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<document id="3EFC911CFA4FE85AACF924DDE16D6571" ID-DOI="10.11646/zootaxa.5047.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5540765" ID-ZooBank="7052707F-306C-49F4-8BD1-C5BAD9F9C128" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632985072705" checkinUser="plazi" docAuthor="Hugel, Sylvain, Warren, Ben H. &amp; Desutter-Grandcolas, Laure" docDate="2021" docId="EA4D6374B956FF9712D258953682FE59" docLanguage="en" docName="zootaxa.5047.3.1.pdf" docOrigin="Zootaxa 5047 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phaeogryllus fuscus Bolivar 1912" docType="treatment" docVersion="3" lastPageNumber="225" masterDocId="16741B0CB940FF8F12455A273669FFD0" masterDocTitle="The Phalangopsidae crickets (Orthoptera, Grylloidea) of the Seychelles Archipelago: Taxonomy of an ecological radiation" masterLastPageNumber="246" masterPageNumber="201" pageNumber="223" updateTime="1699416459910" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="BA9C583167151583A66E5D36354C1F7D">The Phalangopsidae crickets (Orthoptera, Grylloidea) of the Seychelles Archipelago: Taxonomy of an ecological radiation</mods:title>
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<mods:namePart id="08C7BE883BE1592AFB4E33941C61E861">Hugel, Sylvain</mods:namePart>
<mods:affiliation id="9486D8A0D25BC4F7766F410BAC468694">INCI, UPR 3212 CNRS, Université de Strasbourg, Strasbourg, France.</mods:affiliation>
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<mods:namePart id="98CFDF289FC01FADEED02228D2231C4D">Warren, Ben H.</mods:namePart>
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<mods:affiliation id="558DCE2667434130F53F2437D50FF375">Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, Paris, France &amp; https: // orcid. org / 0000 - 0002 - 0758 - 7612</mods:affiliation>
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<mods:namePart id="7BF3DCF5D0E7281B60988F293A2F9973">Desutter-Grandcolas, Laure</mods:namePart>
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<mods:affiliation id="C1E7E5BE097BF1A29AE5736CCDA51847">Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, Paris, France &amp; https: // orcid. org / 0000 - 0002 - 7781 - 3451</mods:affiliation>
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<subSubSection id="2AFE81E9B956FF9912D258953451FD1C" box="[151,568,690,717]" pageId="22" pageNumber="223" type="nomenclature">
<paragraph id="625BD262B956FF9912D258953451FD1C" blockId="22.[151,568,690,754]" box="[151,568,690,717]" pageId="22" pageNumber="223">
<heading id="3913650EB956FF9912D258953451FD1C" bold="true" box="[151,568,690,717]" fontSize="11" level="1" pageId="22" pageNumber="223" reason="1">
<taxonomicName id="A5E4A9E1B956FF9912D258953451FD1C" ID-CoL="6V7SV" authority="Bolivar, 1912" authorityName="Bolivar" authorityYear="1912" box="[151,568,690,717]" class="Insecta" family="Phalangopsidae" genus="Phaeogryllus" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="50900E70B956FF9912D258953451FD1C" bold="true" box="[151,568,690,717]" pageId="22" pageNumber="223">
<emphasis id="50900E70B956FF9912D2589537E5FD1C" bold="true" box="[151,396,690,717]" italics="true" pageId="22" pageNumber="223">Phaeogryllus fuscus</emphasis>
<bibRefCitation id="0675AF93B956FF9913D158953451FD1C" author="Bolivar, I." box="[404,568,690,716]" pageId="22" pageNumber="223" pagination="263 - 292" refId="ref24320" refString="Bolivar, I. (1912) Orthoptera Acrydiidae, Phasgonuridae, Gryllidae of the Seychelles. Transactions of the Linnean Society of London, Series 2, Zoologie, 15, 263 - 292. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00101. x" type="journal article" year="1912">
<collectingRegion id="A0201C80B956FF9913D158953799FD1C" box="[404,496,690,716]" country="Colombia" name="Bolivar" pageId="22" pageNumber="223">Bolivar</collectingRegion>
, 1912
</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="2AFE81E9B956FF9912D258F03469FD22" box="[151,512,727,754]" pageId="22" pageNumber="223" type="description">
<paragraph id="625BD262B956FF9912D258F03469FD22" blockId="22.[151,568,690,754]" box="[151,512,727,754]" pageId="22" pageNumber="223">
(
<figureCitation id="FADFCEE7B956FF9912DA58F03772FD21" box="[159,283,727,753]" captionStart="FIGURE 1" captionStartId="6.[151,250,1896,1920]" captionTargetBox="[151,1436,179,1870]" captionTargetId="figure-21@6.[151,1436,179,1870]" captionTargetPageId="6" captionText="FIGURE 1. Phalangopsidae crickets from the Seychelles in the wild. A, Paragryllodes nitidula (Bolivar, 1912) n. comb., female. B, Phaloria (Papuloria) bolivari Hugel &amp; Desutter-Grandcolas, n. sp., male. C, same, female. D, Phaeogryllus fuscus Bolivar, 1912, male. E, same, female. F, Phalangacris ferlegro Hugel &amp; Desutter-Grandcolas, n. sp. male, G, same, female." figureDoi="http://doi.org/10.5281/zenodo.5540769" httpUri="https://zenodo.org/record/5540769/files/figure.png" pageId="22" pageNumber="223">Figs 1DE</figureCitation>
,
<figureCitation id="FADFCEE7B956FF99136258F0372CFD21" box="[295,325,727,753]" captionStart="FIGURE 10" captionStartId="23.[151,250,1945,1969]" captionTargetBox="[178,1410,241,1921]" captionTargetId="figure-53@23.[164,1423,237,1922]" captionTargetPageId="23" captionText="FIGURE 10. Phaeogryllus fuscus Bolivar, 1912, male specimen MNHN-EO-ENSIF12265:A, habitus, dorsal view; B, face; C, D, head, pronotum and forewings, dorsal (C) and side (D) views; E, hindfemur; F, terminalia, dorsal view. Abbreviations: epc., epiproct; m.p., maxillary palpus; pc, paraproct; sgen., subgenital plate. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540795" httpUri="https://zenodo.org/record/5540795/files/figure.png" pageId="22" pageNumber="223">10</figureCitation>
,
<figureCitation id="FADFCEE7B956FF99131458F03707FD21" box="[337,366,727,753]" captionStart="FIGURE 11" captionStartId="24.[151,250,1890,1914]" captionTargetBox="[170,1406,703,1856]" captionTargetId="figure-181@24.[167,1420,687,1866]" captionTargetPageId="24" captionText="FIGURE 11. Phaeogryllus fuscus Bolivar, 1912, female specimen MNHN-EO-ENSIF12266: A, habitus in dorsal view; B, ovipositor, side view; CE, copulatory papilla in dorsal (C), ventral (D) and side (E) views. Scale bars 2 mm (A, B), 0.1 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.5540799" httpUri="https://zenodo.org/record/5540799/files/figure.png" pageId="22" pageNumber="223">11</figureCitation>
,
<figureCitation id="FADFCEE7B956FF99133F58F037F1FD21" box="[378,408,727,753]" captionStart="FIGURE 12" captionStartId="25.[151,250,1365,1389]" captionTargetBox="[159,1421,196,1339]" captionTargetId="figure-20@25.[151,1436,181,1341]" captionTargetPageId="25" captionText="FIGURE 12. Phaeogryllus fuscus Bolivar, 1912. Male genitalia of specimen MNHN-EO-ENSIF12265 in dorsal (A), ventral (B) and side (C) views. Abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5540801" httpUri="https://zenodo.org/record/5540801/files/figure.png" pageId="22" pageNumber="223">12</figureCitation>
,
<tableCitation id="2F66E7D9B956FF9913E158F0379EFD22" box="[420,503,727,754]" captionStart="TABLE 6" captionStartId="24.[151,239,440,464]" captionText="TABLE 6. Measurements of Phaeogryllus fuscus Bolivar, 1912 (in mm). Bolivar (1912) did not precise if he measured only one female of P. fuscus or both; these measurements thus remain ambiguous." pageId="22" pageNumber="223">Table 6</tableCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="625BD262B956FF9912D2593C3446FCE3" blockId="22.[151,559,795,819]" box="[151,559,795,819]" pageId="22" pageNumber="223">
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<taxonomicName id="A5E4A9E1B956FF9912D2593C3442FCE3" ID-CoL="6V7SV" authority="Bolivar, 1912: 286" authorityName="Bolivar" authorityPageNumber="286" authorityYear="1912" box="[151,555,795,819]" class="Insecta" family="Phalangopsidae" genus="Phaeogryllus" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="50900E70B956FF9912D2593C370DFCE3" box="[151,356,795,819]" italics="true" pageId="22" pageNumber="223">Phaeogryllus fuscus</emphasis>
<treatmentCitation id="E345F473B956FF99132F593C3442FCE3" author="Bolivar, I." box="[362,555,795,819]" page="286" pageId="22" pageNumber="223" year="1912">
<bibRefCitation id="0675AF93B956FF99132F593C3442FCE3" author="Bolivar, I." box="[362,555,795,819]" pageId="22" pageNumber="223" pagination="263 - 292" refId="ref24320" refString="Bolivar, I. (1912) Orthoptera Acrydiidae, Phasgonuridae, Gryllidae of the Seychelles. Transactions of the Linnean Society of London, Series 2, Zoologie, 15, 263 - 292. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00101. x" type="journal article" year="1912">
<collectingRegion id="A0201C80B956FF99132F593C37D1FCE3" box="[362,440,795,819]" country="Colombia" name="Bolivar" pageId="22" pageNumber="223">Bolivar</collectingRegion>
, 1912: 286
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</taxonomicName>
.
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</paragraph>
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<subSubSection id="2AFE81E9B956FF9912D2594637A7FA47" pageId="22" pageNumber="223" type="materials_examined">
<paragraph id="625BD262B956FF9912D2594637A5FC70" blockId="22.[151,1437,865,2008]" pageId="22" pageNumber="223">
<emphasis id="50900E70B956FF9912D25946375EFCAB" bold="true" box="[151,311,865,891]" pageId="22" pageNumber="223">Type locality.</emphasis>
<materialsCitation id="D28CD83FB956FF99137A594637A1FC70" collectingDate="1908-08" collectingDateMax="1909-02" collectingDateMin="1908-08" country="Seychelles" location="Silhouette" pageId="22" pageNumber="223" specimenCount="2" stateProvince="Cascade" typeStatus="lectotype">
<collectingCountry id="1AF392F2B956FF99137A594637D3FCAB" box="[319,442,865,891]" name="Seychelles" pageId="22" pageNumber="223">Seychelles</collectingCountry>
,
<location id="673B84B9B956FF99138359463453FCAB" LSID="urn:lsid:plazi:treatment:EA4D6374B956FF9712D258953682FE59:673B84B9B956FF99138359463453FCAB" box="[454,570,865,891]" country="Seychelles" name="Silhouette" pageId="22" pageNumber="223" stateProvince="Cascade">Silhouette</location>
,
<date id="165AF4A2B956FF991002594634C2FCAB" box="[583,683,865,891]" pageId="22" pageNumber="223" value="1908-08">
<collectingDate id="061E0D4AB956FF991002594634C2FCAB" box="[583,683,865,891]" pageId="22" pageNumber="223" value="1908-08">viii.1908</collectingDate>
</date>
(
<typeStatus id="BD5F6CC0B956FF9910FE59463559FCAB" box="[699,816,865,891]" pageId="22" pageNumber="223" type="lectotype">Lectotype</typeStatus>
, here designated); Mah,
<collectingRegion id="A0201C80B956FF99160C594632CFFCAB" box="[1097,1190,865,891]" country="Seychelles" name="Cascade" pageId="22" pageNumber="223">Cascade</collectingRegion>
Estate,
<date id="165AF4A2B956FF99174759463338FCAB" box="[1282,1361,865,891]" pageId="22" pageNumber="223" value="1909-02">
<collectingDate id="061E0D4AB956FF99174759463338FCAB" box="[1282,1361,865,891]" pageId="22" pageNumber="223" value="1909-02">ii.1909</collectingDate>
</date>
(
<typeStatus id="BD5F6CC0B956FF99172459463768FC4F" pageId="22" pageNumber="223" type="paralectotype">paralectotype</typeStatus>
, here designated)
</materialsCitation>
.
</paragraph>
<paragraph id="625BD262B956FF991282598E3349FB4B" blockId="22.[151,1437,865,2008]" pageId="22" pageNumber="223">
<emphasis id="50900E70B956FF991282598E371CFC13" bold="true" box="[199,373,937,963]" pageId="22" pageNumber="223">Type material.</emphasis>
Two female
<typeStatus id="BD5F6CC0B956FF991043598D3401FC14" box="[518,616,938,964]" pageId="22" pageNumber="223" type="syntype">syntypes</typeStatus>
exist for this species, one from
<collectingCountry id="1AF392F2B956FF991189598E3265FC13" box="[972,1036,937,963]" name="India" pageId="22" pageNumber="223">Mahé</collectingCountry>
(
<collectionCode id="04F54AA7B956FF99165F598E3208FC13" box="[1050,1121,937,963]" pageId="22" pageNumber="223">NHM</collectionCode>
), and the other from Silhouette (
<collectionCode id="04F54AA7B956FF99128A59EA374AFC37" box="[207,291,973,999]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="22" pageNumber="223">CUMZ</collectionCode>
) (see Paris (1991)). Both are identified by Bolivar (manuscript labels), but additional labels not from Bolivar indicate type for the female from Mah, and cotype for the female from Silhouette; Huxley also identified the female from
<collectingCountry id="1AF392F2B956FF9913175E3237FBFBFF" box="[338,402,1045,1071]" name="India" pageId="22" pageNumber="223">Mahé</collectingCountry>
as a “
<typeStatus id="BD5F6CC0B956FF9913905E323428FBFF" box="[469,577,1045,1071]" pageId="22" pageNumber="223" type="lectotype">lectotype</typeStatus>
” but never published it. As no specimen from
<taxonomicName id="A5E4A9E1B956FF9916085E3132C7FBFF" authorityName="I.Bolivar" authorityYear="1912" box="[1101,1198,1045,1071]" class="Insecta" family="Phalangopsidae" genus="Phaeogryllus" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="50900E70B956FF9916085E3132C7FBFF" box="[1101,1198,1045,1071]" italics="true" pageId="22" pageNumber="223">P. fuscus</emphasis>
</taxonomicName>
could be found from Mah, despite intensive search, we designated here the female from Silhouette as the
<typeStatus id="BD5F6CC0B956FF99160C5E1E32D9FB83" box="[1097,1200,1081,1107]" pageId="22" pageNumber="223" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="A5E4A9E1B956FF9916905E1D335FFB83" authorityName="I.Bolivar" authorityYear="1912" box="[1237,1334,1081,1107]" class="Insecta" family="Phalangopsidae" genus="Phaeogryllus" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="50900E70B956FF9916905E1D335FFB83" box="[1237,1334,1081,1107]" italics="true" pageId="22" pageNumber="223">P. fuscus</emphasis>
</taxonomicName>
, in order to stabilize the taxonomy of this species. The female from
<collectingCountry id="1AF392F2B956FF9911755E7A3519FBA7" box="[816,880,1117,1143]" name="India" pageId="22" pageNumber="223">Mahé</collectingCountry>
is here designated as
<typeStatus id="BD5F6CC0B956FF99162C5E7A336AFBA7" box="[1129,1283,1117,1143]" pageId="22" pageNumber="223" type="paralectotype">paralectotype</typeStatus>
, but the presence of
<taxonomicName id="A5E4A9E1B956FF9912B55EA53738FB4B" authorityName="I.Bolivar" authorityYear="1912" box="[240,337,1153,1179]" class="Insecta" family="Phalangopsidae" genus="Phaeogryllus" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="50900E70B956FF9912B55EA53738FB4B" box="[240,337,1153,1179]" italics="true" pageId="22" pageNumber="223">P. fuscus</emphasis>
</taxonomicName>
, or a close species, on
<collectingCountry id="1AF392F2B956FF9910165EA634FAFB4B" box="[595,659,1153,1179]" name="India" pageId="22" pageNumber="223">Mahé</collectingCountry>
island will have to be confirmed by additional field work.
</paragraph>
<paragraph id="625BD262B956FF9912825E823764FA9F" blockId="22.[151,1437,865,2008]" pageId="22" pageNumber="223">
<emphasis id="50900E70B956FF9912825E8237DFFB6F" bold="true" box="[199,438,1189,1215]" pageId="22" pageNumber="223">Examined material.</emphasis>
<materialsCitation id="D28CD83FB956FF9913FA5E823760FA9F" accessionNumber="SIF12268" collectingDate="2010-10-05" collectorName="S. Hugel" country="Seychelles" county="Mt Pot" elevation="400" location="Silhouette" municipality="Jardin Marron" pageId="22" pageNumber="223" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<collectingCountry id="1AF392F2B956FF9913FA5E823453FB6F" box="[447,570,1189,1215]" name="Seychelles" pageId="22" pageNumber="223">Seychelles</collectingCountry>
,
<location id="673B84B9B956FF99100D5E8234D5FB6F" LSID="urn:lsid:plazi:treatment:EA4D6374B956FF9712D258953682FE59:673B84B9B956FF99100D5E8234D5FB6F" box="[584,700,1189,1215]" country="Seychelles" county="Mt Pot" municipality="Jardin Marron" name="Silhouette" pageId="22" pageNumber="223">Silhouette</location>
,
<collectingMunicipality id="823F4818B956FF99108F5E823505FB10" box="[714,876,1189,1216]" pageId="22" pageNumber="223">Jardin Marron</collectingMunicipality>
vers
<collectingCounty id="8B3AAAEEB956FF9911E85E813595FB10" box="[941,1020,1190,1216]" pageId="22" pageNumber="223">Mt Pot</collectingCounty>
à eau,&gt;
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<elevation id="E9C93551B956FF9916295E8232C6FB6F" box="[1132,1199,1189,1215]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="223" unit="m" value="400.0">400m</elevation>
</quantity>
, vue de nuit,
<specimenCount id="74E219EBB956FF99170B5E8233F5FB6F" box="[1358,1436,1189,1215]" pageId="22" pageNumber="223" type="male">1 male</specimenCount>
(2010SEYSH185), MNHN-EO-ENSIF12265; same,
<specimenCount id="74E219EBB956FF9910AC5EEE3523FB33" box="[745,842,1225,1251]" pageId="22" pageNumber="223" type="female">1 female</specimenCount>
(2010SEYSH186), MNHN-EO-ENSIF12266; same,
<specimenCount id="74E219EBB956FF9912D25ECA368EFAD7" box="[151,231,1261,1287]" pageId="22" pageNumber="223" type="male">1 male</specimenCount>
, in alcohol (2010SEYSH192),
<date id="165AF4A2B956FF99100F5ECA34C7FAD8" box="[586,686,1261,1288]" pageId="22" pageNumber="223" value="2010-10-05">
<collectingDate id="061E0D4AB956FF99100F5ECA34C7FAD8" box="[586,686,1261,1288]" pageId="22" pageNumber="223" value="2010-10-05">5.x.2010</collectingDate>
</date>
,
<collectorName id="CF11B7B4B956FF9910FE5ECA3548FAD7" box="[699,801,1261,1287]" pageId="22" pageNumber="223">S. Hugel</collectorName>
, alcohol collection tube 684; MNHN-EO-ENSIF12267; same,
<specimenCount id="74E219EBB956FF9912A75F36372FFAFB" box="[226,326,1297,1323]" pageId="22" pageNumber="223" type="female">1 female</specimenCount>
in alcohol (2010SEYSH193),
<date id="165AF4A2B956FF9910EE5F363566FAFC" box="[683,783,1297,1324]" pageId="22" pageNumber="223" value="2010-10-05">
<collectingDate id="061E0D4AB956FF9910EE5F363566FAFC" box="[683,783,1297,1324]" pageId="22" pageNumber="223" value="2010-10-05">5.x.2010</collectingDate>
</date>
,
<collectorName id="CF11B7B4B956FF99115B5F3635EFFAFB" box="[798,902,1297,1323]" pageId="22" pageNumber="223">S. Hugel</collectorName>
, alcohol collection tube 685; MNHN-EO-EN- SIF12268
</materialsCitation>
.
</paragraph>
<paragraph id="625BD262B956FF9912825F7E37A7FA47" blockId="22.[151,1437,865,2008]" pageId="22" pageNumber="223">
<emphasis id="50900E70B956FF9912825F7E37D4FAA3" bold="true" box="[199,445,1369,1395]" pageId="22" pageNumber="223">Molecular sampling.</emphasis>
LDG
<quantity id="A51C7F87B956FF9910415F7E3422FAA3" box="[516,587,1369,1396]" metricMagnitude="0" metricUnit="m" metricValue="3.429" pageId="22" pageNumber="223" unit="in" value="135.0">135 in</quantity>
<bibRefCitation id="0675AF93B956FF9910175F7E35A9FAA3" author="Chintauan-Marquier, I. C. &amp; Legendre, F. &amp; Hugel, S. &amp; Robillard, T. &amp; Grandcolas, P. &amp; Nel, A. &amp; Zuccon, D. &amp; Desutter-Grandcolas, L." box="[594,960,1369,1395]" pageId="22" pageNumber="223" refId="ref24491" refString="Chintauan-Marquier, I. C., Legendre, F., Hugel, S., Robillard, T., Grandcolas, P., Nel, A., Zuccon, D. &amp; Desutter-Grandcolas, L. (2013) Laying the foundations of evolutionary and systematic studies in crickets (Insecta, Orthoptera): a multilocus phylogenetic analysis. 11 th International Congres of Orthopterology, Kunming, China, 2013, oral communication." type="book" year="2013">
Chintauan-Marquier
<emphasis id="50900E70B956FF9911785F7D351FFAA3" box="[829,886,1369,1395]" italics="true" pageId="22" pageNumber="223">et al.</emphasis>
(2013
</bibRefCitation>
,
<bibRefCitation id="0675AF93B956FF9911895F7E326FFAA4" author="Chintauan-Marquier, I. C. &amp; Legendre, F. &amp; Hugel, S. &amp; Robillard, T. &amp; Grandcolas, P. &amp; Nel, A. &amp; Zuccon, D. &amp; Desutter-Grandcolas, L." box="[972,1030,1369,1396]" pageId="22" pageNumber="223" pagination="54 - 81" refId="ref24572" refString="Chintauan-Marquier, I. C., Legendre, F., Hugel, S., Robillard, T., Grandcolas, P., Nel, A., Zuccon, D. &amp; Desutter-Grandcolas, L. (2016) Laying the foundations of evolutionary and systematic studies in crickets (Insecta, Orthoptera): a multilocus phylogenetic analysis. Cladistics, 32, 54 - 81. https: // doi. org / 10.1111 / cla. 12114" type="journal article" year="2016">2016</bibRefCitation>
) and
<bibRefCitation id="0675AF93B956FF9916065F7D3341FAA3" author="Warren, B. &amp; Hugel, S. &amp; Desutter-Grandcolas, L." box="[1091,1320,1369,1396]" pageId="22" pageNumber="223" refId="ref27041" refString="Warren, B., Hugel, S. &amp; Desutter-Grandcolas, L. (2019) Long-legged cricket phylogeny (Orthoptera, Grylloidea, Phalangopsidae): delimiting new model groups for evolutionary studies. 13 th International Congres of Orthopterology, Agadir (Maroc), 2019, Oral communication." type="book" year="2019">
Warren
<emphasis id="50900E70B956FF9916D95F7D32BDFAA3" box="[1180,1236,1369,1395]" italics="true" pageId="22" pageNumber="223">et al.</emphasis>
(2019)
</bibRefCitation>
, and LDG
<quantity id="A51C7F87B956FF9912D25F5A36B7FA47" box="[151,222,1405,1431]" metricMagnitude="0" metricUnit="m" metricValue="9.4742" pageId="22" pageNumber="223" unit="in" value="373.0">373 in</quantity>
<bibRefCitation id="0675AF93B956FF9912A15F5937A3FA47" author="Warren, B. &amp; Hugel, S. &amp; Desutter-Grandcolas, L." box="[228,458,1405,1432]" pageId="22" pageNumber="223" refId="ref27041" refString="Warren, B., Hugel, S. &amp; Desutter-Grandcolas, L. (2019) Long-legged cricket phylogeny (Orthoptera, Grylloidea, Phalangopsidae): delimiting new model groups for evolutionary studies. 13 th International Congres of Orthopterology, Agadir (Maroc), 2019, Oral communication." type="book" year="2019">
Warren
<emphasis id="50900E70B956FF9913785F59371CFA47" box="[317,373,1405,1431]" italics="true" pageId="22" pageNumber="223">et al.</emphasis>
(2019)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2AFE81E9B956FF9912825F863461FA6C" box="[199,520,1441,1468]" pageId="22" pageNumber="223" type="diagnosis">
<paragraph id="625BD262B956FF9912825F863461FA6C" blockId="22.[151,1437,865,2008]" box="[199,520,1441,1468]" pageId="22" pageNumber="223">
<emphasis id="50900E70B956FF9912825F86372BFA6B" bold="true" box="[199,322,1441,1467]" pageId="22" pageNumber="223">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="2AFE81E9B956FF9712825FE23682FE59" lastPageId="24" lastPageNumber="225" pageId="22" pageNumber="223" type="description">
<paragraph id="625BD262B956FF9912825FE23525FA30" blockId="22.[151,1437,865,2008]" box="[199,844,1477,1504]" pageId="22" pageNumber="223">
<emphasis id="50900E70B956FF9912825FE2371CFA0F" bold="true" box="[199,373,1477,1503]" pageId="22" pageNumber="223">Redescription.</emphasis>
In addition to the characters of the genus:
</paragraph>
<paragraph id="625BD262B956FF9912825FCE3247F92F" blockId="22.[151,1437,865,2008]" pageId="22" pageNumber="223">
<emphasis id="50900E70B956FF9912825FCE37DFF9D3" box="[199,438,1513,1539]" italics="true" pageId="22" pageNumber="223">General morphology.</emphasis>
TIII inner serrulation: no spine in male, zero to one spine in female before isa1; one to two spines in male, three spines in female between isa1 and isa2; three spines in males, four spines in female between isa2 and isa3; 45 spines in male, 57 spines in female between isa3 and isa4; 12 spines in male, 1214 spines in female above isa4. TIII outer serrulation: no spine in male, zero to one spine in female before osa1; three to four spines in male, four to five spines in female between osa1 and osa2; five spines in male, four to six spines in female between osa2 and osa3; seven to eight spines in male, eight to nine spines in female between osa3 and osa4; 1011 spines in male, 1014 spines in female above osa4. Basitarsomeres III with one in male, none in female, inner dorsal spine; four in male, five in female outer dorsal spines, in addition to apical spines.
</paragraph>
<paragraph id="625BD262B956FF9812825D2E32FCFF05" blockId="22.[151,1437,865,2008]" lastBlockId="23.[151,1436,151,213]" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223">
<emphasis id="50900E70B956FF9912825D2E3723F8F3" box="[199,330,1801,1827]" italics="true" pageId="22" pageNumber="223">Coloration.</emphasis>
Face brown (
<figureCitation id="FADFCEE7B956FF9913A75D2E3421F8F3" box="[482,584,1801,1827]" captionStart="FIGURE 10" captionStartId="23.[151,250,1945,1969]" captionTargetBox="[178,1410,241,1921]" captionTargetId="figure-53@23.[164,1423,237,1922]" captionTargetPageId="23" captionText="FIGURE 10. Phaeogryllus fuscus Bolivar, 1912, male specimen MNHN-EO-ENSIF12265:A, habitus, dorsal view; B, face; C, D, head, pronotum and forewings, dorsal (C) and side (D) views; E, hindfemur; F, terminalia, dorsal view. Abbreviations: epc., epiproct; m.p., maxillary palpus; pc, paraproct; sgen., subgenital plate. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540795" httpUri="https://zenodo.org/record/5540795/files/figure.png" pageId="22" pageNumber="223">Fig. 10B</figureCitation>
); a wide yellow line from lower angle of each eye to posterior angle of mandible; a pair of short yellow lines between antennal pits and epistemal suture, widened below and along the eye. Ocelli yellow. Preclypeus with a short, transverse yellow line on each side. Labrum whitish more or less marked with brown. Dorsally, inner margin of eyes marked with yellow, this line prolonged as a thick band toward occiput. Fastigium black with a pair of wide and short yellow lines mediodorsally, not reaching the median ocellus, and separated by a thin black brown line bearing strong dark brown setae. Maxillary palpi brown, their dorsal margin yellow. Femora dark brown (
<figureCitation id="FADFCEE7B957FF9813A55AB0342FFF61" box="[480,582,151,177]" captionStart="FIGURE 10" captionStartId="23.[151,250,1945,1969]" captionTargetBox="[178,1410,241,1921]" captionTargetId="figure-53@23.[164,1423,237,1922]" captionTargetPageId="23" captionText="FIGURE 10. Phaeogryllus fuscus Bolivar, 1912, male specimen MNHN-EO-ENSIF12265:A, habitus, dorsal view; B, face; C, D, head, pronotum and forewings, dorsal (C) and side (D) views; E, hindfemur; F, terminalia, dorsal view. Abbreviations: epc., epiproct; m.p., maxillary palpus; pc, paraproct; sgen., subgenital plate. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540795" httpUri="https://zenodo.org/record/5540795/files/figure.png" pageId="23" pageNumber="224">Fig. 10E</figureCitation>
); tibiae lighter but not annulate; apex of subapical TIII spurs yellow; apical spurs more yellowish, at least in males. Tergites dark brown with brown to yellowish setae.
</paragraph>
<caption id="369B82EAB957FF9812D25DBE327EF829" ID-DOI="http://doi.org/10.5281/zenodo.5540795" ID-Zenodo-Dep="5540795" httpUri="https://zenodo.org/record/5540795/files/figure.png" pageId="23" pageNumber="224" startId="23.[151,250,1945,1969]" targetBox="[178,1410,241,1921]" targetPageId="23">
<paragraph id="625BD262B957FF9812D25DBE327EF829" blockId="23.[151,1436,1945,2042]" pageId="23" pageNumber="224">
<emphasis id="50900E70B957FF9812D25DBE3776F861" bold="true" box="[151,287,1945,1969]" pageId="23" pageNumber="224">FIGURE 10.</emphasis>
<taxonomicName id="A5E4A9E1B957FF9813605DBD34EAF862" authority="Bolivar, 1912" authorityName="Bolivar" authorityYear="1912" box="[293,643,1946,1970]" class="Insecta" family="Phalangopsidae" genus="Phaeogryllus" kingdom="Animalia" order="Orthoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="50900E70B957FF9813605DBD3798F862" box="[293,497,1946,1970]" italics="true" pageId="23" pageNumber="224">Phaeogryllus fuscus</emphasis>
<bibRefCitation id="0675AF93B957FF9813B25DBD34EAF862" author="Bolivar, I." box="[503,643,1946,1970]" pageId="23" pageNumber="224" pagination="263 - 292" refId="ref24320" refString="Bolivar, I. (1912) Orthoptera Acrydiidae, Phasgonuridae, Gryllidae of the Seychelles. Transactions of the Linnean Society of London, Series 2, Zoologie, 15, 263 - 292. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00101. x" type="journal article" year="1912">Bolivar, 1912</bibRefCitation>
</taxonomicName>
, male specimen MNHN-EO-ENSIF12265: A, habitus, dorsal view; B, face; C, D, head, pronotum and forewings, dorsal (C) and side (D) views; E, hindfemur; F, terminalia, dorsal view. Abbreviations: epc., epiproct; m.p., maxillary palpus; pc, paraproct; sgen., subgenital plate. Scale bars 1 mm.
</paragraph>
</caption>
<paragraph id="625BD262B958FF9712825AB0370AFF29" blockId="24.[151,1436,151,393]" pageId="24" pageNumber="225">
<emphasis id="50900E70B958FF9712825AB03761FF61" box="[199,264,151,177]" italics="true" pageId="24" pageNumber="225">Male.</emphasis>
FWs brown, lighter than the body; venation yellowish brown (
<figureCitation id="FADFCEE7B958FF9711875AB03240FF61" box="[962,1065,151,177]" captionStart="FIGURE 10" captionStartId="23.[151,250,1945,1969]" captionTargetBox="[178,1410,241,1921]" captionTargetId="figure-53@23.[164,1423,237,1922]" captionTargetPageId="23" captionText="FIGURE 10. Phaeogryllus fuscus Bolivar, 1912, male specimen MNHN-EO-ENSIF12265:A, habitus, dorsal view; B, face; C, D, head, pronotum and forewings, dorsal (C) and side (D) views; E, hindfemur; F, terminalia, dorsal view. Abbreviations: epc., epiproct; m.p., maxillary palpus; pc, paraproct; sgen., subgenital plate. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540795" httpUri="https://zenodo.org/record/5540795/files/figure.png" pageId="24" pageNumber="225">Fig. 10C</figureCitation>
). FW lateral field with two longitudinal veins (
<figureCitation id="FADFCEE7B958FF9713705A9C37F7FF05" box="[309,414,187,213]" captionStart="FIGURE 10" captionStartId="23.[151,250,1945,1969]" captionTargetBox="[178,1410,241,1921]" captionTargetId="figure-53@23.[164,1423,237,1922]" captionTargetPageId="23" captionText="FIGURE 10. Phaeogryllus fuscus Bolivar, 1912, male specimen MNHN-EO-ENSIF12265:A, habitus, dorsal view; B, face; C, D, head, pronotum and forewings, dorsal (C) and side (D) views; E, hindfemur; F, terminalia, dorsal view. Abbreviations: epc., epiproct; m.p., maxillary palpus; pc, paraproct; sgen., subgenital plate. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540795" httpUri="https://zenodo.org/record/5540795/files/figure.png" pageId="24" pageNumber="225">Fig. 10D</figureCitation>
); dorsal field with six longitudinal veins (
<figureCitation id="FADFCEE7B958FF97112B5A9C35BDFF05" box="[878,980,187,213]" captionStart="FIGURE 10" captionStartId="23.[151,250,1945,1969]" captionTargetBox="[178,1410,241,1921]" captionTargetId="figure-53@23.[164,1423,237,1922]" captionTargetPageId="23" captionText="FIGURE 10. Phaeogryllus fuscus Bolivar, 1912, male specimen MNHN-EO-ENSIF12265:A, habitus, dorsal view; B, face; C, D, head, pronotum and forewings, dorsal (C) and side (D) views; E, hindfemur; F, terminalia, dorsal view. Abbreviations: epc., epiproct; m.p., maxillary palpus; pc, paraproct; sgen., subgenital plate. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540795" httpUri="https://zenodo.org/record/5540795/files/figure.png" pageId="24" pageNumber="225">Fig. 10C</figureCitation>
). Cerci and base of supra-anal plate dirty yellow (
<figureCitation id="FADFCEE7B958FF9712B65AF8373FFF29" box="[243,342,223,249]" captionStart="FIGURE 10" captionStartId="23.[151,250,1945,1969]" captionTargetBox="[178,1410,241,1921]" captionTargetId="figure-53@23.[164,1423,237,1922]" captionTargetPageId="23" captionText="FIGURE 10. Phaeogryllus fuscus Bolivar, 1912, male specimen MNHN-EO-ENSIF12265:A, habitus, dorsal view; B, face; C, D, head, pronotum and forewings, dorsal (C) and side (D) views; E, hindfemur; F, terminalia, dorsal view. Abbreviations: epc., epiproct; m.p., maxillary palpus; pc, paraproct; sgen., subgenital plate. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540795" httpUri="https://zenodo.org/record/5540795/files/figure.png" pageId="24" pageNumber="225">Fig. 10F</figureCitation>
).
</paragraph>
<paragraph id="625BD262B958FF9712825B24345DFECD" blockId="24.[151,1436,151,393]" box="[199,564,259,285]" pageId="24" pageNumber="225">
<emphasis id="50900E70B958FF9712825B24371AFECD" box="[199,371,259,285]" italics="true" pageId="24" pageNumber="225">Male genitalia.</emphasis>
As on
<figureCitation id="FADFCEE7B958FF9713845B243459FECD" box="[449,560,259,285]" captionStart="FIGURE 12" captionStartId="25.[151,250,1365,1389]" captionTargetBox="[159,1421,196,1339]" captionTargetId="figure-20@25.[151,1436,181,1341]" captionTargetPageId="25" captionText="FIGURE 12. Phaeogryllus fuscus Bolivar, 1912. Male genitalia of specimen MNHN-EO-ENSIF12265 in dorsal (A), ventral (B) and side (C) views. Abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5540801" httpUri="https://zenodo.org/record/5540801/files/figure.png" pageId="24" pageNumber="225">Figure 12</figureCitation>
.
</paragraph>
<paragraph id="625BD262B958FF9712825B003488FE91" blockId="24.[151,1436,151,393]" box="[199,737,295,321]" pageId="24" pageNumber="225">
<emphasis id="50900E70B958FF9712825B00374AFE91" box="[199,291,295,321]" italics="true" pageId="24" pageNumber="225">Female.</emphasis>
Ovipositor shorter than FIII (
<figureCitation id="FADFCEE7B958FF97102A5B0034BDFE91" box="[623,724,295,321]" captionStart="FIGURE 11" captionStartId="24.[151,250,1890,1914]" captionTargetBox="[170,1406,703,1856]" captionTargetId="figure-181@24.[167,1420,687,1866]" captionTargetPageId="24" captionText="FIGURE 11. Phaeogryllus fuscus Bolivar, 1912, female specimen MNHN-EO-ENSIF12266: A, habitus in dorsal view; B, ovipositor, side view; CE, copulatory papilla in dorsal (C), ventral (D) and side (E) views. Scale bars 2 mm (A, B), 0.1 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.5540799" httpUri="https://zenodo.org/record/5540799/files/figure.png" pageId="24" pageNumber="225">Fig. 11B</figureCitation>
).
</paragraph>
<paragraph id="625BD262B958FF9712825B6C3682FE59" blockId="24.[151,1436,151,393]" pageId="24" pageNumber="225">
<emphasis id="50900E70B958FF9712825B6C37F9FEB5" box="[199,400,331,357]" italics="true" pageId="24" pageNumber="225">Female genitalia.</emphasis>
Copulatory papilla membranous dorsally and with a distinct ventral process, as on
<figureCitation id="FADFCEE7B958FF9717165B6C368EFE59" captionStart="FIGURE 11" captionStartId="24.[151,250,1890,1914]" captionTargetBox="[170,1406,703,1856]" captionTargetId="figure-181@24.[167,1420,687,1866]" captionTargetPageId="24" captionText="FIGURE 11. Phaeogryllus fuscus Bolivar, 1912, female specimen MNHN-EO-ENSIF12266: A, habitus in dorsal view; B, ovipositor, side view; CE, copulatory papilla in dorsal (C), ventral (D) and side (E) views. Scale bars 2 mm (A, B), 0.1 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.5540799" httpUri="https://zenodo.org/record/5540799/files/figure.png" pageId="24" pageNumber="225">Figure 11CE</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>