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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.245.3416" ID-PMC="PMC3577090" ID-Pensoft-Pub="1313-2970-245-1" ID-Pensoft-UUID="FFFF52503A0AFF882450FFB66D45FF8E" ID-PubMed="23431087" ID-Zenodo-Dep="578462" ModsDocID="1313-2970-245-1" checkinTime="1425642919252" checkinUser="pensoft" docAuthor="Bousquet, Yves" docDate="2012" docId="D4D6FFB5F03CF5682500429906C7BF8A" docLanguage="en" docName="ZooKeys 245: 1-1722" docOrigin="ZooKeys 245" docPubDate="2012-11-28" docSource="http://dx.doi.org/10.3897/zookeys.245.3416" docTitle="Bembidion impotens Casey 1918" docType="treatment" docVersion="9" id="FFFF52503A0AFF882450FFB66D45FF8E" lastPageNumber="388" masterDocId="FFFF52503A0AFF882450FFB66D45FF8E" masterDocTitle="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico" masterLastPageNumber="1722" masterPageNumber="1" pageNumber="388" updateTime="1668154792637" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bousquet, Yves</mods:namePart>
<mods:affiliation>Agriculture and Agri-Food Canada, Central Experimental Farm, Ottawa, Ontario, Canada</mods:affiliation>
<mods:nameIdentifier type="email">bousquety1@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-11-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>245</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>1722</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.245.3416</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.245.3416</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-245-1</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578462</mods:identifier>
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<treatment ID-GBIF-Taxon="127867073" LSID="urn:lsid:plazi:treatment:D4D6FFB5F03CF5682500429906C7BF8A" httpUri="http://treatment.plazi.org/id/D4D6FFB5F03CF5682500429906C7BF8A" lastPageNumber="388" pageId="387" pageNumber="388">
<subSubSection pageId="387" pageNumber="388" type="nomenclature">
<paragraph pageId="387" pageNumber="388">
<taxonomicName LSID="D4D6FFB5-F03C-F568-2500-429906C7BF8A" authority="Casey, 1918" authorityName="Casey" authorityYear="1918" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bembidion impotens" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="impotens">Bembidion impotens Casey, 1918</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="387" pageNumber="388" type="reference_group">
<paragraph pageId="387" pageNumber="388">
<taxonomicName authorityName="LeConte" authorityYear="1847" class="Insecta" family="Carabidae" genus="Ochthedromus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="pictus">Ochthedromus pictus</taxonomicName>
LeConte, 1847: 461 [secondary homonym of
<taxonomicName baseAuthorityName="LeConte" baseAuthorityYear="1847" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="pictum">
<emphasis italics="true" pageId="387" pageNumber="388">Bembidion pictum</emphasis>
</taxonomicName>
(Duftschmid, 1812)]. Type locality: &quot;Rocky Mountains&quot; (original citation). Syntype(s) in MCZ [# 5537].
</paragraph>
<paragraph pageId="387" pageNumber="388">
<taxonomicName authorityName="Motschulky" authorityYear="1858" class="Hexapoda" family="Carabidae" genus="Emphanes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="flavopictus">Emphanes flavopictus</taxonomicName>
Motschulsky, 1858: 153. Replacement name for
<taxonomicName class="Hexapoda" family="Carabidae" genus="Emphanes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="pictus">
<emphasis italics="true" pageId="387" pageNumber="388">Emphanes pictus</emphasis>
</taxonomicName>
(LeConte, 1847). Note. This name was first introduced by Motschulsky (1850a: 13) in a catalogue but the taxon was not described. Subsequently Motschulsky (1858: 153) listed the name in synonymy with
<taxonomicName authorityName="LeConte" authorityYear="1847" class="Insecta" family="Carabidae" genus="Ochthedromus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="pictus">
<emphasis italics="true" pageId="387" pageNumber="388">Ochthedromus pictus</emphasis>
</taxonomicName>
LeConte and pointed out that since
<normalizedToken originalValue="LeContes">LeConte's</normalizedToken>
name was preoccupied, his name could be preserved. In my opinion such statement should be interpreted as the introduction of a replacement name. Casey (1918: 135) eventually described
<taxonomicName baseAuthorityName="Motschulsky" baseAuthorityYear="1858" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="flavopictum">
<emphasis italics="true" pageId="387" pageNumber="388">Bembidion flavopictum</emphasis>
</taxonomicName>
Motschulsky from the &quot;coast region, from S[an]ta Cruz to Humboldt, California.&quot; This name should have precedence over
<taxonomicName authorityName="Casey" authorityYear="1918" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="impotens">
<emphasis italics="true" pageId="387" pageNumber="388">Bembidion impotens</emphasis>
</taxonomicName>
Casey, 1918 but since
<normalizedToken originalValue="Caseys">Casey's</normalizedToken>
name has been used as valid since the 1920s, I believe it should be preserved and the case eventually submitted to the Commission for a ruling. The reversal of precedence (ICZN 1999: Article 23.9.1) could not be applied since
<normalizedToken originalValue="Motschulskys">Motschulsky's</normalizedToken>
name has been used as a valid name after 1899 by Casey (1918: 135).
</paragraph>
<paragraph pageId="387" pageNumber="388">
<taxonomicName authorityName="Casey" authorityYear="1918" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="impotens">Bembidion impotens</taxonomicName>
Casey, 1918: 129. Replacement name for
<taxonomicName baseAuthorityName="LeConte" baseAuthorityYear="1847" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="pictum">
<emphasis italics="true" pageId="387" pageNumber="388">Bembidion pictum</emphasis>
</taxonomicName>
(LeConte, 1847).
</paragraph>
<paragraph pageId="387" pageNumber="388">
<taxonomicName authorityName="Casey" authorityYear="1918" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="aestivum">Bembidion aestivum</taxonomicName>
Casey, 1918: 129. Type locality: &quot;Highland Park [Lake County], northern Illinois&quot; (original citation). Lectotype (♂), designated by Lindroth (1975: 121), in USNM [# 37005]. Synonymy established by Lindroth (1963b: 380).
</paragraph>
<paragraph pageId="387" pageNumber="388">
<taxonomicName authorityName="Casey" authorityYear="1918" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="frugale">Bembidion frugale</taxonomicName>
Casey, 1918: 130. Type locality: &quot;Yuma Reservation, California&quot; (original citation). Lectotype (♂), designated by Erwin (1984a: 174), in USNM [# 37006]. Synonymy established by Erwin (1984a: 174).
</paragraph>
<paragraph pageId="387" pageNumber="388">
<taxonomicName authorityName="Casey" authorityYear="1918" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="imbelle">Bembidion imbelle</taxonomicName>
Casey, 1918: 130. Type locality: &quot;Dallas [Dallas County], Texas&quot; (original citation for the lectotype). Lectotype (♂), designated by Erwin (1984a: 174), in USNM [# 37007]. Synonymy established by Erwin (1984a: 174).
</paragraph>
<paragraph pageId="387" pageNumber="388">
<taxonomicName authorityName="Casey" authorityYear="1918" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="gratuitum">Bembidion gratuitum</taxonomicName>
Casey, 1918: 130. Type locality:
<normalizedToken originalValue="«Arizona»">&quot;Arizona&quot;</normalizedToken>
(original citation). Lectotype (♂), designated by Erwin (1984a: 174), in USNM [# 37004]. Synonymy established by Erwin (1984a: 174).
</paragraph>
<paragraph pageId="387" pageNumber="388">
<taxonomicName authorityName="Casey" authorityYear="1918" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="virgatulum">Bembidion virgatulum</taxonomicName>
Casey, 1918: 131. Type locality: &quot;Reno [Washoe County], Nevada&quot; (original citation for the lectotype). Lectotype (♂), designated by Erwin (1984a: 174), in USNM [# 37003]. Synonymy established by Erwin (1984a: 174).
</paragraph>
<paragraph pageId="387" pageNumber="388">
<taxonomicName authorityName="Casey" authorityYear="1918" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="indigens">Bembidion indigens</taxonomicName>
Casey, 1918: 133. Type locality: &quot;Federal District, Mexico&quot; (original citation). Lectotype (♂), designated by Erwin (1984a: 174), in USNM [# 37017]. Synonymy established by Erwin (1984a: 174).
</paragraph>
<paragraph pageId="387" pageNumber="388">
<taxonomicName authorityName="Casey" authorityYear="1918" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="pullulum">Bembidion pullulum</taxonomicName>
Casey, 1918: 133. Type locality: &quot;Amecameca, Mexico&quot; (original citation). Lectotype (♂), designated by Erwin (1984a: 174), in USNM [# 37018]. Synonymy established by Erwin (1984a: 174).
</paragraph>
</subSubSection>
<subSubSection pageId="387" pageNumber="388" type="distribution">
<paragraph pageId="387" pageNumber="388">Distribution.</paragraph>
<paragraph pageId="387" pageNumber="388">
This species occurs from southwestern New Brunswick (Larochelle and
<normalizedToken originalValue="Larivière">Lariviere</normalizedToken>
1990a: 29, 34) to southern British Columbia (Lindroth 1963b: 381), south to southeastern California (Casey 1918: 130, as
<taxonomicName authorityName="Casey" authorityYear="1918" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="frugale">
<emphasis italics="true" pageId="387" pageNumber="388">Bembidion frugale</emphasis>
</taxonomicName>
; Andrews et al. 1979: 28), the Federal District in Mexico (Casey 1918: 133, as
<taxonomicName authorityName="Casey" authorityYear="1918" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="387" pageNumber="388" phylum="Arthropoda" rank="species" species="indigens">
<emphasis italics="true" pageId="387" pageNumber="388">Bembidion indigens</emphasis>
</taxonomicName>
), and southern Florida (Peck and Thomas 1998: 18).
</paragraph>
</subSubSection>
<subSubSection pageId="387" pageNumber="388" type="records">
<paragraph pageId="387" pageNumber="388">Records.</paragraph>
<paragraph pageId="387" pageNumber="388">
<emphasis bold="true" pageId="387" pageNumber="388">CAN</emphasis>
: BC, MB, NB, ON, QC, SK
<emphasis bold="true" pageId="387" pageNumber="388">USA</emphasis>
: AR, AZ, CA, CO, CT, FL, GA, IA, ID, IL, IN, KS, KY, LA, MA, MD, ME, MI, MN, MO, MS, MT, NC, NE, NJ, NM, NV, OH, OK, OR, PA, SC, SD, TN, TX, UT, VA, VT, WA, WI, WV, WY - Mexico
</paragraph>
</subSubSection>
</treatment>
</document>