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<document id="35A11EF94F9BC7103A1B5E4BBC267DD3" ID-CLB-Dataset="54447" ID-DOI="10.5852/ejt.2023.860.2049" ID-GBIF-Dataset="f3024132-3d29-4ccd-a73d-4378b96c9147" ID-ISSN="2118-9773" ID-Zenodo-Dep="7674396" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677245384703" checkinUser="felipe" docAuthor="Jafari, Farzaneh, Keshavarzi, Maryam, Doostmohammadi, Moslem &amp; Mirtadzadini, Mansour" docDate="2023" docId="BB3C87C0FFB7C464FE6DF969FB805FE3" docLanguage="en" docName="ejt-860_42-61.pdf" docOrigin="European Journal of Taxonomy 860" docSource="http://dx.doi.org/10.5852/ejt.2023.860.2049" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Silene penduliflora F. Jafari, Keshavarzi &amp; Doostm. 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="55" masterDocId="4705FFB8FFBDC469FFD4FF61FF995A0B" masterDocTitle="Two new chasmophytic species of Silene (Caryophyllaceae, sect. Siphonomorpha) from Iran" masterLastPageNumber="61" masterPageNumber="42" pageNumber="52" updateTime="1699475790762" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A68EF0B4642BDAA12124DED166D4596D">Two new chasmophytic species of Silene (Caryophyllaceae, sect. Siphonomorpha) from Iran</mods:title>
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<mods:namePart id="46FB9A3B800A6232DCC9A20DB7CE4024">Jafari, Farzaneh</mods:namePart>
<mods:affiliation id="A62DAC98F7016FFF57D96E75C0562740">Department of Plant Sciences, Faculty of Biological Sciences, Alzahra University, Vanak, 1993893973 Tehran, Iran.</mods:affiliation>
<mods:nameIdentifier id="AD1C6652DE1CE23B7D305567AF78CD44" type="email">f.jafari@alzahra.ac.ir</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="0D23B3B3455037B8C992B003D79831B5">Keshavarzi, Maryam</mods:namePart>
<mods:affiliation id="25247BAD0DF3CB22DF255A7D7835D844">Department of Plant Sciences, Faculty of Biological Sciences, Alzahra University, Vanak, 1993893973 Tehran, Iran.</mods:affiliation>
<mods:nameIdentifier id="C27F17AD3CE82774424B866D67C2C3F8" type="email">m.keshavarzi@alzahra.ac.ir</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="D31AFEE9A2FD30AFB8B8D9784D606E2A">Doostmohammadi, Moslem</mods:namePart>
<mods:affiliation id="D8FB6C5AC3160A16FCEFA1C4D9BF332A">Department of Biology, Faculty of Science, Shahid Bahonar University of Kerman, P. O. Box 76169 - 133, Kerman, Iran.</mods:affiliation>
<mods:nameIdentifier id="83E447EF0CF91102B64B09B66B13F275" type="email">doost@sci.uk.ac.ir</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="F5A4F836553FD71AD9FEAF4024BAD072">Author</mods:roleTerm>
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<mods:namePart id="FBD4E08194506701B85A1128A4125E78">Mirtadzadini, Mansour</mods:namePart>
<mods:affiliation id="09D2C8D126A093A9FC33598A9EDF9F05">Department of Biology, Faculty of Science, Shahid Bahonar University of Kerman, P. O. Box 76169 - 133, Kerman, Iran.</mods:affiliation>
<mods:nameIdentifier id="BAF6934ACBA3C6648CE8169FF2A04429" type="email">mirtadz@uk.ac.ir</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="BE469EF5F422BC56D45862599B8149F6">text</mods:typeOfResource>
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<mods:title id="0E07DFBE2E737A7736068D102689DFD5">European Journal of Taxonomy</mods:title>
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<mods:part id="F02A453990A5D7CEE0F36A6AF64E681E">
<mods:date id="0F59073EAC2BEF902403E35AE276028F">2023</mods:date>
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<mods:number id="2EA447137C12D41EF218C66155350E4D">2023-02-20</mods:number>
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<treatment id="BB3C87C0FFB7C464FE6DF969FB805FE3" ID-DOI="http://doi.org/10.5281/zenodo.7689456" ID-GBIF-Taxon="209384192" ID-Zenodo-Dep="7689456" LSID="urn:lsid:plazi:treatment:BB3C87C0FFB7C464FE6DF969FB805FE3" httpUri="http://treatment.plazi.org/id/BB3C87C0FFB7C464FE6DF969FB805FE3" lastPageId="13" lastPageNumber="55" pageId="10" pageNumber="52">
<subSubSection id="7B8F655DFFB7C463FE6DF969FC5C5C67" pageId="10" pageNumber="52" type="nomenclature">
<paragraph id="332A36D6FFB7C463FE6DF969FBE25C28" blockId="10.[441,1147,1544,1644]" box="[441,1147,1544,1571]" pageId="10" pageNumber="52">
<heading id="686281BAFFB7C463FE6DF969FBE25C28" box="[441,1147,1544,1571]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="52" reason="2">
<taxonomicName id="F4954D55FFB7C463FE6DF969FB835C28" ID-CoL="B6D5P" authority="F.Jafari, Keshavarzi &amp; Doostm." authorityName="F. Jafari, Keshavarzi &amp; Doostm." authorityYear="2023" box="[441,1050,1544,1571]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="52" phylum="Tracheophyta" rank="species" species="penduliflora" status="sp. nov.">
<emphasis id="01E1EAC4FFB7C463FE6DF969FD065C28" bold="true" box="[441,671,1544,1571]" italics="true" pageId="10" pageNumber="52">Silene penduliflora</emphasis>
F.Jafari, Keshavarzi &amp; Doostm.
</taxonomicName>
<taxonomicNameLabel id="1AD257BFFFB7C463FBF5F968FBE25C28" box="[1057,1147,1545,1571]" pageId="10" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="332A36D6FFB7C463FD93F94CFC745C4C" blockId="10.[441,1147,1544,1644]" box="[583,1005,1581,1607]" pageId="10" pageNumber="52">urn:lsid:ipni.org:names:77313984-1</paragraph>
<paragraph id="332A36D6FFB7C463FDBBF933FC5C5C67" blockId="10.[441,1147,1544,1644]" box="[623,965,1618,1644]" pageId="10" pageNumber="52">
<figureCitation id="ABAE2A53FFB7C463FDBBF933FD4E5C67" box="[623,727,1618,1644]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="3.[189,232,1856,1882]" captionStartId-1="4.[189,232,1855,1881]" captionStartId-2="5.[189,232,1892,1918]" captionTargetBox-0="[189,1398,537,1814]" captionTargetBox-1="[189,1399,546,1812]" captionTargetBox-2="[222,1365,957,1853]" captionTargetId-0="figure-103@3.[189,1399,537,1815]" captionTargetId-1="figure-78@4.[189,1399,546,1813]" captionTargetId-2="figure-284@5.[219,1369,955,1854]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Fig. 1. 50% majority-rule consensus tree obtained from the Bayesian inference analysis of the nrDNA ITS sequences in selected species of Silene L. Posterior probabilities (PP) ≥ 0.70 are shown above the branches and Maximum likelihood bootstrap (MLB) values ≥ 70% below. Accessions newly sequenced are indicated by a star. Accessions of S. ruprechtii Schischk. (MN420835 and MK559501) are misidentified by Bahmani et al. (2020)." captionText-1="Fig. 2. 50% majority-rule consensus tree obtained from the Bayesian inference analysis of the cpDNA rps16 sequences in selected species of Silene L. Posterior probabilities (PP) ≥ 0.70 are shown above the branches and Maximum likelihood bootstrap (MLB) values ≥ 70% below. Accessions newly sequenced are indicated by a star. Accessions of S. ruprechtii Schischk. (MN460318 and MN460319) are misidentified by Bahmani et al. (2020)." captionText-2="Fig. 3. Distribution map of S. ghahremaninejadii Hoseini &amp; Assadi (blue dot), S. parrowiana Boiss. &amp; Hausskn. (green dots),S. penduliflora F.Jafari, Keshavarzi &amp; Doostm. sp. nov. (red dots),S. shahrudensis Rech.f. (black dot), and S. thyrsiantha F.Jafari, Mirtadz. &amp; Keshavarzi sp. nov. (purple dot). The map was made with QGIS ver. 3.22.9." figureDoi-0="http://doi.org/10.5281/zenodo.7674398" figureDoi-1="http://doi.org/10.5281/zenodo.7674400" figureDoi-2="http://doi.org/10.5281/zenodo.7674402" httpUri-0="https://zenodo.org/record/7674398/files/figure.png" httpUri-1="https://zenodo.org/record/7674400/files/figure.png" httpUri-2="https://zenodo.org/record/7674402/files/figure.png" pageId="10" pageNumber="52">Figs 1–3</figureCitation>
,
<figureCitation id="ABAE2A53FFB7C463FD30F933FCBE5C67" box="[740,807,1618,1644]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1395,980,1845]" captionTargetId="figure-285@9.[189,1399,980,1845]" captionTargetPageId="9" captionText="Fig. 6. A–C. Silene thyrsiantha F.Jafari, Mirtadz. &amp; Keshavarzi sp. nov. (M. Mirtadzadini 2089; MIR!). A. Seed. B. Calyx. C. Petal. – D–F. Silene penduliflora F.Jafari, Keshavarzi &amp; Doostm. sp. nov. D. Seed (M. Doostmohammadi &amp; A. Ghorbanalizadeh 4054; MIR!). E. Calyx. F. Petal (M. Doostmohammadi 4056; MIR!). Seeds were scanned with a Dino-Lite digital microscope AM413T model." figureDoi="http://doi.org/10.5281/zenodo.7674408" httpUri="https://zenodo.org/record/7674408/files/figure.png" pageId="10" pageNumber="52">6D‒F</figureCitation>
,
<figureCitation id="ABAE2A53FFB7C463FCE7F933FCFB5C67" box="[819,866,1618,1644]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="11.[189,232,1960,1986]" captionStartId-1="12.[189,232,1928,1954]" captionTargetBox-0="[209,1379,265,1931]" captionTargetBox-1="[189,1399,906,1887]" captionTargetId-0="figure-35@11.[209,1379,265,1931]" captionTargetId-1="figure-186@12.[189,1399,906,1887]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="Fig. 7. Silene penduliflora F.Jafari, Keshavarzi &amp; Doostm. sp. nov. (holotype, M. Mirtadzadini 2076; MIR!)." captionText-1="Fig. 8. Silene penduliflora F.Jafari, Keshavarzi &amp; Doostm. sp. nov. (paratypes, M. Doostmohammadi &amp; A. Ghorbanalizadeh 4054; MIR!. M. Doostmohammadi 4056; MIR!). A‒B. Inflorescence at flowering and fruiting times. C. Basal leaves and caudex. D. Habit and habitat. Photos by: M. Doostmohammadi." figureDoi-0="http://doi.org/10.5281/zenodo.7674410" figureDoi-1="http://doi.org/10.5281/zenodo.7674412" httpUri-0="https://zenodo.org/record/7674410/files/figure.png" httpUri-1="https://zenodo.org/record/7674412/files/figure.png" pageId="10" pageNumber="52">7–8</figureCitation>
;
<tableCitation id="7E17036DFFB7C463FCBAF933FC5C5C67" box="[878,965,1618,1644]" captionStart="Table 1" captionStartId="6.[189,258,265,291]" captionTargetPageId="6" captionText="Table 1. Morphological comparison among S. penduliflora F.Jafari, Keshavarzi &amp; Doostm. sp. nov., S. thyrsiantha F.Jafari, Mirtadz. &amp; Keshavarzi sp. nov. and close relatives. Distinctive character states are shown in bold." httpUri="http://table.plazi.org/id/67EA665EFFBBC46FFF69FE68FE0E5B62" pageId="10" pageNumber="52" tableUuid="67EA665EFFBBC46FFF69FE68FE0E5B62">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="7B8F655DFFB7C463FF69F9F4FE7D5D5B" pageId="10" pageNumber="52" type="diagnosis">
<paragraph id="332A36D6FFB7C463FF69F9F4FEAF5CA4" blockId="10.[189,310,1685,1711]" box="[189,310,1685,1711]" pageId="10" pageNumber="52">
<heading id="686281BAFFB7C463FF69F9F4FEAF5CA4" bold="true" box="[189,310,1685,1711]" fontSize="11" level="3" pageId="10" pageNumber="52" reason="3">
<emphasis id="01E1EAC4FFB7C463FF69F9F4FEAF5CA4" bold="true" box="[189,310,1685,1711]" pageId="10" pageNumber="52">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="332A36D6FFB7C463FF69F9A7FE7D5D5B" blockId="10.[189,1398,1734,1872]" pageId="10" pageNumber="52">
<taxonomicName id="F4954D55FFB7C463FF69F9A7FC875CEA" authority="F.Jafari, Keshavarzi &amp; Doostm." authorityName="F. Jafari, Keshavarzi &amp; Doostm." authorityYear="2023" box="[189,798,1734,1761]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="52" phylum="Tracheophyta" rank="species" species="penduliflora" status="sp. nov.">
<emphasis id="01E1EAC4FFB7C463FF69F9A7FE055CEB" box="[189,412,1734,1760]" italics="true" pageId="10" pageNumber="52">Silene penduliflora</emphasis>
F.Jafari, Keshavarzi &amp; Doostm.
</taxonomicName>
,
<taxonomicNameLabel id="1AD257BFFFB7C463FCFFF9A6FC1E5CEA" box="[811,903,1735,1761]" pageId="10" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="F4954D55FFB7C463FBF2F9A7FB465CEB" box="[1062,1247,1734,1760]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="52" phylum="Tracheophyta" rank="species" species="shahrudensis">
<emphasis id="01E1EAC4FFB7C463FBF2F9A7FB465CEB" box="[1062,1247,1734,1760]" italics="true" pageId="10" pageNumber="52">S. shahrudensis</emphasis>
</taxonomicName>
in basal leaf size (11‒60×
<quantity id="F46D9B33FFB7C463FE88F98AFE4D5D0D" box="[348,468,1771,1798]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="10" pageNumber="52" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5‒4 mm</quantity>
vs 15‒35×
<quantity id="F46D9B33FFB7C463FD8AF98AFD405D0D" box="[606,729,1771,1798]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="6.0" metricValueMin="2.5" pageId="10" pageNumber="52" unit="mm" value="4.25" valueMax="6.0" valueMin="2.5">2.5‒6 mm</quantity>
), calyx length (
<quantity id="F46D9B33FFB7C463FC5AF98AFC675D0D" box="[910,1022,1771,1798]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="10" pageNumber="52" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9‒11 mm</quantity>
vs
<quantity id="F46D9B33FFB7C463FBF3F98AFB025D0D" box="[1063,1179,1771,1798]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="10" pageNumber="52" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8‒10 mm</quantity>
), petal limb length (
<quantity id="F46D9B33FFB7C463FF12F871FEB05D20" box="[198,297,1808,1835]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="10" pageNumber="52" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5‒6 mm</quantity>
vs ca
<quantity id="F46D9B33FFB7C463FEA3F871FE595D20" box="[375,448,1808,1835]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="52" unit="mm" value="5.0">5 mm</quantity>
), claw indumentum (glabrous vs ciliate), and capsule size (7.5‒8.5 ×
<quantity id="F46D9B33FFB7C463FB25F871FACD5D20" box="[1265,1364,1808,1835]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="10" pageNumber="52" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4‒5 mm</quantity>
vs 4‒6 ×
<quantity id="F46D9B33FFB7C463FED5F857FEFD5D5B" box="[257,356,1846,1872]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="10" pageNumber="52" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3‒4 mm</quantity>
) (
<tableCitation id="7E17036DFFB7C463FEAFF854FE4A5D5B" box="[379,467,1845,1872]" captionStart="Table 1" captionStartId="6.[189,258,265,291]" captionTargetPageId="6" captionText="Table 1. Morphological comparison among S. penduliflora F.Jafari, Keshavarzi &amp; Doostm. sp. nov., S. thyrsiantha F.Jafari, Mirtadz. &amp; Keshavarzi sp. nov. and close relatives. Distinctive character states are shown in bold." httpUri="http://table.plazi.org/id/67EA665EFFBBC46FFF69FE68FE0E5B62" pageId="10" pageNumber="52" tableUuid="67EA665EFFBBC46FFF69FE68FE0E5B62">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7B8F655DFFB7C463FF69F818FE805DE3" pageId="10" pageNumber="52" type="etymology">
<paragraph id="332A36D6FFB7C463FF69F818FEDD5D98" blockId="10.[189,324,1913,1939]" box="[189,324,1913,1939]" pageId="10" pageNumber="52">
<heading id="686281BAFFB7C463FF69F818FEDD5D98" bold="true" box="[189,324,1913,1939]" fontSize="11" level="3" pageId="10" pageNumber="52" reason="3">
<emphasis id="01E1EAC4FFB7C463FF69F818FEDD5D98" bold="true" box="[189,324,1913,1939]" pageId="10" pageNumber="52">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="332A36D6FFB7C463FF69F8C8FE805DE3" blockId="10.[189,1398,1961,2024]" pageId="10" pageNumber="52">The specific epithet refers to the status of the flowers: flowers are pendulous at flowering time in this species.</paragraph>
</subSubSection>
<caption id="67EA665EFFB6C462FF69F8C9FE965DED" ID-DOI="http://doi.org/10.5281/zenodo.7674410" ID-Zenodo-Dep="7674410" httpUri="https://zenodo.org/record/7674410/files/figure.png" pageId="11" pageNumber="53" startId="11.[189,232,1960,1986]" targetBox="[209,1379,265,1931]" targetPageId="11">
<paragraph id="332A36D6FFB6C462FF69F8C9FE965DED" blockId="11.[189,1398,1960,2022]" pageId="11" pageNumber="53">
<emphasis id="01E1EAC4FFB6C462FF69F8C9FE955DC9" bold="true" box="[189,268,1960,1986]" pageId="11" pageNumber="53">Fig. 7.</emphasis>
<taxonomicName id="F4954D55FFB6C462FEC2F8C9FCE05DC9" authority="F.Jafari, Keshavarzi &amp; Doostm." authorityName="F. Jafari, Keshavarzi &amp; Doostm." authorityYear="2023" box="[278,889,1960,1986]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="53" phylum="Tracheophyta" rank="species" species="penduliflora" status="sp. nov.">
<emphasis id="01E1EAC4FFB6C462FEC2F8C9FE6F5DC9" box="[278,502,1960,1986]" italics="true" pageId="11" pageNumber="53">Silene penduliflora</emphasis>
F.Jafari, Keshavarzi &amp; Doostm.
</taxonomicName>
<taxonomicNameLabel id="1AD257BFFFB6C462FC55F8C9FC475DC9" box="[897,990,1960,1986]" pageId="11" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
(holotype,
<emphasis id="01E1EAC4FFB6C462FBBEF8C8FAF65DC9" box="[1130,1391,1960,1986]" italics="true" pageId="11" pageNumber="53">M. Mirtadzadini 2076</emphasis>
; MIR!).
</paragraph>
</caption>
<subSubSection id="7B8F655DFFB1C465FF69FE69FE82586D" pageId="12" pageNumber="54" type="materials_examined">
<paragraph id="332A36D6FFB1C465FF69FE69FE335B29" blockId="12.[189,426,264,290]" box="[189,426,264,290]" pageId="12" pageNumber="54">
<heading id="686281BAFFB1C465FF69FE69FE335B29" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="12" pageNumber="54" reason="3">
<emphasis id="01E1EAC4FFB1C465FF69FE69FE335B29" bold="true" box="[189,426,264,290]" pageId="12" pageNumber="54">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="332A36D6FFB1C465FF3EFE56FE4B5B93" blockId="12.[189,1399,311,408]" pageId="12" pageNumber="54">
<materialsCitation id="83FD3C8BFFB1C465FF3EFE56FAEB5B7F" ID-GBIF-Occurrence="4040262306" collectingDate="1997-06" collectionCode="MIR" collectorName="M. Mirtadzadini" country="Iran" elevation="3400" location="Jupar Mountain" pageId="12" pageNumber="54" specimenCount="1" stateProvince="Kerman" typeStatus="holotype">
<emphasis id="01E1EAC4FFB1C465FF3EFE56FEBE5B5A" bold="true" box="[234,295,311,337]" pageId="12" pageNumber="54">Type</emphasis>
<collectingCountry id="4B827646FFB1C465FF69FE3BFE9C5B7F" box="[189,261,346,372]" name="Iran" pageId="12" pageNumber="54">IRAN</collectingCountry>
–
<emphasis id="01E1EAC4FFB1C465FEFDFE3BFE085B7F" bold="true" box="[297,401,346,372]" pageId="12" pageNumber="54">
<collectingRegion id="F151F834FFB1C465FEFDFE3BFE085B7F" box="[297,401,346,372]" country="Iran" name="Kerman" pageId="12" pageNumber="54">Kerman</collectingRegion>
</emphasis>
•
<location id="364A600DFFB1C465FE79FE3AFDF55B7F" LSID="urn:lsid:plazi:treatment:BB3C87C0FFB7C464FE6DF969FB805FE3:364A600DFFB1C465FE79FE3AFDF55B7F" box="[429,620,346,373]" country="Iran" name="Jupar Mountain" pageId="12" pageNumber="54" stateProvince="Kerman">Jupar Mountain</location>
; alt.
<quantity id="F46D9B33FFB1C465FD7CFE3BFC9E5B7E" box="[680,775,346,373]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="54" unit="m" value="3400.0">
<elevation id="B8B8D1E5FFB1C465FD7CFE3BFC9E5B7E" box="[680,775,346,373]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="54" unit="m" value="3400.0">3400 m</elevation>
</quantity>
;
<date id="472B1016FFB1C465FCC1FE3AFC145B7F" box="[789,909,346,373]" pageId="12" pageNumber="54" value="1997-06">
<collectingDate id="576FE9FEFFB1C465FCC1FE3AFC145B7F" box="[789,909,346,373]" pageId="12" pageNumber="54" value="1997-06">Jun. 1997</collectingDate>
</date>
;
<emphasis id="01E1EAC4FFB1C465FC4AFE3AFB3A5B7F" box="[926,1187,346,372]" italics="true" pageId="12" pageNumber="54">
<collectorName id="9E605300FFB1C465FC4AFE3AFBC75B7F" box="[926,1118,346,372]" pageId="12" pageNumber="54">M. Mirtadzadini</collectorName>
2076
</emphasis>
;
<typeStatus id="EC2E8874FFB1C465FB61FE3BFAB95B7F" box="[1205,1312,346,372]" pageId="12" pageNumber="54" type="holotype">holotype</typeStatus>
:
<collectionCode id="5584AE13FFB1C465FAF9FE3BFAF75B7F" box="[1325,1390,346,372]" pageId="12" pageNumber="54">MIR</collectionCode>
!
</materialsCitation>
;
<materialsCitation id="83FD3C8BFFB1C465FF6BFE1CFE535B93" ID-GBIF-Occurrence="4040262301" box="[191,458,381,408]" collectingDate="1997-06" collectionCode="MIR, TARI" collectorName="M. Mirtadzadini" country="Iran" elevation="3400" location="Jupar Mountain" pageId="12" pageNumber="54" specimenCount="1" stateProvince="Kerman" typeStatus="isotype">
<typeStatus id="EC2E8874FFB1C465FF6BFE1CFEBD5B9C" box="[191,292,381,407]" pageId="12" pageNumber="54" type="isotype">isotypes</typeStatus>
:
<collectionCode id="5584AE13FFB1C465FEFBFE1FFEF65B93" box="[303,367,382,408]" pageId="12" pageNumber="54">MIR</collectionCode>
!,
<collectionCode id="5584AE13FFB1C465FEAAFE1FFE535B93" box="[382,458,382,408]" pageId="12" pageNumber="54">TARI</collectionCode>
</materialsCitation>
!.
</paragraph>
<paragraph id="332A36D6FFB1C465FF3EFEDEFE82586D" blockId="12.[189,1399,447,614]" pageId="12" pageNumber="54">
<materialsCitation id="83FD3C8BFFB1C465FF3EFEDEFDE85814" ID-GBIF-Occurrence="4040262304" collectingDate="2017-09-15" collectionCode="MIR" collectorName="M. Doostmohammadi" country="Iran" elevation="3300" location="Palvar Mountain" municipality="Mahan" pageId="12" pageNumber="54" specimenCount="1" stateProvince="Kerman" typeStatus="paratype">
<emphasis id="01E1EAC4FFB1C465FF3EFEDEFEFE5BD2" bold="true" box="[234,359,447,473]" pageId="12" pageNumber="54">
<typeStatus id="EC2E8874FFB1C465FF3EFEDEFEFE5BD2" box="[234,359,447,473]" pageId="12" pageNumber="54" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="4B827646FFB1C465FF69FE83FE9C5BF7" box="[189,261,482,508]" name="Iran" pageId="12" pageNumber="54">IRAN</collectingCountry>
–
<emphasis id="01E1EAC4FFB1C465FEEFFE83FE3A5BF7" bold="true" box="[315,419,482,508]" pageId="12" pageNumber="54">
<collectingRegion id="F151F834FFB1C465FEEFFE83FE3A5BF7" box="[315,419,482,508]" country="Iran" name="Kerman" pageId="12" pageNumber="54">Kerman</collectingRegion>
</emphasis>
•
<collectingMunicipality id="D34EACACFFB1C465FE01FE83FDB05BF7" box="[469,553,482,508]" pageId="12" pageNumber="54">Mahan</collectingMunicipality>
,
<location id="364A600DFFB1C465FD96FE83FC8C5BF7" LSID="urn:lsid:plazi:treatment:BB3C87C0FFB7C464FE6DF969FB805FE3:364A600DFFB1C465FD96FE83FC8C5BF7" box="[578,789,482,508]" country="Iran" municipality="Mahan" name="Palvar Mountain" pageId="12" pageNumber="54" stateProvince="Kerman">Palvar Mountain</location>
, western slopes; alt.
<quantity id="F46D9B33FFB1C465FBEEFE83FB3A5BF6" box="[1082,1187,482,509]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="12" pageNumber="54" unit="m" value="3300.0">
<elevation id="B8B8D1E5FFB1C465FBEEFE83FB3A5BF6" box="[1082,1187,482,509]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="12" pageNumber="54" unit="m" value="3300.0">3300 m</elevation>
</quantity>
;
<date id="472B1016FFB1C465FB6FFE83FAEB5BF7" box="[1211,1394,482,509]" pageId="12" pageNumber="54" value="2017-09-15">
<collectingDate id="576FE9FEFFB1C465FB6FFE83FAEB5BF7" box="[1211,1394,482,509]" pageId="12" pageNumber="54" value="2017-09-15">15 Sep. 2017</collectingDate>
</date>
;
<emphasis id="01E1EAC4FFB1C465FF69FD67FD8E5814" box="[189,535,517,543]" italics="true" pageId="12" pageNumber="54">
<collectorName id="9E605300FFB1C465FF69FD67FE515814" box="[189,456,517,543]" pageId="12" pageNumber="54">M. Doostmohammadi</collectorName>
4056
</emphasis>
;
<collectionCode id="5584AE13FFB1C465FDE7FD64FDE85814" box="[563,625,517,543]" pageId="12" pageNumber="54">MIR</collectionCode>
</materialsCitation>
! •
<materialsCitation id="83FD3C8BFFB1C465FD72FD67FE8F586D" ID-GBIF-Occurrence="4040262302" collectingDate="2020-08-14" collectionCode="MIR" collectorName="M. Doostmohammadi &amp; A. Ghorbanalizadeh" country="Iran" elevation="3300" location="Palvar Mountain" municipality="Mahan" pageId="12" pageNumber="54" specimenCount="1" stateProvince="Kerman" typeStatus="paratype">
same collection data as for preceding;
<date id="472B1016FFB1C465FB62FD64FAEB5814" box="[1206,1394,517,543]" pageId="12" pageNumber="54" value="2020-08-14">
<collectingDate id="576FE9FEFFB1C465FB62FD64FAEB5814" box="[1206,1394,517,543]" pageId="12" pageNumber="54" value="2020-08-14">14 Aug. 2020</collectingDate>
</date>
;
<emphasis id="01E1EAC4FFB1C465FF69FD48FC805848" box="[189,793,552,579]" italics="true" pageId="12" pageNumber="54">
<collectorName id="9E605300FFB1C465FF69FD48FD4C5849" box="[189,725,552,579]" pageId="12" pageNumber="54">M. Doostmohammadi &amp; A. Ghorbanalizadeh</collectorName>
4054
</emphasis>
;
<collectionCode id="5584AE13FFB1C465FCFEFD48FCF25848" box="[810,875,553,579]" pageId="12" pageNumber="54">MIR</collectionCode>
!. Genbank:
<accessionNumber id="2CC6AB35FFB1C465FC22FD49FBED5848" box="[1014,1140,552,579]" pageId="12" pageNumber="54">LC710549</accessionNumber>
(ITS) and
<accessionNumber id="2CC6AB35FFB1C465FB2DFD49FAEE5848" box="[1273,1399,552,579]" pageId="12" pageNumber="54">LC710551</accessionNumber>
(
<emphasis id="01E1EAC4FFB1C465FF13FD2AFE90586E" box="[199,265,587,613]" italics="true" pageId="12" pageNumber="54">rps16</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7B8F655DFFB1C464FF69FDECFB575BDD" lastPageId="13" lastPageNumber="55" pageId="12" pageNumber="54" type="description">
<paragraph id="332A36D6FFB1C465FF69FDECFED758AC" blockId="12.[189,334,653,679]" box="[189,334,653,679]" pageId="12" pageNumber="54">
<heading id="686281BAFFB1C465FF69FDECFED758AC" bold="true" box="[189,334,653,679]" fontSize="11" level="3" pageId="12" pageNumber="54" reason="3">
<emphasis id="01E1EAC4FFB1C465FF69FDECFED758AC" bold="true" box="[189,334,653,679]" pageId="12" pageNumber="54">Description</emphasis>
</heading>
</paragraph>
<paragraph id="332A36D6FFB1C464FF69FDDDFB575BDD" blockId="12.[189,1399,699,832]" lastBlockId="13.[189,1400,265,471]" lastPageId="13" lastPageNumber="55" pageId="12" pageNumber="54">
Perennial plants, glabrous, internodes viscid. Stems
<quantity id="F46D9B33FFB1C465FCEDFDDAFC4758DD" box="[825,990,699,726]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="4.05" metricValueMin="2.95" pageId="12" pageNumber="54" unit="cm" value="35.0" valueMax="40.5" valueMin="29.5">29.5‒40.5 cm</quantity>
tall, pale green, glabrous, viscid, internodes
<quantity id="F46D9B33FFB1C465FEEAFDBEFE2558F2" box="[318,444,735,761]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" metricValueMax="8.2" metricValueMin="3.2" pageId="12" pageNumber="54" unit="mm" value="57.0" valueMax="82.0" valueMin="32.0">32–82 mm</quantity>
long. Basal leaves 11‒60×
<quantity id="F46D9B33FFB1C465FD2AFDBEFCE158F2" box="[766,888,735,762]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="12" pageNumber="54" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5‒4 mm</quantity>
, oblanceolate, base attenuate, apex rounded and acute, glabrous, glaucous; cauline leaves oblanceolate to linear, apex acute, 12.5–41 ×
<quantity id="F46D9B33FFB1C465FB36FC63FAEB5916" box="[1250,1394,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" pageId="12" pageNumber="54" unit="mm" value="2.5" valueMax="3.5" valueMin="1.5">1.5–3.5 mm</quantity>
, often glabrous. Synflorescences laxly thyrsoid, pedicels
<quantity id="F46D9B33FFB1C465FCA6FC44FC40594B" box="[882,985,805,832]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="12" pageNumber="54" unit="mm" value="7.0" valueMax="9.0" valueMin="5.0">5‒9 mm</quantity>
long, glabrous. Bracteoles small, lanceolate,
<quantity id="F46D9B33FFB0C464FE97FE68FE225B28" box="[323,443,265,292]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="13" pageNumber="55" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
long. Calyx cylindrical,
<quantity id="F46D9B33FFB0C464FD35FE68FCC85B28" box="[737,849,265,291]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="13" pageNumber="55" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9‒11 mm</quantity>
long, glabrous, coriaceous; teeth membranous at the margin, triangular,
<quantity id="F46D9B33FFB0C464FE3CFE4CFDB45B4C" box="[488,557,301,327]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="55" unit="mm" value="2.0">2 mm</quantity>
long, two teeth acute with narrow transparent margin, other three teeth slightly broader, rounded, and with broad transparent margin. Petals yellowish, claw
<quantity id="F46D9B33FFB0C464FB6DFE30FAAA5B60" box="[1209,1331,337,364]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="13" pageNumber="55" unit="mm" value="6.25" valueMax="6.5" valueMin="6.0">6–6.5 mm</quantity>
long, with broad lateral margins, glabrous, without coronal scales; limb
<quantity id="F46D9B33FFB0C464FC06FE15FBAB5B84" box="[978,1074,372,399]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="13" pageNumber="55" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5‒6 mm</quantity>
long, oblanceolate, bifid to base. Anthophore
<quantity id="F46D9B33FFB0C464FE4DFEF9FE6E5BB8" box="[409,503,408,435]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="55" unit="mm" value="3.5">3.5 mm</quantity>
long, puberulent. Mature capsule 7.5‒9×
<quantity id="F46D9B33FFB0C464FC20FEF9FBEA5BB8" box="[1012,1139,408,435]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="13" pageNumber="55" unit="mm" value="4.75" valueMax="5.5" valueMin="4.0">4‒5.5 mm</quantity>
, ovoid-oblong. Seeds 1–1.1 ×
<quantity id="F46D9B33FFB0C464FEC3FEDCFE3E5BDC" box="[279,423,445,471]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="13" pageNumber="55" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
, basically reniform, seed testa cells elongate with V-shaped margins.
</paragraph>
</subSubSection>
<caption id="67EA665EFFB1C465FF69F8E9FAEE5DE2" ID-DOI="http://doi.org/10.5281/zenodo.7674412" ID-Zenodo-Dep="7674412" httpUri="https://zenodo.org/record/7674412/files/figure.png" pageId="12" pageNumber="54" startId="12.[189,232,1928,1954]" targetBox="[189,1399,906,1887]" targetPageId="12">
<paragraph id="332A36D6FFB1C465FF69F8E9FAEE5DE2" blockId="12.[189,1399,1928,2025]" pageId="12" pageNumber="54">
<emphasis id="01E1EAC4FFB1C465FF69F8E9FE935DA9" bold="true" box="[189,266,1928,1954]" pageId="12" pageNumber="54">Fig. 8.</emphasis>
<taxonomicName id="F4954D55FFB1C465FEC5F8E9FCF35DA8" authority="F.Jafari, Keshavarzi &amp; Doostm." authorityName="F. Jafari, Keshavarzi &amp; Doostm." authorityYear="2023" box="[273,874,1928,1955]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="54" phylum="Tracheophyta" rank="species" species="penduliflora" status="sp. nov.">
<emphasis id="01E1EAC4FFB1C465FEC5F8E9FE765DA9" box="[273,495,1928,1954]" italics="true" pageId="12" pageNumber="54">Silene penduliflora</emphasis>
F.Jafari, Keshavarzi &amp; Doostm.
</taxonomicName>
<taxonomicNameLabel id="1AD257BFFFB1C465FCA4F8E8FC525DA8" box="[880,971,1929,1955]" pageId="12" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
(paratypes,
<emphasis id="01E1EAC4FFB1C465FB8EF8E8FE765DCD" italics="true" pageId="12" pageNumber="54">M. Doostmohammadi &amp; A. Ghorbanalizadeh 4054</emphasis>
; MIR!.
<emphasis id="01E1EAC4FFB1C465FD85F8CDFC0B5DCE" box="[593,914,1963,1989]" italics="true" pageId="12" pageNumber="54">M. Doostmohammadi 4056</emphasis>
; MIR!).
<emphasis id="01E1EAC4FFB1C465FC2AF8CAFBAD5DCE" bold="true" box="[1022,1076,1963,1989]" pageId="12" pageNumber="54">A‒B</emphasis>
. Inflorescence at flowering and fruiting times.
<emphasis id="01E1EAC4FFB1C465FE48F8AFFE285DE3" bold="true" box="[412,433,1998,2024]" pageId="12" pageNumber="54">C</emphasis>
. Basal leaves and caudex.
<emphasis id="01E1EAC4FFB1C465FD39F8AEFC9B5DE2" bold="true" box="[749,770,1999,2025]" pageId="12" pageNumber="54">D</emphasis>
. Habit and habitat. Photos by: M. Doostmohammadi.
</paragraph>
</caption>
<subSubSection id="7B8F655DFFB0C464FF69FE9FFD215867" pageId="13" pageNumber="55" type="distribution">
<paragraph id="332A36D6FFB0C464FF69FE9FFE775813" blockId="13.[189,494,510,537]" box="[189,494,510,537]" pageId="13" pageNumber="55">
<heading id="686281BAFFB0C464FF69FE9FFE775813" bold="true" box="[189,494,510,537]" fontSize="11" level="3" pageId="13" pageNumber="55" reason="3">
<emphasis id="01E1EAC4FFB0C464FF69FE9FFE775813" bold="true" box="[189,494,510,537]" pageId="13" pageNumber="55">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="332A36D6FFB0C464FF69FD4FFD215867" blockId="13.[189,1398,558,620]" pageId="13" pageNumber="55">
<taxonomicName id="F4954D55FFB0C464FF69FD4FFE025843" authority="Jafari &amp; Keshavarzi &amp; Doostmohammadi &amp; Mirtadzadini, 2023" authorityName="Jafari &amp; Keshavarzi &amp; Doostmohammadi &amp; Mirtadzadini" authorityYear="2023" box="[189,411,558,584]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="penduliflora" status="sp. nov.">
<emphasis id="01E1EAC4FFB0C464FF69FD4FFE025843" box="[189,411,558,584]" italics="true" pageId="13" pageNumber="55">Silene penduliflora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AD257BFFFB0C464FE77FD4FFE645843" box="[419,509,558,584]" pageId="13" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
is restricted to rocks in southeast
<collectingCountry id="4B827646FFB0C464FC5FFD4FFC235843" box="[907,954,558,584]" name="Iran" pageId="13" pageNumber="55">Iran</collectingCountry>
on the Jupar and Palvar mountains of
<collectingRegion id="F151F834FFB0C464FF69FD33FE125867" box="[189,395,594,620]" country="Iran" name="Kerman" pageId="13" pageNumber="55">Kerman Province</collectingRegion>
at
<quantity id="F46D9B33FFB0C464FE7BFD33FDC85867" box="[431,593,594,620]" metricMagnitude="3" metricUnit="m" metricValue="3.35" metricValueMax="3.4" metricValueMin="3.3" pageId="13" pageNumber="55" unit="m" value="3350.0" valueMax="3400.0" valueMin="3300.0">3300‒3400 m</quantity>
altitude.
</paragraph>
</subSubSection>
<subSubSection id="7B8F655DFFB0C464FF69FDF5FB805FE3" pageId="13" pageNumber="55" type="discussion">
<paragraph id="332A36D6FFB0C464FF69FDF5FEB658A5" blockId="13.[189,303,660,686]" box="[189,303,660,686]" pageId="13" pageNumber="55">
<heading id="686281BAFFB0C464FF69FDF5FEB658A5" bold="true" box="[189,303,660,686]" fontSize="11" level="3" pageId="13" pageNumber="55" reason="3">
<emphasis id="01E1EAC4FFB0C464FF69FDF5FEB658A5" bold="true" box="[189,303,660,686]" pageId="13" pageNumber="55">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="332A36D6FFB0C464FF69FDA2FE8A5E4F" blockId="13.[189,1399,707,1092]" pageId="13" pageNumber="55">
Although
<taxonomicName id="F4954D55FFB0C464FEECFDA2FCE158D5" authority="F.Jafari, Keshavarzi &amp; Doostm." authorityName="F. Jafari, Keshavarzi &amp; Doostm." authorityYear="2023" box="[312,888,707,734]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="penduliflora" status="sp. nov.">
<emphasis id="01E1EAC4FFB0C464FEECFDA2FE7058D6" box="[312,489,707,733]" italics="true" pageId="13" pageNumber="55">S. penduliflora</emphasis>
F.Jafari, Keshavarzi &amp; Doostm.
</taxonomicName>
<taxonomicNameLabel id="1AD257BFFFB0C464FC50FDA5FC7D58D5" box="[900,996,708,734]" pageId="13" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="F4954D55FFB0C464FBF2FDA2FB7D58D6" box="[1062,1252,707,733]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="shahrudensis">
<emphasis id="01E1EAC4FFB0C464FBF2FDA2FB7D58D6" box="[1062,1252,707,733]" italics="true" pageId="13" pageNumber="55">S. shahrudensis</emphasis>
</taxonomicName>
show more morphological resemblance to each other than to other close relatives, neither their ITS nor
<emphasis id="01E1EAC4FFB0C464FAE0FD86FAEF590A" box="[1332,1398,743,769]" italics="true" pageId="13" pageNumber="55">rps16</emphasis>
sequences reflect a closer relationship, and
<taxonomicName id="F4954D55FFB0C464FD0DFC6AFC12592E" box="[729,907,779,805]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="penduliflora">
<emphasis id="01E1EAC4FFB0C464FD0DFC6AFC12592E" box="[729,907,779,805]" italics="true" pageId="13" pageNumber="55">S. penduliflora</emphasis>
</taxonomicName>
and
<taxonomicName id="F4954D55FFB0C464FC1BFC6AFED85942" authority="F.Jafari, Mirtadz. &amp; Keshavarzi" authorityName="F. Jafari, Mirtadz. &amp; Keshavarzi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="thyrsiantha" status="sp. nov.">
<emphasis id="01E1EAC4FFB0C464FC1BFC6AFBE1592E" box="[975,1144,779,805]" italics="true" pageId="13" pageNumber="55">S. thyrsiantha</emphasis>
F.Jafari, Mirtadz. &amp; Keshavarzi
</taxonomicName>
<taxonomicNameLabel id="1AD257BFFFB0C464FE9BFC4EFE285942" box="[335,433,815,841]" pageId="13" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
form a well-supported clade in the
<emphasis id="01E1EAC4FFB0C464FC51FC4EFC5E5942" box="[901,967,815,841]" italics="true" pageId="13" pageNumber="55">rps16</emphasis>
tree (
<figureCitation id="ABAE2A53FFB0C464FBCEFC4EFBF05942" box="[1050,1129,815,841]" captionStart="Fig" captionStartId="4.[189,232,1855,1881]" captionTargetBox="[189,1399,546,1812]" captionTargetId="figure-78@4.[189,1399,546,1813]" captionTargetPageId="4" captionText="Fig. 2. 50% majority-rule consensus tree obtained from the Bayesian inference analysis of the cpDNA rps16 sequences in selected species of Silene L. Posterior probabilities (PP) ≥ 0.70 are shown above the branches and Maximum likelihood bootstrap (MLB) values ≥ 70% below. Accessions newly sequenced are indicated by a star. Accessions of S. ruprechtii Schischk. (MN460318 and MN460319) are misidentified by Bahmani et al. (2020)." figureDoi="http://doi.org/10.5281/zenodo.7674400" httpUri="https://zenodo.org/record/7674400/files/figure.png" pageId="13" pageNumber="55">Fig. 2</figureCitation>
, PP= 1, MLB= 96%).
<taxonomicName id="F4954D55FFB0C464FF69FC32FE045966" box="[189,413,851,877]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="penduliflora">
<emphasis id="01E1EAC4FFB0C464FF69FC32FE045966" box="[189,413,851,877]" italics="true" pageId="13" pageNumber="55">Silene penduliflora</emphasis>
</taxonomicName>
and
<taxonomicName id="F4954D55FFB0C464FE0AFC32FD005966" box="[478,665,851,877]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="shahrudensis">
<emphasis id="01E1EAC4FFB0C464FE0AFC32FD005966" box="[478,665,851,877]" italics="true" pageId="13" pageNumber="55">S. shahrudensis</emphasis>
</taxonomicName>
differ in pedicel length (
<quantity id="F46D9B33FFB0C464FC11FC33FBB35966" box="[965,1066,850,877]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="13" pageNumber="55" unit="mm" value="7.0" valueMax="9.0" valueMin="5.0">5‒9 mm</quantity>
vs
<quantity id="F46D9B33FFB0C464FB83FC32FB545966" box="[1111,1229,851,877]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="13" pageNumber="55" unit="mm" value="9.0" valueMax="12.0" valueMin="6.0">6‒12 mm</quantity>
), calyx length (
<quantity id="F46D9B33FFB0C464FF12FC16FEA0599A" box="[198,313,887,913]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="13" pageNumber="55" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9‒11 mm</quantity>
vs
<quantity id="F46D9B33FFB0C464FEB3FC16FE46599A" box="[359,479,887,913]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="13" pageNumber="55" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8‒10 mm</quantity>
), petal limb length (
<quantity id="F46D9B33FFB0C464FD02FC17FCA2599A" box="[726,827,886,913]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="13" pageNumber="55" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5‒6 mm</quantity>
vs ca
<quantity id="F46D9B33FFB0C464FC5BFC17FC43599A" box="[911,986,886,913]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="55" unit="mm" value="5.0">5 mm</quantity>
), surface of the claw (glabrous vs ciliate), and capsule size (7.5‒9×
<quantity id="F46D9B33FFB0C464FD98FCFBFD5C59BE" box="[588,709,922,949]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="13" pageNumber="55" unit="mm" value="4.75" valueMax="5.5" valueMin="4.0">4‒5.5 mm</quantity>
vs 4‒6×
<quantity id="F46D9B33FFB0C464FCE0FCFAFC0059BE" box="[820,921,923,949]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="13" pageNumber="55" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3‒4 mm</quantity>
) (
<tableCitation id="7E17036DFFB0C464FC65FCFBFB9359BE" box="[945,1034,922,949]" captionStart="Table 1" captionStartId="6.[189,258,265,291]" captionTargetPageId="6" captionText="Table 1. Morphological comparison among S. penduliflora F.Jafari, Keshavarzi &amp; Doostm. sp. nov., S. thyrsiantha F.Jafari, Mirtadz. &amp; Keshavarzi sp. nov. and close relatives. Distinctive character states are shown in bold." httpUri="http://table.plazi.org/id/67EA665EFFBBC46FFF69FE68FE0E5B62" pageId="13" pageNumber="55" tableUuid="67EA665EFFBBC46FFF69FE68FE0E5B62">Table 1</tableCitation>
). The margin of the seed testa cells is V-shaped in
<taxonomicName id="F4954D55FFB0C464FE7AFCDEFDC259D3" box="[430,603,958,985]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="penduliflora">
<emphasis id="01E1EAC4FFB0C464FE7AFCDEFDC259D3" box="[430,603,958,985]" italics="true" pageId="13" pageNumber="55">S. penduliflora</emphasis>
</taxonomicName>
while it is U-shaped in
<taxonomicName id="F4954D55FFB0C464FCAFFCDEFBAA59D3" box="[891,1075,958,985]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="shahrudensis">
<emphasis id="01E1EAC4FFB0C464FCAFFCDEFBAA59D3" box="[891,1075,958,985]" italics="true" pageId="13" pageNumber="55">S. shahrudensis</emphasis>
</taxonomicName>
. Geographically, these two species are also distant.
<taxonomicName id="F4954D55FFB0C464FE0AFC83FD2059F7" box="[478,697,994,1020]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="sharudensis">
<emphasis id="01E1EAC4FFB0C464FE0AFC83FD2059F7" box="[478,697,994,1020]" italics="true" pageId="13" pageNumber="55">Silene sharudensis</emphasis>
</taxonomicName>
is confined to the central and eastern
<collectingRegion id="F151F834FFB0C464FBA8FC83FB5459F7" box="[1148,1229,994,1020]" country="Iran" name="Alborz" pageId="13" pageNumber="55">Alborz</collectingRegion>
(north central
<collectingCountry id="4B827646FFB0C464FF69FB66FF775E2A" box="[189,238,1031,1057]" name="Iran" pageId="13" pageNumber="55">Iran</collectingCountry>
) while
<taxonomicName id="F4954D55FFB0C464FE98FB66FE655E2B" box="[332,508,1030,1057]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="penduliflora">
<emphasis id="01E1EAC4FFB0C464FE98FB66FE655E2B" box="[332,508,1030,1057]" italics="true" pageId="13" pageNumber="55">S. penduliflora</emphasis>
</taxonomicName>
is endemic to the highlands of
<collectingRegion id="F151F834FFB0C464FC57FB66FC785E2A" box="[899,993,1031,1057]" country="Iran" name="Kerman" pageId="13" pageNumber="55">Kerman</collectingRegion>
in southeast
<collectingCountry id="4B827646FFB0C464FB52FB66FB2C5E2A" box="[1158,1205,1031,1057]" name="Iran" pageId="13" pageNumber="55">Iran</collectingCountry>
about
<quantity id="F46D9B33FFB0C464FAD8FB67FAEF5E2B" box="[1292,1398,1030,1056]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="55" unit="km" value="1000.0">1000 km</quantity>
distant.
</paragraph>
<paragraph id="332A36D6FFB0C464FF69FB0DFDF55F55" blockId="13.[189,1399,1132,1374]" pageId="13" pageNumber="55">
Despite the affinity of
<taxonomicName id="F4954D55FFB0C464FE0FFB0CFB875E8C" authority="F.Jafari, Keshavarzi &amp; Doostm." authorityName="F. Jafari, Keshavarzi &amp; Doostm." authorityYear="2023" box="[475,1054,1132,1159]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="penduliflora" status="sp. nov.">
<emphasis id="01E1EAC4FFB0C464FE0FFB0CFD145E8D" box="[475,653,1132,1159]" italics="true" pageId="13" pageNumber="55">S. penduliflora</emphasis>
F.Jafari, Keshavarzi &amp; Doostm.
</taxonomicName>
<taxonomicNameLabel id="1AD257BFFFB0C464FBFEFB0CFB135E8C" box="[1066,1162,1133,1159]" pageId="13" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="F4954D55FFB0C464FB1AFB0CFDDB5EA1" authority="F.Jafari, Mirtadz. &amp; Keshavarzi" authorityName="F. Jafari, Mirtadz. &amp; Keshavarzi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="thyrsiantha" status="sp. nov.">
<emphasis id="01E1EAC4FFB0C464FB1AFB0CFAEE5E8D" box="[1230,1399,1132,1159]" italics="true" pageId="13" pageNumber="55">S. thyrsiantha</emphasis>
F.Jafari, Mirtadz. &amp; Keshavarzi
</taxonomicName>
<taxonomicNameLabel id="1AD257BFFFB0C464FD84FBF0FD295EA0" box="[592,688,1169,1195]" pageId="13" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
in the phylogenies, they differ in calyx length (9‒11 vs
<quantity id="F46D9B33FFB0C464FF69FBD5FEAC5EC4" box="[189,309,1204,1231]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="13" pageNumber="55" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8‒10 mm</quantity>
), secondary axis inflorescence (one flower vs more than one), capsule size (7.5‒9×4‒5.5 vs 5.5‒6×
<quantity id="F46D9B33FFB0C464FEE9FBB9FE255EF9" box="[317,444,1240,1267]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="13" pageNumber="55" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">3‒3.5 mm</quantity>
), anthophore length (3.5 vs
<quantity id="F46D9B33FFB0C464FCCDFBB9FCFF5EF9" box="[793,870,1240,1266]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="55" unit="mm" value="3.0">3 mm</quantity>
), petal limb shape (oblanceolate vs linear), and seed size (1–1.1× 0.7–0.8 vs 0.8×
<quantity id="F46D9B33FFB0C464FD51FB9DFC815F1D" box="[645,792,1276,1302]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="13" pageNumber="55" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
). Although their calyx lengths overlap, the calyx in
<taxonomicName id="F4954D55FFB0C464FF69FA41FEF85F31" authorityName="F. Jafari, Mirtadz. &amp; Keshavarzi" authorityYear="2023" box="[189,353,1312,1338]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="thyrsiantha">
<emphasis id="01E1EAC4FFB0C464FF69FA41FEF85F31" box="[189,353,1312,1338]" italics="true" pageId="13" pageNumber="55">S. thyrsiantha</emphasis>
</taxonomicName>
is narrower than in
<taxonomicName id="F4954D55FFB0C464FD85FA41FD675F31" box="[593,766,1312,1338]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="penduliflora">
<emphasis id="01E1EAC4FFB0C464FD85FA41FD675F31" box="[593,766,1312,1338]" italics="true" pageId="13" pageNumber="55">S. penduliflora</emphasis>
</taxonomicName>
and both the capsules and seeds of
<taxonomicName id="F4954D55FFB0C464FB7CFA41FAD25F31" authorityName="F. Jafari, Mirtadz. &amp; Keshavarzi" authorityYear="2023" box="[1192,1355,1312,1338]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="thyrsiantha">
<emphasis id="01E1EAC4FFB0C464FB7CFA41FAD25F31" box="[1192,1355,1312,1338]" italics="true" pageId="13" pageNumber="55">S. thyrsiantha</emphasis>
</taxonomicName>
are smaller than those in
<taxonomicName id="F4954D55FFB0C464FE6CFA25FDFD5F55" box="[440,612,1348,1374]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="penduliflora">
<emphasis id="01E1EAC4FFB0C464FE6CFA25FDFD5F55" box="[440,612,1348,1374]" italics="true" pageId="13" pageNumber="55">S. penduliflora</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="332A36D6FFB0C464FF69FAE7FB805FE3" blockId="13.[189,1399,1414,1512]" pageId="13" pageNumber="55">
<taxonomicName id="F4954D55FFB0C464FF69FAE7FCBB5FAB" authority="F.Jafari, Keshavarzi &amp; Doostm." authorityName="F. Jafari, Keshavarzi &amp; Doostm." authorityYear="2023" box="[189,802,1414,1440]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="penduliflora" status="sp. nov.">
<emphasis id="01E1EAC4FFB0C464FF69FAE7FE045FAB" box="[189,413,1414,1440]" italics="true" pageId="13" pageNumber="55">Silene penduliflora</emphasis>
F.Jafari, Keshavarzi &amp; Doostm.
</taxonomicName>
<taxonomicNameLabel id="1AD257BFFFB0C464FCFFFAE7FC115FAB" box="[811,904,1414,1440]" pageId="13" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="F4954D55FFB0C464FBF9FAE7FAB25FAB" box="[1069,1323,1414,1440]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="55" phylum="Tracheophyta" rank="species" species="ghahremaninejadii">
<emphasis id="01E1EAC4FFB0C464FBF9FAE7FAB25FAB" box="[1069,1323,1414,1440]" italics="true" pageId="13" pageNumber="55">S. ghahremaninejadii</emphasis>
</taxonomicName>
in the calyx length (
<quantity id="F46D9B33FFB0C464FE8AFACBFE535FCF" box="[350,458,1450,1476]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="13" pageNumber="55" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9‒11 mm</quantity>
vs
<quantity id="F46D9B33FFB0C464FE24FACBFDCC5FCF" box="[496,597,1450,1476]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="13" pageNumber="55" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7‒9 mm</quantity>
), petal limb length (
<quantity id="F46D9B33FFB0C464FCEFFAC8FC045FCF" box="[827,925,1449,1476]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="13" pageNumber="55" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5‒6 mm</quantity>
vs
<quantity id="F46D9B33FFB0C464FC17FACBFBBE5FCF" box="[963,1063,1450,1476]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="13" pageNumber="55" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2‒3 mm</quantity>
), anthophore length (
<quantity id="F46D9B33FFB0C464FAC9FAC8FAEE5FCF" box="[1309,1399,1449,1476]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="55" unit="mm" value="3.5">3.5 mm</quantity>
vs
<quantity id="F46D9B33FFB0C464FF0AFAACFEC35FE3" box="[222,346,1485,1512]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="13" pageNumber="55" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5‒4 mm</quantity>
), and capsule size (7.5‒9×
<quantity id="F46D9B33FFB0C464FD43FAACFC965FE3" box="[663,783,1485,1512]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="13" pageNumber="55" unit="mm" value="4.75" valueMax="5.5" valueMin="4.0">4‒5.5 mm</quantity>
vs 6.5‒7 ×
<quantity id="F46D9B33FFB0C464FC45FAACFB945FE3" box="[913,1037,1485,1512]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="13" pageNumber="55" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5‒3 mm</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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