497">D. arnaudi</emphasis>
<bibRefCitation id="EFBB4B50FF90F705FAD122E7354DF8AB" author="Muramoto, R." pageId="4" pageNumber="497" pagination="30 - 32" refId="ref10203" refString="Muramoto, R. (2003) A new species of the genus Didrepanephorus from southern Vietnam (Coleoptera, Scarabaeidae, Rutelinae). Kogane, 4, 30 - 32." type="journal article" year="2003">Muramoto, 2003</bibRefCitation>
</taxonomicName>
(South
<collectingCountry id="F33D7631FF90F705FE312303358DF8A8" box="[396,495,1807,1834]" name="Vietnam" pageId="4" pageNumber="497">Vietnam</collectingCountry>
), and
<taxonomicName id="4C2A4D22FF90F705FD8B2303302BF8AB" authority="Muramoto &amp; Kobayashi, 2019" authorityName="Muramoto &amp; Kobayashi" authorityYear="2019" box="[566,1097,1807,1833]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B95EEAB3FF90F705FD8B23033683F8AA" box="[566,737,1807,1832]" italics="true" pageId="4" pageNumber="497">D. vietnamicus</emphasis>
<bibRefCitation id="EFBB4B50FF90F705FD572303302BF8AB" author="Muramoto, R. &amp; Kobayashi H." box="[746,1097,1807,1833]" pageId="4" pageNumber="497" pagination="325 - 330" refId="ref10417" refString="Muramoto, R. &amp; Kobayashi H. (2019) Two new species of the tribe Rutelini, subtribe Didrepanephorina (Coleoptera, Scarabaeidae, Rutelinae) from Vietnam and India. Elytra, New Series, 9 (2), 325 - 330." type="journal article" year="2019">Muramoto &amp; Kobayashi, 2019</bibRefCitation>
</taxonomicName>
(Central and South
<collectingCountry id="F33D7631FF90F705FA89230331F7F8A8" box="[1332,1429,1807,1834]" name="Vietnam" pageId="4" pageNumber="497">Vietnam</collectingCountry>
) can be subsumed in a species group (
<taxonomicName id="4C2A4D22FF90F705FD8A233F36A2F8CF" authorityName="Wood-Mason" authorityYear="1878" box="[567,704,1843,1869]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="bifalcifer">
<emphasis id="B95EEAB3FF90F705FD8A233F36A2F8CF" box="[567,704,1843,1869]" italics="true" pageId="4" pageNumber="497">D. bifalcifer</emphasis>
</taxonomicName>
group) based on the combination of the following morphological characters: lower margin of male mandible completed, without concavity before the basal tooth; upper margin of male mandible with a distinct denticle near base; pronotum of both sexes usually with tufted setae (except for
<taxonomicName id="4C2A4D22FF90F705FA3C2377357FF83A" authorityName="Muramoto &amp; Kobayashi" authorityYear="2019" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B95EEAB3FF90F705FA3C2377357FF83A" italics="true" pageId="4" pageNumber="497">D. vietnamicus</emphasis>
</taxonomicName>
); abdominal ventrites with dense setae; phallobase strongly curved in profile.
</paragraph>
<paragraph id="8B9536A1FF90F704FF7A23CF35A2FDBF" blockId="4.[151,1437,150,2013]" lastBlockId="5.[151,1437,151,573]" lastPageId="5" lastPageNumber="498" pageId="4" pageNumber="497">
<taxonomicName id="4C2A4D22FF90F705FF7A23CF367BF85E" authorityName="Muramoto &amp; Kobayashi" authorityYear="2019" box="[199,537,1987,2013]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B95EEAB3FF90F705FF7A23CF367BF85E" box="[199,537,1987,2013]" italics="true" pageId="4" pageNumber="497">Didrepanephorus vietnamicus</emphasis>
</taxonomicName>
is easily separated from the other three species by short, untufted pronotal setae and unicolor elytra (
<bibRefCitation id="EFBB4B50FF91F704FECB249B36A8FF33" author="Muramoto, R. &amp; Kobayashi H." box="[374,714,151,177]" pageId="5" pageNumber="498" pagination="325 - 330" refId="ref10417" refString="Muramoto, R. &amp; Kobayashi H. (2019) Two new species of the tribe Rutelini, subtribe Didrepanephorina (Coleoptera, Scarabaeidae, Rutelinae) from Vietnam and India. Elytra, New Series, 9 (2), 325 - 330." type="journal article" year="2019">Muramoto &amp; Kobayashi 2019</bibRefCitation>
). While,
<taxonomicName id="4C2A4D22FF91F704FC91249B37B2FF32" authorityName="Arrow" authorityYear="1921" box="[812,976,151,176]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B95EEAB3FF91F704FC91249B37B2FF32" box="[812,976,151,176]" italics="true" pageId="5" pageNumber="498">D. mucronatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A4D22FF91F704FC60249B3007FF33" authorityName="Wood-Mason" authorityYear="1878" box="[989,1125,151,177]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="bifalcifer">
<emphasis id="B95EEAB3FF91F704FC60249B3007FF33" box="[989,1125,151,177]" italics="true" pageId="5" pageNumber="498">D. bifalcifer</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A4D22FF91F704FB24249B3171FF33" authorityName="Muramoto" authorityYear="2003" box="[1177,1299,151,177]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="arnaudi">
<emphasis id="B95EEAB3FF91F704FB24249B3171FF33" box="[1177,1299,151,177]" italics="true" pageId="5" pageNumber="498">D. arnaudi</emphasis>
</taxonomicName>
possess distinct elongated dark patches on distomarginal area and apical umbone, which abruptly discard above distal declivity on elytra. Regarding males of the other three species of this group,
<taxonomicName id="4C2A4D22FF91F704FC0E24D3303BFF7A" authorityName="Arrow" authorityYear="1921" box="[947,1113,223,248]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B95EEAB3FF91F704FC0E24D3303BFF7A" box="[947,1113,223,248]" italics="true" pageId="5" pageNumber="498">D. mucronatus</emphasis>
</taxonomicName>
can be readily differentiated from
<taxonomicName id="4C2A4D22FF91F704FF6B250F3502FE9F" authorityName="Wood-Mason" authorityYear="1878" box="[214,352,259,285]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="bifalcifer">
<emphasis id="B95EEAB3FF91F704FF6B250F3502FE9F" box="[214,352,259,285]" italics="true" pageId="5" pageNumber="498">D. bifalcifer</emphasis>
</taxonomicName>
by the mandibles curved in dorsal view (
<figureCitation id="13112A24FF91F704FC8B250F3719FE9F" box="[822,891,259,285]" captionStart="FIGURES 14" captionStartId="2.[151,264,2001,2025]" captionTargetBox="[213,1380,320,1964]" captionTargetId="figure-84@2.[196,1390,306,1975]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Didrepanephorus mucronatus Arrow, 1921 (dorsal and ventral view). 12, male; 34, female." figureDoi="http://doi.org/10.5281/zenodo.4556622" httpUri="https://zenodo.org/record/4556622/files/figure.png" pageId="5" pageNumber="498">Fig. 1</figureCitation>
; straight in
<taxonomicName id="4C2A4D22FF91F704FBBE250F30EFFE9F" authorityName="Wood-Mason" authorityYear="1878" box="[1027,1165,259,285]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="bifalcifer">
<emphasis id="B95EEAB3FF91F704FBBE250F30EFFE9F" box="[1027,1165,259,285]" italics="true" pageId="5" pageNumber="498">D. bifalcifer</emphasis>
</taxonomicName>
,
<figureCitation id="13112A24FF91F704FB21250F3092FE9F" box="[1180,1264,259,285]" captionStart="FIGURES 4156" captionStartId="15.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1882]" captionTargetId="figure-17@15.[164,1423,181,1885]" captionTargetPageId="15" captionText="FIGURES 4156. Type specimens of Didrepanephorus Wood-Mason, 1878 species. 4146, syntype, labels, and male genitalia of D. bifalcifer Wood-Mason, 1878 (male, BMNH, photograph by Chuan-Bu Gao); 4752, holotype, labels, and male genitalia of D. mucronatus Arrow, 1921 (male, BMNH, photograph by Chuan-Bu Gao); 5356, holotype and labels of D. subvittatus Benderitter, 1922 (female, IRSNB, photograph by Matthias Seidel)." figureDoi="http://doi.org/10.5281/zenodo.4556638" httpUri="https://zenodo.org/record/4556638/files/figure.png" pageId="5" pageNumber="498">Fig. 41</figureCitation>
), the basal half of the side of pronotum distinctly concave (
<figureCitation id="13112A24FF91F704FD3E252B36A5FEC3" box="[643,711,295,321]" captionStart="FIGURES 14" captionStartId="2.[151,264,2001,2025]" captionTargetBox="[213,1380,320,1964]" captionTargetId="figure-84@2.[196,1390,306,1975]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Didrepanephorus mucronatus Arrow, 1921 (dorsal and ventral view). 12, male; 34, female." figureDoi="http://doi.org/10.5281/zenodo.4556622" httpUri="https://zenodo.org/record/4556622/files/figure.png" pageId="5" pageNumber="498">Fig. 1</figureCitation>
; slightly concave in
<taxonomicName id="4C2A4D22FF91F704FC0F252B305EFEC3" authorityName="Wood-Mason" authorityYear="1878" box="[946,1084,295,321]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="bifalcifer">
<emphasis id="B95EEAB3FF91F704FC0F252B305EFEC3" box="[946,1084,295,321]" italics="true" pageId="5" pageNumber="498">D. bifalcifer</emphasis>
</taxonomicName>
,
<figureCitation id="13112A24FF91F704FBF7252B30FFFEC3" box="[1098,1181,295,321]" captionStart="FIGURES 4156" captionStartId="15.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1882]" captionTargetId="figure-17@15.[164,1423,181,1885]" captionTargetPageId="15" captionText="FIGURES 4156. Type specimens of Didrepanephorus Wood-Mason, 1878 species. 4146, syntype, labels, and male genitalia of D. bifalcifer Wood-Mason, 1878 (male, BMNH, photograph by Chuan-Bu Gao); 4752, holotype, labels, and male genitalia of D. mucronatus Arrow, 1921 (male, BMNH, photograph by Chuan-Bu Gao); 5356, holotype and labels of D. subvittatus Benderitter, 1922 (female, IRSNB, photograph by Matthias Seidel)." figureDoi="http://doi.org/10.5281/zenodo.4556638" httpUri="https://zenodo.org/record/4556638/files/figure.png" pageId="5" pageNumber="498">Fig. 44</figureCitation>
), the disc of pronotum with two distinct tufted of dense setae (
<figureCitation id="13112A24FF91F704FDE4254736A5FEE7" box="[601,711,330,357]" captionStart="FIGURES 510" captionStartId="3.[151,264,1215,1239]" captionTargetBox="[155,1430,193,1182]" captionTargetId="figure-17@3.[151,1435,181,1192]" captionTargetPageId="3" captionText="FIGURES 510. Habitus of Didrepanephorus mucronatus Arrow, 1921 (oblique view). 58, males in different body sizes; 910, females in different body sizes." figureDoi="http://doi.org/10.5281/zenodo.4556624" httpUri="https://zenodo.org/record/4556624/files/figure.png" pageId="5" pageNumber="498">Figs. 58</figureCitation>
; indistinct in
<taxonomicName id="4C2A4D22FF91F704FCDE2547378FFEE7" authorityName="Wood-Mason" authorityYear="1878" box="[867,1005,331,357]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="bifalcifer">
<emphasis id="B95EEAB3FF91F704FCDE2547378FFEE7" box="[867,1005,331,357]" italics="true" pageId="5" pageNumber="498">D. bifalcifer</emphasis>
</taxonomicName>
,
<figureCitation id="13112A24FF91F704FC4125473032FEE7" box="[1020,1104,331,357]" captionStart="FIGURES 4156" captionStartId="15.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1882]" captionTargetId="figure-17@15.[164,1423,181,1885]" captionTargetPageId="15" captionText="FIGURES 4156. Type specimens of Didrepanephorus Wood-Mason, 1878 species. 4146, syntype, labels, and male genitalia of D. bifalcifer Wood-Mason, 1878 (male, BMNH, photograph by Chuan-Bu Gao); 4752, holotype, labels, and male genitalia of D. mucronatus Arrow, 1921 (male, BMNH, photograph by Chuan-Bu Gao); 5356, holotype and labels of D. subvittatus Benderitter, 1922 (female, IRSNB, photograph by Matthias Seidel)." figureDoi="http://doi.org/10.5281/zenodo.4556638" httpUri="https://zenodo.org/record/4556638/files/figure.png" pageId="5" pageNumber="498">Fig. 44</figureCitation>
), and the shape of parameres (see
<figureCitation id="13112A24FF91F704FF7125633534FE0B" box="[204,342,366,393]" captionStart="FIGURES 4156" captionStartId="15.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1882]" captionTargetId="figure-17@15.[164,1423,181,1885]" captionTargetPageId="15" captionText="FIGURES 4156. Type specimens of Didrepanephorus Wood-Mason, 1878 species. 4146, syntype, labels, and male genitalia of D. bifalcifer Wood-Mason, 1878 (male, BMNH, photograph by Chuan-Bu Gao); 4752, holotype, labels, and male genitalia of D. mucronatus Arrow, 1921 (male, BMNH, photograph by Chuan-Bu Gao); 5356, holotype and labels of D. subvittatus Benderitter, 1922 (female, IRSNB, photograph by Matthias Seidel)." figureDoi="http://doi.org/10.5281/zenodo.4556638" httpUri="https://zenodo.org/record/4556638/files/figure.png" pageId="5" pageNumber="498">Figs. 45, 51</figureCitation>
). The following features of
<taxonomicName id="4C2A4D22FF91F704FD2E2563376DFE0B" authorityName="Muramoto" authorityYear="2003" box="[659,783,367,393]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="arnaudi">
<emphasis id="B95EEAB3FF91F704FD2E2563376DFE0B" box="[659,783,367,393]" italics="true" pageId="5" pageNumber="498">D. arnaudi</emphasis>
</taxonomicName>
indicate its separation from
<taxonomicName id="4C2A4D22FF91F704FBE925633098FE0A" authorityName="Arrow" authorityYear="1921" box="[1108,1274,367,392]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B95EEAB3FF91F704FBE925633098FE0A" box="[1108,1274,367,392]" italics="true" pageId="5" pageNumber="498">D. mucronatus</emphasis>
</taxonomicName>
: the mandible more strongly curved in dorsal view, the basal half of the left paramere with an additional subtriangular extension, and the body color somewhat darker (
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;
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). The females of the three species are less informative and reliable separations are not available currently. The differential points as indicated by
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,
<emphasis id="B95EEAB3FF91F704FE2B25F335DFFD9A" box="[406,445,511,536]" italics="true" pageId="5" pageNumber="498">viz.</emphasis>
body color and shape of pronotum, are fairly variable when referring to the examined females of
<taxonomicName id="4C2A4D22FF91F704FEA9262F35DBFDBE" authorityName="Arrow" authorityYear="1921" box="[276,441,547,572]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B95EEAB3FF91F704FEA9262F35DBFDBE" box="[276,441,547,572]" italics="true" pageId="5" pageNumber="498">D. mucronatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>