285 lines
30 KiB
XML
285 lines
30 KiB
XML
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<mods:title id="7EBC1605E21B99EF04BB7F9B1FAC88CB">Revising the taxonomy of Darevskia valentini (Boettger, 1892) and Darevskia rudis (Bedriaga, 1886) (Squamata, Lacertidae): a Morpho-Phylogenetic integrated study in a complex Anatolian scenario</mods:title>
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<mods:namePart id="6DF757667EA8E7344BBD8CE3065174C9">Arribas, Oscar</mods:namePart>
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<mods:namePart id="8096C94207ABAE637EA3ADDC94021056">Candan, Kamil</mods:namePart>
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<mods:namePart id="2C2B23F9D7F8BA4A36C8166AE966D4ED">Kumlutaş, Yusuf</mods:namePart>
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<treatment id="038387B04249FE76FF3B499E1013FF56" ID-DOI="http://doi.org/10.5281/zenodo.7517925" ID-GBIF-Taxon="208791114" ID-Zenodo-Dep="7517925" LSID="urn:lsid:plazi:treatment:038387B04249FE76FF3B499E1013FF56" httpUri="http://treatment.plazi.org/id/038387B04249FE76FF3B499E1013FF56" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
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<subSubSection id="C330652D4249FE79FF3B499E129FFF57" pageId="23" pageNumber="24" type="nomenclature">
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<heading id="D0DD81CA4249FE79FF3B499E11DAFF30" box="[151,748,152,178]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="5">
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<taxonomicName id="4C2A4D254249FE79FF3B499E114BFF33" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[151,637,152,178]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="subSpecies" species="spitzenbergerae" status="subsp. nov." subSpecies="wernermayeri">
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<emphasis id="B95EEAB44249FE79FF3B499E114BFF33" box="[151,637,152,178]" italics="true" pageId="23" pageNumber="24">Darevskia spitzenbergerae wernermayeri</emphasis>
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</taxonomicName>
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<emphasis id="B95EEAB44249FE79FD2F499E11DAFF30" bold="true" box="[643,748,152,178]" pageId="23" pageNumber="24">
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<taxonomicNameLabel id="A26D57CF4249FE79FD2F499E11DAFF30" box="[643,748,152,178]" pageId="23" pageNumber="24" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="8B9536A64249FE79FF3B49BD129FFF57" blockId="23.[151,748,152,214]" box="[151,425,187,214]" pageId="23" pageNumber="24">
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<emphasis id="B95EEAB44249FE79FF3B49BD129FFF57" bold="true" box="[151,425,187,214]" pageId="23" pageNumber="24">
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(Appendix 5;
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<figureCitation id="13112A234249FE79FE9749BD12AAFF57" box="[315,412,187,213]" captionStart="FIGURE 12" captionStartId="22.[152,255,955,979]" captionTargetBox="[151,1413,392,928]" captionTargetId="figure-129@22.[151,1435,381,931]" captionTargetPageId="22" captionText="FIGURE 12. a) Darevskia spitzenbergerae wernermayeri ssp. nov. (Paratype; nº 12, Male; Başeğmez Village, Çaldıran, Turkey); b) Darevskia mirabilis stat. nov. (Paratype; nº 5, Female; Ovit Pass, Kaçkar Mountains, Rize, Turkey); c) Darevskia rudis bolkardaghica (Paratype; nº 1, Male; Karagöl, Ulukışla, Niğde, Central Anatolia, Turkey); d) Darevskia rudis lantzicyreni comb. nov. (nº 23, male; Kümbet Village, Zara, Turkey); e) Darevskia josefschmidtleri sp. nov. (Paratype; nº 20, Male; Yukarınarlıca Village, Çatak, Van, Turkey); f) Darevskia valentini (nº 9, Male; Tepeler Village, Ardahan, Turkey) and temporal area of an Armenian specimen (Karvansaray, Martuni District, Armenia); g) Darevskia spitzenbergerae spitzenbergerae stat. et comb. nov. (nº 1, Male; Cilo Sat Mountains, Hakkari, Turkey)- Also, temporal area of other specimen from the same locality. The new nomenclature proposed in the text is used." figureDoi="http://doi.org/10.5281/zenodo.7517939" httpUri="https://zenodo.org/record/7517939/files/figure.png" pageId="23" pageNumber="24">Fig. 12a</figureCitation>
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).
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</emphasis>
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</paragraph>
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<subSubSection id="C330652D4249FE79FF3B48051657FE1F" pageId="23" pageNumber="24" type="description">
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<paragraph id="8B9536A64249FE79FF3B48051288FE9F" blockId="23.[151,446,259,285]" box="[151,446,259,285]" pageId="23" pageNumber="24">
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<emphasis id="B95EEAB44249FE79FF3B48051288FE9F" bold="true" box="[151,446,259,285]" pageId="23" pageNumber="24">Synonymy/Chresonymy:</emphasis>
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</paragraph>
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<paragraph id="8B9536A64249FE79FF3B484F17B1FEE2" blockId="23.[151,1377,328,414]" box="[151,1159,328,352]" pageId="23" pageNumber="24">
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<taxonomicName id="4C2A4D254249FE79FF3B484F12F4FEE2" box="[151,450,328,352]" class="Reptilia" family="Lacertidae" genus="Lacerta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="subSpecies" species="valentini" subSpecies="lantzicyreni">
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<emphasis id="B95EEAB44249FE79FF3B484F12F4FEE2" box="[151,450,328,352]" italics="true" pageId="23" pageNumber="24">Lacerta valentini lantzicyreni</emphasis>
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</taxonomicName>
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;
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<bibRefCitation id="EFBB4B574249FE79FE63484E107EFEE2" author="Eiselt, J. & Darevsky, I. S. & Schmidtler, J. F." box="[463,840,328,352]" pageId="23" pageNumber="24" pagination="1 - 18" refId="ref25991" refString="Eiselt, J., Darevsky, I. S. & Schmidtler, J. F. (1992) Untersuchungen an Felseneidechsen (Lacerta saxicola komplex) in der ostlichen Turkei, I. Lacerta valentini Boettger. Annalen des Naturhistorischen Museums in Wien, 93 (B), 1 - 18." type="journal article" year="1992">Eiselt, Darevsky & Schmidtler, 1992</bibRefCitation>
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. (from “Yukarı Narlıca”-sic.!-).
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</paragraph>
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<paragraph id="8B9536A64249FE79FF3B486110E4FEFD" blockId="23.[151,1377,328,414]" box="[151,978,359,383]" pageId="23" pageNumber="24">
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<taxonomicName id="4C2A4D254249FE79FF3B48611268FEFD" baseAuthorityName="Boettger" baseAuthorityYear="1892" box="[151,350,359,383]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="valentini">
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<emphasis id="B95EEAB44249FE79FF3B48611268FEFD" box="[151,350,359,383]" italics="true" pageId="23" pageNumber="24">Darevskia valentini</emphasis>
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</taxonomicName>
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“Clade A”;
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<bibRefCitation id="EFBB4B574249FE79FE7648611198FEFD" author="Candan, K. & Kornilios, P. & Ayaz, D. & Kumlutas, Y. & Gul, S. & Yildirim-Caynak, E. & Ilgaz, C." box="[474,686,359,383]" pageId="23" pageNumber="24" pagination="665 - 681" refId="ref25168" refString="Candan, K., Kornilios, P., Ayaz, D., Kumlutas, Y., Gul, S., Yildirim-Caynak, E. & Ilgaz, C. (2021) Cryptic genetic structure within Valentin's Lizard, Darevskia valentini (Boettger, 1892) (Squamata, Lacertidae), with implications for systematics and origins of parthenogenesis. Systematics and Biodiversity, 19 (7), 665 - 681. https: // doi. org / 10.1080 / 14772000.2021.1909171" type="journal article" year="2021">
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||
Candan
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||
<emphasis id="B95EEAB44249FE79FD8248611157FEFD" box="[558,609,359,383]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
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(2021)
|
||
</bibRefCitation>
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||
. (same locality as this study)
|
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</paragraph>
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<paragraph id="8B9536A64249FE79FF3B48801657FE1F" blockId="23.[151,1377,328,414]" box="[151,1377,389,414]" pageId="23" pageNumber="24">
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<emphasis id="B95EEAB44249FE79FF3B4880115CFE1F" bold="true" box="[151,618,389,414]" pageId="23" pageNumber="24">ZooBank registration (http://zoobank.org):</emphasis>
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<uri id="FFBB3AA44249FE79FDDC4883166BFE1F" box="[624,1373,389,413]" pageId="23" pageNumber="24">
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urn:lsid:zoobank.org:act:
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<uuid id="FF8C0C734249FE79FCC94883166BFE1F" box="[869,1373,389,413]" pageId="23" pageNumber="24">927894FD-EEFD-450A-81D9-0A68188EDC3B</uuid>
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</uri>
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.
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</paragraph>
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<subSubSection id="C330652D4249FE79FF3B48CA10F8FDAC" pageId="23" pageNumber="24" type="materials_examined">
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<paragraph id="8B9536A64249FE79FF3B48CA122BFD88" blockId="23.[151,1437,459,2034]" pageId="23" pageNumber="24">
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<materialsCitation id="3B423CFB4249FE79FF3B48CA122BFD88" ID-GBIF-Occurrence="4012805381" collectingDate="2015-07-29" collectionCode="ZDEU" collectorName="Yusuf Kumlutas" country="Turkey" location="Yukarinarlica Village" pageId="23" pageNumber="24" specimenCode="ZDEU123" specimenCount="1" stateProvince="Van" typeStatus="holotype">
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<emphasis id="B95EEAB44249FE79FF3B48CA123BFE64" bold="true" box="[151,269,460,486]" pageId="23" pageNumber="24">
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<typeStatus id="549188044249FE79FF3B48CA123EFE64" box="[151,264,460,486]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCode id="DB8C9EDD4249FE79FEBA48CD12BCFE64" box="[278,394,459,486]" collectionCode="ZDEU" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="23" pageNumber="24">ZDEU123</specimenCode>
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/2015 (n.3). ♁,
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<location id="8EF5607D4249FE79FD9748CA1018FE64" LSID="urn:lsid:plazi:treatment:038387B04249FE76FF3B499E1013FF56:8EF5607D4249FE79FD9748CA1018FE64" box="[571,814,460,486]" country="Turkey" name="Yukarinarlica Village" pageId="23" pageNumber="24" stateProvince="Van">Yukarınarlıca Village</location>
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, Çatak,
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<collectingRegion id="49EEF8444249FE79FC2648CA108FFE64" box="[906,953,460,486]" country="Turkey" name="Van" pageId="23" pageNumber="24">Van</collectingRegion>
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,
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<collectingCountry id="F33D76364249FE79FC6848CA1720FE64" box="[964,1046,460,486]" name="Turkey" pageId="23" pageNumber="24">Turkey</collectingCountry>
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. leg.
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<collectorName id="26DF53704249FE79FBFE48CA163AFE64" box="[1106,1292,460,486]" pageId="23" pageNumber="24">Yusuf Kumlutaş</collectorName>
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, Çetin Ilgaz,
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||
<collectingDate id="EFD0E98E4249FE79FF3B48E9122FFD88" box="[151,281,495,522]" pageId="23" pageNumber="24" value="2015-07-29">29.07.2015</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9536A64249FE79FF3B4B1210F8FDAC" blockId="23.[151,1437,459,2034]" box="[151,974,532,560]" pageId="23" pageNumber="24">
|
||
<emphasis id="B95EEAB44249FE79FF3B4B12122EFDAC" bold="true" box="[151,280,532,558]" pageId="23" pageNumber="24">
|
||
<typeStatus id="549188044249FE79FF3B4B121225FDAC" box="[151,275,532,558]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
7 ♁♁,
|
||
<specimenCount id="9D2CFD2F4249FE79FEC04B12128DFDAC" box="[364,443,532,558]" pageId="23" pageNumber="24" type="female">10 ♀♀</specimenCount>
|
||
. Same locality, date and collectors as
|
||
<typeStatus id="549188044249FE79FCC84B1210FCFDAC" box="[868,970,532,558]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330652D4249FE79FF6B4B311096FDF4" pageId="23" pageNumber="24" type="etymology">
|
||
<paragraph id="8B9536A64249FE79FF6B4B311096FDF4" blockId="23.[151,1437,459,2034]" pageId="23" pageNumber="24">
|
||
<emphasis id="B95EEAB44249FE79FF6B4B311291FDD3" bold="true" box="[199,423,567,593]" pageId="23" pageNumber="24">Derivatio nominis:</emphasis>
|
||
The specific epithet refers to Dr. Werner Mayer (1943-2015), for his remarkable work on the knowledge of lacertid genera relationships and species taxonomy.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330652D4249FE79FF6B4B7912A3FC14" pageId="23" pageNumber="24" type="diagnosis">
|
||
<paragraph id="8B9536A64249FE79FF6B4B7912A3FC14" blockId="23.[151,1437,459,2034]" pageId="23" pageNumber="24">
|
||
<emphasis id="B95EEAB44249FE79FF6B4B791272FD1B" bold="true" box="[199,324,639,665]" pageId="23" pageNumber="24">Diagnosis:</emphasis>
|
||
<taxonomicName id="4C2A4D254249FE79FEEB4B861039FD1B" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[327,783,640,666]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="subSpecies" species="spitzenbergerae" status="subsp. nov." subSpecies="wernermayeri">
|
||
<emphasis id="B95EEAB44249FE79FEEB4B861039FD1B" box="[327,783,640,666]" italics="true" pageId="23" pageNumber="24">Darevskia spitzenbergerae wernermayeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB44249FE79FCBF4B861042FD18" bold="true" box="[787,884,640,666]" pageId="23" pageNumber="24">
|
||
<taxonomicNameLabel id="A26D57CF4249FE79FCBF4B861042FD18" box="[787,884,640,666]" pageId="23" pageNumber="24" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from nominate form (
|
||
<taxonomicName id="4C2A4D254249FE79FB014B8616A5FD18" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" baseAuthorityName="Eiselt, Darevsky & Schmidtler" baseAuthorityYear="1992" box="[1197,1427,640,666]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="subSpecies" species="spitzenbergerae" subSpecies="spitzenbergerae">
|
||
<emphasis id="B95EEAB44249FE79FB014B8616A5FD18" box="[1197,1427,640,666]" italics="true" pageId="23" pageNumber="24">D. s. spitzenbergerae</emphasis>
|
||
</taxonomicName>
|
||
) in having a higher number of supraciliar granules (18-31 vs. 14-23), supratemporal (4-6 vs. 3-6), ventrals (29-31 vs. 26-30) (females), preanals (1-3 vs. 1-3), tibial scales (16-21 vs. 15-19), temporal scales 1 (5-7 vs. 2-6) and temporal scales 2 (4-6 vs. 2-6).
|
||
<emphasis id="B95EEAB44249FE79FE0C4BEA11E0FC87" box="[416,726,748,774]" italics="true" pageId="23" pageNumber="24">
|
||
<taxonomicName id="4C2A4D254249FE79FE0C4BEA1124FC84" authorityName="Arribas" authorityYear="1999" box="[416,530,748,774]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="genus">Darevskia</taxonomicName>
|
||
s. wernermayeri
|
||
</emphasis>
|
||
<emphasis id="B95EEAB44249FE79FD4D4BEA107FFC84" bold="true" box="[737,841,748,774]" pageId="23" pageNumber="24">ssp. nov.</emphasis>
|
||
has a relatively smaller head relative length (0.18- 0.21 vs. 0.19-0.24). It also differs by a characteristic color pattern of the body. The main osteological diagnostic characters that differ from the nominate form can be specified as follows: The higher number of maxillary (19-21 vs. 16-18) and dentary teeth (23-24 vs. 20-23). Postorbital greater or equal than postfrontal (greater, rarely equal in
|
||
<taxonomicName id="4C2A4D254249FE79FF3B4A7A12B2FC14" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" baseAuthorityName="Eiselt, Darevsky & Schmidtler" baseAuthorityYear="1992" box="[151,388,892,918]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="subSpecies" species="spitzenbergerae" subSpecies="spitzenbergerae">
|
||
<emphasis id="B95EEAB44249FE79FF3B4A7A12B2FC14" box="[151,388,892,918]" italics="true" pageId="23" pageNumber="24">D. s. spitzenbergerae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330652D4249FE79FF6B4A99176DF84C" pageId="23" pageNumber="24" type="description">
|
||
<paragraph id="8B9536A64249FE79FF6B4A99127EFC5C" blockId="23.[151,1437,459,2034]" pageId="23" pageNumber="24">
|
||
<emphasis id="B95EEAB44249FE79FF6B4A9912D3FC38" bold="true" box="[199,485,927,954]" pageId="23" pageNumber="24">
|
||
Description of
|
||
<typeStatus id="549188044249FE79FED94AA612D6FC38" box="[373,480,928,954]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
An adult male. Tail regenerated (see Appendix 5c). Fixed with ethanol and conserved in 96% ethanol.
|
||
</paragraph>
|
||
<paragraph id="8B9536A64249FE79FF6B4AEE10CBFAC4" blockId="23.[151,1437,459,2034]" pageId="23" pageNumber="24">
|
||
<emphasis id="B95EEAB44249FE79FF6B4AEE120CFB80" box="[199,314,1000,1026]" italics="true" pageId="23" pageNumber="24">Scalation:</emphasis>
|
||
Rostral not in contact with the nostril. Single postnasal on each side. Width of frontonasal (internasal) plate subequal to length, not in contact with rostral. Sutures between prefrontal plates and frontal plate straight. Parietal plates in contact with postorbital plates on each side. Supraciliar granules 13 and 12, interrupted series on left, continuous on right. Supraciliar plates 6 on each side. Supralabial and sublabial plates 4 and 6 on each side, respectively. Plates in supratemporal region 6 on left, 5 on right side. The first supratemporal plate large narrows towards the back, and ends bluntly. Masseteric large, in one piece on left and two pieces on right, separated from the first supratemporal plate by three longitudinal scales on each side. Tympanic obvious, separated from masseteric by three and two rows of small scales on left and right, respectively. Nine flat collaria. Gularia 33. Ventralia contains 6 longitudinal and 28 transverse rows of plates. Preanal scale singular, surrounded by 6 rows of plates. Femoral pores 20 on each side. Subdigital lamellae 28 on each side. Tibialia 20. Dorsalia 51.
|
||
</paragraph>
|
||
<paragraph id="8B9536A64249FE79FF6B4C5612F3FA30" blockId="23.[151,1437,459,2034]" pageId="23" pageNumber="24">
|
||
<emphasis id="B95EEAB44249FE79FF6B4C561201FAEB" box="[199,311,1360,1385]" italics="true" pageId="23" pageNumber="24">Biometry:</emphasis>
|
||
SVL
|
||
<quantity id="4CD29B434249FE79FED74C4912C7FAE8" box="[379,497,1359,1386]" metricMagnitude="-2" metricUnit="m" metricValue="6.357" pageId="23" pageNumber="24" unit="mm" value="63.57">63.57 mm</quantity>
|
||
. Pileus width
|
||
<quantity id="4CD29B434249FE79FD3E4C4911CFFAE8" box="[658,761,1359,1386]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="23" pageNumber="24" unit="mm" value="6.9">6.90 mm</quantity>
|
||
, pileus length
|
||
<quantity id="4CD29B434249FE79FC334C491722FAE8" box="[927,1044,1359,1386]" metricMagnitude="-2" metricUnit="m" metricValue="1.296" pageId="23" pageNumber="24" unit="mm" value="12.96">12.96 mm</quantity>
|
||
, head width
|
||
<quantity id="4CD29B434249FE79FB094C49163BFAE8" box="[1189,1293,1359,1386]" metricMagnitude="-3" metricUnit="m" metricValue="8.26" pageId="23" pageNumber="24" unit="mm" value="8.26">8.26 mm</quantity>
|
||
, head length
|
||
<quantity id="4CD29B434249FE79FF3B4C72123BFA0C" box="[151,269,1396,1422]" metricMagnitude="-2" metricUnit="m" metricValue="1.374" pageId="23" pageNumber="24" unit="mm" value="13.74">13.74 mm</quantity>
|
||
. Length of forelimb
|
||
<quantity id="4CD29B434249FE79FE594C72115CFA0C" box="[501,618,1396,1422]" metricMagnitude="-2" metricUnit="m" metricValue="2.202" pageId="23" pageNumber="24" unit="mm" value="22.02">22.02 mm</quantity>
|
||
, length of hindlimb
|
||
<quantity id="4CD29B434249FE79FCE34C7210F2FA0C" box="[847,964,1396,1422]" metricMagnitude="-2" metricUnit="m" metricValue="3.137" pageId="23" pageNumber="24" unit="mm" value="31.37">31.37 mm</quantity>
|
||
. Anal plate width and length are
|
||
<quantity id="4CD29B434249FE79FA944C7216ABFA0C" box="[1336,1437,1396,1422]" metricMagnitude="-3" metricUnit="m" metricValue="4.12" pageId="23" pageNumber="24" unit="mm" value="4.12">4.12 mm</quantity>
|
||
and
|
||
<quantity id="4CD29B434249FE79FF6B4C9E121BFA30" box="[199,301,1432,1458]" metricMagnitude="-3" metricUnit="m" metricValue="1.84" pageId="23" pageNumber="24" unit="mm" value="1.84">1.84 mm</quantity>
|
||
, respectively.
|
||
</paragraph>
|
||
<paragraph id="8B9536A64249FE79FF6B4CBA11BCF950" blockId="23.[151,1437,459,2034]" pageId="23" pageNumber="24">
|
||
<emphasis id="B95EEAB44249FE79FF6B4CBA1168FA54" box="[199,606,1468,1494]" italics="true" pageId="23" pageNumber="24">Coloration and pattern (in alcohol):</emphasis>
|
||
Ground color of dorsum greenish-brown, with two irregular paravertebral rows of dark spots. Similar dark spotting is present on each side of the body, that show the reminiscent of a reticulate pattern reduced to only isolated irregular spots. Between these two areas of dark spots, located both in the middle of the dorsum and flank, a paler and spot-free double strip area extends from nape to base of the tail (see photos in Appendix 5). There are few pale spots on dark bandings on the flanks, and more on the forelimb. A few spots near the basis of the forelimb are bluish. The belly, along with the head and neck, is whitish (Appendix 5). The background color of head plates is brownish, with a few scattered and small black spots. The first longitudinal row of ventral plates has dark spots on each side.
|
||
</paragraph>
|
||
<paragraph id="8B9536A64249FE79FF6B4FDD176DF84C" blockId="23.[151,1437,459,2034]" pageId="23" pageNumber="24">
|
||
<emphasis id="B95EEAB44249FE79FF6B4FDD12D2F974" bold="true" box="[199,484,1755,1782]" pageId="23" pageNumber="24">
|
||
Variations of
|
||
<typeStatus id="549188044249FE79FEC84FDA12E9F974" box="[356,479,1756,1782]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Descriptive statistics and variation range of the morphometric and scalation characters are given in Table 8. Frontonasal rarely contacts with rostral. The suture between the prefrontal and frontal is usually slanted. The Parietal is in contact with the postorbital in general. Supraciliar granules sometimes are double rows. Tight scales are feeble-keeled. In
|
||
<specimenCount id="9D2CFD2F4249FE79FDBB4E4E11F0F8E0" box="[535,710,1864,1890]" pageId="23" pageNumber="24" type="generic">four specimens</specimenCount>
|
||
, transverse dark spots on ground color are combined with spots on each side, do not form two separate rows. In seven samples, paler spots on the forelimb base are not blue. Belly, along with throat and neck, whitish in
|
||
<specimenCount id="9D2CFD2F4249FE79FDF04E961028F828" box="[604,798,1936,1962]" pageId="23" pageNumber="24" type="generic">seven specimens</specimenCount>
|
||
. In three samples, no dotting on head plates. In fifteen samples, the first longitudinal row of ventral plates contains bluish spots on each side.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330652D4249FE76FF6B4ED11013FF56" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="distribution">
|
||
<paragraph id="8B9536A64249FE79FF6B4ED11740F870" blockId="23.[151,1437,459,2034]" box="[199,1142,2007,2034]" pageId="23" pageNumber="24">
|
||
<emphasis id="B95EEAB44249FE79FF6B4ED11255F873" bold="true" box="[199,355,2007,2033]" pageId="23" pageNumber="24">Distribution:</emphasis>
|
||
Around Narlıca Valley, Çatak (
|
||
<collectingRegion id="49EEF8444249FE79FD694EDE11C5F870" box="[709,755,2008,2034]" country="Turkey" name="Van" pageId="23" pageNumber="24">Van</collectingRegion>
|
||
) in the south of
|
||
<collectingRegion id="49EEF8444249FE79FC044EDE10E2F870" box="[936,980,2008,2034]" country="Turkey" name="Van" pageId="23" pageNumber="24">Van</collectingRegion>
|
||
Lake,
|
||
<collectingCountry id="F33D76364249FE79FB8C4EDE1744F870" box="[1056,1138,2008,2034]" name="Turkey" pageId="23" pageNumber="24">Turkey</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9536A64246FE76FF6B49901013FF56" blockId="24.[151,1437,150,213]" pageId="24" pageNumber="25">
|
||
<emphasis id="B95EEAB44246FE76FF6B4990121AFF32" bold="true" box="[199,300,150,176]" pageId="24" pageNumber="25">Habitat:</emphasis>
|
||
Subalpine-like vegetation of Irano-Turanian Region, on rocky and stony areas,
|
||
<quantity id="4CD29B434246FE76FB614990161EFF33" box="[1229,1320,150,177]" metricMagnitude="3" metricUnit="m" metricValue="2.363" pageId="24" pageNumber="25" unit="m" value="2363.0">2363 m</quantity>
|
||
. No other reptile species could be identified in the area during study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |