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<document id="9AA8E7ED3A6E1254F992CB4756467FAE" ID-CLB-Dataset="55995" ID-DOI="10.1093/zoolinnean/zlab066" ID-GBIF-Dataset="aeb1ef2c-5c50-4d26-aa76-87d90d611e1c" ID-ISSN="0024-4082" ID-Zenodo-Dep="6459236" ID-ZooBank="A80D69C9-219C-40AA-B07E-FE6DEB567497" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649833241178" checkinUser="plazi" docAuthor="Recuero, Ernesto, Rodríguez-Flores, Paula C. &amp; García-París, Mario" docDate="2022" docId="BD486271FF978D593A19FD06FDF0FBBA" docLanguage="en" docName="zlab066.pdf" docOrigin="Zoological Journal of the Linnean Society 194 (4)" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Iberiarmadillidium Recuero &amp; Rodríguez-Flores &amp; García-París 2022, GEN. NOV." docType="treatment" docVersion="8" lastPageNumber="1327" masterDocId="41711A09FF998D563B0EFF97FF86FFAC" masterDocTitle="Homoplasy and morphological stasis revealed through multilocus phylogeny of new myrmecophilous species in Armadillidiidae (Isopoda: Oniscidea)" masterLastPageNumber="1340" masterPageNumber="1312" pageNumber="1326" updateTime="1699328811996" updateUser="ExternalLinkService">
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<mods:title id="CA27CA63467C950B365C0B2073A44B87">Homoplasy and morphological stasis revealed through multilocus phylogeny of new myrmecophilous species in Armadillidiidae (Isopoda: Oniscidea)</mods:title>
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<mods:namePart id="F8A55EAAD195DFCFE1C31ACF5AD621EC">Recuero, Ernesto</mods:namePart>
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<mods:namePart id="D35049C759184263947F96F10497DA2F">Rodríguez-Flores, Paula C.</mods:namePart>
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<mods:namePart id="105886202C89A1767CF29E732EFD1CD6">García-París, Mario</mods:namePart>
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<mods:title id="F2681DA0771D0C1859A91831EE6F52D5">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="5ECD777A0EFF3476912137AF2D0D9978">2022</mods:date>
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<mods:number id="95248918B5105BA8CC006DEEE6131A13">2021-12-04</mods:number>
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<mods:number id="EE2E4B60642BF0D593E9AF37AF81BA0A">194</mods:number>
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<mods:identifier id="4C8BA6EF51DEC6D8144503CA5F3A771A" type="DOI">10.1093/zoolinnean/zlab066</mods:identifier>
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<treatment id="BD486271FF978D593A19FD06FDF0FBBA" ID-DOI="http://doi.org/10.5281/zenodo.6457113" ID-GBIF-Taxon="194594435" ID-Zenodo-Dep="6457113" LSID="urn:lsid:plazi:treatment:BD486271FF978D593A19FD06FDF0FBBA" httpUri="http://treatment.plazi.org/id/BD486271FF978D593A19FD06FDF0FBBA" lastPageId="15" lastPageNumber="1327" pageId="14" pageNumber="1326">
<subSubSection id="7DFB80ECFF978D583A19FD06FE65FC80" pageId="14" pageNumber="1326" type="nomenclature">
<paragraph id="355ED367FF978D583A19FD06FD11FD05" blockId="14.[279,663,657,681]" box="[279,663,657,681]" pageId="14" pageNumber="1326">
<heading id="6E16640BFF978D583A19FD06FD11FD05" box="[279,663,657,681]" centered="true" fontSize="9" level="2" pageId="14" pageNumber="1326" reason="2">
<taxonomicName id="F2E1A8E4FF978D583A19FD06FD9EFD0B" ID-CoL="8TG89" authority="Recuero &amp; Rodríguez-Flores &amp; García-París, 2022" authorityName="Recuero &amp; Rodríguez-Flores &amp; García-París" authorityYear="2022" box="[279,536,657,680]" class="Malacostraca" family="Armadillidiidae" genus="Iberiarmadillidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus" status="gen. nov.">IBERIARMADILLIDIUM</taxonomicName>
<emphasis id="07950F75FF978D583911FD03FD11FD05" bold="true" box="[543,663,657,681]" pageId="14" pageNumber="1326">
<taxonomicNameLabel id="1CA6B20EFF978D583911FD03FD11FD05" box="[543,663,657,681]" pageId="14" pageNumber="1326" rank="genus">GEN. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="355ED367FF978D583BADFD4DFE65FC80" blockId="14.[163,779,729,812]" pageId="14" pageNumber="1326">
<emphasis id="07950F75FF978D583BADFD4DFE65FD43" box="[163,483,729,751]" italics="true" pageId="14" pageNumber="1326">Z o o b a n k r e g i s t r a t i o n:</emphasis>
h t t p: / / z o o b a n k. o r g / u r n: l s i d: z o o b a n k. o r g: a c t:
<uuid id="4147E9B2FF978D58391AFD6FFE65FC80" pageId="14" pageNumber="1326">3 2 B C 4 C A 3 - A 6 4 3 - 4C5F-B69D-815CF20C6F34</uuid>
</paragraph>
</subSubSection>
<subSubSection id="7DFB80ECFF978D593BADFCC7FDF0FBBA" lastPageId="15" lastPageNumber="1327" pageId="14" pageNumber="1326" type="materials_examined">
<paragraph id="355ED367FF978D583BADFCC7FDB9FC28" blockId="14.[163,779,848,900]" pageId="14" pageNumber="1326">
<emphasis id="07950F75FF978D583BADFCC7FD65FCC9" box="[163,739,848,869]" italics="true" pageId="14" pageNumber="1326">
<typeStatus id="EA5A6DC5FF978D583BADFCC7FF5FFCC9" box="[163,217,848,869]" pageId="14" pageNumber="1326">Type</typeStatus>
species:
<taxonomicName id="F2E1A8E4FF978D583A4DFCC7FD65FCC9" authority="Recuero &amp; Rodríguez-Flores &amp; García-París, 2022" authorityName="Recuero &amp; Rodríguez-Flores &amp; García-París" authorityYear="2022" box="[323,739,848,869]" class="Malacostraca" family="Armadillidiidae" genus="Iberiarmadillidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="species" species="psammophilum" status="sp. nov.">Iberiarmadillidium psammophilum</taxonomicName>
</emphasis>
<taxonomicNameLabel id="1CA6B20EFF978D5839E5FCC7FF49FC28" pageId="14" pageNumber="1326" rank="species">sp. nov.</taxonomicNameLabel>
by present original designation.
</paragraph>
<paragraph id="355ED367FF978D583BADFC3AFE9BFBAC" blockId="14.[163,778,941,1024]" pageId="14" pageNumber="1326">
<emphasis id="07950F75FF978D583BADFC3AFEAEFC6E" box="[163,296,941,962]" italics="true" pageId="14" pageNumber="1326">Etymology:</emphasis>
From the Latin
<emphasis id="07950F75FF978D583AF3FC3AFDC3FC6E" box="[509,581,941,962]" italics="true" pageId="14" pageNumber="1326">
<collectingCountry id="4DF693F7FF978D583AF3FC3AFDC3FC6E" box="[509,581,941,962]" name="Georgia" pageId="14" pageNumber="1326">Iberia</collectingCountry>
</emphasis>
,
<collectingCountry id="4DF693F7FF978D583959FC3AFD19FC6F" box="[599,671,941,963]" name="Spain" pageId="14" pageNumber="1326">Spain</collectingCountry>
, and the genus
<taxonomicName id="F2E1A8E4FF978D583BFDFC5BFE2CFC4D" baseAuthorityName="Arcangeli" baseAuthorityYear="1935" box="[243,426,972,993]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583BFDFC5BFE2CFC4D" box="[243,426,972,993]" italics="true" pageId="14" pageNumber="1326">Armadillidium</emphasis>
</taxonomicName>
, which in turn means ‘small armadillo’.
</paragraph>
<paragraph id="355ED367FF978D583BADFBBDFE99F8B3" blockId="14.[163,779,1065,1823]" pageId="14" pageNumber="1326">
<emphasis id="07950F75FF978D583BADFBBDFE99FB93" box="[163,287,1066,1087]" italics="true" pageId="14" pageNumber="1326">Diagnosis:</emphasis>
Endoantennal, euspheric conglobation.
<taxonomicName id="F2E1A8E4FF978D583BADFBDFFF74FBF1" authorityName="Budde-Lund" authorityYear="1885" box="[163,242,1096,1117]" class="Malacostraca" family="Armadillidiidae" genus="Eluma" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583BADFBDFFF74FBF1" box="[163,242,1096,1117]" italics="true" pageId="14" pageNumber="1326">Eluma</emphasis>
</taxonomicName>
-
<typeStatus id="EA5A6DC5FF978D583BF4FBDFFEA8FBF1" box="[250,302,1096,1117]" pageId="14" pageNumber="1326">type</typeStatus>
cephalon, with a frontal line continuing the scutellar upper margin and no trace of a postscutellar line. Scutellum triangular, wider than high, slightly concave in the middle, clearly separated from vertex but not protruding above it. Antennary lobes well developed, obliquely directed frontwards. Eyes small, with four to six pigmented ommatidia. Cephalic shield covered by three or four transverse rows of tubercles, irregularly arranged except for the caudal one. Pereonites with two or three transverse rows of large tubercles. Pleonites with one transverse row of tubercles. Pereonite 1 epimera with a small schisma. Shape of distal part of pereonite 4 and 5 epimera broad, with squarish to broadly rounded angles. Epimera of pereonites 2 or 2 and 3 with sinuous, slightly concave outer margin. Surface of pereon, pleon and epimera mostly covered by round, convex, scale-like papillae with irregular margins. Glandular fields in pereon 2 and 3 epimera with one or two pores, longitudinally elongated, elliptical, submedially located in the lateral margin. Pleopod 1 exopod with concave proximal and distal margins. Pleopod 2 exopod clearly longer (~1.5 times) than wide.
</paragraph>
<paragraph id="355ED367FF978D593BADF8D7FF6AFD48" blockId="14.[163,778,1856,1908]" lastBlockId="15.[145,761,197,1047]" lastPageId="15" lastPageNumber="1327" pageId="14" pageNumber="1326">
<emphasis id="07950F75FF978D583BADF8D7FE93F8F9" box="[163,277,1856,1877]" italics="true" pageId="14" pageNumber="1326">Remarks:</emphasis>
This new genus can be diagnosed from all other
<taxonomicName id="F2E1A8E4FF978D583A1AF8C9FE55F8D8" authorityName="Brandt" authorityYear="1833" box="[276,467,1886,1908]" class="Malacostraca" family="Armadillidiidae" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="family">Armadillidiidae</taxonomicName>
genera by a combination of morphological characters. From
<taxonomicName id="F2E1A8E4FF978D583FE2FF52FC5CFF55" authority="Verhoeff, 1919" authorityName="Verhoeff" authorityYear="1919" class="Malacostraca" family="Armadillidiidae" genus="Alloschizidium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583FE2FF52FA25FF76" box="[1260,1443,197,218]" italics="true" pageId="14" pageNumber="1326">Alloschizidium</emphasis>
<bibRefCitation id="5170AE96FF978D583835FF73FC5CFF55" author="Verhoeff K" box="[827,986,228,250]" pageId="14" pageNumber="1326" pagination="160 - 170" refId="ref19612" refString="Verhoeff K. 1919. Ueber augenlose Armadillidien und kritische Prufung der Familie Armadillidiidae. 25. Isopoden-Aufsatz. Archiv fur Naturgeschichte, Abteilung A 83: 160 - 170." type="journal article" year="1919">Verhoeff, 1919</bibRefCitation>
</taxonomicName>
it differs in having an
<taxonomicName id="F2E1A8E4FF978D583FD5FF73FAA1FF55" authorityName="Budde-Lund" authorityYear="1885" box="[1243,1319,228,249]" class="Malacostraca" family="Armadillidiidae" genus="Eluma" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583FD5FF73FAA1FF55" box="[1243,1319,228,249]" italics="true" pageId="14" pageNumber="1326">Eluma</emphasis>
</taxonomicName>
-
<typeStatus id="EA5A6DC5FF978D583E20FF73FAD9FF55" box="[1326,1375,228,249]" pageId="14" pageNumber="1326">type</typeStatus>
head, a rounded triangular telson [truncate in all species of
<taxonomicName id="F2E1A8E4FF978D583835FEB6FC64FE9A" authorityName="Verhoeff" authorityYear="1919" box="[827,994,289,310]" class="Malacostraca" family="Armadillidiidae" genus="Alloschizidium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583835FEB6FC64FE9A" box="[827,994,289,310]" italics="true" pageId="14" pageNumber="1326">Alloschizidium</emphasis>
</taxonomicName>
except
<emphasis id="07950F75FF978D583F20FEB6FAC3FE9A" box="[1070,1349,289,310]" italics="true" pageId="14" pageNumber="1326">Alloschizidiumracovitzai</emphasis>
(Vandel, 1954)] reduced but well-marked eyes, pigmentation developed, and presence of tubercles in pereon and pleon tergites; from
<taxonomicName id="F2E1A8E4FF978D5838EDFEEAFB12FE3E" baseAuthorityName="Arcangeli" baseAuthorityYear="1935" box="[995,1172,381,402]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D5838EDFEEAFB12FE3E" box="[995,1172,381,402]" italics="true" pageId="14" pageNumber="1326">Armadillidium</emphasis>
</taxonomicName>
it differs in having an
<taxonomicName id="F2E1A8E4FF978D583835FE0BFC0EFE1D" authorityName="Budde-Lund" authorityYear="1885" box="[827,904,412,433]" class="Malacostraca" family="Armadillidiidae" genus="Eluma" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583835FE0BFC0EFE1D" box="[827,904,412,433]" italics="true" pageId="14" pageNumber="1326">Eluma</emphasis>
</taxonomicName>
-
<typeStatus id="EA5A6DC5FF978D583881FE0BFC47FE1D" box="[911,961,412,433]" pageId="14" pageNumber="1326">type</typeStatus>
head, presence of a schisma in pereonite 1 epimera (present in a few
<taxonomicName id="F2E1A8E4FF978D583F53FE2DFADDFE7C" box="[1117,1371,442,464]" pageId="14" pageNumber="1326">
<emphasis id="07950F75FF978D583F53FE2DFA80FE63" box="[1117,1286,442,463]" italics="true" pageId="14" pageNumber="1326">Armadillidium</emphasis>
species
</taxonomicName>
in the
<taxonomicName id="F2E1A8E4FF978D583835FE4EFBC7FE43" authorityName="Racovitza" authorityYear="1907" box="[827,1089,473,495]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="species" species="pruvoti">
<emphasis id="07950F75FF978D583835FE4EFBC7FE43" box="[827,1089,473,495]" italics="true" pageId="14" pageNumber="1326">Armadillidium pruvoti</emphasis>
</taxonomicName>
group) and reduced eyes; from
<taxonomicName id="F2E1A8E4FF978D583835FE6FFC4FFDA1" authorityName="Vandel" authorityYear="1961" box="[827,969,504,525]" class="Malacostraca" family="Armadillidiidae" genus="Ballodillium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583835FE6FFC4FFDA1" box="[827,969,504,525]" italics="true" pageId="14" pageNumber="1326">Ballodillium</emphasis>
</taxonomicName>
it differs in having tubercles in pereon and pleon tergites, reduced eyes, lacking pilose setae covering the body, and absence of a groove along the margin of pereonite 1 epimera. From
<taxonomicName id="F2E1A8E4FF978D583F8CFDC3FAEBFDC5" authorityName="Arcangeli" authorityYear="1935" box="[1154,1389,596,617]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583F8CFDC3FAEBFDC5" box="[1154,1389,596,617]" italics="true" pageId="14" pageNumber="1326">Cristarmadillidium</emphasis>
</taxonomicName>
, the most similar genus morphologically, it differs in having sinuous, slightly concave outer margin of pereonite 2 epimera, pereonite 4 and 5 epimera distally broad, with squarish to broadly rounded angles, tegument covered by round, convex, scale-like papillae, glandular fields in pereon 2 and 3 epimera longitudinally elongated and close to the lateral margin, and pleopod 2 exopod much longer than wide. From
<taxonomicName id="F2E1A8E4FF978D583F44FCDEFA25FCF2" authority="Verhoeff, 1939" authorityName="Verhoeff" authorityYear="1939" box="[1098,1443,841,863]" class="Malacostraca" family="Armadillidiidae" genus="Cyphodillidium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583F44FCDEFB7AFCF2" box="[1098,1276,841,862]" italics="true" pageId="14" pageNumber="1326">Cyphodillidium</emphasis>
Verhoeff, 1939
</taxonomicName>
it differs in having a rounded triangular telson, uropod exopodite about two times as broad as long, reduced eyes, and posterior corner of pereonite 1 epimera not produced acutely backwards. From
<taxonomicName id="F2E1A8E4FF978D583FB2FC54FC5CFC5B" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" class="Malacostraca" family="Armadillidiidae" genus="Echinarmadillidium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583FB2FC54FA25FC74" box="[1212,1443,963,984]" italics="true" pageId="14" pageNumber="1326">Echinarmadillidium</emphasis>
Verhoeff, 1901
</taxonomicName>
it differs in having an
<taxonomicName id="F2E1A8E4FF978D583FD5FC75FAA1FC5B" authorityName="Budde-Lund" authorityYear="1885" box="[1243,1319,994,1015]" class="Malacostraca" family="Armadillidiidae" genus="Eluma" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583FD5FC75FAA1FC5B" box="[1243,1319,994,1015]" italics="true" pageId="14" pageNumber="1326">Eluma</emphasis>
</taxonomicName>
-
<typeStatus id="EA5A6DC5FF978D583E20FC74FAD9FC54" box="[1326,1375,995,1016]" pageId="14" pageNumber="1326">type</typeStatus>
head, a rounded triangular telson, reduced but well-marked eyes, absence of a schisma in pereonite 2 epimera, and absence of a groove along the margin of pereonite 1 epimera. From
<taxonomicName id="F2E1A8E4FF978D5838FCFBCAFAB4FBDE" authority="Racovitza, 1907" authorityName="Racovitza" authorityYear="1907" box="[1010,1330,1117,1139]" class="Malacostraca" family="Armadillidiidae" genus="Eleoniscus" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D5838FCFBCAFBE9FBDE" box="[1010,1135,1117,1138]" italics="true" pageId="14" pageNumber="1326">Eleoniscus</emphasis>
Racovitza, 1907
</taxonomicName>
it differs in having eyes, presence of a schisma in pereonite 1 epimera, pigmentation developed, presence of tubercles in pereon and pleon tergites, uropod exopodite about two times as broad as long, and lacking pilose setae covering the body. From
<taxonomicName id="F2E1A8E4FF978D583F68FB61FB30FAA7" authorityName="Budde-Lund" authorityYear="1885" box="[1126,1206,1270,1291]" class="Malacostraca" family="Armadillidiidae" genus="Eluma" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583F68FB61FB30FAA7" box="[1126,1206,1270,1291]" italics="true" pageId="14" pageNumber="1326">Eluma</emphasis>
</taxonomicName>
it differs in having eyes with several ommatidia instead of the single ommatidium typical of
<taxonomicName id="F2E1A8E4FF978D583F45FAA4FB1FFAE4" authorityName="Budde-Lund" authorityYear="1885" box="[1099,1177,1331,1352]" class="Malacostraca" family="Armadillidiidae" genus="Eluma" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583F45FAA4FB1FFAE4" box="[1099,1177,1331,1352]" italics="true" pageId="14" pageNumber="1326">Eluma</emphasis>
</taxonomicName>
. From
<taxonomicName id="F2E1A8E4FF978D583FE3FAA4FC5EFACB" authority="Verhoeff, 1919" authorityName="Verhoeff" authorityYear="1919" class="Malacostraca" family="Armadillidiidae" genus="Paraschizidium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583FE3FAA4FA25FAE4" box="[1261,1443,1331,1352]" italics="true" pageId="14" pageNumber="1326">Paraschizidium</emphasis>
<bibRefCitation id="5170AE96FF978D583835FAC5FC5EFACB" author="Verhoeff K" box="[827,984,1362,1384]" pageId="14" pageNumber="1326" pagination="160 - 170" refId="ref19612" refString="Verhoeff K. 1919. Ueber augenlose Armadillidien und kritische Prufung der Familie Armadillidiidae. 25. Isopoden-Aufsatz. Archiv fur Naturgeschichte, Abteilung A 83: 160 - 170." type="journal article" year="1919">Verhoeff, 1919</bibRefCitation>
</taxonomicName>
it differs in having a schisma in pereonite 1 epimera, reduced but well-marked eyes, threesegmented antennula, pigmentation developed, and presence of tubercles in pereon and pleon tergites. From
<taxonomicName id="F2E1A8E4FF978D583835FA5AFB28FA4F" authority="Schmalfuss, 1985" authorityName="Schmalfuss" authorityYear="1985" box="[827,1198,1485,1507]" class="Malacostraca" family="Armadillidiidae" genus="Paxodillidium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583835FA5AFC58FA4E" box="[827,990,1485,1506]" italics="true" pageId="14" pageNumber="1326">Paxodillidium</emphasis>
Schmalfuss, 1985
</taxonomicName>
it differs in having a schisma in pereonite 1 epimera, two or three transverse rows of tubercles on each pereon tergite, rectangularshaped uropod exopodite, and a rounded triangular telson. From
<emphasis id="07950F75FF978D5838C1F9D0FBF4F9F0" box="[975,1138,1607,1628]" italics="true" pageId="14" pageNumber="1326">Platanosphera</emphasis>
Strouhal, 1956 it differs in in having a schisma in pereonite 1 epimera, two or three transverse rows of tubercles on each pereon tergite, tegument covered by round, convex, scale-like papillae and male pleopod exopodite 1 with hindlobe. From
<taxonomicName id="F2E1A8E4FF978D583835F976FBDAF95A" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" box="[827,1116,1761,1783]" class="Malacostraca" family="Armadillidiidae" genus="Schizidium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583835F976FC3DF95A" box="[827,955,1761,1782]" italics="true" pageId="14" pageNumber="1326">Schizidium</emphasis>
Verhoeff, 1901
</taxonomicName>
it differs in having a rounded triangular telson and presence of tubercles in pereon and pleon tergites. From
<taxonomicName id="F2E1A8E4FF978D583F2BF889FA24F89F" authority="Schmalfuss, 1989" authorityName="Schmalfuss" authorityYear="1989" box="[1061,1442,1822,1844]" class="Malacostraca" family="Armadillidiidae" genus="Trichodillidium" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583F2BF889FB5EF89F" box="[1061,1240,1822,1843]" italics="true" pageId="14" pageNumber="1326">Trichodillidium</emphasis>
<bibRefCitation id="5170AE96FF978D583FD1F889FA24F89F" author="Schmalfuss H" box="[1247,1442,1822,1844]" pageId="14" pageNumber="1326" pagination="207 - 214" refId="ref18668" refString="Schmalfuss H. 1989. Die Land-Isopoden (Oniscidea) Griechenlands. 10. Beitrag: Gattung Trichodillidium g. n. (Armadillidiidae). Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch- Naturwissenschaftliche Klasse Abteilung I 197: 207 - 214." type="journal article" year="1989">Schmalfuss, 1989</bibRefCitation>
</taxonomicName>
it differs in having an
<taxonomicName id="F2E1A8E4FF978D583F34F8AAFB01F8FE" authorityName="Budde-Lund" authorityYear="1885" box="[1082,1159,1853,1874]" class="Malacostraca" family="Armadillidiidae" genus="Eluma" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1326" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF978D583F34F8AAFB01F8FE" box="[1082,1159,1853,1874]" italics="true" pageId="14" pageNumber="1326">Eluma</emphasis>
</taxonomicName>
-
<typeStatus id="EA5A6DC5FF978D583F80F8AAFB46F8FE" box="[1166,1216,1853,1874]" pageId="14" pageNumber="1326">type</typeStatus>
head, reduced eyes, pigmentation developed, presence of tubercles in pereon and pleon tergites and lacking pilose setae covering the body. From
<taxonomicName id="F2E1A8E4FF968D593A1EFF73FD12FF55" authority="Verhoeff, 1900" authorityName="Verhoeff" authorityYear="1900" box="[272,660,228,250]" class="Malacostraca" family="Armadillidiidae" genus="Troglarmadillidium" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1327" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF968D593A1EFF73FE77FF55" box="[272,497,228,249]" italics="true" pageId="15" pageNumber="1327">Troglarmadillidium</emphasis>
Verhoeff, 1900
</taxonomicName>
it differs in having a schisma in pereonite 1 epimera, reduced but well-marked eyes, pigmentation developed, uropod exopodite about two times as broad as long, presence of a frontal scutellum in the cephalon, presence of a frontal line and antennal lobes and presence of tubercles in pereon and pleon tergites; from
<taxonomicName id="F2E1A8E4FF968D593AE6FE0BFD7FFE1E" authority="Vandel, 1946" authorityName="Vandel" authorityYear="1946" box="[488,761,412,434]" class="Malacostraca" family="Armadillidiidae" genus="Trogleluma" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1327" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF968D593AE6FE0BFDEEFE1D" box="[488,616,412,433]" italics="true" pageId="15" pageNumber="1327">Trogleluma</emphasis>
Vandel, 1946
</taxonomicName>
it differs in having a schisma in pereonite 1 epimera, reduced but well-marked eyes, pigmentation developed, uropod exopodite about two times as broad as long and presence of tubercles in pereon and pleon tergites. From
<taxonomicName id="F2E1A8E4FF968D593B9FFDA2FD99FDE6" authority="Verhoeff, 1900" authorityName="Verhoeff" authorityYear="1900" box="[145,543,565,587]" class="Malacostraca" family="Armadillidiidae" genus="Typhlarmadillidium" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1327" phylum="Arthropoda" rank="genus">
<emphasis id="07950F75FF968D593B9FFDA2FEFFFDE6" box="[145,377,565,586]" italics="true" pageId="15" pageNumber="1327">Typhlarmadillidium</emphasis>
Verhoeff, 1900
</taxonomicName>
it differs in having a schisma in pereonite 1 epimera, reduced but wellmarked eyes, pigmentation developed, uropod exopodite about two times as broad as long, frontal scutellum closed and presence of tubercles in pereon and pleon tergites.
</paragraph>
<paragraph id="355ED367FF968D593BA7FD7AFDF0FBBA" blockId="15.[145,761,197,1047]" pageId="15" pageNumber="1327">
All three
<taxonomicName id="F2E1A8E4FF968D593A24FD7AFD0AFCAF" box="[298,652,749,771]" pageId="15" pageNumber="1327">
<emphasis id="07950F75FF968D593A24FD7AFDA0FCAE" box="[298,550,749,770]" italics="true" pageId="15" pageNumber="1327">Iberiarmadillidium</emphasis>
species
</taxonomicName>
share a similar general habitus (
<figureCitation id="ADDACFE2FF968D593AB1FC9BFDB0FC8E" box="[447,566,780,802]" captionStart="Figure 5" captionStartId="8.[163,244,1675,1697]" captionTargetBox="[285,1321,195,1635]" captionTargetId="figure-26@8.[285,1321,195,1635]" captionTargetPageId="8" captionText="Figure 5. Scanning electron photomicrographs showing the general habitus of Iberiarmadillidium psammophilum (A), Iberiarmadillidium pinicola (B), Iberiarmadillidium sakura (C), Cristarmadillidium myrmecophilum (D) and Cristarmadillidium muricatum (E)." figureDoi="http://doi.org/10.5281/zenodo.6459250" httpUri="https://zenodo.org/record/6459250/files/figure.png" pageId="15" pageNumber="1327">Fig. 5A–C</figureCitation>
). However, they also present high genetic divergences, particularly for the mitochondrial DNA gene
<emphasis id="07950F75FF968D593924FCDEFDE0FCF2" box="[554,614,841,862]" italics="true" pageId="15" pageNumber="1327">Cox1</emphasis>
, with mean interspecific uncorrected
<emphasis id="07950F75FF968D593AD9FCFFFE63FCD1" box="[471,485,872,893]" italics="true" pageId="15" pageNumber="1327">p</emphasis>
-distances of 20–21%. In the case of nuclear markers, with much slower substitution rates than
<emphasis id="07950F75FF968D593AC8FC32FD85FC16" box="[454,515,933,954]" italics="true" pageId="15" pageNumber="1327">Cox1</emphasis>
, mean uncorrected
<emphasis id="07950F75FF968D593B9FFC53FF19FC75" box="[145,159,964,985]" italics="true" pageId="15" pageNumber="1327">p</emphasis>
-distances among species ranged from 0.5 to 0.7% in the analysed fragment of the 18S, from 0.20 to 0.35% in 28S and from 0.7 to 2.3% in
<emphasis id="07950F75FF968D593942FB96FDE8FBBA" box="[588,622,1025,1046]" italics="true" pageId="15" pageNumber="1327">H3</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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