264 lines
33 KiB
XML
264 lines
33 KiB
XML
<document ID-CLB-Dataset="122795" ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-GBIF-Dataset="5f938945-980c-433c-942a-f05d39529a25" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo" docDate="2022" docId="A7E8B110FB875A6CBAAC65042B2B7CE5" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Ayacucho titschacki Roewer 1949" docType="treatment" docVersion="5" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1732651490011" updateUser="ExternalLinkService">
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<mods:title id="1A549C259814A37E7788FA52648DA329">Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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<mods:name id="C22DA61E95D79BFD28EFFE599ADA7D2F" type="personal">
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<mods:namePart id="D4C0F02CD76BF80D410F57E14AC7E968">Benedetti, Alipio Rezende</mods:namePart>
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<mods:affiliation id="C49A565BC0DD5A9E0895330E107D2F12">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil & Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
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<mods:nameIdentifier id="362D44746563D5C229977FC3A71922ED" type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0BCADEDED9A7E87047ADCD66E71BC148">Pinto-da-Rocha, Ricardo</mods:namePart>
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<mods:nameIdentifier id="2AF8A4EFF7EDD417F712027EDEDF98F9" type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
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<mods:affiliation id="D1CCF288535AE242C71015271F3BFA4B">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:title id="C423480C906B9B0100807CC58EC21781">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:part id="178A9255FE6D455FECDC97AA84FF878F">
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<mods:date id="B83307FC2F09B3B8458C524D5965AEB2">2022</mods:date>
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<mods:number id="B6CD22EACCBB72043178086D05A922E5">80</mods:number>
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<treatment id="A7E8B110FB875A6CBAAC65042B2B7CE5" ID-GBIF-Taxon="199468699" LSID="urn:lsid:plazi:treatment:A7E8B110FB875A6CBAAC65042B2B7CE5" httpUri="http://treatment.plazi.org/id/A7E8B110FB875A6CBAAC65042B2B7CE5" lastPageNumber="309" pageId="0" pageNumber="309" scope_family="Metasarcidae" scope_order="Opiliones" scope_subOrder="Laniatores">
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<subSubSection id="FA90B272DCD4334F0F82B955D79C5BED" pageId="0" pageNumber="309" type="nomenclature">
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<paragraph id="3F40AC839A849951C279665D07B7394F" pageId="0" pageNumber="309">
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3.23.
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<taxonomicName id="076CBDB6B8247BE82AEF774828BD7C89" ID-CoL="7S8SN" LSID="A7E8B110-FB87-5A6C-BAAC-65042B2B7CE5" authority="Roewer, 1949" authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho titschacki" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="titschacki">Ayacucho titschacki Roewer, 1949</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="E89F66232BB4A147E346152C52417E72" pageId="0" pageNumber="309" type="description">
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<paragraph id="9AE061197E739716E155C014725069F4" pageId="0" pageNumber="309">
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<figureCitation id="ED590AF022FEC849422114C02CC8F562" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">Figs 3D</figureCitation>
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<figureCitation id="D88DD2AF8A98A6BE79BA8847A9A498D8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">, 7U, V</figureCitation>
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<figureCitation id="62B4CCA97ABE2D3AED4413139AB95A01" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D Ayacucho inermis (Roewer, 1957) comb. nov.; E - F Ayacucho pasolinii sp. nov.; G - H Ayacucho spielbergi sp. nov.; I - J Ayacucho titschacki Roewer, 1949; K - L Ayacucho triarmatus nom. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure12" httpUri="https://binary.pensoft.net/fig/729886" pageId="0" pageNumber="309">, 12I, J</figureCitation>
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<figureCitation id="3BD1DA4A1EF860D9891C16C302A84275" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Live specimens of Ayacucho. A Ayacucho spiniger (Roewer, 1957) comb. nov., male; B female; C A. tapacocha nom. nov., male; D female; E A. titschacki Roewer, 1949, male; F A. vargasllosai sp. nov., male; G A. uniseriatus (Roewer, 1959) comb. nov., male; H: female." figureDoi="10.3897/asp.80.e73829.figure23" httpUri="https://binary.pensoft.net/fig/729897" pageId="0" pageNumber="309">, 23E</figureCitation>
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<figureCitation id="F6D4D88EA27E8895CB1F3FA12F11ADD0" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">, 28</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="565807399C3A397BE4BE8FCBE5DB1575" pageId="0" pageNumber="309" type="reference_group">
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<paragraph id="6CE09EDFCB00D60D308B995196BBA70F" pageId="0" pageNumber="309">
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<taxonomicName id="6BD1D70C9DF67F72F23BABB3A19AAC95" authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho titschacki" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="titschacki">Ayacucho titschacki</taxonomicName>
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Roewer, 1949: 57 (desc.), figs. 114a (dorsal habitus), 114b (ventral habitus), 114c (chelicera, pedipalpus and carapace in lateral view); 114d (detail of coxa IV), 114e (sternum), 114f (chelicera in lateral view);
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<bibRefCitation id="E3B867653FA1B07A85134EC4A774DEAE" author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
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: 144 (cat.).
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</paragraph>
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</subSubSection>
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<subSubSection id="3EFD8DD0C5F3217E4BEB3EEB26E06609" pageId="0" pageNumber="309" type="redescription">
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<paragraph id="C065A839F0E6573BFDE9C24E9B0ACF5C" pageId="0" pageNumber="309">Redescription.</paragraph>
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<paragraph id="CCCF071FEA7CF51EE93F222470A83402" pageId="0" pageNumber="309">
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<emphasis id="AFD69CABEF5C8E78A29444B178676361" bold="true" pageId="0" pageNumber="309">
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MALE:
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<emphasis id="1FFB21837B60131899CF63FE5D5BC816" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
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</emphasis>
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(
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<emphasis id="3004F8F77FC076A8F46297418B6144B2" italics="true" pageId="0" pageNumber="309">n</emphasis>
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=2) DSW: 3.0-4.1; DSL: 4.5-5.4; CL: 1.5-2.2. FIVL: 4.8-5.4. ChL: 2.5-4.0.
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<emphasis id="7D70303567476FF5608C35D0E674553F" bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
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: (Fig.
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<figureCitation id="9059CE246F02B6F07818A5DDD0D09714" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Live specimens of Ayacucho. A Ayacucho spiniger (Roewer, 1957) comb. nov., male; B female; C A. tapacocha nom. nov., male; D female; E A. titschacki Roewer, 1949, male; F A. vargasllosai sp. nov., male; G A. uniseriatus (Roewer, 1959) comb. nov., male; H: female." figureDoi="10.3897/asp.80.e73829.figure23" httpUri="https://binary.pensoft.net/fig/729897" pageId="0" pageNumber="309">23E</figureCitation>
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) Predominantly yellowish, with two longitudinal black stripes in the lateral region of dorsal scutum (carapace and lateral edges of the areas I-IV). Lateral margins of DS yellowish, with small black spots. Posterior margin of the dorsal scutum, free tergites and legs predominantly black. Chelicerae and pedipalpus predominantly dark yellow.
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<emphasis id="FE5B00483F4F4D711A383DB568F88315" bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
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: (Fig.
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<figureCitation id="60FC4D7AA693B624A90FAF652BAA232D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3D</figureCitation>
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) Alpha-type DSS. Anterior margin of carapace with median elevation totally covered with granules. Ocularium with two small acuminate tubercles, absent in some individuals. Areas I-IV with two pairs of small median tubercles; the lateral pair can be greatly reduced in some individuals. Posterior margin of dorsal scutum with a transverse row of 4-5 acuminate tubercles (larger than tubercles of areas I-IV). Free tergites I-III with a row of tubercles similar to the posterior margin of dorsal scutum, increasing the number of free tergite I to III.
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<emphasis id="F01F4159001B0E2882C2967F68B0666A" bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
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: (Fig.
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<figureCitation id="233FF62C958247DAA65DCB7B705C5332" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3D</figureCitation>
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) Swollen or not in males. Segment I densely covered with granules throughout its length. Segment II predominantly smooth, with some hairs on the front face; finger with three teeth. Segment III with two teeth.
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<taxonomicName id="76B43C386BBF23EEF333B628EE679EEE" genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
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<emphasis id="05FA8E21F0C756CBBE17DA636F42FAAD" bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
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</taxonomicName>
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: With very small granules sparsely distributed on the dorsal surface of the femur and patella. Trochanter with a distal ventral setiferous tubercle. Femur with a ventrobasal setiferous tubercle and a row of six small ventromedian setiferous tubercles. Tibia: prolateral IIII, retrolateral IIII. Tarsus: prolateral IIi, retrolateral Iii.
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<emphasis id="CBDDB967105AFF3AC726BC2E43BA4E95" bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
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: Coxa I with dispersed small tubercles. Coxae II-III, predominantly smooth, with sparse granules. Coxa IV with granulation denser than coxae III-IV. Genital area smooth. Free sternites I-IV and anal operculum with small granules sparsely distributed.
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<emphasis id="93A4D921AB19A55AC413AC486E24CA74" bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
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: (Figs
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<figureCitation id="6D1EF8E76FC43743B16D176489133C53" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3D</figureCitation>
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,
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<figureCitation id="215353A70FA1FDBB45A103EF13076AD1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7U, V</figureCitation>
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) Coxa I with a prolateral apophysis; coxa II with a retrolateral and a prolateral apophysis. Coxa IV densely granulate, with larger tubercles distributed more densely in the prolateral portion. Trochanters I-IV with few granules, unarmed. Femora I-III, with granules distributed throughout its length. Femur IV with granules densely distributed in the dorsal and lateral, less heavily on the ventral surface. Tarsal formula: (
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<emphasis id="FFD943D52BEA42D481BD490E60030CAB" italics="true" pageId="0" pageNumber="309">n</emphasis>
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=2) 7, 12, 7, 8.
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<emphasis id="B7786DBDCBE255C8A201B53DA1293E20" bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
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: (Fig.
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<figureCitation id="43095EA0B4CB1AC14A8A4D877236CF54" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D Ayacucho inermis (Roewer, 1957) comb. nov.; E - F Ayacucho pasolinii sp. nov.; G - H Ayacucho spielbergi sp. nov.; I - J Ayacucho titschacki Roewer, 1949; K - L Ayacucho triarmatus nom. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure12" httpUri="https://binary.pensoft.net/fig/729886" pageId="0" pageNumber="309">12I, J</figureCitation>
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) VP rectangular; distal margin slightly concave, with short lateral-apical projections; slightly concave in lateral view. MS C1-C5 (or C6) subapical long and straight; MS A1 median short and straight; MS B1 sub basal long and straight (longer than MS A and shorter than MS C). Lateral sacs short and apically acuminated, with short T3-like microsetae. Stylus slightly thick, apically inflated, dorsally inclined, with apical tiny projections. Dorsal process absent. Promontory convex. -
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<emphasis id="ECCA60A849A99C1C1652ADEA0CC9C7DE" bold="true" pageId="0" pageNumber="309">
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FEMALE:
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<emphasis id="46338C16FA2A898C38E170AE58BFDC7B" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
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</emphasis>
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(
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<emphasis id="341D104A935A4A611486FA069C97071B" italics="true" pageId="0" pageNumber="309">n</emphasis>
|
||
=4) DSW: 3.0-4.4; DSL: 4.5-5.4; CL: 1.9-4.7. FIVL: 3.7-4.5. ChL: 1.7-4.0. Swollen chelicerae in some specimens, like males. Tubercles of femur of pedipalpus slightly smaller. Ocularium tubercles smaller than in male. Granules on femur IV thinner when compared with those of male. Tarsal segmentation: (
|
||
<emphasis id="C5CCBF2466DC2387BFA074B551D31994" italics="true" pageId="0" pageNumber="309">n</emphasis>
|
||
=4) 6-7, 10-11, 7, 7-8.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2BDAF57CCA6ED5EF03B8FB2222D7E21C" pageId="0" pageNumber="309" type="diagnosis">
|
||
<paragraph id="537CAF28387BC612DA6539F5A652E154" pageId="0" pageNumber="309">Diagnosis.</paragraph>
|
||
<paragraph id="EA5AA1A684913AFB13C3C67D8B866455" pageId="0" pageNumber="309">
|
||
Similar to
|
||
<taxonomicName id="CD3728918A5DD22CD614A9CB6E331D7D" authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho glauberrochai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="glauberrochai">
|
||
<emphasis id="383CC8B9817FED72CF67B74F49F4E391" italics="true" pageId="0" pageNumber="309">Ayacucho glauberrochai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C8AD088B6A01DF8982C2E41175DFCE6E" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
||
,
|
||
<taxonomicName id="61F5089E8069CAFF2B2B2C2B56FB550B" lsidName="A. pomacocha" pageId="0" pageNumber="309" rank="species" species="pomacocha">
|
||
<emphasis id="33B1D05A241220F45B42C6B35D8EB960" italics="true" pageId="0" pageNumber="309">A. pomacocha</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1D94144B160B2ACE1F508E5CD455DCB0" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
||
,
|
||
<taxonomicName id="29CB3D77866ED30022347ABC4DDB7DA7" lsidName="A. silvae" pageId="0" pageNumber="309" rank="species" species="silvae">
|
||
<emphasis id="3591402C3C0A3CB6DA7B67F55E90FFB0" italics="true" pageId="0" pageNumber="309">A. silvae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="55C6C1E85A0EFB929D00A8AE44B25268" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
||
and
|
||
<taxonomicName id="D386B4C21CE2F22A15B630DCE6CFC59C" lsidName="A. vargasllosai" pageId="0" pageNumber="309" rank="species" species="vargasllosai">
|
||
<emphasis id="9A9B003EEBC381276B355EFDECB1478D" italics="true" pageId="0" pageNumber="309">A. vargasllosai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0F2934068F7FECBB1252E60644D115B1" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
||
in the following combination of characteristics: dorsal scutum densely granulate; ocularium and areas I-IV of DS unarmed or armed with tiny tubercles, slightly greater than granules; posterior margin of DS and free tergites I-III with median rows of acuminate tubercles (Fig.
|
||
<figureCitation id="606A8B0EB78064CD2DDBFE1CC55E6F11" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3D</figureCitation>
|
||
). It differs from the previously mentioned species in the following combination of characteristics: ocularium with two small acuminate tubercles and densely granulate (Fig.
|
||
<figureCitation id="40D9E001063D1EAB443074EFC4EC3CE2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3D</figureCitation>
|
||
); male femur IV unarmed (Fig.
|
||
<figureCitation id="2B2399E5FA9DBA3E7DB7B4A9D1C87AE4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7U, V</figureCitation>
|
||
; unlike
|
||
<taxonomicName id="2F91210B85E2622641AF5D2B18C36936" lsidName="A. silvae" pageId="0" pageNumber="309" rank="species" species="silvae">
|
||
<emphasis id="4BD962B4A803815E44B7EABEC6E94F1B" italics="true" pageId="0" pageNumber="309">A. silvae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="38410D414ACE4A105BC7CDDAAA69DFE8" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
||
); penis VP rectangular; five-six pairs of subapical MS C; one pair of MS A and B (Fig.
|
||
<figureCitation id="F42B7CBAF3A03395DF2056D3F10FB71B" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D Ayacucho inermis (Roewer, 1957) comb. nov.; E - F Ayacucho pasolinii sp. nov.; G - H Ayacucho spielbergi sp. nov.; I - J Ayacucho titschacki Roewer, 1949; K - L Ayacucho triarmatus nom. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure12" httpUri="https://binary.pensoft.net/fig/729886" pageId="0" pageNumber="309">12I, J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BFEDB3758A6E469BA843185CA2444D26" pageId="0" pageNumber="309" type="remarks">
|
||
<paragraph id="3AD238CE3B5D912C278893E36A9A3890" pageId="0" pageNumber="309">Remarks.</paragraph>
|
||
<paragraph id="5B857A0CF168C4CF851997559715650A" pageId="0" pageNumber="309">The male holotype and paratypes examined by Roewer (SMF RII 8589/20) are actually females.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="075006BF1E1D7047E6137FA62608F3FA" pageId="0" pageNumber="309" type="distribution">
|
||
<paragraph id="E51278EC3D7F74962DA5826DA2749D1E" pageId="0" pageNumber="309">Distribution.</paragraph>
|
||
<paragraph id="8BC274C8AA6D22653DA915AEFACA6C76" pageId="0" pageNumber="309">
|
||
(Fig.
|
||
<figureCitation id="8DAA142BBB492F3DEF6AB8DDDB85610B" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">28</figureCitation>
|
||
) PERU.
|
||
<taxonomicName id="2A9EC1F1A9B27C1E9A9FD6AE14FC9E42" authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Ayacucho</taxonomicName>
|
||
.
|
||
<taxonomicName id="4D6AE273FEE8CE7308515EFEFEC9DE52" authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Ayacucho</taxonomicName>
|
||
, Ocollo and Virgem de Cacharras de Cocha.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9FD0B3EE73D51F8D5C49E5210B840B8F" pageId="0" pageNumber="309" type="materials_examined">
|
||
<paragraph id="B02793ACBD1B25AB87614B01979BC9A7" pageId="0" pageNumber="309">Material examined.</paragraph>
|
||
<paragraph id="6BFDDA5779B143D8C400A952F972C2AE" pageId="0" pageNumber="309">
|
||
<emphasis id="AE5467482AF8A29AF8BED0F70A94BF57" bold="true" pageId="0" pageNumber="309">
|
||
<emphasis id="1559FEAEA242B88082369F11DF4F85B6" italics="true" pageId="0" pageNumber="309">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="B5974A6200D159E32CDBA98C8CBB9D25" ID-GBIF-Occurrence="3889641340" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="PERU" location="PERU" specimenCount="♀" typeStatus="Holotype">
|
||
<typeStatus id="572C560D4DA1B7562B5AE502A558DCB6">Holotype</typeStatus>
|
||
<specimenCount id="10ADABF37DC215F33FC5D19DC55436B4">♀</specimenCount>
|
||
, '
|
||
<collectingCountry id="38256E0473049F6625933D81C6DF86D9" name="Peru">PERU</collectingCountry>
|
||
,
|
||
<taxonomicName id="F3F1288528152241C8C121B9C1C93829" authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Ayacucho</taxonomicName>
|
||
,
|
||
<taxonomicName id="038DF5EEF4AA34A6D5DF5F6D757EFB7F" authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Ayacucho</taxonomicName>
|
||
, without date and leg. (SMF RII 8589/20)
|
||
</materialsCitation>
|
||
</emphasis>
|
||
-
|
||
<emphasis id="B439BB32178CD4FB4E172DCCF8AA25CF" bold="true" pageId="0" pageNumber="309">
|
||
<typeStatus id="897408FDEB4866C40F452C91DDC015BC">Paratype</typeStatus>
|
||
</emphasis>
|
||
<specimenCount id="0FCA4386A304EBE54265A4DF369EA8F5">♀</specimenCount>
|
||
,
|
||
<normalizedToken id="76D390D0166E85C02AEDAC77A2E582D2" originalValue="‘ditto’">'ditto'</normalizedToken>
|
||
(SMF RII 8589/20).
|
||
<emphasis id="D3465F84961FB77B6E4E731569CBA1F2" bold="true" pageId="0" pageNumber="309">
|
||
<emphasis id="0AC712DED83FB0DC6D32619880D70AC5" italics="true" pageId="0" pageNumber="309">Additional material</emphasis>
|
||
.
|
||
</emphasis>
|
||
<materialsCitation id="E965F8F921A5720CB5C33563DBFC28DF" ID-GBIF-Occurrence="3889641355" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="PERU" latitude="-13.029722" location="Virgem de Cacharras de Cocha" longLatPrecision="21" longitude="-73.87056" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
|
||
<specimenCount id="7E4782F5695788D4669C4245DEF85033" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="BDE6854BD0F33537C2027681D1D6DEC7" type="female">2 ♀</specimenCount>
|
||
'
|
||
<collectingCountry id="EB441C20DF9D62349AE67270C636A1CE" name="Peru">PERU</collectingCountry>
|
||
,
|
||
<taxonomicName id="39732478125E82385A15872AE7A959EF" authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Ayacucho</taxonomicName>
|
||
,
|
||
<location id="2E96CA51C1AE6E9164E346767C20A6A2" LSID="urn:lsid:plazi:treatment:A7E8B110FB875A6CBAAC65042B2B7CE5:2E96CA51C1AE6E9164E346767C20A6A2" country="PERU" latitude="-13.029722" longLatPrecision="21" longitude="-73.87056" name="Virgem de Cacharras de Cocha">Virgem de Cacharras de Cocha</location>
|
||
,
|
||
<geoCoordinate id="7BB9435A20048DBE2697267976841062" degrees="13" direction="south" minutes="01" orientation="latitude" precision="15" seconds="47" value="-13.029722">13°01′47″S</geoCoordinate>
|
||
<geoCoordinate id="329A14C1C099DCAF68CEBA44E661D5F7" degrees="73" direction="west" minutes="52" orientation="longitude" precision="15" seconds="14" value="-73.87056">73°52′14″W</geoCoordinate>
|
||
, without date and leg. (MZSP 36972)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="1FC0E40A2A5F6EC3A0BE50E3B1B5E685" ID-GBIF-Occurrence="3889641311" collectingDate="2011-04-29" collectorName="R. Pinto-da-Rocha, A. Benedetti, J. Ochoa, D. Silva" country="PERU" latitude="-13.331388" location="Near Ocollo" longLatPrecision="1" longitude="-74.50489" specimenCount="2" specimenCount-female="2">
|
||
<specimenCount id="2A90FE8678D0473BD544CEE793D95464" type="female">2 ♀</specimenCount>
|
||
'
|
||
<collectingCountry id="8BEBEDAB3D326FED2B2D001DFF40075A" name="Peru">PERU</collectingCountry>
|
||
,
|
||
<taxonomicName id="D3CC4993E69E1B83C4F4C158368C3E0A" authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Ayacucho</taxonomicName>
|
||
,
|
||
<location id="C2719E395FA3B21594C16ABEB1F7B3C0" LSID="urn:lsid:plazi:treatment:A7E8B110FB875A6CBAAC65042B2B7CE5:C2719E395FA3B21594C16ABEB1F7B3C0" country="PERU" latitude="-13.331388" longLatPrecision="1" longitude="-74.50489" name="Near Ocollo">Near Ocollo</location>
|
||
,
|
||
<geoCoordinate id="F4865B7184A54E40903F9B0336B26E60" degrees="13" direction="south" minutes="19" orientation="latitude" precision="1" seconds="53.0" value="-13.331388">13°19′53.0″S</geoCoordinate>
|
||
<geoCoordinate id="1FCBFD44D7F94DBEDDD392D0DB2055E6" degrees="74" direction="west" minutes="30" orientation="longitude" precision="1" seconds="17,6" value="-74.50489">74°30′17,6″ W</geoCoordinate>
|
||
,
|
||
<collectingDate id="A273B268F7823DADB1FD57D884EC3D85" value="2011-04-29">29/IV/2011</collectingDate>
|
||
,
|
||
<collectorName id="991E3F9F7334212BC9809A2CB40D829F">R. Pinto-da-Rocha</collectorName>
|
||
,
|
||
<collectorName id="92792C9DDAC6426DAA3F5647328D4057">A. Benedetti</collectorName>
|
||
,
|
||
<collectorName id="BC2AE800BDA55DC6049408C41314764B">J. Ochoa</collectorName>
|
||
&
|
||
<collectorName id="3ED2E141A3DC8C5141C9DF3D2A99091E">D. Silva</collectorName>
|
||
leg. (MUSM)
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="ACEA8D6027A5F46F0AAF7193C712CB67" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="A8BF898CDCBC0CAF5E766CFCEDAED3D4" type="female">2 ♀</specimenCount>
|
||
,
|
||
<normalizedToken id="E2ABD9D6D755AC76084B32B4FCD6EDF1" originalValue="‘ditto’">'ditto'</normalizedToken>
|
||
(MUBI);
|
||
<specimenCount id="09CE332E6749D2DED5271190EB78FFC9" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="1D06AF98308DB996942B4DD8CCC5B581" type="female">5 ♀</specimenCount>
|
||
<normalizedToken id="48EAF2B510AE5ABC633BA81F9453B083" originalValue="‘ditto’">'ditto'</normalizedToken>
|
||
(MZSP 36973).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |