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<document id="7B0040B4585FBD552DBCD89A1998647A" ID-DOI="10.1206/602.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5389446" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630310581776" checkinUser="carolina" docAuthor="Grimaldi, David A. &amp; Triplehorn, Don M." docDate="2008" docId="A96E8799FFD8DE76198FE233F7ADFDBE" docLanguage="en" docName="N3612.pdf" docOrigin="American Museum Novitates 3612 (1)" docSource="http://www.bioone.org/perlserv/?request=get-abstract&amp;doi=10.1206%2F602.1" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.3:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="3" docTitle="Dasiops Rondani 1856" docType="treatment" docVersion="3" lastPageNumber="14" masterDocId="5557FFE1FFD3DE78193EE429F767FFD0" masterDocTitle="Insects from the Upper Miocene Grubstake Formation of Alaska" masterLastPageNumber="20" masterPageNumber="1" pageNumber="12" updateTime="1699222507036" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="22F31F3BBA0A8FC1C20A4CB6E5E5AACC">Insects from the Upper Miocene Grubstake Formation of Alaska</mods:title>
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<paragraph id="2178368FFFD8DE73198FE233F56BF9E2" blockId="11.[177,524,1562,1586]" box="[177,524,1562,1586]" pageId="11" pageNumber="12">
<heading id="7A3081E3FFD8DE73198FE233F56BF9E2" box="[177,524,1562,1586]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="12" reason="9">
Genus
<taxonomicName id="E6C74D0CFFD8DE73183DE233F56BF9E2" ID-CoL="62Q85" ID-ENA="286456" authority="Rondani, 1856" authorityName="Rondani" authorityYear="1856" box="[259,524,1562,1586]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="13B3EA9DFFD8DE73183DE233F63CF9E2" box="[259,347,1562,1586]" italics="true" pageId="11" pageNumber="12">Dasiops</emphasis>
Rondani, 1856
</taxonomicName>
</heading>
</paragraph>
<paragraph id="2178368FFFD8DE73194CE264F501F90A" blockId="11.[88,614,1613,1754]" pageId="11" pageNumber="12">
SPECIMENS: Specimens UAF-GS9 and UAF- GS26 (
<figureCitation id="B9FC2A0AFFD8DE731997E243F789F952" box="[169,238,1642,1666]" captionStart="Fig" captionStartId="12.[186,224,1412,1434]" captionTargetBox="[149,1207,210,1378]" captionTargetId="figure-138@12.[149,1207,210,1378]" captionTargetPageId="12" captionText="Fig. 9. Photomicrographs of Lonchaeidae (cf. Dasiops) wings. A. UAF-GS9. B. UAF-GS26." figureDoi="http://doi.org/10.5281/zenodo.5389478" httpUri="https://zenodo.org/record/5389478/files/figure.png" pageId="11" pageNumber="12">figs. 9</figureCitation>
,
<figureCitation id="B9FC2A0AFFD8DE73183EE243F67CF952" box="[256,283,1642,1666]" captionStart="Fig" captionStartId="13.[112,150,1376,1398]" captionTargetBox="[99,1176,210,1341]" captionTargetId="figure-128@13.[98,1177,210,1342]" captionTargetPageId="13" captionText="Fig. 10. Camera lucida drawings of Lonchaeidae (cf. Dasiops) wings. A. UAF-GS 9. B. UAF-GS26." figureDoi="http://doi.org/10.5281/zenodo.5389480" httpUri="https://zenodo.org/record/5389480/files/figure.png" pageId="11" pageNumber="12">10</figureCitation>
) are complete and well-preserved wings,
<quantity id="E63F9B6AFFD8DE7319C2E2AEF637F94F" box="[252,336,1671,1695]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="mm" value="3.5">3.5 mm</quantity>
and
<quantity id="E63F9B6AFFD8DE7318ABE2AEF68EF94F" box="[405,489,1671,1695]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="11" pageNumber="12" unit="mm" value="5.6">5.6 mm</quantity>
long. The wings are so well preserved that even the fine veins that comprise the tiny basal cells are intact.
</paragraph>
</subSubSection>
<subSubSection id="69DD6504FFD8DE761B8EE7D1F7ADFDBE" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12" type="discussion">
<paragraph id="2178368FFFD8DE741B8EE7D1F57EF950" blockId="11.[662,1188,1016,1756]" lastBlockId="12.[128,654,1462,1754]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
DISCUSSION: The general facies of the wing is typical of the higher (Cyclorrhaphan) flies, among which the wing venation is rather conservative. Some cyclorrhaphan families with extremely distinctive wing features can be immediately dismissed, such as many of the muscoids (the wing does not have a large calypter),
<taxonomicName id="E6C74D0CFFD8DE731A36E0E1F4C4FB30" box="[776,931,1224,1248]" class="Insecta" family="Agromyzidae" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Agromyzidae</taxonomicName>
,
<taxonomicName id="E6C74D0CFFD8DE731A91E0E1F327FB30" box="[943,1088,1224,1248]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
,
<taxonomicName id="E6C74D0CFFD8DE731D75E0E1F59FFB2E" authorityName="Macquart" authorityYear="1835" class="Insecta" family="Sphaeroceridae" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Sphaeroceridae</taxonomicName>
, many of the tephritoids, etc. Among all other North American cyclorrhaphan genera, these two wings bear closest resemblance to
<taxonomicName id="E6C74D0CFFD8DE731A3DE116F36EFA87" authorityName="Rondani" authorityYear="1856" box="[771,1033,1343,1367]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="13B3EA9DFFD8DE731A3DE116F43CFA87" box="[771,859,1343,1367]" italics="true" pageId="11" pageNumber="12">Dasiops</emphasis>
(Lonchaeidae)
</taxonomicName>
. This is based on: overall shape of the wing, the large subcostal cell, completeness and close proximity of veins Sc and R
<subScript id="BD4334CAFFD8DE731AA1E18AF4CEFA63" attach="left" box="[927,937,1443,1459]" fontSize="7" pageId="11" pageNumber="12">1</subScript>
, closed basal cells bm and cup, and a definite but incomplete vein A
<subScript id="BD4334CAFFD8DE731B94E1F7F5D3FA3E" attach="left" box="[682,692,1502,1518]" fontSize="7" pageId="11" pageNumber="12">1</subScript>
+CuA
<subScript id="BD4334CAFFD8DE731BC7E1F7F464FA3E" attach="left" box="[761,771,1502,1518]" fontSize="7" pageId="11" pageNumber="12">2</subScript>
. Lonchaeids in general have a rather well-developed calypter for acalyptrate flies, but it is possible that some of this region of the wing is not preserved in the fossils. The venation of the
<specimenCount id="37C1FD06FFD8DE731A5AE262F375F9B3" box="[868,1042,1611,1635]" pageId="11" pageNumber="12" type="generic">two specimens</specimenCount>
is virtually identical, the only (slight) differences being that the crossveins (r-m and m-cu) in UAF- GS26 are slightly closer to each other, the apex of R
<subScript id="BD4334CAFFD8DE731A30E2E5F44AF90C" attach="left" box="[782,813,1740,1756]" fontSize="7" pageId="11" pageNumber="12">4+5</subScript>
is slightly more curved toward the apex of M, and the trailing edge of the wing in the distal half is slightly narrower (the tip is narrower). The wing of UAF-GS26 is significantly larger than UAF-GS9 by some 60%, which is extreme for intraspecific variation in acalyptrate flies. Thus, there are probably two species of lonchaeids.
</paragraph>
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<paragraph id="2178368FFFDFDE741984E1ADF3EFFA4A" blockId="12.[186,1160,1412,1434]" box="[186,1160,1412,1434]" pageId="12" pageNumber="13">
Fig. 9. Photomicrographs of
<taxonomicName id="E6C74D0CFFDFDE7418C6E1ADF51DFA4A" authorityName="Rondani" authorityYear="1856" box="[504,634,1412,1434]" class="Insecta" family="Lonchaeidae" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Lonchaeidae</taxonomicName>
(cf.
<taxonomicName id="E6C74D0CFFDFDE741B95E1ACF59DFA4A" authorityName="Rondani" authorityYear="1856" box="[683,762,1413,1434]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="13B3EA9DFFDFDE741B95E1ACF59DFA4A" box="[683,762,1413,1434]" italics="true" pageId="12" pageNumber="13">Dasiops</emphasis>
</taxonomicName>
) wings.
<emphasis id="13B3EA9DFFDFDE741A6DE1ADF40CFA4A" bold="true" box="[851,875,1412,1434]" pageId="12" pageNumber="13">A.</emphasis>
UAF-GS9.
<emphasis id="13B3EA9DFFDFDE741AD2E1ADF365FA4A" bold="true" box="[1004,1026,1412,1434]" pageId="12" pageNumber="13">B.</emphasis>
UAF-GS26.
</paragraph>
</caption>
<paragraph id="2178368FFFDFDE7519A4E2AFF6C1F9B3" blockId="12.[128,654,1462,1754]" lastBlockId="13.[88,614,1433,1754]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
The only account for North America of the genus
<taxonomicName id="E6C74D0CFFDFDE7419F7E28CF646F96D" authorityName="Rondani" authorityYear="1856" box="[201,289,1701,1725]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="13B3EA9DFFDFDE7419F7E28CF646F96D" box="[201,289,1701,1725]" italics="true" pageId="12" pageNumber="13">Dasiops</emphasis>
</taxonomicName>
are descriptions of new species by
<bibRefCitation id="45564B7EFFDFDE74199BE2EBF60CF90A" author="McAlpine, J. F." box="[165,363,1730,1754]" pageId="12" pageNumber="13" pagination="661 - 700" refId="ref7418" refString="McAlpine, J. F. 1964. Descriptions of new Lonchaeidae (Diptera). I. Canadian Entomologist 96: 661 - 700." type="journal article" year="1964">McAlpine (1964)</bibRefCitation>
. The work unfortunately was not a revision, but some cursory distributional data can be gleaned from it.
<taxonomicName id="E6C74D0CFFDFDE741D67E1FDF3D6FA3C" authorityName="Rondani" authorityYear="1856" box="[1113,1201,1492,1516]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="13B3EA9DFFDFDE741D67E1FDF3D6FA3C" box="[1113,1201,1492,1516]" italics="true" pageId="12" pageNumber="13">Dasiops</emphasis>
</taxonomicName>
is distributed throughout North America with more than 50 species, some of which extend into the
<collectingRegion id="E303F86DFFDFDE741A0BE204F367F995" box="[821,1024,1581,1605]" country="Canada" name="Yukon" pageId="12" pageNumber="13">Yukon Territory</collectingRegion>
and
<collectingRegion id="E303F86DFFDFDE741D6DE204F452F9B3" country="Canada" name="Northwest Territories" pageId="12" pageNumber="13">Northwest Territories</collectingRegion>
of
<collectingCountry id="59D0761FFFDFDE741A46E262F4B3F9B3" box="[888,980,1611,1635]" name="Canada" pageId="12" pageNumber="13">Canada</collectingCountry>
. Seven species are recorded from Alaska:
<taxonomicName id="E6C74D0CFFDFDE741AD6E240F3A0F951" authority="McAlpine" authorityName="McAlpine" authorityYear="1964" box="[1000,1223,1641,1665]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="13B3EA9DFFDFDE741AD6E240F359F951" box="[1000,1086,1641,1665]" italics="true" pageId="12" pageNumber="13">borealis</emphasis>
McAlpine
</taxonomicName>
,
<taxonomicName id="E6C74D0CFFDFDE741B83E2AEF468F94F" authorityName="McAlpine" authorityYear="1964" box="[701,783,1671,1695]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="criddlei">
<emphasis id="13B3EA9DFFDFDE741B83E2AEF468F94F" box="[701,783,1671,1695]" italics="true" pageId="12" pageNumber="13">criddlei</emphasis>
</taxonomicName>
McA.,
<emphasis id="13B3EA9DFFDFDE741AB4E2AEF4BBF94F" box="[906,988,1671,1695]" italics="true" pageId="12" pageNumber="13">lineelus</emphasis>
McA.,
<taxonomicName id="E6C74D0CFFDFDE741D69E2AEF3ACF94F" authorityName="McAlpine" authorityYear="1964" box="[1111,1227,1671,1695]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="anepsiosus">
<emphasis id="13B3EA9DFFDFDE741D69E2AEF3ACF94F" box="[1111,1227,1671,1695]" italics="true" pageId="12" pageNumber="13">anepsiosus</emphasis>
</taxonomicName>
McA.,
<taxonomicName id="E6C74D0CFFDFDE741A22E28DF4EBF96C" authorityName="McAlpine" authorityYear="1964" box="[796,908,1700,1724]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="hastulatus">
<emphasis id="13B3EA9DFFDFDE741A22E28DF4EBF96C" box="[796,908,1700,1724]" italics="true" pageId="12" pageNumber="13">hastulatus</emphasis>
</taxonomicName>
McA., and
<taxonomicName id="E6C74D0CFFDFDE741D7CE28DF3ABF96C" authorityName="McAlpine" authorityYear="1964" box="[1090,1228,1700,1724]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="penealbiceps">
<emphasis id="13B3EA9DFFDFDE741D7CE28DF3ABF96C" box="[1090,1228,1700,1724]" italics="true" pageId="12" pageNumber="13">penealbiceps</emphasis>
</taxonomicName>
McA. (the last three in the
<taxonomicName id="E6C74D0CFFDFDE741D2DE2EBF30CF90A" baseAuthorityName="Malloch" baseAuthorityYear="1914" box="[1043,1131,1730,1754]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="albiceps">
<emphasis id="13B3EA9DFFDFDE741D2DE2EBF30CF90A" box="[1043,1131,1730,1754]" italics="true" pageId="12" pageNumber="13">albiceps</emphasis>
</taxonomicName>
group). Species of the genus are not differentiated on the basis of wing venation. According to McAlpine, many species of
<taxonomicName id="E6C74D0CFFDEDE751883E1FDF572FA3C" authorityName="Rondani" authorityYear="1856" box="[445,533,1492,1516]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="13B3EA9DFFDEDE751883E1FDF572FA3C" box="[445,533,1492,1516]" italics="true" pageId="13" pageNumber="14">Dasiops</emphasis>
</taxonomicName>
breed under the bark of dead or dying conifer trees, usually those attacked by bark beetles (
<taxonomicName id="E6C74D0CFFDEDE751961E204F662F995" authorityName="Latreille" authorityYear="1802" box="[95,261,1581,1605]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
:
<taxonomicName id="E6C74D0CFFDEDE75182AE204F6E9F995" box="[276,398,1581,1605]" class="Insecta" family="Scolytinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Scolytinae</taxonomicName>
). Biogeography of this genus is similar to
<taxonomicName id="E6C74D0CFFDEDE75185AE262F6F8F9B3" authorityName="Geoffrey" authorityYear="1762" box="[356,415,1611,1635]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="13B3EA9DFFDEDE75185AE262F6F8F9B3" box="[356,415,1611,1635]" italics="true" pageId="13" pageNumber="14">Bibio</emphasis>
</taxonomicName>
.
</paragraph>
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<paragraph id="2178368FFFDEDE75194EE149F3E8FAA6" blockId="13.[112,1167,1376,1398]" box="[112,1167,1376,1398]" pageId="13" pageNumber="14">
Fig. 10.
<taxonomicName id="E6C74D0CFFDEDE7519E9E149F60AFAA6" box="[215,365,1376,1398]" class="Insecta" family="Ichneumonidae" genus="Camera" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lucida">Camera lucida</taxonomicName>
drawings of
<taxonomicName id="E6C74D0CFFDEDE7518C8E149F51FFAA6" authorityName="Rondani" authorityYear="1856" box="[502,632,1376,1398]" class="Insecta" family="Lonchaeidae" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Lonchaeidae</taxonomicName>
(cf.
<taxonomicName id="E6C74D0CFFDEDE751B97E148F59FFAA6" authorityName="Rondani" authorityYear="1856" box="[681,760,1377,1398]" class="Insecta" family="Lonchaeidae" genus="Dasiops" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="13B3EA9DFFDEDE751B97E148F59FFAA6" box="[681,760,1377,1398]" italics="true" pageId="13" pageNumber="14">Dasiops</emphasis>
</taxonomicName>
) wings.
<emphasis id="13B3EA9DFFDEDE751A6FE149F40EFAA6" bold="true" box="[849,873,1376,1398]" pageId="13" pageNumber="14">A.</emphasis>
UAF-GS 9.
<emphasis id="13B3EA9DFFDEDE751ACCE149F36FFAA6" bold="true" box="[1010,1032,1376,1398]" pageId="13" pageNumber="14">B.</emphasis>
UAF-GS26.
</paragraph>
</caption>
<paragraph id="2178368FFFDEDE75194CE240F33AFA3C" blockId="13.[88,614,1433,1754]" lastBlockId="13.[662,1188,1433,1516]" pageId="13" pageNumber="14">
The only other, putative fossil lonchaeids to have been described are in Baltic amber (
<bibRefCitation id="45564B7EFFDEDE751961E28DF79EF96C" author="Hennig, W." box="[95,249,1700,1724]" pageId="13" pageNumber="14" pagination="1 - 51" refId="ref6902" refString="Hennig, W. 1967. Neue Acalyptratae aus dem baltischen Bernstein (Diptera: Cyclorrhapha). Stuttgarter Beitrage zur Naturkunde 174: 1 - 51." type="journal article" year="1967">Hennig, 1967</bibRefCitation>
),
<emphasis id="13B3EA9DFFDEDE751833E28CF56CF96D" box="[269,523,1701,1725]" italics="true" pageId="13" pageNumber="14">Glaesolonchea electrica</emphasis>
Hennig and
<taxonomicName id="E6C74D0CFFDEDE7519B5E2EBF542F90A" authority="Hennig. However" authorityName="Hennig. However" authorityYear="1967" box="[139,549,1730,1754]" class="Insecta" family="Pallopteridae" genus="Morgea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mcalpinei">
<emphasis id="13B3EA9DFFDEDE7519B5E2EBF636F90A" box="[139,337,1730,1754]" italics="true" pageId="13" pageNumber="14">Morgea mcalpinei</emphasis>
Hennig. However
</taxonomicName>
, both have been found to belong to the
<taxonomicName id="E6C74D0CFFDEDE751D2EE1B0F3C3FA61" authorityName="Loew" authorityYear="1862" box="[1040,1188,1433,1457]" class="Insecta" family="Pallopteridae" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Pallopteridae</taxonomicName>
(
<bibRefCitation id="45564B7EFFDEDE751BA3E19EF455FA1F" author="Morge, G." box="[669,818,1463,1487]" pageId="13" pageNumber="14" pagination="141 - 231" refId="ref7723" refString="Morge, G. 1967. Die Lonchaeidae und Pallopteridae Osterreiches und der Angrezenden Gebiete. Part II. Die Pallopteridae. Naturkunde Jahrbuche der Stadt Linz 1967: 141 - 231." type="journal article" year="1967">Morge, 1967</bibRefCitation>
;
<bibRefCitation id="45564B7EFFDEDE751A7FE19EF49CFA1F" author="McAlpine, J. F." box="[833,1019,1463,1487]" pageId="13" pageNumber="14" pagination="81 - 91" refId="ref7444" refString="McAlpine, J. F. 1981. Morgea friedbergi new species, a living sister-species of the fossil species M. mcalpinei Hennig, and a key to world genera of Pallopteridae (Diptera). Canadian Entomologist 113: 81 - 91." type="journal article" year="1981">McAlpine, 1981</bibRefCitation>
). The Alaskan fossil would be the only fossil lonchaeid.
</paragraph>
<paragraph id="2178368FFFDEDE751B83E219F31BF998" blockId="13.[701,1148,1584,1608]" box="[701,1148,1584,1608]" pageId="13" pageNumber="14">
Order
<taxonomicName id="E6C74D0CFFDEDE751A37E219F31BF998" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[777,1148,1584,1608]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="order">COLEOPTERA Linnaeus, 1758</taxonomicName>
</paragraph>
<paragraph id="2178368FFFDEDE761B8EE240F7ADFDBE" blockId="13.[662,1188,1641,1754]" lastBlockId="14.[128,654,211,622]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Two taxa of beetles are represented only by a specimen each and for which only the elytra remain (
<figureCitation id="B9FC2A0AFFDEDE751BCAE28DF427F96C" box="[756,832,1700,1724]" captionStart="Fig" captionStartId="15.[112,150,1565,1587]" captionTargetBox="[109,1167,210,1531]" captionTargetId="figure-0@15.[109,1167,210,1531]" captionTargetPageId="15" captionText="Fig. 11. Photomicrographs of beetle elytra. A. UAF-GS33: family Curculionidae. B. UAF-GS34: family Carabidae. (Both to the same scale.)" figureDoi="http://doi.org/10.5281/zenodo.5389484" httpUri="https://zenodo.org/record/5389484/files/figure.png" pageId="13" pageNumber="14">fig. 11</figureCitation>
). They are both approximately
<quantity id="E63F9B6AFFDEDE751BA8E2EBF5BFF90A" box="[662,728,1730,1754]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="mm" value="5.0">5 mm</quantity>
in length and have well-developed striae, but the elytra are distinctly different in structure and thus indicative of two distinct taxa. UAF-GS33 consists of both elytra, which strongly taper to an apical point; the elytra have significant relief, suggesting that the beetles body was somewhat dome-shaped. Specimen UAF-GS34 consists of just the left elytron, lateral and medial margins of which are parallel to nearly so, and the elytron is quite flat. Specimen UAF-GS33 is a weevil (family
<taxonomicName id="E6C74D0CFFDDDE7619E0E5D5F6E7FDC4" authorityName="Latreille" authorityYear="1802" box="[222,384,508,532]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
) and UAF-GS34 is a ground beetle (family
<taxonomicName id="E6C74D0CFFDDDE7618B6E633F698FDE2" authorityName="Latreille" authorityYear="1802" box="[392,511,538,562]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
) (identifications courtesy of S. A. Elias, personal commun.).
</paragraph>
</subSubSection>
</treatment>
</document>