252 lines
17 KiB
XML
252 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.878.37845" ID-GBIF-Dataset="faa91fff-ad2f-44bc-8309-388d6755f640" ID-PMC="PMC6791898" ID-Pensoft-Pub="1313-2970-878-33" ID-Pensoft-UUID="C0F938DD0EFE5A18BB267539EC83EFDB" ID-PubMed="31632176" ID-ZooBank="BCE3960BE7C6418FB8802978DF9F099E" ModsDocID="1313-2970-878-33" checkinTime="1570497201903" checkinUser="pensoft" docAuthor="Hopkins, Tapani, Roininen, Heikki, Noort, Simon van, Broad, Gavin R., Kaunisto, Kari & Saeaeksjaervi, Ilari E." docDate="2019" docId="5CA38040F4405B6D928FFCD9E282FED5" docLanguage="en" docName="ZooKeys 878: 33-71" docOrigin="ZooKeys 878" docSource="http://dx.doi.org/10.3897/zookeys.878.37845" docTitle="Epirhyssa quagga Hopkins, Roininen, Noort, Broad, Kaunisto & Saeaeksjaervi, 2019, sp. nov." docType="treatment" docUuid="9BABD7C8-7644-45BA-9985-96408F898402" docUuidSource="ZooBank" docVersion="5" id="C0F938DD0EFE5A18BB267539EC83EFDB" lastPageNumber="33" masterDocId="C0F938DD0EFE5A18BB267539EC83EFDB" masterDocTitle="Extensive sampling and thorough taxonomic assessment of Afrotropical Rhyssinae (Hymenoptera, Ichneumonidae) reveals two new species and demonstrates the limitations of previous sampling efforts" masterLastPageNumber="71" masterPageNumber="33" pageNumber="33" updateTime="1668167851423" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Extensive sampling and thorough taxonomic assessment of Afrotropical Rhyssinae (Hymenoptera, Ichneumonidae) reveals two new species and demonstrates the limitations of previous sampling efforts</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hopkins, Tapani</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Roininen, Heikki</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Noort, Simon van</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Broad, Gavin R.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kaunisto, Kari</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Saeaeksjaervi, Ilari E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>878</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>71</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.878.37845</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.878.37845</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-878-33</mods:identifier>
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<mods:identifier type="ZooBank">BCE3960BE7C6418FB8802978DF9F099E</mods:identifier>
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<mods:identifier type="Pensoft-UUID">C0F938DD0EFE5A18BB267539EC83EFDB</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159654858" LSID="urn:lsid:zoobank.org:act:9BABD7C8-7644-45BA-9985-96408F898402" httpUri="http://treatment.plazi.org/id/5CA38040F4405B6D928FFCD9E282FED5" lastPageNumber="33" pageId="0" pageNumber="33">
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<subSubSection pageId="0" pageNumber="33" type="nomenclature">
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<paragraph pageId="0" pageNumber="33">
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<taxonomicName LSID="5CA38040-F440-5B6D-928F-FCD9E282FED5" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epirhyssa quagga" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="quagga">Epirhyssa quagga</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="33">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 39–45" captionStartId="F10" captionText="Figures 39 - 45. Epirhyssa quagga female (holotype http: // mus. utu. fi / ZMUT. 5788), a new species from Uganda. 39 Habitus 40 face and clypeus 41 frons 42 hypostomal flange 43 mesopleuron dorsal margin 44 apical horn of metasoma 45 tergites 1 - 7. Scale bars: 0.5 mm (40 - 45), 1 mm (39)." figureDoi="10.3897/zookeys.878.37845.figures39-45" httpUri="https://binary.pensoft.net/fig/345026" pageId="0" pageNumber="33" tableDoi="10.3897/zookeys.878.37845.figures39-45">Figs 39-45</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="materials_examined">
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<paragraph pageId="0" pageNumber="33">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" pageId="0" pageNumber="33">Type material</emphasis>
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: UGANDA:
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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• 1 ♀, holotype; Kibale National Park, Kanyawara, Site CC, Malaise trap CCT1;
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<geoCoordinate degrees="0.5497" direction="north" orientation="latitude" precision="5" value="0.5497">0.5497N</geoCoordinate>
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,
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<geoCoordinate degrees="30.3673" direction="east" orientation="longitude" precision="5" value="30.3673">30.3673E</geoCoordinate>
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(WGS84); alt. 1450 m (GPS, WGS84); 11 Aug. 2015-25 Aug. 2015; Tapani Hopkins leg.; ZMUThttp://mus.utu.fi/ZMUT.5788
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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• 1 ♀, paratype; same data as preceding; Site R03, Malaise trap R03T2;
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<geoCoordinate degrees="0.5403" direction="north" orientation="latitude" precision="5" value="0.5403">0.5403N</geoCoordinate>
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,
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<geoCoordinate degrees="30.3608" direction="east" orientation="longitude" precision="5" value="30.3608">30.3608E</geoCoordinate>
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(WGS84); alt. 1490 m (GPS, WGS84); 21 May 2015-4 Jun. 2015; ZMUThttp://mus.utu.fi/ZMUT.1500
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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• 1 ♀, paratype; same data as preceding; Site K31, Malaise trap K31T3;
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<geoCoordinate degrees="0.5360" direction="north" orientation="latitude" precision="5" value="0.536">0.5360N</geoCoordinate>
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,
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<geoCoordinate degrees="30.3469" direction="east" orientation="longitude" precision="5" value="30.3469">30.3469E</geoCoordinate>
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(WGS84); alt. 1450 m (GPS, WGS84); 23 May 2015-4 Jun. 2015; ZMUThttp://mus.utu.fi/ZMUT.4238
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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• 1 ♀, paratype; same data as preceding; Site K30S, Malaise trap K30ST4;
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<geoCoordinate degrees="0.5414" direction="north" orientation="latitude" precision="5" value="0.5414">0.5414N</geoCoordinate>
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,
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<geoCoordinate degrees="30.3755" direction="east" orientation="longitude" precision="5" value="30.3755">30.3755E</geoCoordinate>
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(WGS84); alt. 1420 m (GPS, WGS84); 2 Dec. 2014-15 Dec. 2014; ZMUThttp://mus.utu.fi/ZMUT.5743
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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• 1 ♀, paratype; same data as preceding; Site R01, Malaise trap R01T2;
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<geoCoordinate degrees="0.5501" direction="north" orientation="latitude" precision="5" value="0.5501">0.5501N</geoCoordinate>
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,
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<geoCoordinate degrees="30.3561" direction="east" orientation="longitude" precision="5" value="30.3561">30.3561E</geoCoordinate>
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(WGS84); alt. 1600 m (GPS, WGS84); 27 Mar. 2015-10 Apr. 2015; ZMUThttp://mus.utu.fi/ZMUT.5801
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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• 1 ♂, paratype; Kibale National Park, Kanyawara, Site CC, Malaise trap CCT1;
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<geoCoordinate degrees="0.5497" direction="north" orientation="latitude" precision="5" value="0.5497">0.5497N</geoCoordinate>
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,
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<geoCoordinate degrees="30.3673" direction="east" orientation="longitude" precision="5" value="30.3673">30.3673E</geoCoordinate>
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(WGS84); alt. 1450 m (GPS, WGS84); 2 Jun. 2015-17 Jun. 2015; Tapani Hopkins leg.; ZMUThttp://mus.utu.fi/ZMUT.3074
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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• 1 ♂, paratype; same data as preceding; Site K30S, Malaise trap K30ST2;
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<geoCoordinate degrees="0.5392" direction="north" orientation="latitude" precision="5" value="0.5392">0.5392N</geoCoordinate>
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,
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<geoCoordinate degrees="30.3771" direction="east" orientation="longitude" precision="5" value="30.3771">30.3771E</geoCoordinate>
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(WGS84); alt. 1480 m (GPS, WGS84); 5 May 2015-19 May 2015; ZMUThttp://mus.utu.fi/ZMUT.4000
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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• 1 ♂, paratype; same data as preceding; Site K15, Malaise trap K15T4;
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<geoCoordinate degrees="0.5845" direction="north" orientation="latitude" precision="5" value="0.5845">0.5845N</geoCoordinate>
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,
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<geoCoordinate degrees="30.3674" direction="east" orientation="longitude" precision="5" value="30.3674">30.3674E</geoCoordinate>
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(WGS84); alt. 1510 m (GPS, WGS84); 4 May 2015-20 May 2015; ZMUThttp://mus.utu.fi/ZMUT.5836.
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis italics="true" pageId="0" pageNumber="33">Non-type material</emphasis>
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(only diagnostic characters checked): UGANDA
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</paragraph>
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<paragraph pageId="0" pageNumber="33">• 18 ♀; Kibale National Park, Kanyawara; Tapani Hopkins leg.; ZMUThttp://mus.utu.fi/ZMUT.1259, http://mus.utu.fi/ZMUT.2057, http://mus.utu.fi/ZMUT.2203, http://mus.utu.fi/ZMUT.2358, http://mus.utu.fi/ZMUT.2627, http://mus.utu.fi/ZMUT.3199, http://mus.utu.fi/ZMUT.3649, http://mus.utu.fi/ZMUT.4866, http://mus.utu.fi/ZMUT.5596, http://mus.utu.fi/ZMUT.5647, http://mus.utu.fi/ZMUT.5679, http://mus.utu.fi/ZMUT.5712, http://mus.utu.fi/ZMUT.5717, http://mus.utu.fi/ZMUT.5736, http://mus.utu.fi/ZMUT.5803, http://mus.utu.fi/ZMUT.5804, http://mus.utu.fi/ZMUT.5805, http://mus.utu.fi/ZMUT.5835</paragraph>
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<paragraph pageId="0" pageNumber="33">• 20 ♂; Kibale National Park, Kanyawara; Tapani Hopkins leg.; ZMUThttp://mus.utu.fi/ZMUT.1350, http://mus.utu.fi/ZMUT.1355, http://mus.utu.fi/ZMUT.1401, http://mus.utu.fi/ZMUT.1765, http://mus.utu.fi/ZMUT.2645, http://mus.utu.fi/ZMUT.2657, http://mus.utu.fi/ZMUT.2803, http://mus.utu.fi/ZMUT.3076, http://mus.utu.fi/ZMUT.3081, http://mus.utu.fi/ZMUT.3087, http://mus.utu.fi/ZMUT.3097, http://mus.utu.fi/ZMUT.3761, http://mus.utu.fi/ZMUT.3876, http://mus.utu.fi/ZMUT.4990, http://mus.utu.fi/ZMUT.5143, http://mus.utu.fi/ZMUT.5609, http://mus.utu.fi/ZMUT.5768, http://mus.utu.fi/ZMUT.5787, http://mus.utu.fi/ZMUT.5831, http://mus.utu.fi/ZMUT.6033.</paragraph>
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<caption doi="10.3897/zookeys.878.37845.figures39-45" httpUri="https://binary.pensoft.net/fig/345026" pageId="0" pageNumber="33" start="Figures 39–45" startId="F10">
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" pageId="0" pageNumber="33">Figures 39-45.</emphasis>
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<taxonomicName class="Insecta" family="Ichneumonidae" genus="Epirhyssa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epirhyssa quagga" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="quagga">
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<emphasis italics="true" pageId="0" pageNumber="33">Epirhyssa quagga</emphasis>
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</taxonomicName>
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female (holotype http://mus.utu.fi/ZMUT.5788), a new species from Uganda.
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<emphasis bold="true" pageId="0" pageNumber="33">39</emphasis>
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Habitus
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<emphasis bold="true" pageId="0" pageNumber="33">40</emphasis>
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face and clypeus
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<emphasis bold="true" pageId="0" pageNumber="33">41</emphasis>
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frons
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<emphasis bold="true" pageId="0" pageNumber="33">42</emphasis>
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hypostomal flange
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<emphasis bold="true" pageId="0" pageNumber="33">43</emphasis>
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mesopleuron dorsal margin
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<emphasis bold="true" pageId="0" pageNumber="33">44</emphasis>
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apical horn of metasoma
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<emphasis bold="true" pageId="0" pageNumber="33">45</emphasis>
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tergites 1-7. Scale bars: 0.5 mm (
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<emphasis bold="true" pageId="0" pageNumber="33">40-45</emphasis>
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), 1 mm (
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<emphasis bold="true" pageId="0" pageNumber="33">39</emphasis>
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).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" pageId="0" pageNumber="33">Known material</emphasis>
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: 46 specimens (23 ♀, 23 ♂, Ugandan specimens, data above).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="diagnosis">
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<paragraph pageId="0" pageNumber="33">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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This species can be distinguished from other Afrotropical
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<taxonomicName lsidName="" pageId="0" pageNumber="33" rank="subfamily" subfamily="Rhyssinae">Rhyssinae</taxonomicName>
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by the distinctive pattern of striation on the frons and the densely striate tergite 3. No other species has the same colour pattern.
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Head</emphasis>
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: frons without median carinae, without lateral carinae; hypostomal carina raised into a low flange, its height slightly less than or equivalent to the maximum width of the second maxillary palp segment.
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Mesosoma</emphasis>
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: subalar prominence without a lateral flange; mesopleuron without a flange along the dorsal margin; epicnemial carina reaches the approximate height of the mesopleural pit.
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Metasoma</emphasis>
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: tip of apical horn elliptical in posterior view; tergite 3 densely striate.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="description">
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<paragraph pageId="0" pageNumber="33">Description (female).</paragraph>
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<paragraph pageId="0" pageNumber="33">
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Body length 8.5
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<normalizedToken originalValue="mm–">mm-</normalizedToken>
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14.3 mm (holotype 12.3 mm).
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Head</emphasis>
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: Frons without clear median carinae, without lateral carinae, with faint median rugae that fan out towards ocelli. Occipital carina interrupted dorsally. Hypostomal carina raised into a low flange, its height slightly less than or equivalent to maximum width of second maxillary palp segment. Face densely and deeply punctate. Clypeus with little or no punctation, with a median apical tubercle. Antenna with 30-33 flagellar segments (32 in holotype).
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Mesosoma</emphasis>
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: Subalar prominence without a lateral flange. Mesopleuron without a flange along dorsal margin. Epicnemial carina reaches approximate height of mesopleural pit. Fore wing with 2m-cu varying from clearly distal to opposite rs-m.
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Metasoma</emphasis>
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: Tip of apical horn elliptical in posterior view. Tergites 1-5 with dense, light, predominantly longitudinal striation and punctation, 6-7 smoother and more pubescent, tergite 1 1.9-2.4 times as long as apically wide (1.9 in holotype).
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Colour</emphasis>
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: General colour mottled black and white with metasoma orange testaceous from tergite 3 onwards. Hind tibia dark brown. Antennae black. Ovipositor sheaths black to dark testaceous. Wings hyaline.
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Variation</emphasis>
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: Colour of the dark patches of the legs varies from entirely black to testaceous, the black and white metasomal colour extends onto tergite 3 in some individuals.
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" pageId="0" pageNumber="33">Male.</emphasis>
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</paragraph>
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<paragraph pageId="0" pageNumber="33">
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Similar to female. Body length 7.9
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<normalizedToken originalValue="mm–">mm-</normalizedToken>
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10.4 mm. T1 3.2-3.6. Antenna with 29-33 flagellar segments. Males are smaller than females on average.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="etymology">
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<paragraph pageId="0" pageNumber="33">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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Refers to the colour pattern which is reminiscent of the plains zebra, especially its extinct subspecies, the
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<taxonomicName lsidName="quagga" pageId="0" pageNumber="33" rank="species" species="quagga">quagga</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="distribution">
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<paragraph pageId="0" pageNumber="33">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="33">Uganda.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="biology_ecology">
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<paragraph pageId="0" pageNumber="33">Biology.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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In Uganda, this species was most abundantly caught during the dry season (
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<bibRefCitation DOI="https://doi.org/10.1098/rsos.190913" author="Hopkins, T" journalOrPublisher="African Entomology" pageId="0" pageNumber="33" refId="B7" refString="Hopkins, T, Roininen, H, Saeaeksjaervi, IE, 2019b. . https://doi.org/10.1098/rsos.190913" url="https://doi.org/10.1098/rsos.190913" year="2019 b">Hopkins et al. 2019b</bibRefCitation>
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). It has not been caught outside the forest. It appears to be attracted to decaying wood (although the sample size is too small for statistical significance).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |