treatments-xml/data/03/83/87/038387B7FF91F709FF2A26823611F849.xml

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<mods:title id="5566F6E51C9B644C002FE42C6EDE5A6C">Description of a new species of the genus Didrepanephorus Wood-Mason, 1878 (Coleoptera: Scarabaeidae: Rutelinae) from China with notes on allied species</mods:title>
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<mods:namePart id="69324CF89F7E33B0E8B7A72F77108A34">Qiu, Jian-Yue</mods:namePart>
<mods:affiliation id="0553E5527B46DCF3A6D93F805B4BE451">Engineering Research Center for Forest and Grassland Disaster Prevention and Reduction, Mianyang Normal University, 166 Mi- anxing West Road, Mianyang, Sichuan 621000, P. R. China.</mods:affiliation>
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<mods:namePart id="CC35FFB10F361DDBA514211321D07D2C">Zhao, Ming-Zhi</mods:namePart>
<mods:affiliation id="D8C8AF8B1E739BF5DA61FDFE13AF1B21">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, P. R. China. zhaomzhai @ 126. com; https: // orcid. org // 0000 - 0002 - 3158 - 6177</mods:affiliation>
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<mods:namePart id="B968178E0B0E1296917ABEA162255BBD">Xu, Hao</mods:namePart>
<mods:affiliation id="699FD8EB7ACE0CE25EF043141FEA262B">Engineering Research Center for Forest and Grassland Disaster Prevention and Reduction, Mianyang Normal University, 166 Mi- anxing West Road, Mianyang, Sichuan 621000, P. R. China.</mods:affiliation>
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<taxonomicName id="4C2A4D22FF91F704FF2A268236B0FD2B" authority="Qiu, Zhao &amp; Xu" authorityName="Qiu, Zhao &amp; Xu" authorityYear="2021" box="[151,722,654,681]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="heterocolor" status="sp. nov.">
<emphasis id="B95EEAB3FF91F704FF2A2682359FFD2B" bold="true" box="[151,509,654,681]" italics="true" pageId="5" pageNumber="498">Didrepanephorus heterocolor</emphasis>
Qiu, Zhao &amp; Xu
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-•镰'•龟
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(
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, 
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, 
<figureCitation id="13112A24FF91F704FE3B26BE35ACFD4F" box="[390,462,690,717]" captionStart="FIGURES 57–64" captionStartId="16.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1885]" captionTargetId="figure-23@16.[164,1423,181,1885]" captionTargetPageId="16" captionText="FIGURES 57–64. Habitat and different stages of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species. 57, forest in Maolan Nature Reserve (Guizhou, China); 58–59, a closer view of habitat and pupal cells in rotten wood (both photograph by Ri-Xin Jiang); 60, a larva under the bark of decayed wood in wild; 61–62, pupa and newly emerged female in pupal cells in artificial conditions; 63, male in a pupal cell in rotten wood (photograph by Ri-Xin Jiang); 64, egg in artificial conditions." figureDoi="http://doi.org/10.5281/zenodo.4556640" httpUri="https://zenodo.org/record/4556640/files/figure.png" pageId="5" pageNumber="498">57–64</figureCitation>
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<figureCitation id="13112A24FF91F704FE6726BF3595FD4F" box="[474,503,691,717]" captionStart="FIGURE 70" captionStartId="18.[151,250,1036,1060]" captionTargetBox="[151,1435,181,1012]" captionTargetId="figure-23@18.[151,1435,181,1012]" captionTargetPageId="18" captionText="FIGURE 70. Known distribution of four Didrepanephorus Wood-Mason, 1878 species. Type locality of D. mucronatus Arrow, 1921 is missing because the precise location is unknown." figureDoi="http://doi.org/10.5281/zenodo.4556644" httpUri="https://zenodo.org/record/4556644/files/figure.png" pageId="5" pageNumber="498">70</figureCitation>
)
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<emphasis id="B95EEAB3FF91F704FF2A26F73524FC96" bold="true" box="[151,326,762,789]" pageId="5" pageNumber="498">Type material.</emphasis>
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JIANG leg. // pupa in 
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: 
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</quantity>
, 
<location id="8EF5607AFF91F704FF5027873559FC27" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FF5027873559FC27" box="[237,315,907,933]" country="China" county="Libo County" latitude="25.297499" longLatPrecision="21" longitude="108.07444" municipality="Maolan Nature Reserve" name="Ri-Xin" pageId="5" pageNumber="498" stateProvince="Guizhou">Ri-Xin</location>
JIANG leg.; 4♂♂
</materialsCitation>
, 
<materialsCitation id="3B423CFCFF91F704FDA62786368EFC49" ID-GBIF-Occurrence="3050816320" collectionCode="SHNU" country="China" location="Ri-Xin" municipality="Maolan Nature Reserve" pageId="5" pageNumber="498" specimenCode="F1" specimenCount="5" specimenCount-female="5" typeStatus="paratype">
<specimenCount id="9D2CFD28FF91F704FDA627863631FC27" box="[539,595,906,933]" pageId="5" pageNumber="498" type="female">5♀♀</specimenCount>
(
<collectionCode id="ED3BAE64FF91F704FDDF278736D6FC27" box="[610,692,907,933]" pageId="5" pageNumber="498">SHNU</collectionCode>
), 
<specimenCode id="DB8C9EDAFF91F704FD7527873682FC29" box="[712,736,907,939]" collectionCode="SHNU" pageId="5" pageNumber="498">
F 
<subScript id="17AE34E4FF91F704FD6527913682FC29" attach="left" box="[728,736,925,939]" fontSize="6" pageId="5" pageNumber="498">1</subScript>
</specimenCode>
generation of one female from 
<collectingMunicipality id="6BF1ACDBFF91F704FBFE27873133FC27" box="[1091,1361,907,933]" pageId="5" pageNumber="498">Maolan Nature Reserve</collectingMunicipality>
, 2017, 
<location id="8EF5607AFF91F704FF2A27A33487FC4B" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FF2A27A33487FC4B" box="[151,229,943,969]" country="China" municipality="Maolan Nature Reserve" name="Ri-Xin" pageId="5" pageNumber="498">Ri-Xin</location>
JIANG leg. // adult emerged in 2018.II; 2♂♂
</materialsCitation>
, 
<materialsCitation id="3B423CFCFF91F704FD4627A33497FB93" ID-GBIF-Occurrence="3050816315" collectingDate="2012-08-11" collectionCode="MYNU" collectorName="Jian-Yue &amp; Hao XU" country="China" county="Libo County" elevation="800" location="La'nei" municipality="Maolan Nature Reserve" pageId="5" pageNumber="498" specimenCount="2" specimenCount-female="2" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="9D2CFD28FF91F704FD4627A33751FC4B" box="[763,819,943,969]" pageId="5" pageNumber="498" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED3BAE64FF91F704FCFF27A337FFFC4B" box="[834,925,943,969]" pageId="5" pageNumber="498">MYNU</collectionCode>
), 
<collectingMunicipality id="6BF1ACDBFF91F704FC0C27A330A2FC4B" box="[945,1216,943,969]" pageId="5" pageNumber="498">Maolan Nature Reserve</collectingMunicipality>
, 
<location id="8EF5607AFF91F704FB7627A33175FC4B" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FB7627A33175FC4B" box="[1227,1303,943,969]" country="China" county="Libo County" municipality="Maolan Nature Reserve" name="La'nei" pageId="5" pageNumber="498" stateProvince="Guizhou">La’nei</location>
, 
<location id="8EF5607AFF91F704FA9E27A331FAFC4B" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FA9E27A331FAFC4B" box="[1315,1432,943,969]" country="China" county="Libo County" municipality="Maolan Nature Reserve" name="Weng'ang" pageId="5" pageNumber="498" stateProvince="Guizhou">Weng’ang</location>
, 
<collectingCounty id="62F44E2DFF91F704FF2A27DF3545FC6F" box="[151,295,979,1005]" pageId="5" pageNumber="498">Libo County</collectingCounty>
, 
<collectingRegion id="49EEF843FF91F704FE8F27DF35F7FC6F" box="[306,405,979,1005]" country="China" name="Guizhou" pageId="5" pageNumber="498">Guizhou</collectingRegion>
, alt. 
<quantity id="4CD29B44FF91F704FE7127DF3677FC6F" box="[460,533,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="498" unit="m" value="800.0">
<elevation id="0007D192FF91F704FE7127DF3677FC6F" box="[460,533,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="498" unit="m" value="800.0">800 m</elevation>
</quantity>
, 
<collectorName id="26DF5377FF91F704FD9C27DF36E1FC6F" box="[545,643,979,1005]" pageId="5" pageNumber="498">Jian-Yue</collectorName>
QIU &amp; 
<collectorName id="26DF5377FF91F704FD5C27DF375DFC6F" box="[737,831,979,1005]" pageId="5" pageNumber="498">Hao XU</collectorName>
leg. // mature larva in 
<date id="FF941061FF91F704FBFD27DF30B6FC6F" box="[1088,1236,979,1005]" pageId="5" pageNumber="498" value="2012-08-11">
<collectingDate id="EFD0E989FF91F704FBFD27DF30B6FC6F" box="[1088,1236,979,1005]" pageId="5" pageNumber="498" value="2012-08-11">2012.VIII.11</collectingDate>
</date>
, adult emerged in 2013.IV
</materialsCitation>
; 
<materialsCitation id="3B423CFCFF91F704FEBD27FB367DFBB5" ID-GBIF-Occurrence="3050816313" collectionCode="MYNU" collectorName="Hao XU" country="China" location="Jian-Yue" municipality="Maolan Nature Reserve" pageId="5" pageNumber="498" specimenCode="F1" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="9D2CFD28FF91F704FEBD27FB3555FB93" box="[256,311,1015,1041]" pageId="5" pageNumber="498" type="female">3♀♀</specimenCount>
(
<collectionCode id="ED3BAE64FF91F704FEFB27FB35C3FB93" box="[326,417,1015,1041]" pageId="5" pageNumber="498">MYNU</collectionCode>
), 
<specimenCode id="DB8C9EDAFF91F704FE0E27FB35A9FB95" box="[435,459,1015,1047]" collectionCode="MYNU" pageId="5" pageNumber="498">
F 
<subScript id="17AE34E4FF91F704FE7E200535A9FB95" attach="left" box="[451,459,1033,1047]" fontSize="6" pageId="5" pageNumber="498">1</subScript>
</specimenCode>
generation of one female from 
<collectingMunicipality id="6BF1ACDBFF91F704FC9527FB3051FB93" box="[808,1075,1015,1041]" pageId="5" pageNumber="498">Maolan Nature Reserve</collectingMunicipality>
, 2012, 
<location id="8EF5607AFF91F704FB3E27FB3087FB93" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FB3E27FB3087FB93" box="[1155,1253,1015,1041]" country="China" municipality="Maolan Nature Reserve" name="Jian-Yue" pageId="5" pageNumber="498">Jian-Yue</location>
QIU &amp; 
<collectorName id="26DF5377FF91F704FA8227FB31FEFB93" box="[1343,1436,1015,1041]" pageId="5" pageNumber="498">Hao XU</collectorName>
leg. // adult emerged in 2014.V; 1♂
</materialsCitation>
, 
<materialsCitation id="3B423CFCFF91F704FD93201730D3FBDB" ID-GBIF-Occurrence="3050816308" collectingDate="2013-07-25" collectionCode="MYNU" collectorName="Jian-Yue &amp; Hao XU" country="China" county="Libo County" location="La'nei" municipality="Maolan Nature Reserve" pageId="5" pageNumber="498" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
<specimenCount id="9D2CFD28FF91F704FD9320173604FBB7" box="[558,614,1051,1077]" pageId="5" pageNumber="498" type="female">4♀♀</specimenCount>
(
<collectionCode id="ED3BAE64FF91F704FDC8201736B2FBB7" box="[629,720,1051,1077]" pageId="5" pageNumber="498">MYNU</collectionCode>
), 
<collectingMunicipality id="6BF1ACDBFF91F704FD5E2017378DFBB7" box="[739,1007,1051,1077]" pageId="5" pageNumber="498">Maolan Nature Reserve</collectingMunicipality>
, 
<location id="8EF5607AFF91F704FC4420173027FBB7" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FC4420173027FBB7" box="[1017,1093,1051,1077]" country="China" county="Libo County" municipality="Maolan Nature Reserve" name="La'nei" pageId="5" pageNumber="498">La’nei</location>
, 
<location id="8EF5607AFF91F704FBF2201730A6FBB7" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FBF2201730A6FBB7" box="[1103,1220,1051,1077]" country="China" county="Libo County" municipality="Maolan Nature Reserve" name="Weng'ang" pageId="5" pageNumber="498">Weng’ang</location>
, 
<collectingCounty id="62F44E2DFF91F704FB722017313DFBB7" box="[1231,1375,1051,1077]" pageId="5" pageNumber="498">Libo County</collectingCounty>
, Guizhou, 
<collectorName id="26DF5377FF91F704FF612033355CFBDB" box="[220,318,1087,1113]" pageId="5" pageNumber="498">Jian-Yue</collectorName>
QIU &amp; 
<collectorName id="26DF5377FF91F704FE262033359BFBDB" box="[411,505,1087,1113]" pageId="5" pageNumber="498">Hao XU</collectorName>
leg. // mature larva in 
<date id="FF941061FF91F704FD44203237E6FBDB" box="[761,900,1086,1113]" pageId="5" pageNumber="498" value="2013-07-25">
<collectingDate id="EFD0E989FF91F704FD44203237E6FBDB" box="[761,900,1086,1113]" pageId="5" pageNumber="498" value="2013-07-25">2013.VII.25</collectingDate>
</date>
, adult emerged in 2014.IV
</materialsCitation>
; 
<materialsCitation id="3B423CFCFF91F704FB01203331F7FBFD" ID-GBIF-Occurrence="3050816318" collectingDate="2018-06-12" collectionCode="MYNU" collectorName="Hao XU" country="China" county="Libo County" elevation="790" location="Dongduo" municipality="Maolan Nature Reserve" pageId="5" pageNumber="498" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="9D2CFD28FF91F704FB01203330BDFBDB" box="[1212,1247,1087,1113]" pageId="5" pageNumber="498" type="female">1♀</specimenCount>
(
<collectionCode id="ED3BAE64FF91F704FB5220333128FBDB" box="[1263,1354,1087,1113]" pageId="5" pageNumber="498">MYNU</collectionCode>
), 
<date id="FF941061FF91F704FAE0203334B8FBFF" pageId="5" pageNumber="498" value="2018-06-12">
<collectingDate id="EFD0E989FF91F704FAE0203334B8FBFF" pageId="5" pageNumber="498" value="2018-06-12">2018. VI.12</collectingDate>
</date>
, 
<collectingMunicipality id="6BF1ACDBFF91F704FF5B206F3591FBFF" box="[230,499,1123,1149]" pageId="5" pageNumber="498">Maolan Nature Reserve</collectingMunicipality>
, 
<location id="8EF5607AFF91F704FE42206F3608FBFF" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FE42206F3608FBFF" box="[511,618,1123,1149]" country="China" county="Libo County" municipality="Maolan Nature Reserve" name="Dongduo" pageId="5" pageNumber="498" stateProvince="Guizhou">Dongduo</location>
, 
<collectingCounty id="62F44E2DFF91F704FDCB206F3764FBFF" box="[630,774,1123,1149]" pageId="5" pageNumber="498">Libo County</collectingCounty>
, 
<collectingRegion id="49EEF843FF91F704FCAC206F3716FBFF" box="[785,884,1123,1149]" country="China" name="Guizhou" pageId="5" pageNumber="498">Guizhou</collectingRegion>
, alt. 
<quantity id="4CD29B44FF91F704FC17206F3791FBFF" box="[938,1011,1123,1149]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="5" pageNumber="498" unit="m" value="790.0">
<elevation id="0007D192FF91F704FC17206F3791FBFF" box="[938,1011,1123,1149]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="5" pageNumber="498" unit="m" value="790.0">790 m</elevation>
</quantity>
, 
<location id="8EF5607AFF91F704FC42206F3003FBFF" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FC42206F3003FBFF" box="[1023,1121,1123,1149]" country="China" county="Libo County" municipality="Maolan Nature Reserve" name="Jian-Yue" pageId="5" pageNumber="498" stateProvince="Guizhou">Jian-Yue</location>
QIU &amp; 
<collectorName id="26DF5377FF91F704FB02206F317FFBFF" box="[1215,1309,1123,1149]" pageId="5" pageNumber="498">Hao XU</collectorName>
leg.; 4♂♂
</materialsCitation>
, 
<materialsCitation id="3B423CFCFF91F704FF2A208B3692FB6B" ID-GBIF-Occurrence="3050816324" collectingDate="1988-05-24" collectingDateMax="1990-05-14" collectingDateMin="1988-05-24" collectionCode="GFGY" country="China" county="Libo County" elevation="540" location="Banzhai" municipality="Maolan Nature Reserve" pageId="5" pageNumber="498" specimenCount="3" specimenCount-female="3" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="9D2CFD28FF91F704FF2A208B34ADFB23" box="[151,207,1159,1185]" pageId="5" pageNumber="498" type="female">3♀♀</specimenCount>
(
<collectionCode id="ED3BAE64FF91F704FF63208B3552FB23" box="[222,304,1159,1185]" pageId="5" pageNumber="498">GFGY</collectionCode>
), 
<date id="FF941061FF91F704FEFF208B35D5FB23" box="[322,439,1159,1185]" pageId="5" pageNumber="498" value="1988-05-24">
<collectingDate id="EFD0E989FF91F704FEFF208B35D5FB23" box="[322,439,1159,1185]" pageId="5" pageNumber="498" value="1988-05-24">1988.V.24</collectingDate>
</date>
, 
<collectingMunicipality id="6BF1ACDBFF91F704FE7F208B36ACFB23" box="[450,718,1159,1185]" pageId="5" pageNumber="498">Maolan Nature Reserve</collectingMunicipality>
, 
<location id="8EF5607AFF91F704FD64208B375AFB23" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FD64208B375AFB23" box="[729,824,1159,1185]" country="China" county="Libo County" municipality="Maolan Nature Reserve" name="Banzhai" pageId="5" pageNumber="498" stateProvince="Guizhou">Banzhai</location>
, 
<collectingCounty id="62F44E2DFF91F704FCFF208B37B3FB23" box="[834,977,1159,1185]" pageId="5" pageNumber="498">Libo County</collectingCounty>
, 
<collectingRegion id="49EEF843FF91F704FC66208B305CFB23" box="[987,1086,1159,1185]" country="China" name="Guizhou" pageId="5" pageNumber="498">Guizhou</collectingRegion>
, alt. 
<quantity id="4CD29B44FF91F704FBCE208A30DEFB23" box="[1139,1212,1158,1185]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="5" pageNumber="498" unit="m" value="540.0">
<elevation id="0007D192FF91F704FBCE208A30DEFB23" box="[1139,1212,1158,1185]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="5" pageNumber="498" unit="m" value="540.0">540 m</elevation>
</quantity>
; 1♂ (
<collectionCode id="ED3BAE64FF91F704FB44208B3129FB23" box="[1273,1355,1159,1185]" pageId="5" pageNumber="498">GFGY</collectionCode>
), 
<date id="FF941061FF91F704FAE0208B34AFFB47" pageId="5" pageNumber="498" value="1990-05-14">
<collectingDate id="EFD0E989FF91F704FAE0208B34AFFB47" pageId="5" pageNumber="498" value="1990-05-14">1990. V.14</collectingDate>
</date>
, 
<location id="8EF5607AFF91F704FF6620A7358EFB47" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FF6620A7358EFB47" box="[219,492,1195,1221]" country="China" county="Libo County" municipality="Maolan Nature Reserve" name="Maolan Nature Reserve" pageId="5" pageNumber="498" stateProvince="Guizhou">Maolan Nature Reserve</location>
, 
<location id="8EF5607AFF91F704FE4420A73637FB47" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FE4420A73637FB47" box="[505,597,1195,1221]" country="China" county="Libo County" municipality="Maolan Nature Reserve" name="Banzhai" pageId="5" pageNumber="498" stateProvince="Guizhou">Banzhai</location>
to 
<location id="8EF5607AFF91F704FDC020A7376BFB47" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FDC020A7376BFB47" box="[637,777,1195,1221]" country="China" county="Libo County" municipality="Maolan Nature Reserve" name="Limingguan" pageId="5" pageNumber="498" stateProvince="Guizhou">Limingguan</location>
, 
<location id="8EF5607AFF91F704FCAB20A737CAFB47" LSID="urn:lsid:plazi:treatment:038387B7FF91F709FF2A26823611F849:8EF5607AFF91F704FCAB20A737CAFB47" box="[790,936,1195,1221]" country="China" county="Libo County" municipality="Maolan Nature Reserve" name="Libo County" pageId="5" pageNumber="498" stateProvince="Guizhou">Libo County</location>
, 
<collectingRegion id="49EEF843FF91F704FC0820A7307AFB47" box="[949,1048,1195,1221]" country="China" name="Guizhou" pageId="5" pageNumber="498">Guizhou</collectingRegion>
, alt. 
<quantity id="4CD29B44FF91F704FBEF20A630FCFB47" box="[1106,1182,1194,1221]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="5" pageNumber="498" unit="m" value="570.0">
<elevation id="0007D192FF91F704FBEF20A630FCFB47" box="[1106,1182,1194,1221]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="5" pageNumber="498" unit="m" value="570.0">570 m</elevation>
</quantity>
; 1♂ (
<collectionCode id="ED3BAE64FF91F704FB6220A73153FB47" box="[1247,1329,1195,1221]" pageId="5" pageNumber="498">GFGY</collectionCode>
), without label but should be also from the same nature reserve
</materialsCitation>
.
</paragraph>
<paragraph id="8B9536A1FF91F704FF7A20FE369DFAB3" blockId="5.[151,1437,762,2013]" pageId="5" pageNumber="498">
<materialsCitation id="3B423CFCFF91F704FF7A20FE3699FAB3" ID-GBIF-Occurrence="3050816303" collectingDate="2017-04-29" collectionCode="MYNU" county="Libo County" elevation="679" latitude="25.297499" location="Maolan Nature Reserve" longLatPrecision="21" longitude="108.07444" municipality="Ri-Xin" pageId="5" pageNumber="498" specimenCount="3" specimenCount-female="3">
<emphasis id="B95EEAB3FF91F704FF7A20FE365AFA8E" bold="true" box="[199,568,1266,1292]" pageId="5" pageNumber="498">Additional material examined.</emphasis>
<specimenCount id="9D2CFD28FF91F704FDFC20FF361BFA8F" box="[577,633,1267,1293]" pageId="5" pageNumber="498" type="female">3♀♀</specimenCount>
(
<collectionCode id="ED3BAE64FF91F704FD3720FF3687FA8F" box="[650,741,1267,1293]" pageId="5" pageNumber="498">MYNU</collectionCode>
), same label data as holotype. These females were deformed during eclosion and thus removed from the type series
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330652AFF91F707FF7A21363082F98B" lastPageId="6" lastPageNumber="499" pageId="5" pageNumber="498" type="description">
<paragraph id="8B9536A1FF91F707FF7A213631E7FB93" blockId="5.[151,1437,762,2013]" lastBlockId="6.[151,1437,150,2013]" lastPageId="6" lastPageNumber="499" pageId="5" pageNumber="498">
<emphasis id="B95EEAB3FF91F704FF7A213636B5FAD6" bold="true" box="[199,727,1338,1365]" pageId="5" pageNumber="498">
Description of the 
<typeStatus id="54918803FF91F704FE182137366EFAD7" box="[421,524,1339,1365]" pageId="5" pageNumber="498" type="holotype">holotype</typeStatus>
(male). General:
</emphasis>
Body oval and convex, hairs and setae yellowish-brown (
<figureCitation id="13112A24FF91F704FAD921373482FAFB" captionStart="FIGURES 11–14" captionStartId="7.[151,264,1944,1968]" captionTargetBox="[182,1406,185,1905]" captionTargetId="figure-17@7.[169,1418,181,1920]" captionTargetPageId="7" captionText="FIGURES 11–14. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (dorsal and ventral view). 11–12, male holotype; 13–14, female allotype." figureDoi="http://doi.org/10.5281/zenodo.4556626" httpUri="https://zenodo.org/record/4556626/files/figure.png" pageId="5" pageNumber="498">Figs. 11–12</figureCitation>
). 
<emphasis id="B95EEAB3FF91F704FF482153355CFAFB" bold="true" box="[245,318,1375,1401]" pageId="5" pageNumber="498">Head:</emphasis>
Clypeus reddish-brown, other portions dark brown. Clypeus flat, trapezoidal, anterior margin feebly arcuate; anterior angles rounded, sides strongly convergent anteriad and expanded in basal half, concave near middle, then almost subparallel in apical half; with dense large punctures, coarse but not punctured near anterior margin; anterior margin with some short setae. Frontal-clypeal suture almost absent, represented by a short carina at each side. Frons and vertex with dense large punctures and dense, erect long setae. Eye-canthus wide and truncate at outer margin, not extends beyond outermost point of eye, anterior angle nearly right angle. Antennal scapus reddish-brown, base of antennal club dark brown, gradually changed fulvous to apex, length of antennal club distinctly longer than antennomeres 2–7 combined. Labrum exposed, anterior margin deflected and concave at middle, dorsal surface sparsely microsculptured, with sparse, short and long setae. Mandible dark brown, broadest at base, lower margin deeply constricted before basal tooth, apex sharply bended upward. Maxillary palpi and labial palpi dark brown, simple. 
<collectingRegion id="49EEF843FF91F704FEF922CB35C1F963" box="[324,419,1735,1761]" country="Myanmar" name="Chin" pageId="5" pageNumber="498">Mentum</collectingRegion>
with dense punctures and long erect setae, anterior margin slightly concave in middle. Gula dark brown, glabrous and matt. 
<emphasis id="B95EEAB3FF91F704FE4422E73619F887" bold="true" box="[505,635,1771,1797]" pageId="5" pageNumber="498">Pronotum:</emphasis>
Dark brown, each side with an indistinct orange-brown patch near middle. Strongly convex, 1.45 times as wide as long, widest near middle. Anterior margin distinctly bisinuate; anterior marginal membrane distinct, obsolete before reaching anterior angle. Sides feebly concave and slightly convergent posteriad in basal half, roundly and broadly curved at middle, strongly convergent anteriad in apical half. Posterior margin broadly protruding in middle. Basal marginal line interrupted before scutellum, all other marginal lines completed. Anterior angle moderately protruding, posterior angle slightly protruding, both blunt at apex. Surface with dense small punctures, rugopunctate along lateral margin, disc with a longitudinal medial line. Most areas of disc with rather dense semierect long setae, setae on lateral portions being shorter. 
<emphasis id="B95EEAB3FF92F707FB84249A30D8FF32" bold="true" box="[1081,1210,150,176]" pageId="6" pageNumber="499">Scutellum:</emphasis>
Black. Nearly short triangular, lateral margin slightly curved, apex rounded. Surface with dense large punctures and short setae except for midline and margin. 
<emphasis id="B95EEAB3FF92F707FE1224D33666FF7B" bold="true" box="[431,516,223,249]" pageId="6" pageNumber="499">Elytra:</emphasis>
Convex. Margin, suture and apical umbone black. Almost as wide as long, widest at middle. Primary stria 1 clearly defined by a row of strial punctures, other intervals unrecognizable. Humeral umbone and apical umbone convex. Surface coarse, with sparse, shallow, round large punctures and dense, short, semierect setae, alternated by rather sparse, short erect setae. Epipleura glabrous. 
<emphasis id="B95EEAB3FF92F707FB8C2547313EFEE6" bold="true" box="[1073,1372,330,357]" pageId="6" pageNumber="499">Ventral thoracic surface:</emphasis>
Dark brown, posterior part of ventral mesothoracic surface reddish-brown (
<figureCitation id="13112A24FF92F707FC212563378DFE0B" box="[924,1007,367,393]" captionStart="FIGURES 11–14" captionStartId="7.[151,264,1944,1968]" captionTargetBox="[182,1406,185,1905]" captionTargetId="figure-17@7.[169,1418,181,1920]" captionTargetPageId="7" captionText="FIGURES 11–14. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (dorsal and ventral view). 11–12, male holotype; 13–14, female allotype." figureDoi="http://doi.org/10.5281/zenodo.4556626" httpUri="https://zenodo.org/record/4556626/files/figure.png" pageId="6" pageNumber="499">Fig. 12</figureCitation>
). Base of ventral prothoracic and anterior half of ventral metathoracic surfaces with dense long hairs; anterior half of ventral prothoracic and posterior half of ventral mesothoracic surfaces with erect short setae. Prosternal process small, horizontally broad, shield-liked with an medial carina. Punctures on ventral mesothoracic surface round and gradually changed into sinuous striolae laterad. Metasternum moderately bulging along glabrous midline. 
<emphasis id="B95EEAB3FF92F707FCC725F23042FD9A" bold="true" box="[890,1056,510,536]" pageId="6" pageNumber="499">Propygidium:</emphasis>
Dark brown. Surface coarse, with dense, semierect short setae. 
<emphasis id="B95EEAB3FF92F707FE55262E3607FDBE" bold="true" box="[488,613,546,572]" pageId="6" pageNumber="499">Pygidium:</emphasis>
Convex. Dark brown. Disc coarse, lateral and posterior portions indistinctly and transversely puncatate. Setosus as in propygidium. 
<emphasis id="B95EEAB3FF92F707FCE4264A37B5FDE2" bold="true" box="[857,983,582,608]" pageId="6" pageNumber="499">Abdomen:</emphasis>
Dark brown, distinctly curved in lateral view, abdominal ventrites 1–4 abbreviated (
<figureCitation id="13112A24FF92F707FD38266736B8FD07" box="[645,730,619,645]" captionStart="FIGURES 11–14" captionStartId="7.[151,264,1944,1968]" captionTargetBox="[182,1406,185,1905]" captionTargetId="figure-17@7.[169,1418,181,1920]" captionTargetPageId="7" captionText="FIGURES 11–14. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (dorsal and ventral view). 11–12, male holotype; 13–14, female allotype." figureDoi="http://doi.org/10.5281/zenodo.4556626" httpUri="https://zenodo.org/record/4556626/files/figure.png" pageId="6" pageNumber="499">Fig. 12</figureCitation>
). With dense round punctures and semierect long setae. Posterior margin of abdominal ventrite 6 broadly and distinctly concave medially, with a row of dense, semierect long setae. 
<emphasis id="B95EEAB3FF92F707FF6026BF357CFD4F" bold="true" box="[221,286,691,717]" pageId="6" pageNumber="499">Legs:</emphasis>
Dark brown, with moderately dense, erect long setae. Femora with dense punctures; tibiae with dense, coarse and irregular punctures. Inner protarsal claw simple and thickened, outer protarsal claw smaller and sharp at apex. Outer meso- and metatarsal claws widely and deeply spilt into two branches respectively, the upper branches thinner and sharper; in outer mesotarsal claw the two branches equal in length, in outer metatarsal claw the upper branch slightly longer than the lower branch. Inner meso- and metatarsal claws more curved than the outer ones. Protibia with three distinct and acute teeth. Protibial spur situates at the level of middle protibial tooth, vertically orient forwards. Meso- and metatibiae bidentate at apical edges, the upper tooth in mesotibia distinctly protruding. Protarsus strongly thickened; protarsomeres 2–4 abbreviated; protarsomere 4 ventrally with a short blunt protrusion; meso- and metatarsomeres 4 each with an acute ventroapical protrusion, each protrusion with a pair of spines at base; tarsomeres 5 of all legs with a small and blunt ventral protrusion in basal third. 
<emphasis id="B95EEAB3FF92F707FBEC27FA30ACFB92" bold="true" box="[1105,1230,1014,1040]" pageId="6" pageNumber="499">Aedeagus:</emphasis>
As 
<figureCitation id="13112A24FF92F707FB4427FB31E3FB93" box="[1273,1409,1014,1041]" captionStart="FIGURES 33–40" captionStartId="13.[151,264,1635,1659]" captionTargetBox="[156,1393,201,1596]" captionTargetId="figure-17@13.[151,1435,181,1611]" captionTargetPageId="13" captionText="FIGURES 33–40. Male genitalia of Didrepanephorus Wood-Mason, 1878 species (showing parameres in dorsal and ventral views). 33–34, D. mucronatus Arrow, 1921; 35–36, D. heterocolor new species (holotype); 37–38, D. nishiyamai Muramoto, 2006; 39–40, D. subvittatus Benderitter, 1922." figureDoi="http://doi.org/10.5281/zenodo.4556636" httpUri="https://zenodo.org/record/4556636/files/figure.png" pageId="6" pageNumber="499">Figs. 35–36</figureCitation>
.
</paragraph>
<paragraph id="8B9536A1FF92F707FF7A20173657FA43" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="499">
<emphasis id="B95EEAB3FF92F707FF7A201735F0FBB7" bold="true" box="[199,402,1051,1077]" pageId="6" pageNumber="499">
<typeStatus id="54918803FF92F707FF7A20173523FBB7" box="[199,321,1051,1077]" pageId="6" pageNumber="499" type="paratype">Paratypes</typeStatus>
. Male:
</emphasis>
Larger males have longer mandibles (
<figureCitation id="13112A24FF92F707FC85201737A2FBB7" box="[824,960,1050,1077]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1323,1347]" captionTargetBox="[155,1432,191,1289]" captionTargetId="figure-23@8.[151,1435,181,1299]" captionTargetPageId="8" captionText="FIGURES 15–20. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (oblique view). 15–18, males in different body sizes (16 holotype); 19–20, females in different elytral colorations (19 allotype)." figureDoi="http://doi.org/10.5281/zenodo.4556628" httpUri="https://zenodo.org/record/4556628/files/figure.png" pageId="6" pageNumber="499">Figs. 15–18</figureCitation>
). 
<emphasis id="B95EEAB3FF92F707FC6F20173051FBB7" bold="true" box="[978,1075,1051,1077]" pageId="6" pageNumber="499">Female:</emphasis>
Body color darker; head, pronotum, scutellum, pygidium, ventral surface, and legs black; the anterior 1/2 to 3/4 of elytron fulvous, and the posterior 1/2 to 1/4 dark brown or black (
<figureCitation id="13112A24FF92F707FDB7206F3608FBFF" box="[522,618,1123,1149]" captionStart="FIGURES 11–14" captionStartId="7.[151,264,1944,1968]" captionTargetBox="[182,1406,185,1905]" captionTargetId="figure-17@7.[169,1418,181,1920]" captionTargetPageId="7" captionText="FIGURES 11–14. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (dorsal and ventral view). 11–12, male holotype; 13–14, female allotype." figureDoi="http://doi.org/10.5281/zenodo.4556626" httpUri="https://zenodo.org/record/4556626/files/figure.png" pageId="6" pageNumber="499">Figs. 13</figureCitation>
, 
<figureCitation id="13112A24FF92F707FDC5206F36A1FBFF" box="[632,707,1123,1149]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1323,1347]" captionTargetBox="[155,1432,191,1289]" captionTargetId="figure-23@8.[151,1435,181,1299]" captionTargetPageId="8" captionText="FIGURES 15–20. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (oblique view). 15–18, males in different body sizes (16 holotype); 19–20, females in different elytral colorations (19 allotype)." figureDoi="http://doi.org/10.5281/zenodo.4556628" httpUri="https://zenodo.org/record/4556628/files/figure.png" pageId="6" pageNumber="499">19–20</figureCitation>
). Mandible small, short, nearly triangular; outer edge bidentate, apical tooth acute and bends upwards, second tooth broadly rounded. Clypeus longer than in male; anterior margin straight or feebly emarginated, distinctly convex. Labrum not visible in dorsal view. Antennal club slightly shorter. Intervals defined by rows of close and large punctures; three inner primary striae feebly convex, with scattered small punctures; interstices with dense, irregular large punctures. Punctures on dorsal surface of head, pronutum, scutellum and elytra less but larger, deeper and coarser. Setae and hairs on the whole body surface rather sparser. Abdominal ventrites convex; abdominal ventrite 1 abbreviated medially, abdominal ventrites 2–4 almost equal in length (
<figureCitation id="13112A24FF92F707FF5021533523FAFB" box="[237,321,1375,1401]" captionStart="FIGURES 11–14" captionStartId="7.[151,264,1944,1968]" captionTargetBox="[182,1406,185,1905]" captionTargetId="figure-17@7.[169,1418,181,1920]" captionTargetPageId="7" captionText="FIGURES 11–14. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (dorsal and ventral view). 11–12, male holotype; 13–14, female allotype." figureDoi="http://doi.org/10.5281/zenodo.4556626" httpUri="https://zenodo.org/record/4556626/files/figure.png" pageId="6" pageNumber="499">Fig. 14</figureCitation>
); posterior margin of abdominal ventrite 6 straight medially. Protarsus smaller, but metatibia stronger. Tarsomeres 4 of all legs each with an acute ventroapical protrusion, each protrusion ofmeso- and metatarsomeres 4 with a pair of spines at base.
</paragraph>
<paragraph id="8B9536A1FF92F707FF7A21C73082F98B" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="499">
<emphasis id="B95EEAB3FF92F707FF7A21C7351CFA67" bold="true" box="[199,382,1483,1509]" pageId="6" pageNumber="499">Measurements.</emphasis>
Body length: male 15.0–17.8 mm (
<typeStatus id="54918803FF92F707FCA921C7371AFA67" box="[788,888,1483,1509]" pageId="6" pageNumber="499" type="holotype">holotype</typeStatus>
23.5 mm), female 14.2–18.1 mm (
<typeStatus id="54918803FF92F707FABA21C73101FA67" box="[1287,1379,1483,1509]" pageId="6" pageNumber="499" type="allotype">allotype</typeStatus>
18.1 mm); body width: male 7.8–9.1 mm (
<typeStatus id="54918803FF92F707FD8621E336FDF98B" box="[571,671,1519,1545]" pageId="6" pageNumber="499" type="holotype">holotype</typeStatus>
9.7 mm), female 8.1–10.0 mm (
<typeStatus id="54918803FF92F707FBB421E33007F98B" box="[1033,1125,1519,1545]" pageId="6" pageNumber="499" type="allotype">allotype</typeStatus>
10.0 mm).
</paragraph>
</subSubSection>
<subSubSection id="C330652AFF92F707FF7A221E3141F8F3" pageId="6" pageNumber="499" type="diagnosis">
<paragraph id="8B9536A1FF92F707FF7A221E3141F8F3" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="499">
<emphasis id="B95EEAB3FF92F707FF7A221E35ADF9AE" bold="true" box="[199,463,1554,1580]" pageId="6" pageNumber="499">Differential diagnosis.</emphasis>
<taxonomicName id="4C2A4D22FF92F707FE65221F3745F9AF" authority="Qiu &amp; Zhao &amp; Xu, 2021" authorityName="Qiu &amp; Zhao &amp; Xu" authorityYear="2021" box="[472,807,1555,1581]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="heterocolor" status="sp. nov.">
<emphasis id="B95EEAB3FF92F707FE65221F3745F9AF" box="[472,807,1555,1581]" italics="true" pageId="6" pageNumber="499">Didrepanephorus heterocolor</emphasis>
</taxonomicName>
<emphasis id="B95EEAB3FF92F707FC92221F37D8F9AE" bold="true" box="[815,954,1554,1581]" pageId="6" pageNumber="499">
<taxonomicNameLabel id="A26D57C8FF92F707FC92221F37D8F9AE" box="[815,954,1554,1581]" pageId="6" pageNumber="499" rank="species">new species</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="4C2A4D22FF92F707FBE9221F3090F9AF" authorityName="Muramoto" authorityYear="2006" box="[1108,1266,1555,1581]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="nishiyamai">
<emphasis id="B95EEAB3FF92F707FBE9221F3090F9AF" box="[1108,1266,1555,1581]" italics="true" pageId="6" pageNumber="499">D. nishiyamai</emphasis>
</taxonomicName>
and 
<emphasis id="B95EEAB3FF92F707FA91221F34AFF9D2" italics="true" pageId="6" pageNumber="499">D. subvittatus</emphasis>
, but the body color of the latter two species is reddish-brown. This new species can be separated from the latter two also by the following characters: antennal club bicolor (base dark brown and gradually changed fulvous to apex, 
<figureCitation id="13112A24FF92F707FF6722733500F91B" box="[218,354,1662,1689]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1323,1347]" captionTargetBox="[155,1432,191,1289]" captionTargetId="figure-23@8.[151,1435,181,1299]" captionTargetPageId="8" captionText="FIGURES 15–20. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (oblique view). 15–18, males in different body sizes (16 holotype); 19–20, females in different elytral colorations (19 allotype)." figureDoi="http://doi.org/10.5281/zenodo.4556628" httpUri="https://zenodo.org/record/4556628/files/figure.png" pageId="6" pageNumber="499">Figs. 15–18</figureCitation>
), while unicolor in 
<taxonomicName id="4C2A4D22FF92F707FD83227336B9F91B" authorityName="Muramoto" authorityYear="2006" box="[574,731,1663,1689]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="nishiyamai">
<emphasis id="B95EEAB3FF92F707FD83227336B9F91B" box="[574,731,1663,1689]" italics="true" pageId="6" pageNumber="499">D. nishiyamai</emphasis>
</taxonomicName>
(
<figureCitation id="13112A24FF92F707FD5622733713F91B" box="[747,881,1663,1689]" captionStart="FIGURES 29–32" captionStartId="12.[151,264,966,990]" captionTargetBox="[183,1407,191,937]" captionTargetId="figure-23@12.[177,1411,181,942]" captionTargetPageId="12" captionText="FIGURES 29–32. Habitus of Didrepanephorus Wood-Mason, 1878 species (oblique view). 29–30, D. nishiyamai Muramoto, 2006: 29, male; 30, female. 31–32, D. subvittatus Benderitter, 1922: 31, male; 32, female." figureDoi="http://doi.org/10.5281/zenodo.4556634" httpUri="https://zenodo.org/record/4556634/files/figure.png" pageId="6" pageNumber="499">Figs. 29–30</figureCitation>
) and 
<taxonomicName id="4C2A4D22FF92F707FC0C2273302EF91B" authorityName="Benderitter" authorityYear="1922" box="[945,1100,1663,1689]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="subvittatus">
<emphasis id="B95EEAB3FF92F707FC0C2273302EF91B" box="[945,1100,1663,1689]" italics="true" pageId="6" pageNumber="499">D. subvittatus</emphasis>
</taxonomicName>
(
<figureCitation id="13112A24FF92F707FBE122733087F91B" box="[1116,1253,1663,1689]" captionStart="FIGURES 29–32" captionStartId="12.[151,264,966,990]" captionTargetBox="[183,1407,191,937]" captionTargetId="figure-23@12.[177,1411,181,942]" captionTargetPageId="12" captionText="FIGURES 29–32. Habitus of Didrepanephorus Wood-Mason, 1878 species (oblique view). 29–30, D. nishiyamai Muramoto, 2006: 29, male; 30, female. 31–32, D. subvittatus Benderitter, 1922: 31, male; 32, female." figureDoi="http://doi.org/10.5281/zenodo.4556634" httpUri="https://zenodo.org/record/4556634/files/figure.png" pageId="6" pageNumber="499">Figs. 31–32</figureCitation>
); length of male antennal club as long as antennomeres 2–7 combined (
<figureCitation id="13112A24FF92F707FD4C22AF3715F93F" box="[753,887,1699,1725]" captionStart="FIGURES 11–14" captionStartId="7.[151,264,1944,1968]" captionTargetBox="[182,1406,185,1905]" captionTargetId="figure-17@7.[169,1418,181,1920]" captionTargetPageId="7" captionText="FIGURES 11–14. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (dorsal and ventral view). 11–12, male holotype; 13–14, female allotype." figureDoi="http://doi.org/10.5281/zenodo.4556626" httpUri="https://zenodo.org/record/4556626/files/figure.png" pageId="6" pageNumber="499">Figs. 11–12</figureCitation>
), while longer in 
<taxonomicName id="4C2A4D22FF92F707FB8A22AF30B1F93F" authorityName="Muramoto" authorityYear="2006" box="[1079,1235,1699,1725]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="nishiyamai">
<emphasis id="B95EEAB3FF92F707FB8A22AF30B1F93F" box="[1079,1235,1699,1725]" italics="true" pageId="6" pageNumber="499">D. nishiyamai</emphasis>
</taxonomicName>
(
<figureCitation id="13112A24FF92F707FB5C22AF3104F93F" box="[1249,1382,1699,1725]" captionStart="FIGURES 21–24" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[162,1422,194,1903]" captionTargetId="figure-23@10.[151,1435,181,1920]" captionTargetPageId="10" captionText="FIGURES 21–24. Habitus of Didrepanephorus nishiyamai Muramoto, 2006 (dorsal and ventral view). 21–22, male (Mt. Dayaoshan); 23–24, female (Mt. Damingshan)." figureDoi="http://doi.org/10.5281/zenodo.4556630" httpUri="https://zenodo.org/record/4556630/files/figure.png" pageId="6" pageNumber="499">Figs. 21–22</figureCitation>
) and shorter in 
<taxonomicName id="4C2A4D22FF92F707FEBA22CB35C3F963" authorityName="Benderitter" authorityYear="1922" box="[263,417,1735,1761]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="subvittatus">
<emphasis id="B95EEAB3FF92F707FEBA22CB35C3F963" box="[263,417,1735,1761]" italics="true" pageId="6" pageNumber="499">D. subvittatus</emphasis>
</taxonomicName>
(
<figureCitation id="13112A24FF92F707FE1222CB365AF963" box="[431,568,1734,1761]" captionStart="FIGURES 25–28" captionStartId="11.[151,264,1944,1968]" captionTargetBox="[177,1412,187,1905]" captionTargetId="figure-17@11.[162,1426,181,1920]" captionTargetPageId="11" captionText="FIGURES 25–28. Habitus of Didrepanephorus subvittatus Benderitter, 1922 (dorsal and ventral view). 25–26, male; 27–28, female. (both from Tam Dao)." figureDoi="http://doi.org/10.5281/zenodo.4556632" httpUri="https://zenodo.org/record/4556632/files/figure.png" pageId="6" pageNumber="499">Figs. 25–26</figureCitation>
); male mandibles deeply constricted before base tooth (
<figureCitation id="13112A24FF92F707FB2322CB3092F963" box="[1182,1264,1734,1761]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1323,1347]" captionTargetBox="[155,1432,191,1289]" captionTargetId="figure-23@8.[151,1435,181,1299]" captionTargetPageId="8" captionText="FIGURES 15–20. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (oblique view). 15–18, males in different body sizes (16 holotype); 19–20, females in different elytral colorations (19 allotype)." figureDoi="http://doi.org/10.5281/zenodo.4556628" httpUri="https://zenodo.org/record/4556628/files/figure.png" pageId="6" pageNumber="499">Fig. 15</figureCitation>
), while slightly constricted in 
<taxonomicName id="4C2A4D22FF92F707FE8722E735B5F887" authorityName="Benderitter" authorityYear="1922" box="[314,471,1771,1797]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="subvittatus">
<emphasis id="B95EEAB3FF92F707FE8722E735B5F887" box="[314,471,1771,1797]" italics="true" pageId="6" pageNumber="499">D. subvittatus</emphasis>
</taxonomicName>
(
<figureCitation id="13112A24FF92F707FE5522E7362AF887" box="[488,584,1771,1797]" captionStart="FIGURES 29–32" captionStartId="12.[151,264,966,990]" captionTargetBox="[183,1407,191,937]" captionTargetId="figure-23@12.[177,1411,181,942]" captionTargetPageId="12" captionText="FIGURES 29–32. Habitus of Didrepanephorus Wood-Mason, 1878 species (oblique view). 29–30, D. nishiyamai Muramoto, 2006: 29, male; 30, female. 31–32, D. subvittatus Benderitter, 1922: 31, male; 32, female." figureDoi="http://doi.org/10.5281/zenodo.4556634" httpUri="https://zenodo.org/record/4556634/files/figure.png" pageId="6" pageNumber="499">Figs. 31</figureCitation>
); anterior margin of male mentum slightly depressed in middle, while distinctly depressed in 
<taxonomicName id="4C2A4D22FF92F707FEC82303366DF8AB" authorityName="Benderitter" authorityYear="1922" box="[373,527,1807,1833]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="subvittatus">
<emphasis id="B95EEAB3FF92F707FEC82303366DF8AB" box="[373,527,1807,1833]" italics="true" pageId="6" pageNumber="499">D. subvittatus</emphasis>
</taxonomicName>
; setae on male pronotum longer (
<figureCitation id="13112A24FF92F707FC3F23033065F8AB" box="[898,1031,1806,1833]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1323,1347]" captionTargetBox="[155,1432,191,1289]" captionTargetId="figure-23@8.[151,1435,181,1299]" captionTargetPageId="8" captionText="FIGURES 15–20. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (oblique view). 15–18, males in different body sizes (16 holotype); 19–20, females in different elytral colorations (19 allotype)." figureDoi="http://doi.org/10.5281/zenodo.4556628" httpUri="https://zenodo.org/record/4556628/files/figure.png" pageId="6" pageNumber="499">Figs. 15–18</figureCitation>
) than that of 
<taxonomicName id="4C2A4D22FF92F707FB2423033157F8AB" authorityName="Muramoto" authorityYear="2006" box="[1177,1333,1807,1833]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="nishiyamai">
<emphasis id="B95EEAB3FF92F707FB2423033157F8AB" box="[1177,1333,1807,1833]" italics="true" pageId="6" pageNumber="499">D. nishiyamai</emphasis>
</taxonomicName>
(
<figureCitation id="13112A24FF92F707FAF9230331F6F8AB" box="[1348,1428,1807,1833]" captionStart="FIGURES 29–32" captionStartId="12.[151,264,966,990]" captionTargetBox="[183,1407,191,937]" captionTargetId="figure-23@12.[177,1411,181,942]" captionTargetPageId="12" captionText="FIGURES 29–32. Habitus of Didrepanephorus Wood-Mason, 1878 species (oblique view). 29–30, D. nishiyamai Muramoto, 2006: 29, male; 30, female. 31–32, D. subvittatus Benderitter, 1922: 31, male; 32, female." figureDoi="http://doi.org/10.5281/zenodo.4556634" httpUri="https://zenodo.org/record/4556634/files/figure.png" pageId="6" pageNumber="499">Fig. 29</figureCitation>
) and 
<taxonomicName id="4C2A4D22FF92F707FF7A233F3501F8CF" authorityName="Benderitter" authorityYear="1922" box="[199,355,1843,1869]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="subvittatus">
<emphasis id="B95EEAB3FF92F707FF7A233F3501F8CF" box="[199,355,1843,1869]" italics="true" pageId="6" pageNumber="499">D. subvittatus</emphasis>
</taxonomicName>
(
<figureCitation id="13112A24FF92F707FECE233F35A5F8CF" box="[371,455,1843,1869]" captionStart="FIGURES 29–32" captionStartId="12.[151,264,966,990]" captionTargetBox="[183,1407,191,937]" captionTargetId="figure-23@12.[177,1411,181,942]" captionTargetPageId="12" captionText="FIGURES 29–32. Habitus of Didrepanephorus Wood-Mason, 1878 species (oblique view). 29–30, D. nishiyamai Muramoto, 2006: 29, male; 30, female. 31–32, D. subvittatus Benderitter, 1922: 31, male; 32, female." figureDoi="http://doi.org/10.5281/zenodo.4556634" httpUri="https://zenodo.org/record/4556634/files/figure.png" pageId="6" pageNumber="499">Fig. 31</figureCitation>
); female with dense and coarse punctures on head and pronotum (
<figureCitation id="13112A24FF92F707FB0C233F3158F8CF" box="[1201,1338,1843,1869]" captionStart="FIGURES 11–14" captionStartId="7.[151,264,1944,1968]" captionTargetBox="[182,1406,185,1905]" captionTargetId="figure-17@7.[169,1418,181,1920]" captionTargetPageId="7" captionText="FIGURES 11–14. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (dorsal and ventral view). 11–12, male holotype; 13–14, female allotype." figureDoi="http://doi.org/10.5281/zenodo.4556626" httpUri="https://zenodo.org/record/4556626/files/figure.png" pageId="6" pageNumber="499">Figs. 13–14</figureCitation>
, 
<figureCitation id="13112A24FF92F707FAFB233F31EDF8CF" box="[1350,1423,1843,1869]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1323,1347]" captionTargetBox="[155,1432,191,1289]" captionTargetId="figure-23@8.[151,1435,181,1299]" captionTargetPageId="8" captionText="FIGURES 15–20. Habitus of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species (oblique view). 15–18, males in different body sizes (16 holotype); 19–20, females in different elytral colorations (19 allotype)." figureDoi="http://doi.org/10.5281/zenodo.4556628" httpUri="https://zenodo.org/record/4556628/files/figure.png" pageId="6" pageNumber="499">19–20</figureCitation>
), while those punctures sparser and finer in 
<taxonomicName id="4C2A4D22FF92F707FDCC235B376CF8F3" authorityName="Muramoto" authorityYear="2006" box="[625,782,1879,1905]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="nishiyamai">
<emphasis id="B95EEAB3FF92F707FDCC235B376CF8F3" box="[625,782,1879,1905]" italics="true" pageId="6" pageNumber="499">D. nishiyamai</emphasis>
</taxonomicName>
(
<figureCitation id="13112A24FF92F707FCA0235B3718F8F3" box="[797,890,1879,1905]" captionStart="FIGURES 21–24" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[162,1422,194,1903]" captionTargetId="figure-23@10.[151,1435,181,1920]" captionTargetPageId="10" captionText="FIGURES 21–24. Habitus of Didrepanephorus nishiyamai Muramoto, 2006 (dorsal and ventral view). 21–22, male (Mt. Dayaoshan); 23–24, female (Mt. Damingshan)." figureDoi="http://doi.org/10.5281/zenodo.4556630" httpUri="https://zenodo.org/record/4556630/files/figure.png" pageId="6" pageNumber="499">Figs. 23</figureCitation>
, 
<figureCitation id="13112A24FF92F707FC3B235B37C1F8F3" box="[902,931,1879,1905]" captionStart="FIGURES 29–32" captionStartId="12.[151,264,966,990]" captionTargetBox="[183,1407,191,937]" captionTargetId="figure-23@12.[177,1411,181,942]" captionTargetPageId="12" captionText="FIGURES 29–32. Habitus of Didrepanephorus Wood-Mason, 1878 species (oblique view). 29–30, D. nishiyamai Muramoto, 2006: 29, male; 30, female. 31–32, D. subvittatus Benderitter, 1922: 31, male; 32, female." figureDoi="http://doi.org/10.5281/zenodo.4556634" httpUri="https://zenodo.org/record/4556634/files/figure.png" pageId="6" pageNumber="499">30</figureCitation>
) and 
<taxonomicName id="4C2A4D22FF92F707FC5F235B301FF8F3" authorityName="Benderitter" authorityYear="1922" box="[994,1149,1879,1905]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="subvittatus">
<emphasis id="B95EEAB3FF92F707FC5F235B301FF8F3" box="[994,1149,1879,1905]" italics="true" pageId="6" pageNumber="499">D. subvittatus</emphasis>
</taxonomicName>
(
<figureCitation id="13112A24FF92F707FB30235B3088F8F3" box="[1165,1258,1879,1905]" captionStart="FIGURES 25–28" captionStartId="11.[151,264,1944,1968]" captionTargetBox="[177,1412,187,1905]" captionTargetId="figure-17@11.[162,1426,181,1920]" captionTargetPageId="11" captionText="FIGURES 25–28. Habitus of Didrepanephorus subvittatus Benderitter, 1922 (dorsal and ventral view). 25–26, male; 27–28, female. (both from Tam Dao)." figureDoi="http://doi.org/10.5281/zenodo.4556632" httpUri="https://zenodo.org/record/4556632/files/figure.png" pageId="6" pageNumber="499">Figs. 27</figureCitation>
, 
<figureCitation id="13112A24FF92F707FB4B235B3174F8F3" box="[1270,1302,1879,1905]" captionStart="FIGURES 29–32" captionStartId="12.[151,264,966,990]" captionTargetBox="[183,1407,191,937]" captionTargetId="figure-23@12.[177,1411,181,942]" captionTargetPageId="12" captionText="FIGURES 29–32. Habitus of Didrepanephorus Wood-Mason, 1878 species (oblique view). 29–30, D. nishiyamai Muramoto, 2006: 29, male; 30, female. 31–32, D. subvittatus Benderitter, 1922: 31, male; 32, female." figureDoi="http://doi.org/10.5281/zenodo.4556634" httpUri="https://zenodo.org/record/4556634/files/figure.png" pageId="6" pageNumber="499">32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C330652AFF92F707FF7A237730FBF83B" pageId="6" pageNumber="499" type="etymology">
<paragraph id="8B9536A1FF92F707FF7A237730FBF83B" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="499">
<emphasis id="B95EEAB3FF92F707FF7A2377352CF817" bold="true" box="[199,334,1915,1941]" pageId="6" pageNumber="499">Etymology.</emphasis>
The specific epithet “ 
<taxonomicName id="4C2A4D22FF92F707FDFE237736A7F817" authorityName="Qiu &amp; Zhao &amp; Xu" authorityYear="2021" box="[579,709,1915,1941]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="heterocolor">
<emphasis id="B95EEAB3FF92F707FDFE237736A7F817" box="[579,709,1915,1941]" italics="true" pageId="6" pageNumber="499">heterocolor</emphasis>
</taxonomicName>
” is a combination of the Greek “ 
<emphasis id="B95EEAB3FF92F707FB88237730E4F817" box="[1077,1158,1915,1941]" italics="true" pageId="6" pageNumber="499">heteros</emphasis>
” and Latin noun “ 
<emphasis id="B95EEAB3FF92F707FAE8237731F2F817" box="[1365,1424,1915,1941]" italics="true" pageId="6" pageNumber="499">color</emphasis>
” meaning different colors, in allusion to the significant sexual dichroism of this new species.
</paragraph>
</subSubSection>
<subSubSection id="C330652AFF92F709FF7A23CE37CEF94D" lastPageId="8" lastPageNumber="501" pageId="6" pageNumber="499" type="distribution">
<paragraph id="8B9536A1FF92F707FF7A23CE3643F85F" blockId="6.[151,1437,150,2013]" box="[199,545,1986,2013]" pageId="6" pageNumber="499">
<emphasis id="B95EEAB3FF92F707FF7A23CE3503F85E" bold="true" box="[199,353,1986,2012]" pageId="6" pageNumber="499">Distribution.</emphasis>
<collectingCountry id="F33D7631FF92F707FEDA23CF35CCF85F" box="[359,430,1987,2013]" name="China" pageId="6" pageNumber="499">China</collectingCountry>
: 
<collectingRegion id="49EEF843FF92F707FE0423CF367FF85F" box="[441,541,1987,2013]" country="China" name="Guizhou" pageId="6" pageNumber="499">Guizhou</collectingRegion>
.
</paragraph>
<caption id="DF556629FF93F706FF2A23943647F856" ID-DOI="http://doi.org/10.5281/zenodo.4556626" ID-Zenodo-Dep="4556626" httpUri="https://zenodo.org/record/4556626/files/figure.png" pageId="7" pageNumber="500" startId="7.[151,264,1944,1968]" targetBox="[182,1406,185,1905]" targetPageId="7">
<paragraph id="8B9536A1FF93F706FF2A23943647F856" blockId="7.[151,1436,1944,2004]" pageId="7" pageNumber="500">
<emphasis id="B95EEAB3FF93F706FF2A23943536F832" bold="true" box="[151,340,1944,1968]" pageId="7" pageNumber="500">FIGURES 11–14.</emphasis>
Habitus of 
<taxonomicName id="4C2A4D22FF93F706FE6D239437D6F82D" authority="Qiu, Zhao &amp; Xu" authorityName="Qiu, Zhao &amp; Xu" authorityYear="2021" box="[464,948,1944,1968]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="500" phylum="Arthropoda" rank="species" species="heterocolor" status="sp. nov.">
<emphasis id="B95EEAB3FF93F706FE6D2394369DF832" box="[464,767,1944,1968]" italics="true" pageId="7" pageNumber="500">Didrepanephorus heterocolor</emphasis>
Qiu, Zhao &amp; Xu
</taxonomicName>
, 
<emphasis id="B95EEAB3FF93F706FC7D2394305DF832" bold="true" box="[960,1087,1944,1968]" pageId="7" pageNumber="500">
<taxonomicNameLabel id="A26D57C8FF93F706FC7D2394305DF832" box="[960,1087,1944,1968]" pageId="7" pageNumber="500" rank="species">new species</taxonomicNameLabel>
</emphasis>
(dorsal and ventral view). 11–12, male holotype; 13–14, female allotype.
</paragraph>
</caption>
<caption id="DF556629FF9CF709FF2A2127305DFAE5" ID-DOI="http://doi.org/10.5281/zenodo.4556628" ID-Zenodo-Dep="4556628" httpUri="https://zenodo.org/record/4556628/files/figure.png" pageId="8" pageNumber="501" startId="8.[151,264,1323,1347]" targetBox="[155,1432,191,1289]" targetPageId="8">
<paragraph id="8B9536A1FF9CF709FF2A2127305DFAE5" blockId="8.[151,1437,1323,1383]" pageId="8" pageNumber="501">
<emphasis id="B95EEAB3FF9CF709FF2A21273536FAC1" bold="true" box="[151,340,1323,1347]" pageId="8" pageNumber="501">FIGURES 15–20.</emphasis>
Habitus of 
<taxonomicName id="4C2A4D22FF9CF709FE77212737C6FAC1" authority="Qiu, Zhao &amp; Xu" authorityName="Qiu, Zhao &amp; Xu" authorityYear="2021" box="[458,932,1323,1347]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="heterocolor" status="sp. nov.">
<emphasis id="B95EEAB3FF9CF709FE7721273694FAC1" box="[458,758,1323,1347]" italics="true" pageId="8" pageNumber="501">Didrepanephorus heterocolor</emphasis>
Qiu, Zhao &amp; Xu
</taxonomicName>
, 
<emphasis id="B95EEAB3FF9CF709FC1321273049FAC1" bold="true" box="[942,1067,1323,1347]" pageId="8" pageNumber="501">
<taxonomicNameLabel id="A26D57C8FF9CF709FC1321273049FAC1" box="[942,1067,1323,1347]" pageId="8" pageNumber="501" rank="species">new species</taxonomicNameLabel>
</emphasis>
(oblique view). 15–18, males in different body sizes (16 holotype); 19–20, females in different elytral colorations (19 allotype).
</paragraph>
</caption>
<paragraph id="8B9536A1FF9CF709FF7A219937CEF94D" blockId="8.[151,1437,1429,1996]" pageId="8" pageNumber="501">
<emphasis id="B95EEAB3FF9CF709FF7A219935EAFA2D" bold="true" box="[199,392,1429,1455]" pageId="8" pageNumber="501">Natural history.</emphasis>
This species is only known from the Maolan Nature Reserve in southern 
<collectingRegion id="49EEF843FF9CF709FB5C21993126FA2D" box="[1249,1348,1429,1455]" country="China" name="Guizhou" pageId="8" pageNumber="501">Guizhou</collectingRegion>
, 
<collectingCountry id="F33D7631FF9CF709FAEF219931FAFA2D" box="[1362,1432,1429,1455]" name="China" pageId="8" pageNumber="501">China</collectingCountry>
, where has a subtropical forest of karst landform at elevation of 
<quantity id="4CD29B44FF9CF709FCD021B5378FFA56" box="[877,1005,1465,1492]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.5" metricValueMin="5.5" pageId="8" pageNumber="501" unit="m" value="700.0" valueMax="850.0" valueMin="550.0">550–850 m</quantity>
(
<figureCitation id="13112A24FF9CF709FC4321B530EBFA56" box="[1022,1161,1465,1492]" captionStart="FIGURES 57–64" captionStartId="16.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1885]" captionTargetId="figure-23@16.[164,1423,181,1885]" captionTargetPageId="16" captionText="FIGURES 57–64. Habitat and different stages of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species. 57, forest in Maolan Nature Reserve (Guizhou, China); 58–59, a closer view of habitat and pupal cells in rotten wood (both photograph by Ri-Xin Jiang); 60, a larva under the bark of decayed wood in wild; 61–62, pupa and newly emerged female in pupal cells in artificial conditions; 63, male in a pupal cell in rotten wood (photograph by Ri-Xin Jiang); 64, egg in artificial conditions." figureDoi="http://doi.org/10.5281/zenodo.4556640" httpUri="https://zenodo.org/record/4556640/files/figure.png" pageId="8" pageNumber="501">Figs. 57–58</figureCitation>
). The mature larvae and newly emerged adults have been found in humid rotten wood in later April and early May (
<figureCitation id="13112A24FF9CF709FB1421D1313CFA7A" box="[1193,1374,1501,1528]" captionStart="FIGURES 57–64" captionStartId="16.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1885]" captionTargetId="figure-23@16.[164,1423,181,1885]" captionTargetPageId="16" captionText="FIGURES 57–64. Habitat and different stages of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species. 57, forest in Maolan Nature Reserve (Guizhou, China); 58–59, a closer view of habitat and pupal cells in rotten wood (both photograph by Ri-Xin Jiang); 60, a larva under the bark of decayed wood in wild; 61–62, pupa and newly emerged female in pupal cells in artificial conditions; 63, male in a pupal cell in rotten wood (photograph by Ri-Xin Jiang); 64, egg in artificial conditions." figureDoi="http://doi.org/10.5281/zenodo.4556640" httpUri="https://zenodo.org/record/4556640/files/figure.png" pageId="8" pageNumber="501">Figs. 59–60, 63</figureCitation>
), and an exhausted female was collected by a light trap in early 
<date id="FF941061FF9CF709FC9B220E37FFF999" box="[806,925,1537,1564]" pageId="8" pageNumber="501" value="2018-06">June 2018</date>
. It is suggested the peak of activity for adults should be mid and later May. In the field, third instar larvae of the new generation can be excavated in rotten wood of broad-leaved trees as early as late July (
<figureCitation id="13112A24FF9CF709FD3F224536BAF9E6" box="[642,728,1609,1636]" captionStart="FIGURES 57–64" captionStartId="16.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1885]" captionTargetId="figure-23@16.[164,1423,181,1885]" captionTargetPageId="16" captionText="FIGURES 57–64. Habitat and different stages of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species. 57, forest in Maolan Nature Reserve (Guizhou, China); 58–59, a closer view of habitat and pupal cells in rotten wood (both photograph by Ri-Xin Jiang); 60, a larva under the bark of decayed wood in wild; 61–62, pupa and newly emerged female in pupal cells in artificial conditions; 63, male in a pupal cell in rotten wood (photograph by Ri-Xin Jiang); 64, egg in artificial conditions." figureDoi="http://doi.org/10.5281/zenodo.4556640" httpUri="https://zenodo.org/record/4556640/files/figure.png" pageId="8" pageNumber="501">Fig. 60</figureCitation>
). Wild-collected larvae were raised in small plastic containers filled with pieces of wet, rotten oak wood at the temperature below 26°C. Females have been bred for reproduction under artificial conditions (
<figureCitation id="13112A24FF9CF709FE46229D36CCF92E" box="[507,686,1681,1708]" captionStart="FIGURES 57–64" captionStartId="16.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1885]" captionTargetId="figure-23@16.[164,1423,181,1885]" captionTargetPageId="16" captionText="FIGURES 57–64. Habitat and different stages of Didrepanephorus heterocolor Qiu, Zhao &amp; Xu, new species. 57, forest in Maolan Nature Reserve (Guizhou, China); 58–59, a closer view of habitat and pupal cells in rotten wood (both photograph by Ri-Xin Jiang); 60, a larva under the bark of decayed wood in wild; 61–62, pupa and newly emerged female in pupal cells in artificial conditions; 63, male in a pupal cell in rotten wood (photograph by Ri-Xin Jiang); 64, egg in artificial conditions." figureDoi="http://doi.org/10.5281/zenodo.4556640" httpUri="https://zenodo.org/record/4556640/files/figure.png" pageId="8" pageNumber="501">Figs. 61–62, 64</figureCitation>
), and several larvae were successfully developed to the adult stage (personal observation, Jian-Yue Qiu, 2012–2013; Jun Qi, 2017–2018).
</paragraph>
</subSubSection>
<subSubSection id="C330652AFF9CF709FF7A22D53611F849" pageId="8" pageNumber="501" type="discussion">
<paragraph id="8B9536A1FF9CF709FF7A22D53611F849" blockId="8.[151,1437,1429,1996]" pageId="8" pageNumber="501">
<emphasis id="B95EEAB3FF9CF709FF7A22D53559F971" bold="true" box="[199,315,1753,1779]" pageId="8" pageNumber="501">Remarks.</emphasis>
The body color of this new species is sexually dichroic, the male is dark brown, while the female has a black body with bicolor elytra. This phenomenon is distinctive in this genus, and 
<taxonomicName id="4C2A4D22FF9CF709FBF322F230A9F895" authorityName="Muramoto" authorityYear="2003" box="[1102,1227,1789,1815]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="arnaudi">
<emphasis id="B95EEAB3FF9CF709FBF322F230A9F895" box="[1102,1227,1789,1815]" italics="true" pageId="8" pageNumber="501">D. arnaudi</emphasis>
</taxonomicName>
was the only case before. In general appearance, 
<taxonomicName id="4C2A4D22FF9CF709FE51232E36F2F8B9" authority="Qiu &amp; Zhao &amp; Xu, 2021" authorityName="Qiu &amp; Zhao &amp; Xu" authorityYear="2021" box="[492,656,1825,1851]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="heterocolor" status="sp. nov.">
<emphasis id="B95EEAB3FF9CF709FE51232E36F2F8B9" box="[492,656,1825,1851]" italics="true" pageId="8" pageNumber="501">D. heterocolor</emphasis>
</taxonomicName>
<emphasis id="B95EEAB3FF9CF709FD28232D377CF8B9" bold="true" box="[661,798,1825,1851]" pageId="8" pageNumber="501">
<taxonomicNameLabel id="A26D57C8FF9CF709FD28232D377CF8B9" box="[661,798,1825,1851]" pageId="8" pageNumber="501" rank="species">new species</taxonomicNameLabel>
</emphasis>
is close to 
<taxonomicName id="4C2A4D22FF9CF709FC25232E3057F8B9" authorityName="Muramoto" authorityYear="2006" box="[920,1077,1825,1851]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="nishiyamai">
<emphasis id="B95EEAB3FF9CF709FC25232E3057F8B9" box="[920,1077,1825,1851]" italics="true" pageId="8" pageNumber="501">D. nishiyamai</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2A4D22FF9CF709FBD4232E3161F8B9" authorityName="Benderitter" authorityYear="1922" box="[1129,1283,1825,1851]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="subvittatus">
<emphasis id="B95EEAB3FF9CF709FBD4232E3161F8B9" box="[1129,1283,1825,1851]" italics="true" pageId="8" pageNumber="501">D. subvittatus</emphasis>
</taxonomicName>
, and they can be attributed to a species group (
<taxonomicName id="4C2A4D22FF9CF709FDBD234A36F8F8DD" authorityName="Benderitter" authorityYear="1922" box="[512,666,1861,1887]" class="Insecta" family="Rutelidae" genus="Didrepanephorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="subvittatus">
<emphasis id="B95EEAB3FF9CF709FDBD234A36F8F8DD" box="[512,666,1861,1887]" italics="true" pageId="8" pageNumber="501">D. subvittatus</emphasis>
</taxonomicName>
group) characterized by the combination of the following characters: lower margin of male mandible with a distinct concavity before basal tooth; pronotum evenly covered with untufted dense setae in male, with sparse short setae in female; abdominal ventrites with sparse setae; phallobase evenly curved in profile; parameres fused basally.
</paragraph>
</subSubSection>
</treatment>
</document>