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<document id="27DCF44215C07611393B9F11FF29B630" ID-DOI="10.11646/zootaxa.3599.3.2" ID-GBIF-Dataset="65cc8b22-f862-47b5-91cb-5d1e70d182cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="216117" ID-ZooBank="DCE30D81-4C5B-415E-B458-A5C332DA2D0D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460567401535" checkinUser="plazi" docAuthor="Salgado, Luiz Gustavo Vargas, Carvalho, Alcimar Do Lago &amp; Pinto, Ângelo Parise" docDate="2013" docId="DF7A8C5E3777FFFAA1CFFF092DFEF970" docLanguage="en" docName="zt03599p245.pdf" docOrigin="Zootaxa 3599 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrothemis hemichlora Burmeister 1839" docType="treatment" docVersion="7" lastPageNumber="238" masterDocId="2343F426377EFFF3A158FF9E2D0BFFF2" masterDocTitle="Larval taxonomy of Macrothemis Hagen, 1868 (Odonata: Libellulidae), with descriptions of four larvae and a key to the fourteen known species" masterLastPageNumber="245" masterPageNumber="229" pageNumber="238" updateTime="1698321284514" updateUser="plazi">
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<mods:title id="3B14D740B681719A713A8FF9DD529B13">Larval taxonomy of Macrothemis Hagen, 1868 (Odonata: Libellulidae), with descriptions of four larvae and a key to the fourteen known species</mods:title>
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<mods:namePart id="CE72A087582FC01EC995A8A5CBCC5C9D">Salgado, Luiz Gustavo Vargas</mods:namePart>
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<treatment id="DF7A8C5E3777FFFAA1CFFF092DFEF970" ID-DOI="http://doi.org/10.5281/zenodo.5658973" ID-GBIF-Taxon="119557982" ID-Zenodo-Dep="5658973" LSID="urn:lsid:plazi:treatment:DF7A8C5E3777FFFAA1CFFF092DFEF970" httpUri="http://treatment.plazi.org/id/DF7A8C5E3777FFFAA1CFFF092DFEF970" lastPageNumber="238" pageId="9" pageNumber="238">
<subSubSection id="1FC96EC33777FFFAA1CFFF092C41FF20" pageId="9" pageNumber="238" type="nomenclature">
<paragraph id="576C3D483777FFFAA1CFFF092FBCFF43" blockId="9.[151,695,151,210]" box="[151,695,151,177]" pageId="9" pageNumber="238">
<heading id="0C248A243777FFFAA1CFFF092FBCFF43" bold="true" box="[151,695,151,177]" fontSize="11" level="1" pageId="9" pageNumber="238" reason="1">
<taxonomicName id="90D346CB3777FFFAA1CFFF092FBCFF43" ID-CoL="3XBXK" authority="Burmeister, 1839" authorityName="Burmeister" authorityYear="1839" box="[151,695,151,177]" class="Insecta" family="Libellulidae" genus="Macrothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="hemichlora">
<emphasis id="65A7E15A3777FFFAA1CFFF092FBCFF43" bold="true" box="[151,695,151,177]" pageId="9" pageNumber="238">
<emphasis id="65A7E15A3777FFFAA1CFFF092CC3FF43" bold="true" box="[151,456,151,177]" italics="true" pageId="9" pageNumber="238">Macrothemis hemichlora</emphasis>
(Burmeister, 1839)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="576C3D483777FFFAA1CFFF242C41FF20" blockId="9.[151,695,151,210]" box="[151,330,186,210]" pageId="9" pageNumber="238">
(
<figureCitation id="CFE821CD3777FFFAA1C6FF242C48FF20" box="[158,323,186,210]" captionStart="FIGURES 21 29" captionStartId="10.[151,269,1885,1909]" captionTargetBox="[202,1381,223,1858]" captionTargetId="figure@10.[177,1411,194,1863]" captionTargetPageId="10" captionText="FIGURES 21 29. Macrothemis hemichlora (Burmeister, 1839), ultimate stadium larva, 3: 21. right mandible, inner view; 22. left mandible, inner view; 23. right labial palp, dorsal view; 24. distal border of right labial palp, ventral view; 25. prementum, dorsal view; 26. abdomen, dorsal view; 27. abdomen, lateral view; 28. caudal appendages, dorsal view; 29. caudal appendages, lateral view." httpUri="https://zenodo.org/record/216120/files/figure.png" pageId="9" pageNumber="238">Figures 2129</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1FC96EC33777FFFAA1CFFE9C2DFEF970" pageId="9" pageNumber="238" type="materials_examined">
<paragraph id="576C3D483777FFFAA1CFFE9C2DE8FE97" blockId="9.[151,1436,258,1666]" pageId="9" pageNumber="238">
Material.
<collectingCountry id="2FC47DD83777FFFAA050FE9D2C79FEE9" box="[264,370,259,283]" name="Brazil" pageId="9" pageNumber="238">BRAZIL</collectingCountry>
, Rio de Janeiro State: Maricá municipality, Rio Ubatiba [
<geoCoordinate id="32E75B8F3777FFFAA54BFE9C29CEFEE8" box="[1043,1221,258,282]" direction="south" orientation="latitude" pageId="9" pageNumber="238" precision="1" value="-22.87145">22º5217.22S</geoCoordinate>
,
<geoCoordinate id="32E75B8F3777FFFAA58CFE9C2893FEE8" box="[1236,1432,258,282]" direction="west" orientation="longitude" pageId="9" pageNumber="238" precision="1" value="-42.783566">42º47 00.84W</geoCoordinate>
;
<quantity id="902B90AD3777FFFAA1CFFEB92DDEFECD" box="[151,213,295,319]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="238" unit="m" value="36.0">36 m</quantity>
a.s.l.],
<date id="236D1B883777FFFAA06DFEB92CA6FECD" box="[309,429,295,319]" pageId="9" pageNumber="238" value="2000-05-24">24.V.2000</date>
, Equipe Entomologia
<collectionCode id="31C2A58D3777FFFAA3E2FEB62FF6FECD" box="[698,765,296,319]" pageId="9" pageNumber="238">UFRJ</collectionCode>
leg., ̩ F-0, adult emergence
<date id="236D1B883777FFFAA528FEB929FEFECD" box="[1136,1269,295,319]" pageId="9" pageNumber="238" value="2000-06-04">04.VI.2000</date>
. Deposited in
<collectionCode id="31C2A58D3777FFFAA1CFFED02DD4FE97" box="[151,223,334,357]" pageId="9" pageNumber="238">DZRJ</collectionCode>
.
</paragraph>
<paragraph id="576C3D483777FFFAA1CFFEED29C2FE78" blockId="9.[151,1436,258,1666]" box="[151,1225,370,395]" pageId="9" pageNumber="238">
Larva very similar to
<taxonomicName id="90D346CB3777FFFAA0D7FEEA2F6DFE79" box="[399,614,372,395]" class="Insecta" family="Libellulidae" genus="Macrothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="238" phylum="Arthropoda" rank="subSpecies" species="imitans" subSpecies="imitans">
<emphasis id="65A7E15A3777FFFAA0D7FEEA2F6DFE79" box="[399,614,372,395]" italics="true" pageId="9" pageNumber="238">M. imitans imitans</emphasis>
</taxonomicName>
described above, differing by the following features:
</paragraph>
<paragraph id="576C3D483777FFFAA19FFE092ED5FE5D" blockId="9.[151,1436,258,1666]" box="[199,990,407,431]" pageId="9" pageNumber="238">Posterior areas of head and thorax with short setae flattened apically.</paragraph>
<paragraph id="576C3D483777FFFAA19FFE232861FC48" blockId="9.[151,1436,258,1666]" pageId="9" pageNumber="238">
Head (
<figureCitation id="CFE821CD3777FFFAA04CFE232CAAFE27" box="[276,417,445,469]" captionStart="FIGURES 21 29" captionStartId="10.[151,269,1885,1909]" captionTargetBox="[202,1381,223,1858]" captionTargetId="figure@10.[177,1411,194,1863]" captionTargetPageId="10" captionText="FIGURES 21 29. Macrothemis hemichlora (Burmeister, 1839), ultimate stadium larva, 3: 21. right mandible, inner view; 22. left mandible, inner view; 23. right labial palp, dorsal view; 24. distal border of right labial palp, ventral view; 25. prementum, dorsal view; 26. abdomen, dorsal view; 27. abdomen, lateral view; 28. caudal appendages, dorsal view; 29. caudal appendages, lateral view." httpUri="https://zenodo.org/record/216120/files/figure.png" pageId="9" pageNumber="238">Figs. 2125</figureCitation>
). About 2.5x as wide as long in dorsal view; occipital lobes covered with short flattened and long filiform setae. Vertex with short filiform setae, not especially concentrated. Occiput bearing short robust setae flattened distally. Lateral portion of eyes rather pronounced forward and laterally. Proportional length of antennomeres: 60, 87, 100, 65, 90, 86, 70. Mandibular formula L 12234 abb / R 1234 y- abd (
<figureCitation id="CFE821CD3777FFFAA43AFDB32DE8FD98" captionStart="FIGURES 21 29" captionStartId="10.[151,269,1885,1909]" captionTargetBox="[202,1381,223,1858]" captionTargetId="figure@10.[177,1411,194,1863]" captionTargetPageId="10" captionText="FIGURES 21 29. Macrothemis hemichlora (Burmeister, 1839), ultimate stadium larva, 3: 21. right mandible, inner view; 22. left mandible, inner view; 23. right labial palp, dorsal view; 24. distal border of right labial palp, ventral view; 25. prementum, dorsal view; 26. abdomen, dorsal view; 27. abdomen, lateral view; 28. caudal appendages, dorsal view; 29. caudal appendages, lateral view." httpUri="https://zenodo.org/record/216120/files/figure.png" pageId="9" pageNumber="238">Figs. 2122</figureCitation>
); left mandible: molars and incisors 1 and 4 with obtuse apex, 22 and 3 with acute apex, size bb&gt;a, size 3&gt;4&gt;1&gt;2, prominence 3&gt;4&gt;2&gt;1; right mandible: molars a and d and incisors 13 with acute apex, b and 4 obtuse, size b&gt;d&gt;a, size 4&gt;1&gt;3&gt;2, prominence 4&gt;3&gt;2&gt;1, y- acute. Labium when folded reaching level between pro- and mesocoxae; prementum longer than width in ventral view; ligula projected 0.25, forming an angle of about 100º (
<figureCitation id="CFE821CD3777FFFAA071FD792C89FD0D" box="[297,386,743,767]" captionStart="FIGURES 21 29" captionStartId="10.[151,269,1885,1909]" captionTargetBox="[202,1381,223,1858]" captionTargetId="figure@10.[177,1411,194,1863]" captionTargetPageId="10" captionText="FIGURES 21 29. Macrothemis hemichlora (Burmeister, 1839), ultimate stadium larva, 3: 21. right mandible, inner view; 22. left mandible, inner view; 23. right labial palp, dorsal view; 24. distal border of right labial palp, ventral view; 25. prementum, dorsal view; 26. abdomen, dorsal view; 27. abdomen, lateral view; 28. caudal appendages, dorsal view; 29. caudal appendages, lateral view." httpUri="https://zenodo.org/record/216120/files/figure.png" pageId="9" pageNumber="238">Fig. 23</figureCitation>
); margin of ligula bearing a submarginal row of about 30 setae, with about 10 of these distinctly larger; premental setae 78; external margins of prementum bearing a row of about 10 short and robust setae. Labial palps with 5 palpal setae (
<figureCitation id="CFE821CD3777FFFAA30DFCAC2FA1FCB8" box="[597,682,818,842]" captionStart="FIGURES 21 29" captionStartId="10.[151,269,1885,1909]" captionTargetBox="[202,1381,223,1858]" captionTargetId="figure@10.[177,1411,194,1863]" captionTargetPageId="10" captionText="FIGURES 21 29. Macrothemis hemichlora (Burmeister, 1839), ultimate stadium larva, 3: 21. right mandible, inner view; 22. left mandible, inner view; 23. right labial palp, dorsal view; 24. distal border of right labial palp, ventral view; 25. prementum, dorsal view; 26. abdomen, dorsal view; 27. abdomen, lateral view; 28. caudal appendages, dorsal view; 29. caudal appendages, lateral view." httpUri="https://zenodo.org/record/216120/files/figure.png" pageId="9" pageNumber="238">Fig. 24</figureCitation>
); proximal portion with setella and a group of 68 short spiniform setae; proximal 0.66 of outer margin with a row of about 1112 short and robust setae; distal margin with 9 crenations (
<figureCitation id="CFE821CD3777FFFAA045FCE32C78FC67" box="[285,371,893,917]" captionStart="FIGURES 21 29" captionStartId="10.[151,269,1885,1909]" captionTargetBox="[202,1381,223,1858]" captionTargetId="figure@10.[177,1411,194,1863]" captionTargetPageId="10" captionText="FIGURES 21 29. Macrothemis hemichlora (Burmeister, 1839), ultimate stadium larva, 3: 21. right mandible, inner view; 22. left mandible, inner view; 23. right labial palp, dorsal view; 24. distal border of right labial palp, ventral view; 25. prementum, dorsal view; 26. abdomen, dorsal view; 27. abdomen, lateral view; 28. caudal appendages, dorsal view; 29. caudal appendages, lateral view." httpUri="https://zenodo.org/record/216120/files/figure.png" pageId="9" pageNumber="238">Fig. 25</figureCitation>
); central crenations with 35 setae,
<quantity id="902B90AD3777FFFAA248FCE32E52FC67" box="[784,857,893,917]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="9" pageNumber="238" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">13 in</quantity>
those closest to outer margin; inner margin with a row of about 1217 setae; ventral surface of palps with small dark spots not coincident with setae insertion.
</paragraph>
<paragraph id="576C3D483777FFFAA19FFC592FF8FBF7" blockId="9.[151,1436,258,1666]" pageId="9" pageNumber="238">Thorax. Pronotum with short setae; posterolateral margins with short robust setae distally flattened; inverted “V” spot barely visible in the only examined exuvia.</paragraph>
<paragraph id="576C3D483777FFFAA19FFB8C2EEEFB87" blockId="9.[151,1436,258,1666]" pageId="9" pageNumber="238">
Abdomen (
<figureCitation id="CFE821CD3777FFFAA01FFB8C2CDEFBD8" box="[327,469,1042,1066]" captionStart="FIGURES 21 29" captionStartId="10.[151,269,1885,1909]" captionTargetBox="[202,1381,223,1858]" captionTargetId="figure@10.[177,1411,194,1863]" captionTargetPageId="10" captionText="FIGURES 21 29. Macrothemis hemichlora (Burmeister, 1839), ultimate stadium larva, 3: 21. right mandible, inner view; 22. left mandible, inner view; 23. right labial palp, dorsal view; 24. distal border of right labial palp, ventral view; 25. prementum, dorsal view; 26. abdomen, dorsal view; 27. abdomen, lateral view; 28. caudal appendages, dorsal view; 29. caudal appendages, lateral view." httpUri="https://zenodo.org/record/216120/files/figure.png" pageId="9" pageNumber="238">Figs. 2629</figureCitation>
). Tergites with spiniform setae of lateral margins equally distributed along abdomen (
<figureCitation id="CFE821CD3777FFFAA1C7FBA92C26FBBD" box="[159,301,1079,1103]" captionStart="FIGURES 21 29" captionStartId="10.[151,269,1885,1909]" captionTargetBox="[202,1381,223,1858]" captionTargetId="figure@10.[177,1411,194,1863]" captionTargetPageId="10" captionText="FIGURES 21 29. Macrothemis hemichlora (Burmeister, 1839), ultimate stadium larva, 3: 21. right mandible, inner view; 22. left mandible, inner view; 23. right labial palp, dorsal view; 24. distal border of right labial palp, ventral view; 25. prementum, dorsal view; 26. abdomen, dorsal view; 27. abdomen, lateral view; 28. caudal appendages, dorsal view; 29. caudal appendages, lateral view." httpUri="https://zenodo.org/record/216120/files/figure.png" pageId="9" pageNumber="238">Figs. 2627</figureCitation>
); lateral spines on S89, as long as 0.7 and 0.8 mediodorsal length of respective segment. Cerci ca. 0.8 of epiproct length in lateral view (
<figureCitation id="CFE821CD3777FFFAA312FBC32FDCFB87" box="[586,727,1117,1141]" captionStart="FIGURES 21 29" captionStartId="10.[151,269,1885,1909]" captionTargetBox="[202,1381,223,1858]" captionTargetId="figure@10.[177,1411,194,1863]" captionTargetPageId="10" captionText="FIGURES 21 29. Macrothemis hemichlora (Burmeister, 1839), ultimate stadium larva, 3: 21. right mandible, inner view; 22. left mandible, inner view; 23. right labial palp, dorsal view; 24. distal border of right labial palp, ventral view; 25. prementum, dorsal view; 26. abdomen, dorsal view; 27. abdomen, lateral view; 28. caudal appendages, dorsal view; 29. caudal appendages, lateral view." httpUri="https://zenodo.org/record/216120/files/figure.png" pageId="9" pageNumber="238">Figs. 2829</figureCitation>
); apex of cerci straight.
</paragraph>
<paragraph id="576C3D483777FFFAA19FFB1D2946FADD" blockId="9.[151,1436,258,1666]" pageId="9" pageNumber="238">Measurements (in mm; n=1). Total length 14.22; head mediodorsal length 1.85; head max. width 4.35; antenna total length 1.40; length of antennomeres 0.15, 0.22, 0.25, 0.16, 0.23, 0.22, 0.17; prementum length 3.37; prementum maximum width 3.05; hind wing sheath length 4.73; metafemur length 3.06; metatibia length 4.00; abdomen length (incl. appendages) 7.12; abdomen max. width (S6) 5.20; epiproct length in lateral view 0.81; paraproct max. length in lateral view 0.76; cercus length in lateral view 0.65.</paragraph>
<paragraph id="576C3D483777FFFAA19FFAA32DFEF970" blockId="9.[151,1436,258,1666]" pageId="9" pageNumber="238">
Diagnosis. The F-0 larva of
<taxonomicName id="90D346CB3777FFFAA352FAA02FB8FAA7" box="[522,691,1341,1365]" class="Insecta" family="Libellulidae" genus="Macrothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="hemichlora">
<emphasis id="65A7E15A3777FFFAA352FAA02FB8FAA7" box="[522,691,1341,1365]" italics="true" pageId="9" pageNumber="238">M. hemichlora</emphasis>
</taxonomicName>
can be correctly identified as
<taxonomicName id="90D346CB3777FFFAA554FAA329A9FAA7" box="[1036,1186,1341,1365]" class="Insecta" family="Libellulidae" genus="Macrothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="65A7E15A3777FFFAA554FAA329A9FAA7" box="[1036,1186,1341,1365]" italics="true" pageId="9" pageNumber="238">Macrothemis</emphasis>
</taxonomicName>
in the majority of the keys examined, except that of Rodríguez
<bibRefCitation id="334240B93777FFFAA337FAFC2E2EFA88" author="Capitulo" box="[623,805,1378,1402]" pageId="9" pageNumber="238" refString="Rodrigues Capitulo, A. (1992) Los Odonata de la republica Argentina. Fauna de agua dulce de la Republica Argentina, 34, 1 - 91." type="journal article" year="1992">Capítulo (1992)</bibRefCitation>
(
<tableCitation id="1A5108F33777FFFAA26BFAFC2EBEFA88" box="[819,949,1378,1402]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="6.[119,197,521,541]" captionStartId-1="8.[151,242,162,186]" captionTargetBox-1="[167,1392,216,673]" captionTargetPageId-1="8" captionText-0="TABLE 1. Determinations in genera for all described larvae of Macrothemis, Brechmorhoga, Gynothemis and Scapanea species using six of the most used keys. Numbers in the genera line correspond respectively to correct, possible correct (dubious), incorrect, possible incorrect (dubious), error (not applied characters) identification. Names without asterisk (s) corresponding to potential unambiguous determinations to genus." captionText-1="TABLE 2. Summary of the results of identification attempts shown in Table 1." httpUri-1="http://table.plazi.org/id/03AC6DC03776FFFBA1CFFF3C2EE8FF4B" pageId="9" pageNumber="238" tableUuid-1="03AC6DC03776FFFBA1CFFF3C2EE8FF4B">Tables 12</tableCitation>
). The following combination of characters separates it from the other described larvae of the genus: antennomeres 57 distinctly longer than half of 3; ligula moderately prominent, with lateral margins forming an angle of ca. 100o (
<figureCitation id="CFE821CD3777FFFAA566FA2E299FFA3A" box="[1086,1172,1456,1480]" captionStart="FIGURES 11 20" captionStartId="5.[151,269,1915,1939]" captionTargetBox="[182,1413,182,1903]" captionTargetId="figure@5.[161,1446,172,1911]" captionTargetPageId="5" captionText="FIGURES 11 20. Macrothemis declivata Calvert, 1909, ultimate stadium larva, 3: 11. right mandible, inner view; 12. left mandible, inner view; 13. right labial palp, dorsal view; 14. distal border of right labial palp, ventral view; 15. prementum, dorsal view; 16. abdomen, dorsal view; 17. abdomen, lateral view; 18. abdominal segment 9, ventral view; 19 caudal appendages, dorsal view; 20. caudal appendages, lateral view." httpUri="https://zenodo.org/record/216119/files/figure.png" pageId="9" pageNumber="238">Fig. 15</figureCitation>
); premental setae 78; rows of premental setae not clearly divided in two groups; labial palps with semicircular crenations, notches deep (regular
<typeStatus id="886883EA3777FFFAA06FFA642C61F9E0" box="[311,362,1530,1554]" pageId="9" pageNumber="238">type</typeStatus>
) (
<figureCitation id="CFE821CD3777FFFAA0DBFA642F1BF9E0" box="[387,528,1530,1554]" captionStart="FIGURES 11 20" captionStartId="5.[151,269,1915,1939]" captionTargetBox="[182,1413,182,1903]" captionTargetId="figure@5.[161,1446,172,1911]" captionTargetPageId="5" captionText="FIGURES 11 20. Macrothemis declivata Calvert, 1909, ultimate stadium larva, 3: 11. right mandible, inner view; 12. left mandible, inner view; 13. right labial palp, dorsal view; 14. distal border of right labial palp, ventral view; 15. prementum, dorsal view; 16. abdomen, dorsal view; 17. abdomen, lateral view; 18. abdominal segment 9, ventral view; 19 caudal appendages, dorsal view; 20. caudal appendages, lateral view." httpUri="https://zenodo.org/record/216119/files/figure.png" pageId="9" pageNumber="238">Figs. 1314</figureCitation>
); 5 palpal setae; setae of distal margin of labial palps cylindrical; dorsal hooks on S39 (
<figureCitation id="CFE821CD3777FFFAA05EF9BE2C7EF9CA" box="[262,373,1568,1592]" captionStart="FIGURES 1 10" captionStartId="3.[151,269,1887,1911]" captionTargetBox="[208,1403,211,1858]" captionTargetId="figure@3.[179,1413,195,1864]" captionTargetPageId="3" captionText="FIGURES 1 10. Macrothemis imitans imitans Karsch, 1890, ultimate stadium larva, 3: 1 head, dorsal view; 2. right mandible, inner view; 3. left mandible, inner view; 4. right labial palp, dorsal view; 5. distal border of right labial palp, ventral view; 6. prementum, dorsal view; 7. abdomen, dorsal view; 8. abdomen, lateral view; 9. caudal appendages, dorsal view; 10. caudal appendages, lateral view. “ a ” and “ b ”, measurements used to represent the projection of ligula." httpUri="https://zenodo.org/record/216118/files/figure.png" pageId="9" pageNumber="238">Figs. 78</figureCitation>
); dorsal hook on S5 not reduced, larger than those of preceding segments; dorsal hook of S9 shorter and less pronounced distally than corresponding lateral spines in dorsal view; lateral spines of S9 straight.
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</subSubSection>
</treatment>
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