306 lines
34 KiB
XML
306 lines
34 KiB
XML
<document id="ECCF5C64E14B87B71AC4EF622E84579E" ID-DOI="10.11646/zootaxa.3709.5.6" ID-GBIF-Dataset="d06ae2dd-7e83-4c8d-aa6d-ee3e849c4c39" ID-ISSN="1175-5326" ID-Zenodo-Dep="216025" ID-ZooBank="F7C29934-8467-4D16-9F44-B4E98F6052FD" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460674669436" checkinUser="plazi" docAuthor="Lee, Jimin, Kim, Dongsung & Chang, Cheon Young" docDate="2013" docId="E64D1645FFDAFFF156B3A6A3FB62842C" docLanguage="en" docName="zt03709p493.pdf" docOrigin="Zootaxa 3709 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tetranchyroderma anisoankyrum Lee, Kim & Chang, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="486" masterDocId="1A746E3DFFDBFFF25624A527FFAE8259" masterDocTitle="Description of three new Tetranchyroderma gastrotrichs (Macrodasyida: Thaumastodermatidae) from South Korea" masterLastPageNumber="493" masterPageNumber="483" pageNumber="484" updateTime="1698349599323" updateUser="plazi">
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<mods:mods id="12F3441DDEE2ED72611C19811EAFF28D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C6925C93CBB0E048A27D7C234FB4B4CF">
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<mods:title id="CD25F37777756E4FDCAA454836AD175D">Description of three new Tetranchyroderma gastrotrichs (Macrodasyida: Thaumastodermatidae) from South Korea</mods:title>
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<mods:name id="9469FA923E699B34A8363A6AC146FA19" type="personal">
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<mods:role id="211906C00B4BE5E3654D8889D61EA886">
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<mods:roleTerm id="F0557239B7505AF8255AD3E33AE085EB">Author</mods:roleTerm>
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<mods:namePart id="C4BBA76C50F9F02F91145EACA0CF8BF0">Lee, Jimin</mods:namePart>
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<mods:name id="AF933FE41C30950095F1E87E35BC3B49" type="personal">
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<mods:roleTerm id="9549F6D3A9BD6952C9972D039E84DF91">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="47C56A3602BC3CB32D61DB55B8BDBA69">Kim, Dongsung</mods:namePart>
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</mods:name>
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<mods:name id="4463B8B1D6E9F9FD32F6990EECBAB041" type="personal">
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<mods:role id="98B47ED4ADE6A03060E13D25D84B7504">
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<mods:roleTerm id="A5A3DF45E981BB4B3AAB456B2CB739C5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="05C34D6D3199279614845D650EF960CE">Chang, Cheon Young</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="36ECA93C392675B65A9FDBD4C945B841">text</mods:typeOfResource>
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<mods:titleInfo id="F8B9FB205D9AF2A606DBB91B21C18B6E">
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<mods:title id="D423E88ECCEFE593DD2DBB566AA8A180">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="EECA8573C55B8CEFA8C622AE39D78667">
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<mods:date id="AD5BBDE14898123BFE9EAEDFB96664B4">2013</mods:date>
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<mods:detail id="690AE0E571971E0E31E3308A0FD7248D" type="volume">
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<mods:number id="8188CB7D9BFA17B5CFB8D89E929831A9">3709</mods:number>
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</mods:detail>
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<mods:detail id="89A571B9D47601390AF54C866690EC62" type="issue">
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<mods:number id="121BC3D4E9BA98540A11570D85F3259D">5</mods:number>
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<mods:start id="16185A08802DE7A9A111774A1B2EA8B6">483</mods:start>
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<mods:end id="1AC64C9E097E65862E87493204746624">493</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="8D0D6D8F944EF1512A19863C51B3E74E">journal article</mods:classification>
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<mods:identifier id="DE804F6329DEA58639BF9033D6F2CED4" type="DOI">10.11646/zootaxa.3709.5.6</mods:identifier>
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<mods:identifier id="00C09068EC8FB50FEAA682A9B96B70C0" type="GBIF-Dataset">d06ae2dd-7e83-4c8d-aa6d-ee3e849c4c39</mods:identifier>
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<mods:identifier id="7796C1AA9A2963B64BDC56554DA4E694" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="CA8FD9021874311CF169B124919D64B6" type="Zenodo-Dep">216025</mods:identifier>
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<mods:identifier id="DADC341966FAC0FB2759872D3189F735" type="ZooBank">F7C29934-8467-4D16-9F44-B4E98F6052FD</mods:identifier>
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</mods:mods>
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<treatment id="E64D1645FFDAFFF156B3A6A3FB62842C" ID-DOI="http://doi.org/10.5281/zenodo.6148068" ID-GBIF-Taxon="119580432" ID-Zenodo-Dep="6148068" LSID="urn:lsid:plazi:treatment:E64D1645FFDAFFF156B3A6A3FB62842C" httpUri="http://treatment.plazi.org/id/E64D1645FFDAFFF156B3A6A3FB62842C" lastPageId="3" lastPageNumber="486" pageId="1" pageNumber="484">
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<subSubSection id="26FEF4D8FFDAFFF356B3A6A3FF4581E6" pageId="1" pageNumber="484" type="nomenclature">
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<paragraph id="6E5BA753FFDAFFF356B3A6A3FD3F81C7" blockId="1.[151,657,900,959]" box="[151,657,900,926]" pageId="1" pageNumber="484">
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<heading id="3513103FFFDAFFF356B3A6A3FD3F81C7" bold="true" box="[151,657,900,926]" fontSize="11" level="1" pageId="1" pageNumber="484" reason="1">
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<emphasis id="5C907B41FFDAFFF356B3A6A3FD3F81C7" bold="true" box="[151,657,900,926]" pageId="1" pageNumber="484">
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<taxonomicName id="A9E4DCD0FFDAFFF356B3A6A3FD8281C7" ID-CoL="7C57G" box="[151,556,900,926]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="1" pageNumber="484" phylum="Gastrotricha" rank="species" species="anisoankyrum" status="sp. nov.">
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<emphasis id="5C907B41FFDAFFF356B3A6A3FD8281C7" bold="true" box="[151,556,900,926]" italics="true" pageId="1" pageNumber="484">Tetranchyroderma anisoankyrum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="47A3C63AFFDAFFF35417A6A3FD3F81C7" box="[563,657,900,926]" pageId="1" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="6E5BA753FFDAFFF356B3A681FF4581E6" blockId="1.[151,657,900,959]" box="[151,235,934,959]" pageId="1" pageNumber="484">
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(
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<figureCitation id="F6DFBBD6FFDAFFF356BBA681FF4E81E6" box="[159,224,934,959]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1986]" captionTargetBox="[214,1354,217,1941]" captionTargetId="figure@2.[198,1370,195,1942]" captionTargetPageId="2" captionText="FIGURE 1. Tetranchyroderma anisoankyrum n. sp. A, habitus, dorsal view; B, habitus, ventral view; C, pentancre. Scale bars = 50 µm (A, B), 10 µm (C)." httpUri="https://zenodo.org/record/216026/files/figure.png" pageId="1" pageNumber="484">Fig. 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="26FEF4D8FFDAFFF356B3A6C9FB368616" pageId="1" pageNumber="484" type="materials_examined">
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<paragraph id="6E5BA753FFDAFFF356B3A6C9FB918673" blockId="1.[151,1436,1006,2003]" pageId="1" pageNumber="484">
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<emphasis id="5C907B41FFDAFFF356B3A6C9FEEC865E" bold="true" box="[151,322,1006,1031]" pageId="1" pageNumber="484">
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<typeStatus id="B15F19F1FFDAFFF356B3A6C9FF7C865E" box="[151,210,1006,1031]" pageId="1" pageNumber="484">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="B15F19F1FFDAFFF35772A6C9FE10865E" box="[342,446,1006,1031]" pageId="1" pageNumber="484" type="holotype">Holotype</typeStatus>
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(NIBRIV0000279610) and 15
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<typeStatus id="B15F19F1FFDAFFF35516A6C8FC33865E" box="[818,925,1007,1031]" pageId="1" pageNumber="484" type="paratype">paratypes</typeStatus>
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(KIOSTG0101–0113, NIBRIV0000279611, NIBRIV0000279612) mounted in glycerin on H-S slides,
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<date id="1A5A8193FFDAFFF35505A134FC178672" box="[801,953,1042,1067]" pageId="1" pageNumber="484" value="2006-06-29">29 June 2006</date>
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,
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<emphasis id="5C907B41FFDAFFF355E0A135FC488672" box="[964,998,1042,1067]" italics="true" pageId="1" pageNumber="484">leg</emphasis>
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. J. Lee.
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</paragraph>
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<paragraph id="6E5BA753FFDAFFF356E3A111FB368616" blockId="1.[151,1436,1006,2003]" box="[199,1176,1078,1103]" pageId="1" pageNumber="484">
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<emphasis id="5C907B41FFDAFFF356E3A111FEF38616" bold="true" box="[199,349,1078,1103]" pageId="1" pageNumber="484">
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<typeStatus id="B15F19F1FFDAFFF356E3A111FEAC8616" box="[199,258,1078,1103]" pageId="1" pageNumber="484">Type</typeStatus>
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locality
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</emphasis>
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. Baegripo Beach, Taean,
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<collectingCountry id="16F3E7C3FFDAFFF3545AA110FD6C8617" box="[638,706,1079,1102]" name="South Korea" pageId="1" pageNumber="484">Korea</collectingCountry>
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(36°48′43″N, 126°09′14″E;
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<quantity id="A91C0AB6FFDAFFF35220A110FBE58617" box="[1028,1099,1079,1103]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="484" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
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deep).
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</paragraph>
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</subSubSection>
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<subSubSection id="26FEF4D8FFDAFFF356E3A17DFEB486E2" pageId="1" pageNumber="484" type="etymology">
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<paragraph id="6E5BA753FFDAFFF356E3A17DFEB486E2" blockId="1.[151,1436,1006,2003]" pageId="1" pageNumber="484">
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<emphasis id="5C907B41FFDAFFF356E3A17DFEE8862A" bold="true" box="[199,326,1114,1139]" pageId="1" pageNumber="484">Etymology</emphasis>
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. The proposed specific name,
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<taxonomicName id="A9E4DCD0FFDAFFF3548FA17DFCE4862A" box="[683,842,1114,1139]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="1" pageNumber="484" phylum="Gastrotricha" rank="species" species="anisoankyrum">
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<emphasis id="5C907B41FFDAFFF3548FA17DFCE4862A" box="[683,842,1114,1139]" italics="true" pageId="1" pageNumber="484">anisoankyrum</emphasis>
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</taxonomicName>
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, is from the Greeks
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<emphasis id="5C907B41FFDAFFF35218A17BFBF6862A" box="[1084,1112,1116,1139]" italics="true" pageId="1" pageNumber="484">an</emphasis>
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, meaning ‘not’,
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<emphasis id="5C907B41FFDAFFF3533EA17BFAE8862A" box="[1306,1350,1116,1139]" italics="true" pageId="1" pageNumber="484">isos</emphasis>
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‘equal, like’, and
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<emphasis id="5C907B41FFDAFFF3572FA159FEF786CE" box="[267,345,1150,1175]" italics="true" pageId="1" pageNumber="484">ankyra</emphasis>
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‘anchor’, which refers to the different cuticular armature of pentancres with central tine longer than others.
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</paragraph>
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</subSubSection>
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<subSubSection id="26FEF4D8FFDAFFF356E3A1E1FB4187EE" pageId="1" pageNumber="484" type="diagnosis">
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<paragraph id="6E5BA753FFDAFFF356E3A1E1FB4187EE" blockId="1.[151,1436,1006,2003]" pageId="1" pageNumber="484">
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<emphasis id="5C907B41FFDAFFF356E3A1E1FEEF8686" bold="true" box="[199,321,1222,1247]" pageId="1" pageNumber="484">Diagnosis.</emphasis>
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A small
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<taxonomicName id="A9E4DCD0FFDAFFF35790A1E1FD2C8686" box="[436,642,1222,1247]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="1" pageNumber="484" phylum="Gastrotricha" rank="genus">
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<emphasis id="5C907B41FFDAFFF35790A1E1FD2C8686" box="[436,642,1222,1247]" italics="true" pageId="1" pageNumber="484">Tetranchyroderma</emphasis>
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</taxonomicName>
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with adult length upto 312 µm; pharynx length to 72 µm; with 12 papillae on anterior border of head; a pair of rod-like cephalic tentacles arising from papillary basis; pestle organ absent; cuticular armature of pentancres with central tine longer than the others; 2 pairs of dorsolateral cirratumtype tubes located just behind PhJIn and at mid-intestinal region, respectively; 13–14 lateral cirratum-type tubes per side distributed along whole body length; adhesive tubes: 4 TbA per side; a pair of TbL in mid-trunk region; 13 TbVL per side in intestinal region; foot-type TbV absent; 4 pairs of TbP, forming trifid pedicles with 2 distal tubes and 1 elongate cirratum-type tube, flanked by 1 medial tube; copulatory organ slender pear-shaped.
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</paragraph>
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</subSubSection>
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<subSubSection id="26FEF4D8FFDAFFF156E3A0E5FB62842C" lastPageId="3" lastPageNumber="486" pageId="1" pageNumber="484" type="description">
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<paragraph id="6E5BA753FFDAFFF356E3A0E5FE8A847A" blockId="1.[151,1436,1006,2003]" pageId="1" pageNumber="484">
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<emphasis id="5C907B41FFDAFFF356E3A0E5FDBB8782" bold="true" box="[199,533,1474,1499]" pageId="1" pageNumber="484">
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Description of the
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<typeStatus id="B15F19F1FFDAFFF35783A0E5FDBE8782" box="[423,528,1474,1499]" pageId="1" pageNumber="484" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Body small, Lt 255 µm; broad head, both sides nearly parallel in trunk region, narrowing posteriorly; widths of head/neck/PhJIn/trunk/caudal base 57/31/33/39/17µm at U09/U33/U43/U67/U95, respectively.
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</paragraph>
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<paragraph id="6E5BA753FFDAFFF356E3A309FDED848E" blockId="1.[151,1436,1006,2003]" pageId="1" pageNumber="484">Head protruding anteriorly with scalloping edge, armed with 12 large papillae on anterior border of oral hood, each bearing 1–2 sensilla at tip, except for two papillae each with short, rod-like cephalic tentacle (7 µm long), at U04. Thirteen short sensory hairs (3–8 µm long) scattered along anterior margin of oral hood, which lacks cuticular armature of pentancres; 22–25 hairs (11–16 µm long) per side aligned dorsolaterally and laterally throughout nearly whole body length (U11–U95).</paragraph>
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<paragraph id="6E5BA753FFDAFFF356E3A3C5FC038546" blockId="1.[151,1436,1006,2003]" pageId="1" pageNumber="484">Epidermal glands, 5–6 per side, with irregular shape and size (4–6 µm in diameter), asymmetrically distributed from anterior pharyngeal region to posterior trunk region at U15–U81.</paragraph>
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<paragraph id="6E5BA753FFDAFFF356E3A20DFC9D858A" blockId="1.[151,1436,1006,2003]" pageId="1" pageNumber="484">
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Cuticular armature with pentancres only (
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<figureCitation id="F6DFBBD6FFDAFFF35489A20DFD5F851A" box="[685,753,1834,1859]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1986]" captionTargetBox="[214,1354,217,1941]" captionTargetId="figure@2.[198,1370,195,1942]" captionTargetPageId="2" captionText="FIGURE 1. Tetranchyroderma anisoankyrum n. sp. A, habitus, dorsal view; B, habitus, ventral view; C, pentancre. Scale bars = 50 µm (A, B), 10 µm (C)." httpUri="https://zenodo.org/record/216026/files/figure.png" pageId="1" pageNumber="484">Fig. 1</figureCitation>
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C), covering dorsal and lateral surfaces nearly completely, except for most anterior part of oral hood and caudal lobes; with 10–12 columns in mid-trunk region, each column comprising 33–37 ancres; central tine (10 µm long) 1.4–1.7 times longer than the others (7 µm long) in mid-trunk region; ancres 11 µm in diagonal length in most parts of trunk, but smaller on anterior part of oral hood (4 µm in diagonal length) and caudal base (5 µm in diagonal length).
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</paragraph>
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<caption id="3A9BF7DBFFD9FFF056B3A28BFE1E85B9" httpUri="https://zenodo.org/record/216026/files/figure.png" pageId="2" pageNumber="485" targetBox="[214,1354,217,1941]" targetPageId="2">
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<paragraph id="6E5BA753FFD9FFF056B3A28BFE1E85B9" blockId="2.[151,1435,1964,2016]" pageId="2" pageNumber="485">
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<emphasis id="5C907B41FFD9FFF056B3A28BFEBD8598" bold="true" box="[151,275,1964,1986]" pageId="2" pageNumber="485">FIGURE 1.</emphasis>
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<taxonomicName id="A9E4DCD0FFD9FFF0573DA28BFDC7859B" box="[281,617,1964,1986]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="2" pageNumber="485" phylum="Gastrotricha" rank="species" species="anisoankyrum" status="sp. nov.">
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<emphasis id="5C907B41FFD9FFF0573DA28BFDC7859B" box="[281,617,1964,1986]" italics="true" pageId="2" pageNumber="485">Tetranchyroderma anisoankyrum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="47A3C63AFFD9FFF05454A28BFD078598" box="[624,681,1964,1985]" pageId="2" pageNumber="485" rank="species">
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<emphasis id="5C907B41FFD9FFF05454A28BFD078598" bold="true" box="[624,681,1964,1985]" pageId="2" pageNumber="485">n. sp.</emphasis>
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</taxonomicNameLabel>
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A, habitus, dorsal view; B, habitus, ventral view; C, pentancre. Scale bars = 50 µm (A, B), 10 µm (C).
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</paragraph>
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</caption>
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<paragraph id="6E5BA753FFD8FFF156E3A5BFFB6882A1" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="486">Two pairs of dorsolateral cirratum-type tubes slender, frontal tube (11–14 µm long) located just behind PhJIn (U43) and rear one (16 µm long) at mid-intestinal region (U67). Lateral cirratum-type tubes, 13–14 per side, ranging 11–16 µm long, more or less evenly spaced along whole body length from U20 to U90.</paragraph>
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<paragraph id="6E5BA753FFD8FFF156E3A423FD6B8389" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="486">Adhesive tubes: TbA 4 per side, comprising 1 small medial tube, 5 µm long at U16 and 3 ventrolateral tubes forming small arc, 7 µm long at U14 to U15. A pair of TbL (15 µm long), more slender than TbVL, situated behind first dorsolateral cirratum-type tube at U49. TbVL, 13 per side, robust, ranging 9–14 µm long, aligned in intestinal region (U45–U83) densely except for backmost tube behind anus at U90. Foot-type TbV absent. TbP 4 per side, forming a feeble, trifid pedicle with 2 distal tubes (7 µm long) and 1 elongate cirratum-type tube (10 µm long), accompanied by 1 small medial tube (4 µm long).</paragraph>
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<paragraph id="6E5BA753FFD8FFF156E3A4FBFB6083AC" blockId="3.[151,1436,151,1653]" box="[199,1230,476,501]" pageId="3" pageNumber="486">Ventral ciliation aligned in single column from just behind TbA to caudal base at U17–U95.</paragraph>
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<paragraph id="6E5BA753FFD8FFF156E3A4D8FCFD8039" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="486">Digestive tract: mouth opening broad (51 µm wide), with oral hood extending forward from U0 to U15; pharynx 72 µm long (measured from ventral border of oral opening to PhJIn), pharyngeal pores at U41; intestine broad and narrowing toward posterior end; anus opens at U88.</paragraph>
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<paragraph id="6E5BA753FFD8FFF156E3A74BFC0A80A9" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="486">
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Reproductive system: simultaneous hermaphrodite; single testis extending behind PhJIn at U54, posterior end bent to copulatory organ. A large ovum (
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<date id="1A5A8193FFD8FFF15440A7A8FD1F80F1" box="[612,689,655,680]" pageId="3" pageNumber="486" value="1945-10-30">30X45</date>
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µm in diameter) located dorsally in mid-intestinal region at U55– U76. Seminal receptacle small, located between ovum and copulatory organ at U78–U80. Copulatory organ slender, pyriform with truncated anterior border, located at U80–U89.
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</paragraph>
|
||
<paragraph id="6E5BA753FFD8FFF156E3A7DBFB288105" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="486">
|
||
<emphasis id="5C907B41FFD8FFF156E3A7DBFE8A814C" bold="true" box="[199,292,764,789]" pageId="3" pageNumber="486">Ecology</emphasis>
|
||
. Specimens occurred rarely in fine to medium sublittoral sands (
|
||
<quantity id="A91C0AB6FFD8FFF15222A7DBFBE0814D" box="[1030,1102,764,788]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="3" pageNumber="486" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">1–3 m</quantity>
|
||
in depth), often co-occurring together with
|
||
<taxonomicName id="A9E4DCD0FFD8FFF15719A638FC018161" authority="Lee & Chang, 2007" authorityName="Lee & Chang" authorityYear="2007" box="[317,943,799,824]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="multicirratum">
|
||
<emphasis id="5C907B41FFD8FFF15719A638FD1D8161" box="[317,691,799,824]" italics="true" pageId="3" pageNumber="486">Tetranchyroderma multicirratum</emphasis>
|
||
Lee & Chang, 2007
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A9E4DCD0FFD8FFF1559BA606FAD98161" authority="Lee & Chang, 2007" authorityName="Lee & Chang" authorityYear="2007" box="[959,1399,799,824]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="monokerosum">
|
||
<emphasis id="5C907B41FFD8FFF1559BA606FBD58161" box="[959,1147,799,824]" italics="true" pageId="3" pageNumber="486">T. monokerosum</emphasis>
|
||
Lee & Chang, 2007
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A9E4DCD0FFD8FFF153A3A606FDEF8105" authority="Lee & Chang, 2012" authorityName="Lee & Chang" authorityYear="2012" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="megabitubulatum">
|
||
<emphasis id="5C907B41FFD8FFF153A3A606FEF58104" italics="true" pageId="3" pageNumber="486">T. megabitubulatum</emphasis>
|
||
Lee & Chang, 2012
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="A9E4DCD0FFD8FFF15458A663FB2D8105" authority="Lee & Chang, 2003" authorityName="Lee & Chang" authorityYear="2003" box="[636,1155,836,861]" family="Thaumastodermatidae" genus="Ptychostomella" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="orientalis">
|
||
<emphasis id="5C907B41FFD8FFF15458A663FC348104" box="[636,922,836,861]" italics="true" pageId="3" pageNumber="486">Ptychostomella orientalis</emphasis>
|
||
Lee & Chang, 2003
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6E5BA753FFD8FFF156E3A640FBD481FD" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="486">
|
||
<emphasis id="5C907B41FFD8FFF156E3A640FD8181D9" bold="true" box="[199,559,871,896]" pageId="3" pageNumber="486">Measurements and variability</emphasis>
|
||
. Body lengths of
|
||
<specimenCount id="78E26CDAFFD8FFF154D9A640FCF781D9" box="[765,857,871,896]" pageId="3" pageNumber="486" type="adult">16 adult</specimenCount>
|
||
specimens ranged from 234 to 312 µm (mean 280 µm, SD 24), maximum widths 35–45 µm (14U–16U) when measured in glycerin mount.
|
||
</paragraph>
|
||
<paragraph id="6E5BA753FFD8FFF156B3A688FD218601" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="486">
|
||
The number and arrangement of TbL, TbP and dorsolateral cirratum-type tubes were consistent, while TbVL and lateral cirratum-type tubes were somewhat variable, ranging from
|
||
<date id="1A5A8193FFD8FFF155B8A6F3FBE581B4" box="[924,1099,980,1005]" pageId="3" pageNumber="486" value="1916-09-18" valueMax="1916-09-08" valueMin="1916-09-18">9–18 and 8–16</date>
|
||
per side, respectively. Three specimens showed a ‘foot-type’ lateral cirratum-type tube consisting of two tubes confluent at its base only on one side. TbA arrangement was relatively consistent as one medial and three lateral TbA per side, except for two specimens with four lateral TbA on one side.
|
||
</paragraph>
|
||
<paragraph id="6E5BA753FFD8FFF156E3A143FD0186B1" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="486">
|
||
<emphasis id="5C907B41FFD8FFF156E3A143FE138624" bold="true" box="[199,445,1124,1149]" pageId="3" pageNumber="486">Taxonomic affinities</emphasis>
|
||
. Among 42
|
||
<taxonomicName id="A9E4DCD0FFD8FFF15470A143FC8C8624" box="[596,802,1124,1149]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="genus">
|
||
<emphasis id="5C907B41FFD8FFF15470A143FC8C8624" box="[596,802,1124,1149]" italics="true" pageId="3" pageNumber="486">Tetranchyroderma</emphasis>
|
||
</taxonomicName>
|
||
species currently described with cuticular armature of pentanceres only,
|
||
<taxonomicName id="A9E4DCD0FFD8FFF15746A1AEFDB386F9" box="[354,541,1159,1184]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="anisoankyrum" status="sp. nov.">
|
||
<emphasis id="5C907B41FFD8FFF15746A1AEFDB386F9" box="[354,541,1159,1184]" italics="true" pageId="3" pageNumber="486">T. anisoankyrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="47A3C63AFFD8FFF15401A1A0FD2E86F9" box="[549,640,1159,1184]" pageId="3" pageNumber="486" rank="species">
|
||
<emphasis id="5C907B41FFD8FFF15401A1A0FD2E86F9" bold="true" box="[549,640,1159,1184]" pageId="3" pageNumber="486">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, in bearing ancres with different length, is allied with
|
||
<taxonomicName id="A9E4DCD0FFD8FFF152CEA1AEFEE1869D" authority="Remane, 1927" authorityName="Remane" authorityYear="1927" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="polyacanthus">
|
||
<emphasis id="5C907B41FFD8FFF152CEA1AEFA3286F9" box="[1258,1436,1159,1184]" italics="true" pageId="3" pageNumber="486">T. polyacanthus</emphasis>
|
||
(Remane, 1927)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A9E4DCD0FFD8FFF15778A18AFBFB869C" authority="Hummon, Todaro, Balsamo & Tongiorgi, 1996" authorityName="Hummon, Todaro, Balsamo & Tongiorgi" authorityYear="1996" box="[348,1109,1196,1221]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="tanymesatherum">
|
||
<emphasis id="5C907B41FFD8FFF15778A18AFD9D869C" box="[348,563,1196,1221]" italics="true" pageId="3" pageNumber="486">T. tanymesatherum</emphasis>
|
||
Hummon, Todaro, Balsamo & Tongiorgi, 1996
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A9E4DCD0FFD8FFF15246A18AFE8086B1" authority="Nicholas & Todaro, 2006" authorityName="Nicholas & Todaro" authorityYear="2006" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="pentaspersus">
|
||
<emphasis id="5C907B41FFD8FFF15246A18AFABF869D" box="[1122,1297,1197,1220]" italics="true" pageId="3" pageNumber="486">T. pentaspersus</emphasis>
|
||
Nicholas & Todaro, 2006
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="A9E4DCD0FFD8FFF1574DA1F6FD0486B1" authority="Hummon, 2011" authorityName="Hummon" authorityYear="2011" box="[361,682,1231,1256]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="corallium">
|
||
<emphasis id="5C907B41FFD8FFF1574DA1F6FE5F86B1" box="[361,497,1231,1256]" italics="true" pageId="3" pageNumber="486">T. corallium</emphasis>
|
||
Hummon, 2011
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6E5BA753FFD8FFF156E3A1D3FC548721" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="486">
|
||
<taxonomicName id="A9E4DCD0FFD8FFF156E3A1D3FD928754" box="[199,572,1268,1293]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="anisoankyrum" status="sp. nov.">
|
||
<emphasis id="5C907B41FFD8FFF156E3A1D3FD928754" box="[199,572,1268,1293]" italics="true" pageId="3" pageNumber="486">Tetranchyroderma anisoankyrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="47A3C63AFFD8FFF15462A1D3FD0C8754" box="[582,674,1268,1293]" pageId="3" pageNumber="486" rank="species">
|
||
<emphasis id="5C907B41FFD8FFF15462A1D3FD0C8754" bold="true" box="[582,674,1268,1293]" pageId="3" pageNumber="486">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is most characteristic and easily distinguished from them, for it is the only species that has a pair of rod-like cephalic tentacles on the anterior border of oral hood. Moreover,
|
||
<taxonomicName id="A9E4DCD0FFD8FFF153A3A03EFE98870C" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="anisoankyrum" status="sp. nov.">
|
||
<emphasis id="5C907B41FFD8FFF153A3A03EFE98870C" italics="true" pageId="3" pageNumber="486">T. anisoankyrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="47A3C63AFFD8FFF15763A01BFE05870C" box="[327,427,1340,1365]" pageId="3" pageNumber="486" rank="species">
|
||
<emphasis id="5C907B41FFD8FFF15763A01BFE05870C" bold="true" box="[327,427,1340,1365]" pageId="3" pageNumber="486">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs clearly from three of the above four allied species (
|
||
<taxonomicName id="A9E4DCD0FFD8FFF15297A01AFAC0870C" box="[1203,1390,1340,1365]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="polyacanthus">
|
||
<emphasis id="5C907B41FFD8FFF15297A01AFAC0870C" box="[1203,1390,1340,1365]" italics="true" pageId="3" pageNumber="486">T. polyacanthus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A9E4DCD0FFD8FFF153A3A01AFEFE8721" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="tanymesatherum">
|
||
<emphasis id="5C907B41FFD8FFF153A3A01AFEFE8721" italics="true" pageId="3" pageNumber="486">T. tanymesatherum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A9E4DCD0FFD8FFF157A2A046FD9D8721" box="[390,563,1377,1400]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="pentaspersus">
|
||
<emphasis id="5C907B41FFD8FFF157A2A046FD9D8721" box="[390,563,1377,1400]" italics="true" pageId="3" pageNumber="486">T. pentaspersus</emphasis>
|
||
</taxonomicName>
|
||
) by the presence of cirratum-type tubes.
|
||
</paragraph>
|
||
<paragraph id="6E5BA753FFD8FFF156E3A0A3FB1887BC" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="486">
|
||
<taxonomicName id="A9E4DCD0FFD8FFF156E3A0A3FD9A87C4" box="[199,564,1412,1437]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="polyacanthus">
|
||
<emphasis id="5C907B41FFD8FFF156E3A0A3FD9A87C4" box="[199,564,1412,1437]" italics="true" pageId="3" pageNumber="486">Tetranchyroderma polyacanthus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A9E4DCD0FFD8FFF15455A0A2FCE687C4" box="[625,840,1412,1437]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="tanymesatherum">
|
||
<emphasis id="5C907B41FFD8FFF15455A0A2FCE687C4" box="[625,840,1412,1437]" italics="true" pageId="3" pageNumber="486">T. tanymesatherum</emphasis>
|
||
</taxonomicName>
|
||
have an extremely long central tine more than two times longer than peripheral tines, 4–5 times and 2–3 times longer, respectively, (cf. the key to the species of
|
||
<taxonomicName id="A9E4DCD0FFD8FFF156B3A0EBFECB87BC" box="[151,357,1484,1509]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="genus">
|
||
<emphasis id="5C907B41FFD8FFF156B3A0EBFECB87BC" box="[151,357,1484,1509]" italics="true" pageId="3" pageNumber="486">Tetranchyroderma</emphasis>
|
||
</taxonomicName>
|
||
in Todaro 2002), whereas 1.4–1.7 times longer in
|
||
<taxonomicName id="A9E4DCD0FFD8FFF155BFA0EAFBFA87BC" box="[923,1108,1484,1509]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="anisoankyrum" status="sp. nov.">
|
||
<emphasis id="5C907B41FFD8FFF155BFA0EAFBFA87BC" box="[923,1108,1484,1509]" italics="true" pageId="3" pageNumber="486">T. anisoankyrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="47A3C63AFFD8FFF15278A0EBFB1887BC" box="[1116,1206,1484,1509]" pageId="3" pageNumber="486" rank="species">
|
||
<emphasis id="5C907B41FFD8FFF15278A0EBFB1887BC" bold="true" box="[1116,1206,1484,1509]" pageId="3" pageNumber="486">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="6E5BA753FFD8FFF156E3A0C8FB62842C" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="486">
|
||
<taxonomicName id="A9E4DCD0FFD8FFF156E3A0C8FD808451" box="[199,558,1519,1544]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="pentaspersus">
|
||
<emphasis id="5C907B41FFD8FFF156E3A0C8FD808451" box="[199,558,1519,1544]" italics="true" pageId="3" pageNumber="486">Tetranchyroderma pentaspersus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A9E4DCD0FFD8FFF15441A0D6FD408451" box="[613,750,1519,1544]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="corallium">
|
||
<emphasis id="5C907B41FFD8FFF15441A0D6FD408451" box="[613,750,1519,1544]" italics="true" pageId="3" pageNumber="486">T. corallium</emphasis>
|
||
</taxonomicName>
|
||
have a central tine less than two times as in
|
||
<taxonomicName id="A9E4DCD0FFD8FFF152C6A0D6FA328451" box="[1250,1436,1519,1544]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="anisoankyrum" status="sp. nov.">
|
||
<emphasis id="5C907B41FFD8FFF152C6A0D6FA328451" box="[1250,1436,1519,1544]" italics="true" pageId="3" pageNumber="486">T. anisoankyrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="47A3C63AFFD8FFF156B3A333FF5F8474" box="[151,241,1556,1581]" pageId="3" pageNumber="486" rank="species">
|
||
<emphasis id="5C907B41FFD8FFF156B3A333FF5F8474" bold="true" box="[151,241,1556,1581]" pageId="3" pageNumber="486">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
However, the former
|
||
<taxonomicName id="A9E4DCD0FFD8FFF157CDA332FD388475" box="[489,662,1557,1580]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="pentaspersus">
|
||
<emphasis id="5C907B41FFD8FFF157CDA332FD388475" box="[489,662,1557,1580]" italics="true" pageId="3" pageNumber="486">T. pentaspersus</emphasis>
|
||
</taxonomicName>
|
||
was different from the new species by a large, thin-walled oral hood with a smooth edge, bifid pedicle and lacking TbDL, and the latter
|
||
<taxonomicName id="A9E4DCD0FFD8FFF155A4A31EFBA68409" box="[896,1032,1591,1616]" family="Thaumastodermatidae" genus="Tetranchyroderma" kingdom="Animalia" order="Macrodasyida" pageId="3" pageNumber="486" phylum="Gastrotricha" rank="species" species="corallium">
|
||
<emphasis id="5C907B41FFD8FFF155A4A31EFBA68409" box="[896,1032,1591,1616]" italics="true" pageId="3" pageNumber="486">T. corallium</emphasis>
|
||
</taxonomicName>
|
||
by truncated head, larger number of TbDL, dorsolateral cirratum-type tube and medial TbP between pedicles and a presence of TbV.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |