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<mods:title id="BAA5EC29978016CE84C7B5C7FC3381E8">A taxonomic review of Korean Acrotona Thomson with a description of new species (Coleoptera, Staphylinidae, Aleocharinae)</mods:title>
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<emphasis id="0B9DEABFFFD75B4EFF30F9FAFE8FF9FA" bold="true" box="[151,263,1629,1655]" italics="true" pageId="1" pageNumber="248">Acrotona</emphasis>
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<emphasis id="0B9DEABFFFD75B4EFEBFF9FAFE00F9FA" bold="true" box="[280,392,1629,1655]" italics="true" pageId="1" pageNumber="248">Acrotona</emphasis>
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<emphasis id="0B9DEABFFFD75B4EFE3DF9FAFD8AF9FA" bold="true" box="[410,514,1629,1655]" italics="true" pageId="1" pageNumber="248">aterrima</emphasis>
(
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(
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<taxonomicName id="FEE94D2EFFD75B4EFF30F96FFDC3F953" ID-CoL="8XTD9" authority="Gravenhorst, 1802: 83" authorityName="Gravenhorst. This" authorityPageNumber="83" authorityYear="1802" box="[151,587,1736,1758]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="0B9DEABFFFD75B4EFF30F96FFED7F953" box="[151,351,1736,1758]" italics="true" pageId="1" pageNumber="248">Aleochara aterrima</emphasis>
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<taxonomicName id="FEE94D2EFFD75B4EFF30F941FDC7F971" ID-CoL="MDL7" authority="Mannerheim, 1830: 84" authorityName="Mannerheim" authorityPageNumber="84" authorityYear="1830" box="[151,591,1766,1788]" class="Insecta" family="Staphylinidae" genus="Bolitochara" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="fuscula">
<emphasis id="0B9DEABFFFD75B4EFF30F941FED7F971" box="[151,351,1766,1788]" italics="true" pageId="1" pageNumber="248">Bolitochara fuscula</emphasis>
<bibRefCitation id="5D784B5CFFD75B4EFEC2F941FDC7F971" author="Mannerheim, C. G." box="[357,591,1766,1788]" pageId="1" pageNumber="248" refId="ref11637" refString="Mannerheim, C. G. (1830) Precis d'un nouvel arrangement de la famille des brachelytres, de l'ordre des insectes coleopteres. St. Petersbourg, 87 pp." type="book" year="1830">Mannerheim, 1830: 84</bibRefCitation>
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<treatmentCitationGroup id="19F91183FFD75B4EFF30F8A2FD2FF896" box="[151,679,1797,1819]" pageId="1" pageNumber="248">
<treatmentCitation id="B84810BCFFD75B4EFF30F8A2FD2BF896" author="Sahlberg, C. R." box="[151,675,1797,1819]" page="380" pageId="1" pageNumber="248" year="1831">
<taxonomicName id="FEE94D2EFFD75B4EFF30F8A2FD2BF896" ID-CoL="BNPG" authority="C. R. Sahlberg, 1831: 380" authorityName="C. R. Sahlberg" authorityPageNumber="380" authorityYear="1831" box="[151,675,1797,1819]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="mannerheimii">
<emphasis id="0B9DEABFFFD75B4EFF30F8A2FE1AF896" box="[151,402,1797,1819]" italics="true" pageId="1" pageNumber="248">Aleochara mannerheimii</emphasis>
C. R.
<bibRefCitation id="5D784B5CFFD75B4EFE73F8A2FD2BF896" author="Sahlberg, C. R." box="[468,675,1797,1819]" pageId="1" pageNumber="248" pagination="361 - 408" refId="ref12147" refString="Sahlberg, C. R. (1831) Insecta Fennica, dissertationibus academicis. A. 1817 - 1834 editis. A. Frenckelliana, Helsingforsiae, 48 pp. [pp. 361 - 408]" type="book chapter" year="1831">Sahlberg, 1831: 380</bibRefCitation>
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<taxonomicName id="FEE94D2EFFD75B4EFF30F883FDCBF8B7" ID-CoL="573RN" authority="Stephens, 1832: 133" authorityName="Stephens" authorityPageNumber="133" authorityYear="1832" box="[151,579,1828,1850]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="brunnipes">
<emphasis id="0B9DEABFFFD75B4EFF30F883FEE3F8B7" box="[151,363,1828,1850]" italics="true" pageId="1" pageNumber="248">Aleochara brunnipes</emphasis>
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<taxonomicName id="FEE94D2EFFD75B4EFF30F8E5FDACF8D5" ID-CoL="BNP6" authority="Stephens, 1832: 136" authorityName="Stephens" authorityPageNumber="136" authorityYear="1832" box="[151,548,1858,1880]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="luripes">
<emphasis id="0B9DEABFFFD75B4EFF30F8E5FEC4F8D5" box="[151,332,1858,1880]" italics="true" pageId="1" pageNumber="248">Aleochara luripes</emphasis>
<bibRefCitation id="5D784B5CFFD75B4EFEF4F8E5FDACF8D5" author="Stephens, J. F." box="[339,548,1858,1880]" pageId="1" pageNumber="248" refId="ref12555" refString="Stephens, J. F. (1832) Illustrations of British entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable Printed for the author; published by Baldwin and Cradock, London, 447 pp." type="book" year="1832">Stephens, 1832: 136</bibRefCitation>
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<taxonomicName id="FEE94D2EFFD75B4EFF30F8C6FDCBF8FA" ID-CoL="BNR7" authority="Stephens, 1832: 137" authorityName="Stephens" authorityPageNumber="137" authorityYear="1832" box="[151,579,1889,1911]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="obscurata">
<emphasis id="0B9DEABFFFD75B4EFF30F8C6FEE3F8FA" box="[151,363,1889,1911]" italics="true" pageId="1" pageNumber="248">Aleochara obscurata</emphasis>
<bibRefCitation id="5D784B5CFFD75B4EFED5F8C6FDCBF8FA" author="Stephens, J. F." box="[370,579,1889,1911]" pageId="1" pageNumber="248" refId="ref12555" refString="Stephens, J. F. (1832) Illustrations of British entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable Printed for the author; published by Baldwin and Cradock, London, 447 pp." type="book" year="1832">Stephens, 1832: 137</bibRefCitation>
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<taxonomicName id="FEE94D2EFFD75B4EFF30F827FDCCF81B" ID-CoL="8Y4LC" authority="Kiesenwetter, 1844: 319" authorityName="Kiesenwetter" authorityPageNumber="319" authorityYear="1844" box="[151,580,1920,1942]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="lugens">
<emphasis id="0B9DEABFFFD75B4EFF30F827FECCF81B" box="[151,324,1920,1942]" italics="true" pageId="1" pageNumber="248">Homalota lugens</emphasis>
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<treatmentCitation id="B84810BCFFD75B4EFF30F839FD09F839" author="Brundin, L." box="[151,641,1950,1973]" page="121" pageId="1" pageNumber="248" year="1952">
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<emphasis id="0B9DEABFFFD75B4EFF30F839FF51F839" box="[151,217,1950,1972]" italics="true" pageId="1" pageNumber="248">Atheta</emphasis>
(
<emphasis id="0B9DEABFFFD75B4EFF4FF838FECDF839" box="[232,325,1951,1972]" italics="true" pageId="1" pageNumber="248">Acrotona</emphasis>
)
<emphasis id="0B9DEABFFFD75B4EFEF4F807FE24F838" box="[339,428,1952,1973]" italics="true" pageId="1" pageNumber="248">aterrima</emphasis>
:
<bibRefCitation id="5D784B5CFFD75B4EFE1DF839FD09F839" author="Brundin, L." box="[442,641,1950,1972]" pageId="1" pageNumber="248" pagination="93 - 145" refId="ref11202" refString="Brundin, L. (1952) Acrotona - Studien. (Gattung Atheta, Col., Staphylinidae). Entomologisk Tidskrift, 73, 93 - 145." type="journal article" year="1952">Brundin, 1952: 121</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="B84810BCFFD75B4EFD2BF839FCBEF839" author="Palm, T." box="[652,822,1950,1972]" page="288" pageId="1" pageNumber="248" year="1970">
<bibRefCitation id="5D784B5CFFD75B4EFD2BF839FCBEF839" author="Palm, T." box="[652,822,1950,1972]" pageId="1" pageNumber="248" pagination="117 - 296" refId="ref12070" refString="Palm, T. (1970) Svensk Insektfauna: Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Underfam. Aleocharinae (Atheta). Hafte 6. Entomologiska Foreningen I, Stockholm, 180 pp. [pp. 117 - 296]" type="book chapter" year="1970">Palm, 1970: 288</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="B84810BCFFD75B4EFCE6F839FBD4F839" author="Benick, G. &amp; Lohse, G. A." box="[833,1116,1950,1972]" page="187" pageId="1" pageNumber="248" year="1974">
<bibRefCitation id="5D784B5CFFD75B4EFCE6F839FBD4F839" author="Benick, G. &amp; Lohse, G. A." box="[833,1116,1950,1972]" pageId="1" pageNumber="248" pagination="72 - 220" refId="ref11125" refString="Benick, G. &amp; Lohse, G. A. (1974) [Tribus 14 (Callicerini)]. In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5. Staphylinidae II (Hypocyptinae und Aleocharinae), Pselaphidae. Goecke and Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse, 1974: 187</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="B84810BCFFD75B4EFBC1F839FAFAF839" author="Yosii, R. &amp; Sawada, K." box="[1126,1394,1950,1972]" page="94" pageId="1" pageNumber="248" year="1976">
<bibRefCitation id="5D784B5CFFD75B4EFBC1F839FAFAF839" author="Yosii, R. &amp; Sawada, K." box="[1126,1394,1950,1972]" pageId="1" pageNumber="248" pagination="11 - 140" refId="ref12754" refString="Yosii, R. &amp; Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of genera and subgenera with description of representative species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140." type="journal article" year="1976">Yosii &amp; Sawada, 1976: 94</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="B84810BCFFD75B4EFF30F819FD91F85E" author="Pasnik, G." box="[151,537,1981,2003]" page="202" pageId="1" pageNumber="248" year="2001">
<taxonomicName id="FEE94D2EFFD75B4EFF30F819FD91F85E" ID-CoL="9QY9" authority=": Pasnik, 2001: 202" authorityName="Pasnik" authorityPageNumber="202" authorityYear="2001" box="[151,537,1981,2003]" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="0B9DEABFFFD75B4EFF30F819FEDBF85E" box="[151,339,1982,2003]" italics="true" pageId="1" pageNumber="248">Acrotona aterrima</emphasis>
:
<bibRefCitation id="5D784B5CFFD75B4EFEC7F81AFD91F85E" author="Pasnik, G." box="[352,537,1981,2003]" pageId="1" pageNumber="248" pagination="185 - 234" refId="ref12116" refString="Pasnik, G. (2001) The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeography. Acta Zoologica Cracoviensia, 44, 185 - 234." type="journal article" year="2001">Paśnik, 2001: 202</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="B84810BCFFD75B4EFD83F81AFD79F85E" author="Smetana, A." box="[548,753,1981,2003]" page="362" pageId="1" pageNumber="248" year="2004">
<bibRefCitation id="5D784B5CFFD75B4EFD83F81AFD79F85E" author="Smetana, A." box="[548,753,1981,2003]" pageId="1" pageNumber="248" pagination="353 - 494" refId="ref12505" refString="Smetana, A. (2004) Family Staphylinidae: Aleocharinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea-Histeroidea-Staphylinoidea. Apollo Books, Stenstrup, pp. 353 - 494." type="book chapter" year="2004">Smetana, 2004: 362</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="395636ADFFD75B4EFF30F87BFC74F87F" blockId="1.[151,1399,1736,2034]" box="[151,1020,2012,2034]" pageId="1" pageNumber="248">
<treatmentCitationGroup id="19F91183FFD75B4EFF30F87BFC74F87F" box="[151,1020,2012,2034]" pageId="1" pageNumber="248">
<treatmentCitation id="B84810BCFFD75B4EFF30F87BFC9BF87F" author="Schulke, M. &amp; Smetana, A." box="[151,787,2012,2034]" page="506" pageId="1" pageNumber="248" year="2015">
<taxonomicName id="FEE94D2EFFD75B4EFF30F87BFC9BF87F" ID-CoL="9QY9" authority=": Schulke &amp; Smetana, 2015: 506" authorityName=": Schulke &amp; Smetana" authorityPageNumber="506" authorityYear="2015" box="[151,787,2012,2034]" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="aterrima" subGenus="Acrotona">
<emphasis id="0B9DEABFFFD75B4EFF30F87BFF7CF87C" box="[151,244,2012,2033]" italics="true" pageId="1" pageNumber="248">Acrotona</emphasis>
(
<emphasis id="0B9DEABFFFD75B4EFEA4F87BFEE8F87C" box="[259,352,2012,2033]" italics="true" pageId="1" pageNumber="248">Acrotona</emphasis>
)
<emphasis id="0B9DEABFFFD75B4EFEC9F87AFE4FF87F" box="[366,455,2013,2034]" italics="true" pageId="1" pageNumber="248">aterrima</emphasis>
:
<bibRefCitation id="5D784B5CFFD75B4EFE72F87BFC9BF87F" author="Schulke, M. &amp; Smetana, A." box="[469,787,2012,2034]" pageId="1" pageNumber="248" pagination="304 - 1134" refId="ref12294" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae Latreille, 1802. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 2 / 1. Revised and updated edition. Hydrophiloidea-Staphylinoidea. Brill, Leiden, pp. 304 - 1134." type="book chapter" year="2015">Schülke &amp; Smetana, 2015: 506</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="B84810BCFFD75B4EFCB9F87BFC70F87F" author="Ahn, K. - J. &amp; Cho, Y. B. &amp; Kim, Y. - H. &amp; Yoo, I. - S." box="[798,1016,2012,2034]" page="292" pageId="1" pageNumber="248" year="2017">
<bibRefCitation id="5D784B5CFFD75B4EFCB9F87BFC70F87F" author="Ahn, K. - J. &amp; Cho, Y. B. &amp; Kim, Y. - H. &amp; Yoo, I. - S." box="[798,1016,2012,2034]" pageId="1" pageNumber="248" pagination="279 - 336" refId="ref11023" refString="Ahn, K. - J., Cho, Y. B., Kim, Y. - H. &amp; Yoo, I. - S. (2017) Checklist of the Staphylinidae (Coleoptera) in Korea. Journal of Asia- Pacific Biodiversity, 10 (3), 279 - 336." type="journal article" year="2017">
Ahn
<emphasis id="0B9DEABFFFD75B4EFCF7F87AFCF4F87F" box="[848,892,2012,2034]" italics="true" pageId="1" pageNumber="248">et al</emphasis>
., 2017: 292
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="6D966625FFD45B4DFF30FA3FFC69FA40" ID-DOI="http://doi.org/10.5281/zenodo.3769617" ID-Zenodo-Dep="3769617" httpUri="https://zenodo.org/record/3769617/files/figure.png" pageId="2" pageNumber="249" startId="2.[151,250,1432,1454]" targetBox="[336,1264,208,1376]" targetPageId="2">
<paragraph id="395636ADFFD45B4DFF30FA3FFC69FA40" blockId="2.[151,1436,1432,1485]" pageId="2" pageNumber="249">
<emphasis id="0B9DEABFFFD45B4DFF30FA3FFE91FA20" bold="true" box="[151,281,1432,1454]" pageId="2" pageNumber="249">FIGURE 1.</emphasis>
Habitus. A,
<taxonomicName id="FEE94D2EFFD45B4DFE09FA3EFD64FA23" authorityName=": Schulke &amp; Smetana" authorityYear="2015" box="[430,748,1432,1454]" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="249" phylum="Arthropoda" rank="species" species="aterrima" subGenus="Acrotona">
<emphasis id="0B9DEABFFFD45B4DFE09FA3EFD83FA23" box="[430,523,1433,1454]" italics="true" pageId="2" pageNumber="249">Acrotona</emphasis>
(
<emphasis id="0B9DEABFFFD45B4DFD87FA3EFDF5FA23" box="[544,637,1433,1454]" italics="true" pageId="2" pageNumber="249">Acrotona</emphasis>
)
<emphasis id="0B9DEABFFFD45B4DFD34FA3EFD64FA23" box="[659,748,1433,1454]" italics="true" pageId="2" pageNumber="249">aterrima</emphasis>
</taxonomicName>
; B,
<emphasis id="0B9DEABFFFD45B4DFC83FA3EFCBBFA23" box="[804,819,1433,1454]" italics="true" pageId="2" pageNumber="249">A</emphasis>
.
<taxonomicName id="FEE94D2EFFD45B4DFCE1FA3FFC44FA23" authority="Lee &amp; Ahn, 2018" authorityName="Lee &amp; Ahn" authorityYear="2018" box="[838,972,1432,1454]" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="249" phylum="Arthropoda" rank="species" species="koreana" status="sp. nov." subGenus="Acrotona">
(
<emphasis id="0B9DEABFFFD45B4DFCE8FA3EFCD6FA23" box="[847,862,1433,1454]" italics="true" pageId="2" pageNumber="249">A</emphasis>
.)
<emphasis id="0B9DEABFFFD45B4DFCDDFA3FFC44FA23" box="[890,972,1432,1454]" italics="true" pageId="2" pageNumber="249">koreana</emphasis>
</taxonomicName>
<emphasis id="0B9DEABFFFD45B4DFC7EFA3EFBB8FA23" bold="true" box="[985,1072,1433,1454]" pageId="2" pageNumber="249">
<taxonomicNameLabel id="10AE57C4FFD45B4DFC7EFA3EFBB8FA23" box="[985,1072,1433,1454]" pageId="2" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; C,
<emphasis id="0B9DEABFFFD45B4DFBCEFA3EFBF0FA23" box="[1129,1144,1433,1454]" italics="true" pageId="2" pageNumber="249">A</emphasis>
.
<taxonomicName id="FEE94D2EFFD45B4DFB2CFA3FFA91FA23" baseAuthorityName="Sharp" baseAuthorityYear="1888" box="[1163,1305,1432,1454]" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="249" phylum="Arthropoda" rank="species" species="lutulenta" subGenus="Acrotona">
(
<emphasis id="0B9DEABFFFD45B4DFB33FA3EFB2BFA23" box="[1172,1187,1433,1454]" italics="true" pageId="2" pageNumber="249">A</emphasis>
.)
<emphasis id="0B9DEABFFFD45B4DFB18FA3FFA91FA23" box="[1215,1305,1432,1454]" italics="true" pageId="2" pageNumber="249">lutulenta</emphasis>
</taxonomicName>
; D,
<emphasis id="0B9DEABFFFD45B4DFAF4FA3EFAEAFA23" box="[1363,1378,1433,1454]" italics="true" pageId="2" pageNumber="249">A</emphasis>
.
<taxonomicName id="FEE94D2EFFD45B4DFAD2FA3FFE97FA40" baseAuthorityName="Cameron" baseAuthorityYear="1933" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="249" phylum="Arthropoda" rank="species" species="pseudotenera" subGenus="Acrotona">
(
<emphasis id="0B9DEABFFFD45B4DFAD9FA3EFA05FA23" box="[1406,1421,1433,1454]" italics="true" pageId="2" pageNumber="249">A</emphasis>
.)
<emphasis id="0B9DEABFFFD45B4DFF30FA10FE97FA40" box="[151,287,1463,1485]" italics="true" pageId="2" pageNumber="249">pseudotenera</emphasis>
</taxonomicName>
; E,
<emphasis id="0B9DEABFFFD45B4DFEEEFA1FFED0FA40" box="[329,344,1464,1485]" italics="true" pageId="2" pageNumber="249">A</emphasis>
.
<taxonomicName id="FEE94D2EFFD45B4DFEC2FA10FE50FA40" baseAuthorityName="Brundin" baseAuthorityYear="1952" box="[357,472,1463,1485]" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="249" phylum="Arthropoda" rank="species" species="setaria" subGenus="Acrotona">
(
<emphasis id="0B9DEABFFFD45B4DFECAFA1FFEF4FA40" box="[365,380,1464,1485]" italics="true" pageId="2" pageNumber="249">A</emphasis>
.)
<emphasis id="0B9DEABFFFD45B4DFE35FA1FFE50FA40" box="[402,472,1464,1485]" italics="true" pageId="2" pageNumber="249">setaria</emphasis>
</taxonomicName>
; F,
<emphasis id="0B9DEABFFFD45B4DFE59FA1FFD85FA40" box="[510,525,1464,1485]" italics="true" pageId="2" pageNumber="249">A</emphasis>
.
<taxonomicName id="FEE94D2EFFD45B4DFDBDFA10FD39FA40" authorityName="Cameron" authorityYear="1939" box="[538,689,1463,1485]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="249" phylum="Arthropoda" rank="species" species="suspiciosa" subGenus="Acrotona">
(
<emphasis id="0B9DEABFFFD45B4DFD84FA1FFDBAFA40" box="[547,562,1464,1485]" italics="true" pageId="2" pageNumber="249">A</emphasis>
.)
<emphasis id="0B9DEABFFFD45B4DFDE0FA1FFD39FA40" box="[583,689,1464,1485]" italics="true" pageId="2" pageNumber="249">suspiciosa</emphasis>
</taxonomicName>
<taxonomicName id="FEE94D2EFFD45B4DFD10FA1FFCA8FA40" authorityName="Cameron" authorityYear="1939" box="[695,800,1464,1485]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="249" phylum="Arthropoda" rank="species" species="suspiciosa">
<emphasis id="0B9DEABFFFD45B4DFD10FA1FFCA8FA40" box="[695,800,1464,1485]" italics="true" pageId="2" pageNumber="249">suspiciosa</emphasis>
</taxonomicName>
; G,
<emphasis id="0B9DEABFFFD45B4DFCEDFA1FFCD1FA40" box="[842,857,1464,1485]" italics="true" pageId="2" pageNumber="249">A</emphasis>
.
<taxonomicName id="FEE94D2EFFD45B4DFCC1FA10FC53FA40" baseAuthorityName="Kraatz" baseAuthorityYear="1859" box="[870,987,1463,1485]" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="249" phylum="Arthropoda" rank="species" species="vicaria" subGenus="Acrotona">
(
<emphasis id="0B9DEABFFFD45B4DFCC9FA1FFCF5FA40" box="[878,893,1464,1485]" italics="true" pageId="2" pageNumber="249">A</emphasis>
.)
<emphasis id="0B9DEABFFFD45B4DFC34FA1FFC53FA40" box="[915,987,1464,1485]" italics="true" pageId="2" pageNumber="249">vicaria</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="71F36526FFD45B4CFF30FA53FE50FEE9" lastPageId="3" lastPageNumber="250" pageId="2" pageNumber="249" type="description">
<paragraph id="395636ADFFD45B4CFF30FA53FE50FEE9" blockId="2.[151,1437,1524,2017]" lastBlockId="3.[151,1437,151,717]" lastPageId="3" lastPageNumber="250" pageId="2" pageNumber="249">
<emphasis id="0B9DEABFFFD45B4DFF30FA53FEA1F980" bold="true" box="[151,297,1524,1549]" pageId="2" pageNumber="249">Description.</emphasis>
Length 2.0
<quantity id="FE119B48FFD45B4DFE1FFA53FD98F981" box="[440,528,1524,1549]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="249" unit="mm" value="2.5">2.5 mm</quantity>
. Body (
<figureCitation id="A1D22A28FFD45B4DFDC0FA53FD36F980" box="[615,702,1524,1549]" captionStart="FIGURE 1" captionStartId="2.[151,250,1432,1454]" captionTargetBox="[336,1264,208,1376]" captionTargetId="figure@2.[313,1273,195,1408]" captionTargetPageId="2" captionText="FIGURE 1. Habitus. A, Acrotona (Acrotona) aterrima; B, A. (A.) koreana sp. nov.; C, A. (A.) lutulenta; D, A. (A.) pseudotenera; E, A. (A.) setaria; F, A. (A.) suspiciosa suspiciosa; G, A. (A.) vicaria." figureDoi="http://doi.org/10.5281/zenodo.3769617" httpUri="https://zenodo.org/record/3769617/files/figure.png" pageId="2" pageNumber="249">Fig. 1A</figureCitation>
) surface glossy and densely pubescent, with fine microsculpture. Body dark brown to black; antennae, elytra and legs slightly paler than other parts.
<emphasis id="0B9DEABFFFD45B4DFBF8F9BFFB14F9BC" box="[1119,1180,1560,1585]" italics="true" pageId="2" pageNumber="249">Head</emphasis>
. Subcircular, about as wide as long, widest at middle, distinctly narrower than pronotum; eyes large and slightly prominent, about as long as temples; gular sutures moderately separated, diverged basally. Antennae long and slender, slightly longer than head and pronotum combined; antennomeres compactly close together, 13 elongate, 1 longest, 2 slightly longer than 3, 410 quadrate to subquadrate, 11 longer than wide, about as long as preceding two combined.
<emphasis id="0B9DEABFFFD45B4DFAE3F90FFA1DF94C" box="[1348,1429,1704,1729]" italics="true" pageId="2" pageNumber="249">Thorax</emphasis>
. Pronotum approximately 1.5 times as wide as long, widest in basal third to half, pubescence directed posteriorly in midline. Elytra slightly transverse, slightly wider than pronotum; elytron approximately 1.5 times as long as wide, pubescence directed posteriorly and postero-laterally; postero-lateral margin slightly sinuate; hind wings fully developed.
<emphasis id="0B9DEABFFFD45B4DFEBFF89DFEC5F8DC" box="[280,333,1850,1873]" italics="true" pageId="2" pageNumber="249">Legs</emphasis>
. Slender and long, with pubescence and macrosetae; tibiae with two spurs at apex; metatarsomere 1 about as long as 2.
<emphasis id="0B9DEABFFFD45B4DFE2AF8FBFE7FF8F8" box="[397,503,1884,1909]" italics="true" pageId="2" pageNumber="249">Abdomen</emphasis>
. Widest in segments IIIIV; surface glossy and densely pubescent, with narrowly imbricate microsculpture; macrochaetal arrangement 01-13-13-13-13; male tergite VIII with four macrosetae on each side of midline, posterior margin truncate; male sternite VIII with seven macrosetae on each side of midline, posterior margin convex, slightly round, with inconspicuous and long marginal setae; posterior margin of female tergite VIII similar to males; female sternite VIII round, with long and short marginal setae.
<emphasis id="0B9DEABFFFD55B4CFB6EFF30FABEFF3D" box="[1225,1334,151,176]" italics="true" pageId="3" pageNumber="250">Aedeagus</emphasis>
. Median lobe narrowly ovate and widest in basal third, apical process short and convergent at apex in ventral aspect (
<bibRefCitation id="5D784B5CFFD55B4CFF39FF78FEC3FF75" author="Brundin, L." box="[158,331,223,248]" pageId="3" pageNumber="250" pagination="93 - 145" refId="ref11202" refString="Brundin, L. (1952) Acrotona - Studien. (Gattung Atheta, Col., Staphylinidae). Entomologisk Tidskrift, 73, 93 - 145." type="journal article" year="1952">Brundin, 1952</bibRefCitation>
: figs. 4041;
<bibRefCitation id="5D784B5CFFD55B4CFE56FF78FDF1FF75" author="Palm, T." box="[497,633,223,248]" pageId="3" pageNumber="250" pagination="117 - 296" refId="ref12070" refString="Palm, T. (1970) Svensk Insektfauna: Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Underfam. Aleocharinae (Atheta). Hafte 6. Entomologiska Foreningen I, Stockholm, 180 pp. [pp. 117 - 296]" type="book chapter" year="1970">Palm, 1970</bibRefCitation>
: PL. XX, 194;
<bibRefCitation id="5D784B5CFFD55B4CFC91FF78FBCBFF75" author="Yosii, R. &amp; Sawada, K." box="[822,1091,223,248]" pageId="3" pageNumber="250" pagination="11 - 140" refId="ref12754" refString="Yosii, R. &amp; Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of genera and subgenera with description of representative species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140." type="journal article" year="1976">Yosii &amp; Sawada, 1976</bibRefCitation>
: figs. 39JL). Apical lobe of paramerites short, with four setae; a-seta slightly longer than b- and d-setae, c-seta short and close together (
<bibRefCitation id="5D784B5CFFD55B4CFAC7FEA3FEDCFECD" author="Yosii, R. &amp; Sawada, K." pageId="3" pageNumber="250" pagination="11 - 140" refId="ref12754" refString="Yosii, R. &amp; Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of genera and subgenera with description of representative species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140." type="journal article" year="1976">Yosii &amp; Sawada, 1976</bibRefCitation>
: figs. 39M).
<emphasis id="0B9DEABFFFD55B4CFE46FE80FDFBFECD" box="[481,627,295,320]" italics="true" pageId="3" pageNumber="250">Spermatheca</emphasis>
. Umbilicus absent; duct densely coiled (
<bibRefCitation id="5D784B5CFFD55B4CFB90FE80FB34FECD" author="Palm, T." box="[1079,1212,295,320]" pageId="3" pageNumber="250" pagination="117 - 296" refId="ref12070" refString="Palm, T. (1970) Svensk Insektfauna: Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Underfam. Aleocharinae (Atheta). Hafte 6. Entomologiska Foreningen I, Stockholm, 180 pp. [pp. 117 - 296]" type="book chapter" year="1970">Palm, 1970</bibRefCitation>
: PL. XX, 194;
<bibRefCitation id="5D784B5CFFD55B4CFAC5FE80FEDDFEE8" author="Yosii, R. &amp; Sawada, K." pageId="3" pageNumber="250" pagination="11 - 140" refId="ref12754" refString="Yosii, R. &amp; Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of genera and subgenera with description of representative species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140." type="journal article" year="1976">Yosii &amp; Sawada, 1976</bibRefCitation>
: figs. 39O).
</paragraph>
</subSubSection>
<subSubSection id="71F36526FFD55B4CFF60FEC8FCD6FDB1" pageId="3" pageNumber="250" type="materials_examined">
<paragraph id="395636ADFFD55B4CFF60FEC8FE3BFE79" blockId="3.[151,1437,151,717]" pageId="3" pageNumber="250">
<emphasis id="0B9DEABFFFD55B4CFF60FEC8FEF0FE05" bold="true" box="[199,376,367,392]" pageId="3" pageNumber="250">Type material.</emphasis>
<typeStatus id="E652880FFFD55B4CFE26FED7FE7CFE05" box="[385,500,368,392]" pageId="3" pageNumber="250" type="lectotype">Lectotype</typeStatus>
,
<specimenCount id="2FEFFD24FFD55B4CFDA5FED7FD91FE0A" box="[514,537,368,391]" pageId="3" pageNumber="250" type="male"></specimenCount>
, labeled as follows:
<collectingCountry id="41FE763DFFD55B4CFCAAFEC8FC09FE05" box="[781,897,367,392]" name="Germany" pageId="3" pageNumber="250">Germania</collectingCountry>
Nr. 5506,
<emphasis id="0B9DEABFFFD55B4CFBA7FED6FBB7FE05" box="[1024,1087,369,392]" italics="true" pageId="3" pageNumber="250">Typus</emphasis>
, Zool. Mus.
<collectingRegion id="FB2DF84FFFD55B4CFB7DFEC8FAABFE05" box="[1242,1315,367,392]" country="Germany" name="Berlin" pageId="3" pageNumber="250">Berlin</collectingRegion>
,
<collectionCode id="5FF8AE68FFD55B4CFA96FED7FAFFFE0A" box="[1329,1399,368,391]" pageId="3" pageNumber="250">TYPE</collectionCode>
<taxonomicName id="FEE94D2EFFD55B4CFA26FED7FEB3FE21" authority="Grav." authorityName="Grav." class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="250" phylum="Arthropoda" rank="species" species="aterrima">A. aterrima Grav.</taxonomicName>
LECTO-TYPE
<collectionCode id="5FF8AE68FFD55B4CFE51FE32FD82FE21" box="[502,522,405,428]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="250" type="Museum">V</collectionCode>
. Mahler des. 1986,
<taxonomicName id="FEE94D2EFFD55B4CFD48FE33FCFEFE21" authority="Er." authorityName="Er." box="[751,886,404,429]" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="250" phylum="Arthropoda" rank="species" species="aterrima">aterrima Er.</taxonomicName>
at. aterrima gr.,
<taxonomicName id="FEE94D2EFFD55B4CFB90FE33FEF7FE5D" authority="(Grav.) Det. Klimaszewski 2004" authorityName="Det. Klimaszewski" authorityYear="2004" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="250" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="0B9DEABFFFD55B4CFB90FE33FA82FE21" box="[1079,1290,404,428]" italics="true" pageId="3" pageNumber="250">Acrotona aterrima</emphasis>
(Grav.) Det. Klimaszewski 2004
</taxonomicName>
.
<typeStatus id="E652880FFFD55B4CFE37FE1FFE72FE5D" box="[400,506,440,464]" pageId="3" pageNumber="250" type="syntype">Syntypes</typeStatus>
, 4 exx.,
<collectingCountry id="41FE763DFFD55B4CFDF8FE10FD45FE5D" box="[607,717,439,464]" name="Germany" pageId="3" pageNumber="250">Germania</collectingCountry>
Nr. 5506,
<emphasis id="0B9DEABFFFD55B4CFCECFE1EFC02FE5D" box="[843,906,441,464]" italics="true" pageId="3" pageNumber="250">Typus</emphasis>
, Zool.Mus.
<collectingRegion id="FB2DF84FFFD55B4CFBBDFE10FBEBFE5D" box="[1050,1123,439,464]" country="Germany" name="Berlin" pageId="3" pageNumber="250">Berlin</collectingRegion>
,
<taxonomicName id="FEE94D2EFFD55B4CFBD7FE1FFE27FE79" authority="(Grav.) Det. Klimaszewski 2004" authorityName="Det. Klimaszewski" authorityYear="2004" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="250" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="0B9DEABFFFD55B4CFBD7FE1FFACBFE5D" box="[1136,1347,440,464]" italics="true" pageId="3" pageNumber="250">Acrotona aterrima</emphasis>
(Grav.) Det. Klimaszewski 2004
</taxonomicName>
.
</paragraph>
<paragraph id="395636ADFFD55B4CFF60FE58FCD6FDB1" blockId="3.[151,1437,151,717]" pageId="3" pageNumber="250">
<emphasis id="0B9DEABFFFD55B4CFF60FE58FE3AFD95" bold="true" box="[199,434,511,536]" pageId="3" pageNumber="250">Material examined.</emphasis>
<materialsCitation id="89813CF0FFD55B4CFE1CFE58FC3FFD95" ID-GBIF-Occurrence="2608960677" box="[443,951,511,536]" collectingDate="1983-09" country="North Korea" location="NORTH KOREA" pageId="3" pageNumber="250" specimenCount="1">
<emphasis id="0B9DEABFFFD55B4CFE1CFE58FD11FD95" bold="true" box="[443,665,511,536]" pageId="3" pageNumber="250">
<collectingCountry id="41FE763DFFD55B4CFE1CFE58FD1CFD95" box="[443,660,511,536]" name="North Korea" pageId="3" pageNumber="250">NORTH KOREA</collectingCountry>
:
</emphasis>
5 exx., Corea
<date id="4D57106DFFD55B4CFCE4FDA7FC3FFD95" box="[835,951,512,536]" pageId="3" pageNumber="250" value="1983-09">
<collectingDate id="5D13E985FFD55B4CFCE4FDA7FC3FFD95" box="[835,951,512,536]" pageId="3" pageNumber="250" value="1983-09">sept. 1983</collectingDate>
</date>
</materialsCitation>
<materialsCitation id="89813CF0FFD55B4CFC67FE58FCD2FDB1" ID-GBIF-Occurrence="2608960679" collectingDate="1983-09" collectionCode="ISEA" collectorName="Myohyang Mts. Exp. Inst. Zool. Cr." country="North Korea" location="Myohyang Mts." municipality="Mts. Exp. Inst. Zool. Cr." pageId="3" pageNumber="250" specimenCount="1">
<collectorName id="941C537BFFD55B4CFC67FE58FAC1FD95" box="[960,1353,511,536]" pageId="3" pageNumber="250">
<location id="3C366076FFD55B4CFC67FE58FBFDFD9A" LSID="urn:lsid:plazi:treatment:B14087BBFFD75B4CFF30F9FAFE86FD40:3C366076FFD55B4CFC67FE58FBFDFD9A" box="[960,1141,511,536]" country="North Korea" municipality="Mts. Exp. Inst. Zool. Cr." name="Myohyang Mts." pageId="3" pageNumber="250">
Myohyang
<collectingMunicipality id="D932ACD7FFD55B4CFBE5FDA7FBFDFD9A" box="[1090,1141,512,535]" pageId="3" pageNumber="250">Mts.</collectingMunicipality>
</location>
Exp. Inst.Zool.Cr.
</collectorName>
[North Korea, Pyeonganbuk Prov., Mt. Myohyangsan,
<date id="4D57106DFFD55B4CFD0BFD83FC8CFDB1" box="[684,772,548,573]" pageId="3" pageNumber="250" value="1983-09">
<collectingDate id="5D13E985FFD55B4CFD0BFD83FC8CFDB1" box="[684,772,548,573]" pageId="3" pageNumber="250" value="1983-09">ix 1983</collectingDate>
</date>
,
<collectionCode id="5FF8AE68FFD55B4CFCA8FD83FCDBFDB1" box="[783,851,548,572]" pageId="3" pageNumber="250">ISEA</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="71F36526FFD55B4CFF60FDE0FACFFDD2" box="[199,1351,583,608]" pageId="3" pageNumber="250" type="distribution">
<paragraph id="395636ADFFD55B4CFF60FDE0FACFFDD2" blockId="3.[151,1437,151,717]" box="[199,1351,583,608]" pageId="3" pageNumber="250">
<emphasis id="0B9DEABFFFD55B4CFF60FDE0FEE8FDED" bold="true" box="[199,352,583,608]" pageId="3" pageNumber="250">Distribution.</emphasis>
<collectingCountry id="41FE763DFFD55B4CFEC0FDEFFE23FDD2" box="[359,427,584,607]" name="South Korea" pageId="3" pageNumber="250">Korea</collectingCountry>
(North),
<collectingCountry id="41FE763DFFD55B4CFDB3FDE0FDD7FDED" box="[532,607,583,608]" name="Russia" pageId="3" pageNumber="250">Russia</collectingCountry>
(East Siberia, West Siberia),
<collectingCountry id="41FE763DFFD55B4CFC0EFDE0FBA7FDED" box="[937,1071,583,608]" name="Kazakhstan" pageId="3" pageNumber="250">Kazakhstan</collectingCountry>
,
<collectingCountry id="41FE763DFFD55B4CFB9EFDE0FB3FFDED" box="[1081,1207,583,608]" name="Uzbekistan" pageId="3" pageNumber="250">Uzbekistan</collectingCountry>
and Europe.
</paragraph>
</subSubSection>
<subSubSection id="71F36526FFD55B4CFF60FDCBFE86FD40" pageId="3" pageNumber="250" type="discussion">
<paragraph id="395636ADFFD55B4CFF60FDCBFE86FD40" blockId="3.[151,1437,151,717]" pageId="3" pageNumber="250">
<emphasis id="0B9DEABFFFD55B4CFF60FDCBFEB3FD08" bold="true" box="[199,315,620,645]" pageId="3" pageNumber="250">Remarks.</emphasis>
This species was recorded by
<bibRefCitation id="5D784B5CFFD55B4CFD06FDCBFCC8FD09" author="Pasnik, G." box="[673,832,620,645]" pageId="3" pageNumber="250" pagination="185 - 234" refId="ref12116" refString="Pasnik, G. (2001) The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeography. Acta Zoologica Cracoviensia, 44, 185 - 234." type="journal article" year="2001">Paśnik (2001)</bibRefCitation>
in
<collectingCountry id="41FE763DFFD55B4CFCCEFDCBFC70FD09" box="[873,1016,620,645]" name="North Korea" pageId="3" pageNumber="250">North Korea</collectingCountry>
but no specimen has been collected from
<collectingCountry id="41FE763DFFD55B4CFF7EFD28FEE6FD2A" box="[217,366,655,680]" name="South Korea" pageId="3" pageNumber="250">South Korea</collectingCountry>
. We could not describe the mouthparts in detail because specimens were not available for dissection.
</paragraph>
</subSubSection>
</treatment>
</document>