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<document id="4F19A8AA95B6B2AF368BEEB2F893BBB7" ID-CLB-Dataset="7964" ID-DOI="10.11646/zootaxa.4903.2.1" ID-GBIF-Dataset="37ab0d60-c931-4718-b85a-00d5087fca50" ID-ISSN="1175-5326" ID-Zenodo-Dep="4422874" ID-ZooBank="339B8883-E0FA-4310-968B-81E7AAC76376" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1610012447887" checkinUser="plazi" docAuthor="Yeshwanth, H. M., Chérot, F. &amp; Henry, T. J." docDate="2021" docId="C24787C5BF59FFFCFF74F3CD6847633A" docLanguage="en" docName="zootaxa.4903.2.1.pdf" docOrigin="Zootaxa 4903 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Isometopus (Jehania) mahal Distant 1911" docType="treatment" docVersion="11" lastPageNumber="161" masterDocId="3E7EFFBDBF53FFF7FFE3F062697E627B" masterDocTitle="The Isometopinae (Hemiptera: Heteroptera: Miridae) of India and Sri Lanka: A Review of the Subfamily, with Descriptions of Six New Species" masterLastPageNumber="193" masterPageNumber="151" pageNumber="161" updateTime="1698920153217" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="97D8121EB8D4DD8CF280220E17D8932F">The Isometopinae (Hemiptera: Heteroptera: Miridae) of India and Sri Lanka: A Review of the Subfamily, with Descriptions of Six New Species</mods:title>
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<mods:roleTerm id="11F01B3920BF35058872800803241C73">Author</mods:roleTerm>
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<mods:namePart id="46463BDE5112FB3570E56F7A09FC9FAC">Yeshwanth, H. M.</mods:namePart>
<mods:nameIdentifier id="43CE3F9C11EE36554EF4E12AB79955B2" type="ORCID">0000-0002-5104-3401</mods:nameIdentifier>
<mods:affiliation id="3971180730D0594EFDC18CC9E6266B09">Department of Entomology, University of Agricultural Sciences, GKVK, Bengaluru 560 065, India.</mods:affiliation>
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<mods:namePart id="63C924D67D529A723B10E52CF65D9088">Chérot, F.</mods:namePart>
<mods:nameIdentifier id="0A2AC75E4158FF03BCE0F072E641739B" type="ORCID">0000-0001-7918-7012</mods:nameIdentifier>
<mods:affiliation id="B9799046B1AEEB268D39C9067DE852FE">Département de l’Etude du Milieu Naturel et Agricole, Service Public de Wallonie, Gembloux, BE- 5030, Belgium. frederic. cherot @ spw. wallonie. be; https: // orcid. org / 0000 - 0001 - 7918 - 7012</mods:affiliation>
<mods:nameIdentifier id="D0486A0DCFFC3222BDE2BB1B11BE6441" type="email">frederic.cherot@spw.wallonie.be</mods:nameIdentifier>
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<mods:namePart id="D83A3523DCE34922A82D2467F6E787FF">Henry, T. J.</mods:namePart>
<mods:nameIdentifier id="E30BA439D6299F7F0307DAC4D9B315BA" type="ORCID">0000-0002-0653-7728</mods:nameIdentifier>
<mods:affiliation id="8ABAA416004ADE2B57A44196B69A0C3C">Systematic Entomology Laboratory, Agricultural Research Service, U. S. Department of Agriculture, c / o National Museum of Natural History, MRC- 168, Smithsonian Institution, Washington, DC 20013 - 7013 USA.</mods:affiliation>
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<mods:title id="40EBC4AD7BD805DA074F3B5104AE76D9">Zootaxa</mods:title>
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<mods:date id="F402426DCE69C1F237BB5904CA069C87">2021</mods:date>
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<mods:number id="03D88C258D17D8C4A0621E01B29960FC">2021-01-07</mods:number>
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<mods:number id="15863F401D461A240CB2A32F24949150">4903</mods:number>
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<treatment id="C24787C5BF59FFFCFF74F3CD6847633A" ID-DOI="http://doi.org/10.5281/zenodo.4569769" ID-GBIF-Taxon="179981705" ID-Zenodo-Dep="4569769" LSID="urn:lsid:plazi:treatment:C24787C5BF59FFFCFF74F3CD6847633A" httpUri="http://treatment.plazi.org/id/C24787C5BF59FFFCFF74F3CD6847633A" lastPageId="11" lastPageNumber="161" pageId="10" pageNumber="161">
<subSubSection id="02F46558BF59FFFDFF74F3CD6BE361B3" box="[151,669,942,969]" pageId="10" pageNumber="161" type="nomenclature">
<paragraph id="4A5136D3BF59FFFDFF74F3CD6BE361B3" blockId="10.[151,669,942,1005]" box="[151,669,942,969]" pageId="10" pageNumber="161">
<heading id="111981BFBF59FFFDFF74F3CD6BE361B3" bold="true" box="[151,669,942,969]" fontSize="11" level="1" pageId="10" pageNumber="161" reason="1">
<taxonomicName id="8DEE4D50BF59FFFDFF74F3CD6BE361B3" ID-CoL="3QNXR" authority="Distant, 1911" authorityName="Distant" authorityYear="1911" box="[151,669,942,969]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="mahal" subGenus="Jehania">
<emphasis id="789AEAC1BF59FFFDFF74F3CD6BE361B3" bold="true" box="[151,669,942,969]" pageId="10" pageNumber="161">
<emphasis id="789AEAC1BF59FFFDFF74F3CD688D61B2" bold="true" box="[151,499,942,969]" italics="true" pageId="10" pageNumber="161">Isometopus (Jehania) mahal</emphasis>
<bibRefCitation id="2E7F4B22BF59FFFDFE19F3CC6BE361B3" author="Distant, W. L." box="[506,669,942,968]" pageId="10" pageNumber="161" refId="ref18092" refString="Distant, W. L. (1911) The Fauna of British India, Including Ceylan and Burma. Rhynchota. Vol. V. Heteroptera, appendix. Taylor &amp; Francis, London, xii + 362 pp. [1910, dating follows Dolling (1991)]" type="book" year="1911">Distant, 1911</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="02F46558BF59FFFDFF74F3B16B5A6196" box="[151,548,979,1005]" pageId="10" pageNumber="161" type="description">
<paragraph id="4A5136D3BF59FFFDFF74F3B16B5A6196" blockId="10.[151,669,942,1005]" box="[151,548,979,1005]" pageId="10" pageNumber="161">
(
<figureCitation id="D2D52A56BF59FFFDFF7CF3B1687C6196" box="[159,258,979,1005]" captionStart="FIGURES 2" captionStartId="8.[151,264,1759,1783]" captionTargetBox="[199,1412,212,1679]" captionTargetId="figure-22@8.[154,1434,179,1739]" captionTargetPageId="8" captionText="FIGURES 2. Indian Isometopinae: Isometopini. Habitus in dorsal view.A. Isometopus cuneatus (Distant, 1904). B. Isometopus mahal Distant, 1911. C. Isometopus webbi, n. sp. D. Isometopus wolskii, n. sp. E. Isometopus yoshizawai Yasunaga and Duwal, 2006. F. Paloniella feana (Distant, 1904)." figureDoi="http://doi.org/10.5281/zenodo.4422882" httpUri="https://zenodo.org/record/4422882/files/figure.png" pageId="10" pageNumber="161">Figs. 2B</figureCitation>
,
<figureCitation id="D2D52A56BF59FFFDFEEEF3B1684C6196" box="[269,306,979,1005]" captionStart="FIGURES 6" captionStartId="23.[151,264,1863,1887]" captionTargetBox="[156,1420,192,1827]" captionTargetId="figure-19@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 6. Indian Isometopinae: Gigantometopini, Isometopini and Myiommini. Head in frontal view.A. Astroscopometopus hesaraghattaensis, n. sp. B. Nymph of Astroscopometopus hesaraghattaensis, n sp. C. Isometopus cuneatus (Distant, 1904). D. Isometopus mahal Distant, 1911. E. Isometopus webbi, n. sp. F. Isometopus wolskii, n. sp. G. Isometopus yoshizawai Yasunaga and Duwal, 2006. H. Paloniella feana (Distant, 1904). I. Paloniella parallela Yasunaga &amp; Hayashi, 2002. J. Isometopidea viraktamathi, n. sp. K. Myiomma belavadii, n. sp. L. Myiomma ramamurthyi, n sp." figureDoi="http://doi.org/10.5281/zenodo.4422892" httpUri="https://zenodo.org/record/4422892/files/figure.png" pageId="10" pageNumber="161">6D</figureCitation>
,
<figureCitation id="D2D52A56BF59FFFDFEDDF3B168356196" box="[318,331,979,1005]" captionStart="FIGURES 9" captionStartId="26.[151,264,1909,1933]" captionTargetBox="[163,1423,193,1874]" captionTargetId="figure-22@26.[151,1436,181,1886]" captionTargetPageId="26" captionText="FIGURES 9. Historical types of Isometopinae Isometopini and Myiommini mentioned in the text. A. Holotype of Isometopus cuneatus (Distant, 1904) in dorsal view. B. Labels. C. Holotype of Isometopus mahal Distant, 1911 in dorsal view. D. Labels. E–G. Holotype of Isometopus feanus Distant, 1904. E. Dorsal view. F. Ventral view. G. Lateral view. H. Label. I–J. Holotype of Alcerocoris globosus Carvalho, 1951 in dorsal view. J. Labels." figureDoi="http://doi.org/10.5281/zenodo.4422900" httpUri="https://zenodo.org/record/4422900/files/figure.png" pageId="10" pageNumber="161">9</figureCitation>
C–D, 10A, 19D–F)
</paragraph>
</subSubSection>
<subSubSection id="02F46558BF59FFFDFF74F4756C7566F0" pageId="10" pageNumber="161" type="reference_group">
<paragraph id="4A5136D3BF59FFFDFF74F475684A6635" blockId="10.[151,1436,1047,1164]" pageId="10" pageNumber="161">
<treatmentCitationGroup id="6AFE11FDBF59FFFDFF74F475684A6635" pageId="10" pageNumber="161">
<treatmentCitation id="CB4F10C2BF59FFFDFF74F475688E6654" author="Distant, W. L." box="[151,496,1047,1071]" page="294" pageId="10" pageNumber="161" year="1911">
<taxonomicName id="8DEE4D50BF59FFFDFF74F475688E6654" ID-CoL="3QNXR" authority="Distant, 1911: 294" authorityName="Distant" authorityPageNumber="294" authorityYear="1911" box="[151,496,1047,1071]" class="Insecta" family="Miridae" genus="Jehania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="mahal">
<emphasis id="789AEAC1BF59FFFDFF74F47568536654" box="[151,301,1047,1071]" italics="true" pageId="10" pageNumber="161">Jehania mahal</emphasis>
<bibRefCitation id="2E7F4B22BF59FFFDFED7F475688E6654" author="Distant, W. L." box="[308,496,1047,1071]" pageId="10" pageNumber="161" refId="ref18092" refString="Distant, W. L. (1911) The Fauna of British India, Including Ceylan and Burma. Rhynchota. Vol. V. Heteroptera, appendix. Taylor &amp; Francis, London, xii + 362 pp. [1910, dating follows Dolling (1991)]" type="book" year="1911">Distant, 1911: 294</bibRefCitation>
</taxonomicName>
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(as new species, DV). Synonymized with
<taxonomicName id="8DEE4D50BF59FFFDFC42F47A6C67664B" ID-CoL="6MZ5S" authority="Mulsant &amp; Rey" authorityName="Mulsant &amp; Rey" authorityYear="1879" box="[929,1305,1047,1072]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="mirificus">
<emphasis id="789AEAC1BF59FFFDFC42F47A6D0B6654" box="[929,1141,1047,1071]" italics="true" pageId="10" pageNumber="161">Isometopus mirificus</emphasis>
Mulsant &amp; Rey
</taxonomicName>
by
<treatmentCitation id="CB4F10C2BF59FFFDFADCF475684E6635" author="Carvalho, J. C. M." page="391" pageId="10" pageNumber="161" year="1951">
<bibRefCitation id="2E7F4B22BF59FFFDFADCF475684E6635" author="Carvalho, J. C. M." pageId="10" pageNumber="161" pagination="381 - 391" refId="ref17892" refString="Carvalho, J. C. M. (1951) New genera and species of Isometopinae in the collection of the British Museum of Natural History (Hemiptera). Anais da Academia Brasilerira de Ciencias, 23 (4), 381 - 391. https: // doi. org / 10.1080 / 00222935108654138" type="journal article" year="1951">Carvalho 1951: 391</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="4A5136D3BF59FFFDFF74F4376C7566F0" blockId="10.[151,1436,1047,1164]" pageId="10" pageNumber="161">
<treatmentCitationGroup id="6AFE11FDBF59FFFDFF74F4376C7566F0" pageId="10" pageNumber="161">
<treatmentCitation id="CB4F10C2BF59FFFDFF74F4376BB86617" author="Akingbohungbe, A. E." box="[151,710,1108,1133]" page="62" pageId="10" pageNumber="161" year="1996">
<taxonomicName id="8DEE4D50BF59FFFDFF74F4376BB86617" ID-CoL="3QNXR" authority=": Akingbohungbe 1996: 62" authorityName="Akingbohungbe" authorityPageNumber="62" authorityYear="1996" box="[151,710,1108,1133]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="mahal" subGenus="Jehania">
<emphasis id="789AEAC1BF59FFFDFF74F43768746617" box="[151,266,1109,1132]" italics="true" pageId="10" pageNumber="161">Isometopus</emphasis>
(
<emphasis id="789AEAC1BF59FFFDFEFAF43668176617" box="[281,361,1108,1132]" italics="true" pageId="10" pageNumber="161">Jehania</emphasis>
)
<emphasis id="789AEAC1BF59FFFDFE9BF43668C96617" box="[376,439,1108,1132]" italics="true" pageId="10" pageNumber="161">mahal</emphasis>
:
<bibRefCitation id="2E7F4B22BF59FFFDFE27F4366BB86617" author="Akingbohungbe, A. E." box="[452,710,1108,1132]" pageId="10" pageNumber="161" refId="ref17705" refString="Akingbohungbe, A. E. (1996) The Isometopinae (Heteroptera: Miridae) of Africa, Europe and Middle East. Delar Tertiary Publisher, Ibadan, 174 pp" type="book" year="1996">Akingbohungbe 1996: 62</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(resurrected from synonymy with
<taxonomicName id="8DEE4D50BF59FFFDFBC4F4376DE86616" ID-CoL="6MZ5S" authorityName="Mulsant &amp; Rey" authorityYear="1879" box="[1063,1174,1109,1133]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="mirificus">
<emphasis id="789AEAC1BF59FFFDFBC4F4376DE86616" box="[1063,1174,1109,1133]" italics="true" pageId="10" pageNumber="161">I. mirificus</emphasis>
</taxonomicName>
); Yasunaga &amp; H a y a s h
<date id="3E501013BF59FFFDFA76F436698066F0" pageId="10" pageNumber="161" value="2002-01">i 2002</date>
: 99, 100 (recorded from
<collectingCountry id="32F97643BF59FFFDFE1AF4166B4B66F0" box="[505,565,1140,1163]" name="Japan" pageId="10" pageNumber="161">Japan</collectingCountry>
, DV);
<bibRefCitation id="2E7F4B22BF59FFFDFD9AF4116B8C66F0" author="Schuh, R. T." box="[633,754,1139,1163]" pageId="10" pageNumber="161" refId="ref19473" refString="Schuh, R. T. (2002 - 2013) On-line Systematic Catalogue of Plant Bugs (Insecta: Heteroptera: Miridae). Available from: http: // research. amnh. org / pbi / catalogue / (accessed 28 October 2020)" type="url" year="2002">Schuh 2002</bibRefCitation>
–2013 (online catalogue, as synonym of
<taxonomicName id="8DEE4D50BF59FFFDFB6EF4166D8266F0" ID-CoL="6MZ5S" authorityName="Mulsant &amp; Rey" authorityYear="1879" box="[1165,1276,1139,1163]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="mirificus">
<emphasis id="789AEAC1BF59FFFDFB6EF4166D8266F0" box="[1165,1276,1139,1163]" italics="true" pageId="10" pageNumber="161">I. mirificus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="02F46558BF59FFFDFF74F4DB6C4167B4" pageId="10" pageNumber="161" type="diagnosis">
<paragraph id="4A5136D3BF59FFFDFF74F4DB6C4167B4" blockId="10.[151,1437,1209,2028]" pageId="10" pageNumber="161">
<emphasis id="789AEAC1BF59FFFDFF74F4DB686C66A8" bold="true" box="[151,274,1209,1235]" pageId="10" pageNumber="161">Diagnosis.</emphasis>
Small, greyish-white species, length
<quantity id="8D169B36BF59FFFDFD56F4DB6A7066AF" box="[693,782,1209,1236]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="161" unit="mm" value="2.2">2.2 mm</quantity>
.
<emphasis id="789AEAC1BF59FFFDFCFAF4DB6ADC66A8" box="[793,930,1209,1235]" italics="true" pageId="10" pageNumber="161">Colouration</emphasis>
: Head shiny with a reddish-brown patch medially on the frons. Ocelli reddish. Antennal segments I and II white dorsally and pale brown ventrally; segment III and IV pale brown. Pronotum with anterior part medially and lateral margins yellowish to pale brown, remainder darker brown. Mesoscutum dark brown, with lateral margins brown. Scutellum pale brown on basal 1/3
<superScript id="BD9B9B9BBF59FFFDFAF8F5476C576748" attach="left" box="[1307,1321,1317,1331]" fontSize="6" pageId="10" pageNumber="161">rd</superScript>
and white toward apex. Clavus near apex pale brown; corium with more than basal half brown; cuneus white; membrane pale brown. Legs with basal 3/4th of fore and mid femur white, apical 1/3
<superScript id="BD9B9B9BBF59FFFDFC7BF50F6AD86700" attach="left" box="[920,934,1389,1403]" fontSize="6" pageId="10" pageNumber="161">rd</superScript>
pale brown; hind femur white with a median pale brown marking. Head shiny, smooth; pronotum and hemelytra having prominent punctures, with semierect setae; cuneus broadly triangular, with prominent cuneal commissure. Detailed measurements: See
<tableCitation id="076C0368BF59FFFDFB06F5D76C4767B4" box="[1253,1337,1461,1487]" captionStart="TABLE 1" captionStartId="4.[151,239,872,896]" captionTargetPageId="4" captionText="TABLE 1. Measurements of Indian Isometopinae (species collected during present work)" httpUri="http://table.plazi.org/id/1E91665BBF57FFF3FF74F30A6D2A61FB" pageId="10" pageNumber="161" tableUuid="1E91665BBF57FFF3FF74F30A6D2A61FB">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="02F46558BF59FFFDFF24F5BB6D5564F8" pageId="10" pageNumber="161" type="materials_examined">
<paragraph id="4A5136D3BF59FFFDFF24F5BB6A1F646C" blockId="10.[151,1437,1209,2028]" pageId="10" pageNumber="161">
<emphasis id="789AEAC1BF59FFFDFF24F5BB68926788" bold="true" box="[199,492,1497,1523]" pageId="10" pageNumber="161">Type material examined</emphasis>
.
<materialsCitation id="FA863C8EBF59FFFDFE1FF5BB6A23646C" ID-GBIF-Occurrence="3043834331" collectionCode="BMNH" collectorName=" Annandale N. " country="India" elevation="122" location="West Bengal" pageId="10" pageNumber="161" specimenCount="1" specimenCount-female="1" stateProvince="West Bengal" typeStatus="syntype">
<collectingCountry id="32F97643BF59FFFDFE1FF5BB6B446788" box="[508,570,1497,1523]" name="India" pageId="10" pageNumber="161">India</collectingCountry>
:
<typeStatus id="95558871BF59FFFDFDA5F5BB6BDC6788" box="[582,674,1497,1523]" pageId="10" pageNumber="161" type="syntype">Syntype</typeStatus>
(
<specimenCount id="5CE8FD5ABF59FFFDFD55F5BB6BB76788" box="[694,713,1497,1523]" pageId="10" pageNumber="161" type="female">♀</specimenCount>
),
<collectingRegion id="882AF831BF59FFFDFD01F5B86A0F6788" box="[738,881,1497,1524]" country="India" name="West Bengal" pageId="10" pageNumber="161">West Bengal</collectingRegion>
(as “
<collectorName id="E71B5305BF59FFFDFC53F5BB6D2F678F" box="[944,1105,1497,1524]" pageId="10" pageNumber="161">E. Himalayas</collectorName>
”),
<collectorName id="E71B5305BF59FFFDFB8CF5BB6D966788" box="[1135,1256,1497,1523]" pageId="10" pageNumber="161">Darjeeling</collectorName>
,
<collectorName id="E71B5305BF59FFFDFB16F5B86C1B678F" box="[1269,1381,1498,1524]" pageId="10" pageNumber="161">Kurseong</collectorName>
,
<quantity id="8D169B36BF59FFFDFA91F5BB69BA646C" metricMagnitude="2" metricUnit="m" metricValue="1.2192" pageId="10" pageNumber="161" unit="ft" value="400.0">
<elevation id="C1C3D1E0BF59FFFDFA91F5BB69BA646C" metricMagnitude="2" metricUnit="m" metricValue="1.2192" pageId="10" pageNumber="161" unit="ft" value="400.0">400 feet</elevation>
</quantity>
, among lichens on tree-trunk,
<collectorName id="E71B5305BF59FFFDFDFBF59F6B926463" box="[536,748,1533,1560]" pageId="10" pageNumber="161">
<emphasis id="789AEAC1BF59FFFDFDFBF59F6BCD646C" box="[536,691,1533,1559]" italics="true" pageId="10" pageNumber="161">Annandale N.</emphasis>
Leg.
</collectorName>
(
<collectionCode id="2CFFAE16BF59FFFDFD19F59F6A2A646C" box="[762,852,1533,1559]" pageId="10" pageNumber="161">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="4A5136D3BF59FFFDFF24F6436D5564F8" blockId="10.[151,1437,1209,2028]" pageId="10" pageNumber="161">
<emphasis id="789AEAC1BF59FFFDFF24F6436B756440" bold="true" box="[199,523,1569,1595]" pageId="10" pageNumber="161">Other specimens examined</emphasis>
:
<materialsCitation id="FA863C8EBF59FFFDFDFFF64069926424" ID-GBIF-Occurrence="3043834315" collectingDate="2011-07-21" collectionCode="GKVK" collectorName="Yeshwanth, H. M." country="India" location="Bangalore" pageId="10" pageNumber="161" specimenCount="2" specimenCount-female="2" stateProvince="Karnataka">
<subScript id="D66A3496BF59FFFDFDFFF6406B286442" attach="left" box="[540,598,1570,1596]" fontSize="7" pageId="10" pageNumber="161">
<collectingCountry id="32F97643BF59FFFDFDFFF6406B286442" box="[540,598,1570,1596]" name="India" pageId="10" pageNumber="161">InDIa</collectingCountry>
</subScript>
:
<specimenCount id="5CE8FD5ABF59FFFDFD85F6436BF56440" box="[614,651,1569,1595]" pageId="10" pageNumber="161" type="female">2♀</specimenCount>
,
<collectingRegion id="882AF831BF59FFFDFD7AF6436A716440" box="[665,783,1569,1595]" country="India" name="Karnataka" pageId="10" pageNumber="161">Karnataka</collectingRegion>
,
<location id="4F316008BF59FFFDFCF8F6436AEF6440" LSID="urn:lsid:plazi:treatment:C24787C5BF59FFFCFF74F3CD6847633A:4F316008BF59FFFDFCF8F6436AEF6440" box="[795,913,1569,1595]" country="India" name="Bangalore" pageId="10" pageNumber="161" stateProvince="Karnataka">Bangalore</location>
,
<collectionCode id="2CFFAE16BF59FFFDFC7EF6436A8F6440" box="[925,1009,1569,1595]" pageId="10" pageNumber="161">GKVK</collectionCode>
,
<date id="3E501013BF59FFFDFC1DF6436D016440" box="[1022,1151,1569,1595]" pageId="10" pageNumber="161" value="2011-07-21">
<collectingDate id="2E14E9FBBF59FFFDFC1DF6436D016440" box="[1022,1151,1569,1595]" pageId="10" pageNumber="161" value="2011-07-21">21.vii.2011</collectingDate>
</date>
, ex: bark,
<collectorName id="E71B5305BF59FFFDFB15F64369C9641B" pageId="10" pageNumber="161">Yeshwanth, H. M.</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="FA863C8EBF59FFFDFF1AF6276C126424" ID-GBIF-Occurrence="3043834329" box="[249,1388,1605,1632]" collectingDate="2015-07-18" collectionCode="UASB" collectorName="Yeshwanth, H. M." country="InDIa" location="Nandi Hills" municipality="Chikkaballapura" pageId="10" pageNumber="161" specimenCount="2" specimenCount-female="2">
<specimenCount id="5CE8FD5ABF59FFFDFF1AF62768616424" box="[249,287,1605,1631]" pageId="10" pageNumber="161" type="female">2♀</specimenCount>
:
<collectingMunicipality id="AA35ACA9BF59FFFDFECFF62768966424" box="[300,488,1605,1631]" pageId="10" pageNumber="161">Chikkaballapura</collectingMunicipality>
,
<location id="4F316008BF59FFFDFE17F6276B096424" LSID="urn:lsid:plazi:treatment:C24787C5BF59FFFCFF74F3CD6847633A:4F316008BF59FFFDFE17F6276B096424" box="[500,631,1605,1631]" country="InDIa" municipality="Chikkaballapura" name="Nandi Hills" pageId="10" pageNumber="161">Nandi Hills</location>
,
<date id="3E501013BF59FFFDFD64F6276A77641B" box="[647,777,1605,1632]" pageId="10" pageNumber="161" value="2015-07-18">
<collectingDate id="2E14E9FBBF59FFFDFD64F6276A77641B" box="[647,777,1605,1632]" pageId="10" pageNumber="161" value="2015-07-18">18.vii.2015</collectingDate>
</date>
, ex:
<taxonomicName id="8DEE4D50BF59FFFDFCA3F6246AED641B" box="[832,915,1606,1632]" class="Magnoliopsida" family="Malvaceae" genus="Grewia" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="genus">
<emphasis id="789AEAC1BF59FFFDFCA3F6246AED641B" box="[832,915,1606,1632]" italics="true" pageId="10" pageNumber="161">Grewia</emphasis>
</taxonomicName>
sp. bark,
<collectorName id="E71B5305BF59FFFDFBE7F6276DAC641B" box="[1028,1234,1605,1632]" pageId="10" pageNumber="161">Yeshwanth, H. M.</collectorName>
leg. (
<collectionCode id="2CFFAE16BF59FFFDFAF7F6276C1B6424" box="[1300,1381,1605,1631]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="10" pageNumber="161">UASB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="FA863C8EBF59FFFDFA9AF6276D5964F8" ID-GBIF-Occurrence="3043834327" collectingDate="2020-01-23" collectionCode="GKVK" collectorName="Yeshwanth, H. M." country="InDIa" location="Bengaluru" pageId="10" pageNumber="161" specimenCount="6" specimenCount-female="6">
<specimenCount id="5CE8FD5ABF59FFFDFA9AF6276CE26424" box="[1401,1436,1605,1631]" pageId="10" pageNumber="161" type="female">6♀</specimenCount>
<location id="4F316008BF59FFFDFF74F60B687164F8" LSID="urn:lsid:plazi:treatment:C24787C5BF59FFFCFF74F3CD6847633A:4F316008BF59FFFDFF74F60B687164F8" box="[151,271,1641,1667]" country="InDIa" name="Bengaluru" pageId="10" pageNumber="161">Bengaluru</location>
,
<collectionCode id="2CFFAE16BF59FFFDFEF9F60B681064F8" box="[282,366,1641,1667]" pageId="10" pageNumber="161">GKVK</collectionCode>
,
<date id="3E501013BF59FFFDFE9AF60B689B64F8" box="[377,485,1641,1667]" pageId="10" pageNumber="161" value="2020-01-23">
<collectingDate id="2E14E9FBBF59FFFDFE9AF60B689B64F8" box="[377,485,1641,1667]" pageId="10" pageNumber="161" value="2020-01-23">23.i.2020</collectingDate>
</date>
,
<collectorName id="E71B5305BF59FFFDFE0CF60B6BC564FF" box="[495,699,1641,1668]" pageId="10" pageNumber="161">Yeshwanth, H. M.</collectorName>
leg. ex:
<taxonomicName id="8DEE4D50BF59FFFDFCF8F6086A9564F8" box="[795,1003,1642,1667]" class="Magnoliopsida" family="Fabaceae" genus="Pongamia" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="species" species="pinnata">
<emphasis id="789AEAC1BF59FFFDFCF8F6086A9564F8" box="[795,1003,1642,1667]" italics="true" pageId="10" pageNumber="161">Pongamia pinnata</emphasis>
</taxonomicName>
bark
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="02F46558BF59FFFDFF24F6EF6B8764D3" box="[199,761,1677,1704]" pageId="10" pageNumber="161" type="distribution">
<paragraph id="4A5136D3BF59FFFDFF24F6EF6B8764D3" blockId="10.[151,1437,1209,2028]" box="[199,761,1677,1704]" pageId="10" pageNumber="161">
<emphasis id="789AEAC1BF59FFFDFF24F6EF682764DC" bold="true" box="[199,345,1677,1703]" pageId="10" pageNumber="161">Distribution</emphasis>
.
<collectingCountry id="32F97643BF59FFFDFE84F6EF68DF64DC" box="[359,417,1677,1703]" name="India" pageId="10" pageNumber="161">India</collectingCountry>
(
<collectingRegion id="882AF831BF59FFFDFE4CF6EF6B5A64DC" box="[431,548,1677,1703]" country="India" name="Karnataka" pageId="10" pageNumber="161">Karnataka</collectingRegion>
and
<collectingRegion id="882AF831BF59FFFDFDB9F6EC6B9364D3" box="[602,749,1677,1704]" country="India" name="West Bengal" pageId="10" pageNumber="161">West Bengal</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="02F46558BF59FFFDFF24F6D36BF26494" pageId="10" pageNumber="161" type="biology_ecology">
<paragraph id="4A5136D3BF59FFFDFF24F6D36BF26494" blockId="10.[151,1437,1209,2028]" pageId="10" pageNumber="161">
<emphasis id="789AEAC1BF59FFFDFF24F6D3685F64B0" bold="true" box="[199,289,1713,1739]" pageId="10" pageNumber="161">Biology</emphasis>
. Nymphs and adults were collected on
<taxonomicName id="8DEE4D50BF59FFFDFD39F6D06A5364B7" box="[730,813,1714,1740]" class="Magnoliopsida" family="Malvaceae" genus="Grewia" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="genus">
<emphasis id="789AEAC1BF59FFFDFD39F6D06A5364B7" box="[730,813,1714,1740]" italics="true" pageId="10" pageNumber="161">Grewia</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="8DEE4D50BF59FFFDFC81F6D36AA164B7" box="[866,991,1713,1740]" class="Magnoliopsida" family="Malvaceae" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="family">Malvaceae</taxonomicName>
) and
<taxonomicName id="8DEE4D50BF59FFFDFBFFF6D06D9264B0" box="[1052,1260,1714,1739]" class="Magnoliopsida" family="Fabaceae" genus="Pongamia" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="species" species="pinnata">
<emphasis id="789AEAC1BF59FFFDFBFFF6D06D9264B0" box="[1052,1260,1714,1739]" italics="true" pageId="10" pageNumber="161">Pongamia pinnata</emphasis>
</taxonomicName>
(L.) Pierre (
<taxonomicName id="8DEE4D50BF59FFFDFA96F6D36996648B" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
), with lichens on bark (
<figureCitation id="D2D52A56BF59FFFDFE0DF6B76B366494" box="[494,584,1749,1775]" captionStart="FIGURES 19" captionStartId="36.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-22@36.[151,1436,181,1894]" captionTargetPageId="36" captionText="FIGURES 19. Indian Isometopinae Isometopini alive. A–C. Isometopus yoshizawai Yasunaga and Duwal, 2006. D–F. Nymph (D) and adults (D, F) of Isometopus mahal Distant, 1911." figureDoi="http://doi.org/10.5281/zenodo.4422929" httpUri="https://zenodo.org/record/4422929/files/figure.png" pageId="10" pageNumber="161">Figs. 19</figureCitation>
D–F).
</paragraph>
</subSubSection>
<subSubSection id="02F46558BF59FFFCFF24F69B6847633A" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161" type="discussion">
<paragraph id="4A5136D3BF59FFFCFF24F69B6847633A" blockId="10.[151,1437,1209,2028]" lastBlockId="11.[151,1437,151,321]" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161">
<emphasis id="789AEAC1BF59FFFDFF24F69B683B6568" bold="true" box="[199,325,1785,1811]" pageId="10" pageNumber="161">Discussion</emphasis>
. Although the habitus and, particularly, the colour pattern of both species are similar,
<taxonomicName id="8DEE4D50BF59FFFDFAEBF6986879654C" authority="Distant, 1911" authorityName="Distant" authorityYear="1911" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="mahal">
<emphasis id="789AEAC1BF59FFFDFAEBF6986C1B6568" box="[1288,1381,1785,1811]" italics="true" pageId="10" pageNumber="161">I. mahal</emphasis>
<bibRefCitation id="2E7F4B22BF59FFFDFA8FF69B6879654C" author="Distant, W. L." pageId="10" pageNumber="161" refId="ref18092" refString="Distant, W. L. (1911) The Fauna of British India, Including Ceylan and Burma. Rhynchota. Vol. V. Heteroptera, appendix. Taylor &amp; Francis, London, xii + 362 pp. [1910, dating follows Dolling (1991)]" type="book" year="1911">Distant, 1911</bibRefCitation>
</taxonomicName>
differs from
<taxonomicName id="8DEE4D50BF59FFFDFE7EF77C6A6C654C" authority="Mulsant &amp; Rey, 1872" authorityName="Mulsant &amp; Rey" authorityYear="1872" box="[413,786,1821,1848]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="mirificus">
<emphasis id="789AEAC1BF59FFFDFE7EF77C6B67654C" box="[413,537,1821,1847]" italics="true" pageId="10" pageNumber="161">I. mirificus</emphasis>
Mulsant &amp; Rey, 1872
</taxonomicName>
by the absence of a short gula in females and the vertical head in frontal view ––two states “typical” of the subgenus
<taxonomicName id="8DEE4D50BF59FFFDFCD4F7236AEE6520" authorityName="Distant" authorityYear="1910" box="[823,912,1857,1883]" class="Insecta" family="Miridae" genus="Jehania" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="genus">
<emphasis id="789AEAC1BF59FFFDFCD4F7236AEE6520" box="[823,912,1857,1883]" italics="true" pageId="10" pageNumber="161">Jehania</emphasis>
</taxonomicName>
, according to
<bibRefCitation id="2E7F4B22BF59FFFDFBCCF7236C466527" author="Akingbohungbe, A. E." box="[1071,1336,1857,1884]" pageId="10" pageNumber="161" refId="ref17705" refString="Akingbohungbe, A. E. (1996) The Isometopinae (Heteroptera: Miridae) of Africa, Europe and Middle East. Delar Tertiary Publisher, Ibadan, 174 pp" type="book" year="1996">Akingbohungbe (1996)</bibRefCitation>
, but also by the less strongly explanate and arcuate pronotal lateral margins and the rounded anterior angles of pronotum, barely produced anteriorly and never partially surrounding eyes laterally. The non-type Indian specimens agrees with
<taxonomicName id="8DEE4D50BF59FFFDFF32F7CC685165BC" authorityName="Distant" authorityYear="1911" box="[209,303,1965,1991]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="mahal">
<emphasis id="789AEAC1BF59FFFDFF32F7CC685165BC" box="[209,303,1965,1991]" italics="true" pageId="10" pageNumber="161">I. mahal</emphasis>
</taxonomicName>
in these character states but have a different pronotal and hemelytral color pattern than the drawing of
<taxonomicName id="8DEE4D50BF59FFFDFF54F7B068646590" authorityName="Distant" authorityYear="1911" box="[183,282,2001,2027]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="mahal">
<emphasis id="789AEAC1BF59FFFDFF54F7B068646590" box="[183,282,2001,2027]" italics="true" pageId="10" pageNumber="161">J. mahal</emphasis>
</taxonomicName>
by
<bibRefCitation id="2E7F4B22BF59FFFDFEA4F7B368946590" author="Distant, W. L." box="[327,490,2001,2027]" pageId="10" pageNumber="161" refId="ref18092" refString="Distant, W. L. (1911) The Fauna of British India, Including Ceylan and Burma. Rhynchota. Vol. V. Heteroptera, appendix. Taylor &amp; Francis, London, xii + 362 pp. [1910, dating follows Dolling (1991)]" type="book" year="1911">Distant (1911)</bibRefCitation>
and photographs of
<taxonomicName id="8DEE4D50BF59FFFDFD3BF7B06A466590" authorityName="Distant" authorityYear="1911" box="[728,824,2001,2027]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="mahal">
<emphasis id="789AEAC1BF59FFFDFD3BF7B06A466590" box="[728,824,2001,2027]" italics="true" pageId="10" pageNumber="161">I. mahal</emphasis>
</taxonomicName>
from
<collectingCountry id="32F97643BF59FFFDFC63F7B06ABE6597" box="[896,960,2002,2028]" name="Japan" pageId="10" pageNumber="161">Japan</collectingCountry>
by
<bibRefCitation id="2E7F4B22BF59FFFDFC0FF7B06C4C6590" author="Yasunaga, T. &amp; Hayashi, M." box="[1004,1330,2001,2028]" pageId="10" pageNumber="161" pagination="95 - 101" refId="ref20082" refString="Yasunaga, T. &amp; Hayashi, M. (2002) New or little known isometopine plant bugs from Japan (Heteroptera: Miridae). Tijdschrift voor Entomologie, 145, 95 - 101. https: // doi. org / 10.1163 / 22119434 - 900000103" type="journal article" year="2002">Yasunaga &amp; Hayashi (2002)</bibRefCitation>
. The sig-nificance of such differences remains difficult to interpret. It could be only intraspecific variability. However, we cannot exclude the presence of multiple, closely related species that are impossible to clearly separate based on the material available to us. The relationships between
<taxonomicName id="8DEE4D50BF58FFFCFD36F0BD6A4C6282" authorityName="Distant" authorityYear="1911" box="[725,818,223,249]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="mahal">
<emphasis id="789AEAC1BF58FFFCFD36F0BD6A4C6282" box="[725,818,223,249]" italics="true" pageId="11" pageNumber="162">I. mahal</emphasis>
</taxonomicName>
and
<taxonomicName id="8DEE4D50BF58FFFCFC8BF0BD6A9A6282" authorityName="Mulsant &amp; Rey" authorityYear="1872" box="[872,996,223,249]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="mirificus">
<emphasis id="789AEAC1BF58FFFCFC8BF0BD6A9A6282" box="[872,996,223,249]" italics="true" pageId="11" pageNumber="162">I. mirificus</emphasis>
</taxonomicName>
(particularly if the differences between both sexes of European
<taxonomicName id="8DEE4D50BF58FFFCFE46F1616B5F6366" authorityName="Mulsant &amp; Rey" authorityYear="1872" box="[421,545,259,285]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="mirificus">
<emphasis id="789AEAC1BF58FFFCFE46F1616B5F6366" box="[421,545,259,285]" italics="true" pageId="11" pageNumber="162">I. mirificus</emphasis>
</taxonomicName>
are considered) and the taxonomic validity of the subgenus
<taxonomicName id="8DEE4D50BF58FFFCFB24F1616C5E6366" authorityName="Distant" authorityYear="1910" box="[1223,1312,259,285]" class="Insecta" family="Miridae" genus="Jehania" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="789AEAC1BF58FFFCFB24F1616C5E6366" box="[1223,1312,259,285]" italics="true" pageId="11" pageNumber="162">Jehania</emphasis>
</taxonomicName>
need more detailed study.
</paragraph>
</subSubSection>
</treatment>
</document>
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