128 lines
14 KiB
XML
128 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.395.7110" ID-GBIF-Dataset="d9627d89-aad8-4b48-b024-c7a97d580231" ID-PMC="PMC3978236" ID-Pensoft-Pub="1313-2970-395-33" ID-PubMed="24715788" ID-ZBK="B4D4D101C9AE47F09B7B320735B69D1D" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-395-33" ModsDocOrigin="ZooKeys 395" ModsDocTitle="Six new species of Philiris Röber, 1891 (Lepidoptera, Lycaenidae) from Papua New Guinea" checkinTime="1451246046685" checkinUser="pensoft" docAuthor="Mueller, Chris J." docDate="2014" docId="CB2C0F566492A7F2E346F664B48D791A" docLanguage="en" docName="ZooKeys 395: 33-55" docOrigin="ZooKeys 395" docSource="http://dx.doi.org/10.3897/zookeys.395.7110" docTitle="Philiris parsonsi Mueller, sp. n." docType="treatment" docUuid="92B7442D-AE55-430F-97EF-D716A2FC17D9" docUuidSource="ZooBank" docVersion="4" lastPageNumber="46" masterDocId="FFEBBE73FFD8D16FFFC64B5A751FFFB3" masterDocTitle="Six new species of Philiris Roeber, 1891 (Lepidoptera, Lycaenidae) from Papua New Guinea" masterLastPageNumber="55" masterPageNumber="33" pageNumber="45" updateTime="1668158131885" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Six new species of Philiris Roeber, 1891 (Lepidoptera, Lycaenidae) from Papua New Guinea</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Mueller, Chris J.</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2014</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>395</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>33</mods:start>
|
|
<mods:end>55</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.395.7110</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.395.7110</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-395-33</mods:identifier>
|
|
<mods:identifier type="ZBK">B4D4D101C9AE47F09B7B320735B69D1D</mods:identifier>
|
|
<mods:identifier type="ZooBank">B4D4D101C9AE47F09B7B320735B69D1D</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-GBIF-Taxon="152052283" LSID="urn:lsid:zoobank.org:act:92B7442D-AE55-430F-97EF-D716A2FC17D9" httpUri="http://treatment.plazi.org/id/CB2C0F566492A7F2E346F664B48D791A" lastPageId="13" lastPageNumber="46" pageId="12" pageNumber="45">
|
|
<subSubSection pageId="12" pageNumber="45" type="nomenclature">
|
|
<paragraph pageId="12" pageNumber="45">
|
|
<taxonomicName LSID="http://zoobank.org/92B7442D-AE55-430F-97EF-D716A2FC17D9" authority="Mueller" class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris parsonsi" order="Lepidoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="parsonsi">
|
|
Philiris parsonsi
|
|
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel pageId="12" pageNumber="45">sp. n.</taxonomicNameLabel>
|
|
Figs 56-58, 73
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="12" pageNumber="45" type="reference_group">
|
|
<paragraph pageId="12" pageNumber="45">
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris" order="Lepidoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="genus">Philiris</taxonomicName>
|
|
sp. c (
|
|
<bibRefCitation author="Parsons, MJ" journalOrPublisher="Academic Pr., London" pageId="15" pageNumber="48" title="The butterflies of Papua New Guinea. Their systematics and biology." year="1998">Parsons 1998</bibRefCitation>
|
|
, p. 378, Plates 53, XII, XXVI)
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="12" pageNumber="45" type="type material">
|
|
<paragraph pageId="12" pageNumber="45">Type material.</paragraph>
|
|
<paragraph pageId="12" pageNumber="45">Holotype ♂ (Figs 56-58, 73): "On Creek nr. Kupeng, Nr. Koinambe Mission, W. Highlands Prov; PNG, M. J. Parsons, 18-11-1981, I. F. T. A. coll. Alt 1500m", "Philiris sp. nov. ♂, M. J. Parsons det. 1982", "Brit. Mus. 1987-194", B.M.(N.H.) Rhopalocera No. (V) 1065". No Paratypes.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="13" lastPageNumber="46" pageId="12" pageNumber="45" type="diagnosis">
|
|
<paragraph pageId="12" pageNumber="45">Diagnosis.</paragraph>
|
|
<paragraph pageId="13" pageNumber="46">
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris parsonsi" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="parsonsi">
|
|
<pageBreakToken pageId="13" pageNumber="46" start="start">Philiris</pageBreakToken>
|
|
parsonsi
|
|
</taxonomicName>
|
|
is a distinctive species, presently known only from the unique male holotype, which
|
|
<bibRefCitation author="Parsons, MJ" journalOrPublisher="Academic Pr., London" pageId="15" pageNumber="48" title="The butterflies of Papua New Guinea. Their systematics and biology." year="1998">Parsons (1998)</bibRefCitation>
|
|
illustrated as an undescribed species. He compared the external facies and male genitalia to
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris angabunga" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="angabunga">Philiris angabunga</taxonomicName>
|
|
Bethune-Baker, 1908 (Holotype; Figs 59, 60), within the
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris refusa" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="refusa">Philiris refusa</taxonomicName>
|
|
(Grose-Smith, 1894) group. However, the shape of the forewing and the colouration and maculation of both wing surfaces are unlike any of the described species within that species group. The forewing of
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris parsonsi" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="parsonsi">Philiris parsonsi</taxonomicName>
|
|
is more elongated than in other species and the dark border is much narrower, with the exception of
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris biplaga" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="biplaga">Philiris biplaga</taxonomicName>
|
|
and
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris pagwi" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="pagwi">Philiris pagwi</taxonomicName>
|
|
Sands, 1979, which have borders parallel to the termen and broader at the tornus, respectively, while that in
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris parsonsi" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="parsonsi">Philiris parsonsi</taxonomicName>
|
|
is widest at the apex. The upperside ground colour in
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris parsonsi" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="parsonsi">Philiris parsonsi</taxonomicName>
|
|
is an unusual pale lilac colour and there are white scales present in the median area of the forewing, not present in other species. The underside configuration of black spots is more complex than in other species and bears a total of ten individual markings, whereas a maximum of five spots are present in any other species (that for
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris maculata" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="maculata">Philiris maculata</taxonomicName>
|
|
Sands, 1981).
|
|
</paragraph>
|
|
<paragraph pageId="13" pageNumber="46">
|
|
The male genitalia of
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris parsonsi" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="parsonsi">Philiris parsonsi</taxonomicName>
|
|
(Fig. 73) is also distinctive, the tegumen ring being very squat and squared dorsally when compared with that of
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris angabunga" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="angabunga">Philiris angabunga</taxonomicName>
|
|
(Fig. 74). The socii in
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris parsonsi" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="parsonsi">Philiris parsonsi</taxonomicName>
|
|
are widely spaced and the valvae are smaller than those of
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris angabunga" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="angabunga">Philiris angabunga</taxonomicName>
|
|
and rounded, whereas those of
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris angabunga" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="angabunga">Philiris angabunga</taxonomicName>
|
|
are triangular-shaped.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="13" pageNumber="46" type="description">
|
|
<paragraph pageId="13" pageNumber="46">Description.</paragraph>
|
|
<paragraph pageId="13" pageNumber="46">♂ (Figs 56-57): Forewing length 13.5 mm, antenna 7 mm (holotype). Head, palpus and thorax dark grey dorsally, white ventrally, abdomen dark grey dorsally, white ventrally, frons grey with white eye ring; legs white with black areas on tibiae; shaft black, ringed conspicuously with white between segments, apex of club brown.</paragraph>
|
|
<paragraph pageId="13" pageNumber="46">Fore wing termen very slightly convex, inner margin very slightly bowed near base, apex slightly rounded; upperside pale lilac-blue, a scattering of white scales in median area, largely concentrated, but not restricted to, the area between veins 2 and 4, apical area broadly dark brown-black, border 5 mm wide at apex, narrowing evenly to meet tornus, cilia dark brown; underside dull white, cell with a large black spot (approximately 1.5 mm wide) about two thirds from base, two smaller obscure brown-black spots surrounding this spot, slightly closer to base and junction of cubitus with vein 3, discocellulars black, forming an elongated spot, cilia white, dark brown-black at vein ends.</paragraph>
|
|
<paragraph pageId="13" pageNumber="46">Hind wing rounded; upperside pale lilac-blue, costa and inner margin dark, termen very narrowly dark brown (hairline thickness), cilia dark brown; underside dull white, two large (approximately 1.5 mm diameter) black spots in cell, about two thirds from base, a much smaller spot close to upper spot on basal side, a large black spot between veins 1a and 1b and another obscure dark brown spot of similar size between veins 1b and 2, both spots about one third from base, discocellulars black, forming an elongated spot, cilia as in forewing underside.</paragraph>
|
|
<paragraph pageId="13" pageNumber="46">Male genitalia (Fig. 73): Vinculum and tegumen ring approximately rectangular-shaped, expanded at sociuncus, sociuncus broad, socii with lateral margin rounded, dorsally socii widely separated by U-shaped sinus, saccus short and squared dorsally, brachium short and abruptly terminated; valva slightly asymmetrical, with right valva larger than left valva, valva rounded with a small protrusion apically; phallus with pre-zonal section shorter than post-zonal section, tapered apically.</paragraph>
|
|
<paragraph pageId="13" pageNumber="46">♀. Unknown.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="13" pageNumber="46" type="etymology">
|
|
<paragraph pageId="13" pageNumber="46">Etymology.</paragraph>
|
|
<paragraph pageId="13" pageNumber="46">Named after the collector of the holotype, Dr Michael Parsons, California.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="13" pageNumber="46" type="distribution">
|
|
<paragraph pageId="13" pageNumber="46">Distribution.</paragraph>
|
|
<paragraph pageId="13" pageNumber="46">Western Highlands Province, Papua New Guinea.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="13" pageNumber="46" type="remarks">
|
|
<paragraph pageId="13" pageNumber="46">Remarks.</paragraph>
|
|
<paragraph pageId="13" pageNumber="46">
|
|
The unique holotype was taken in a creek by Parsons (pers. comm. 2013), flying together with
|
|
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris angabunga" order="Lepidoptera" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="angabunga">Philiris angabunga</taxonomicName>
|
|
. Although not listed on the specimen label,
|
|
<bibRefCitation author="Parsons, MJ" journalOrPublisher="Academic Pr., London" pageId="15" pageNumber="48" title="The butterflies of Papua New Guinea. Their systematics and biology." year="1998">Parsons (1998)</bibRefCitation>
|
|
noted that the creek is known as Wara [Tok Pisin for River] Pimbi.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |