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<document id="F262CE5D4974C233CB2CCCA03225AEEB" ID-CLB-Dataset="34190" ID-DOI="http://dx.doi.org/10.3897/zookeys.555.6011" ID-GBIF-Dataset="ebc8fa61-040b-464e-aef4-5efa9dc3c820" ID-PMC="PMC4740823" ID-Pensoft-Pub="1313-2970-555-91" ID-PubMed="26877688" ID-ZooBank="E1446FEB806142E79C952CD6E56E6E9C" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-555-91" ModsDocOrigin="ZooKeys 555" ModsDocTitle="A new species of Liolaemus related to L.nigroviridis from the Andean highlands of Central Chile (Iguania, Liolaemidae)" checkinTime="1453318032677" checkinUser="pensoft" docAuthor="Troncoso-Palacios, Jaime, Elorza, Alvaro A., Puas, German I. &amp; Alfaro-Pardo, Edmundo" docDate="2016" docId="A47F38B704327B1FA8E6D9E99C525368" docLanguage="en" docName="ZooKeys 555: 91-114" docOrigin="ZooKeys 555" docSource="http://dx.doi.org/10.3897/zookeys.555.6011" docTitle="Liolaemus uniformis Troncoso-Palacios, Elorza, Puas &amp; Alfaro-Pardo, 2016, sp. n." docType="treatment" docUuid="B412BEF2-C337-4472-A4CE-9AFD73876B07" docUuidSource="ZooBank" docVersion="7" lastPageNumber="101" masterDocId="FFD1030AFF8C83074F38FFFCFF9BFFE7" masterDocTitle="A new species of Liolaemus related to L. nigroviridis from the Andean highlands of Central Chile (Iguania, Liolaemidae)" masterLastPageNumber="114" masterPageNumber="91" pageNumber="95" updateTime="1701383682364" updateUser="ExternalLinkService">
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<mods:title id="51B498D72D2CED040A31A08B4A53ED45">A new species of Liolaemus related to L. nigroviridis from the Andean highlands of Central Chile (Iguania, Liolaemidae)</mods:title>
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<mods:namePart id="8F4690188288728708D354A8EDE10230">Troncoso-Palacios, Jaime</mods:namePart>
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<mods:namePart id="01AFDB0062D5D3B67AC6DACA867DA5FC">Elorza, Alvaro A.</mods:namePart>
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<mods:namePart id="140B625394C7C8C094EBBEF8E7152CD8">Puas, German I.</mods:namePart>
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<mods:namePart id="8C661429F43E05FE81A52EE352BA45C4">Alfaro-Pardo, Edmundo</mods:namePart>
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<mods:date id="64AD44726F5424BF2D2B853A45E569F2">2016</mods:date>
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<treatment id="A47F38B704327B1FA8E6D9E99C525368" ID-GBIF-Taxon="127888038" LSID="urn:lsid:zoobank.org:act:B412BEF2-C337-4472-A4CE-9AFD73876B07" httpUri="http://treatment.plazi.org/id/A47F38B704327B1FA8E6D9E99C525368" lastPageId="11" lastPageNumber="101" pageId="4" pageNumber="95">
<subSubSection id="B53A00951617D466663B7A375753A70E" pageId="4" pageNumber="95" type="multiple">
<paragraph id="53F6C93F81BD57B84D7E635CB1C25A87" pageId="4" pageNumber="95">
<pageBreakToken id="31B5DE63F4FAE43596393FD68DE7D0FA" pageId="4" pageNumber="95" start="start">Taxon</pageBreakToken>
classification Animalia ORDO FAMILIA
</paragraph>
</subSubSection>
<subSubSection id="E07F683D01AAAC601095C0D67BDDAE15" pageId="4" pageNumber="95" type="nomenclature">
<paragraph id="4032E746CBB62921C3FD10599D1A7C18" pageId="4" pageNumber="95">
<taxonomicName id="F0383FFA975C0A16BD3541DEBAAE25B3" ID-CoL="88K92" LSID="http://zoobank.org/B412BEF2-C337-4472-A4CE-9AFD73876B07" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
<taxonomicNameLabel id="A9167A92784B56F42846BE4D3C7A97C2" pageId="4" pageNumber="95">sp. n.</taxonomicNameLabel>
Fig. 2A, B
</paragraph>
</subSubSection>
<subSubSection id="9216DB805DAA34C955FCD4F24366E6D8" pageId="4" pageNumber="95" type="reference_group">
<paragraph id="3F85903ED5F9BFD97681C7EA4345289E" pageId="4" pageNumber="95">
<taxonomicName id="EE12F6FAB43A173275FB7870B306A5DB" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus altissimus" pageId="4" pageNumber="95" phylum="Chordata" rank="subspecies" species="altissimus" subspecies="altissimus">Liolaemus altissimus altissimus</taxonomicName>
(in part?),
<bibRefCitation id="595F83689B69AF60C46B836211E8D111" author="Mella, JE" editor="Penaloza, APG" journalOrPublisher="Ediciones del Centro de Ecologia Aplicada Ltda" pageId="16" pageNumber="107" title="Guia de Campo Reptiles de Chile: Zona Central" year="2005">Mella. 2005</bibRefCitation>
,
<normalizedToken id="5F925D3CAE1E8A66FD1C1AB22A01F213" originalValue="Guía">Guia</normalizedToken>
Camp. Rep. Chil. Zon. Cent., p. 38.
</paragraph>
<paragraph id="D86EA5875860C8B3E937E547B760A7C7" pageId="4" pageNumber="95">
<taxonomicName id="F0C0981D2EA173B8ECFDAC801D884B80" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus monticola" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="monticola">Liolaemus monticola</taxonomicName>
?,
<bibRefCitation id="E5FA48156D39E29943D952C4E3914263" author="Nunez, H" journalOrPublisher="Boleti ́ n del Museo Nacional de Historia Natural de Chile" pageId="16" pageNumber="107" pagination="41 - 74" title="Nuevas especies de Phymaturus (grupo palluma) para la zona Cordillerana Central de Chile (Reptilia, Sauria, Liolaemidae)." volume="59" year="2010">
<normalizedToken id="EDC6C7C9371425F27CD6CB7893F2CC31" originalValue="Núñez">Nunez</normalizedToken>
et al. 2010
</bibRefCitation>
, Bol. Mus. Nac. Hist. Nat., p. 57.
</paragraph>
</subSubSection>
<subSubSection id="CD17FAF5248FA3D48238E0FE7F5BB71B" pageId="4" pageNumber="95" type="holotype">
<paragraph id="6BDBCD1F893FC6C06F9EE23C9EE7307E" pageId="4" pageNumber="95">Holotype.</paragraph>
<paragraph id="26CC6E7E28379229583FB253ACC5D098" pageId="4" pageNumber="95">
SSUC Re 674. Adult male. Collected in the west shore of the Chepical Lagoon (
<geoCoordinate id="490129714D1B39284701A6FA8F1DA066" direction="south" orientation="latitude" precision="925" value="-32.25">32°15'S</geoCoordinate>
-
<geoCoordinate id="440973D2C784F21DA050863A39B66F04" direction="west" orientation="longitude" precision="925" value="-70.5">70°30'W</geoCoordinate>
), approximately 30 km NE Alicahue, San Felipe de Aconcagua Province,
<normalizedToken id="5E97C23F26B1C66E4C48E73C2FDCDC3B" originalValue="Valparaíso">Valparaiso</normalizedToken>
Region, Chile. Collectors: J. Troncoso-Palacios and E. Alfaro. December, 2012.
</paragraph>
</subSubSection>
<subSubSection id="E89DC49AC435AA843906B897744D1F62" pageId="4" pageNumber="95" type="paratypes">
<paragraph id="B837A64DF4C37EEE66817AF0F8494A86" pageId="4" pageNumber="95">Paratypes</paragraph>
<paragraph id="A1E9D00D7FE98C87C28F1CB723F0B19E" pageId="4" pageNumber="95">(Fig. 2C, D, E, F). SSUC Re 675, male. SSUC Re 676-78, three females. SSUC Re 679, juvenile. The same data as the holotype.</paragraph>
<caption id="594A359AE2075340DA5210F5AF23FE12" pageId="4" pageNumber="95">
<paragraph id="1133A8B392F4F7F37667221C41F4EADE" pageId="4" pageNumber="95">
Figure 2.
<taxonomicName id="624B17D2025822EC244E002A2771BF1F" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
sp. n. A, B Holotype, male C, D Paratype, female E Paratype, male F Paratype, juvenile (unknown sex). All from the type locality.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="EF06D44A81EBA63DBFD1DC1F1A4E7EF4" pageId="4" pageNumber="95" type="etymology">
<paragraph id="A6E8B66D6B4234D3C4E738103AE9DF27" pageId="4" pageNumber="95">Etymology.</paragraph>
<paragraph id="6F0828A5789B7C22101713886629723D" pageId="4" pageNumber="95">
The species name &quot;
<taxonomicName id="D03AAD5E14F90287C16AD725A98F0A67" lsidName="uniformis" pageId="4" pageNumber="95" rank="species" species="uniformis">uniformis</taxonomicName>
&quot; (Latin) refers to the lack of dorsal pattern and uniform color found for both males and females.
</paragraph>
</subSubSection>
<subSubSection id="F3BAB14B841002BE1D33EBA1BF334E65" lastPageId="8" lastPageNumber="99" pageId="4" pageNumber="95" type="diagnosis">
<paragraph id="E7C96AC642288CC8CAEE34F9E6D2C7A2" pageId="4" pageNumber="95">Diagnosis.</paragraph>
<paragraph id="5EAC2D3C0D9083A438C60BA8655771A6" pageId="4" pageNumber="95">
<taxonomicName id="1E0CD057873ED3090E4DF652A8997441" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
is larger than
<taxonomicName id="FFED1689AE546B0C2011D2FDF29E0755" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus constanzae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="constanzae">Liolaemus constanzae</taxonomicName>
(
<normalizedToken id="9FB564BABD56B94F0E99F2F8EF970F6D" originalValue="Mann–Whitney">Mann-Whitney</normalizedToken>
U = 0.5, P &lt;0.01, Table 1).
<taxonomicName id="7052CBB462B9E6F3A5C107CC61D5B484" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus constanzae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="constanzae">Liolaemus constanzae</taxonomicName>
has sexual dichromatism, a feature absent in
<taxonomicName id="E747EFE02F5FDA04DAB8F8F5B1C10507" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
. Males of
<taxonomicName id="76D757633B2E7DD11740E56E5C091A1B" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus constanzae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="constanzae">Liolaemus constanzae</taxonomicName>
have a black vertebral line and black spots on the paravertebral fields (Fig. 3A), whereas
<taxonomicName id="37C890EF2077D69A9FB7781ACB3E5065" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
has no dorsal pattern. Additionally, the southern distributional limit of
<taxonomicName id="C125848F2887ABE68510BC66E0398A05" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus constanzae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="constanzae">Liolaemus constanzae</taxonomicName>
in Agua Verde, Antofagasta Region, Chile (
<bibRefCitation id="81A9FAA03247744D4DA0855AF10C1E5F" author="Ortiz, JC" journalOrPublisher="Anales del Museo de Historia Natural de Valparaiso" pageId="16" pageNumber="107" pagination="62 - 65" title="Una nueva especie de lagartija chilena, Liolaemusdonosoi sp. nov. (Squamata-Iguanidae)." volume="8" year="1975">Ortiz 1975</bibRefCitation>
), is more than 750 km north of the type locality recorded for
<taxonomicName id="5A2B27350E41DCE62B4ABBB7FACFA542" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
.
</paragraph>
<caption id="AD3D529B56A78C46989056BF94D050C9" pageId="4" pageNumber="95">
<paragraph id="8C6859A73F2E93DF6379B637DD7306B9" pageId="4" pageNumber="95">
Figure 3. Chilean species of the
<taxonomicName id="D67E2A7B3FDF895353E71B0A6D395DD0" lsidName="nigroviridis" pageId="4" pageNumber="95" rank="species" species="nigroviridis">nigroviridis</taxonomicName>
group (with the exception of
<taxonomicName id="E33DDD9EBC97874EBCDAB7B59A335F66" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
), ordered from north to south. A
<taxonomicName id="BE10EF66508A638ED3EAB1DCCB8626C7" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus constanzae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="constanzae">Liolaemus constanzae</taxonomicName>
, male from vicinity of San Pedro (picture by JTP) B
<taxonomicName id="BEA7996CD8A5D27A0A5370BA53B83021" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus melanopleurus" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="melanopleurus">Liolaemus melanopleurus</taxonomicName>
, male from Atacama (picture by JTP) C
<taxonomicName id="7ADC4C4B456862C91EA35F812A4C7C11" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus isabelae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="isabelae">Liolaemus isabelae</taxonomicName>
, male from
<normalizedToken id="9EA4371E2805E03B7767BCEA40CAA176" originalValue="Montandón">Montandon</normalizedToken>
(picture by JTP) D
<taxonomicName id="BC28AF6C9A0EA8865148BF486A224303" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus juanortizi" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="juanortizi">Liolaemus juanortizi</taxonomicName>
, unknown sex specimen from road to Negro Francisco (picture by F. de Grotee) E
<taxonomicName id="7C0C2CA2226953C28CE9833EC4250865" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus lorenzmuelleri" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="lorenzmuelleri">Liolaemus lorenzmuelleri</taxonomicName>
, unknown sex specimen from Embalse La Laguna (picture by A. Labra) F
<taxonomicName id="F44EE5B3A5E5DC77C90E615DFD973582" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus maldonadae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="maldonadae">Liolaemus maldonadae</taxonomicName>
, male from vicinity of Alcohuaz (picture by JTP).
</paragraph>
</caption>
<caption id="D84B5C4D005E004459584172FF42AF26" pageId="4" pageNumber="95">
<paragraph id="5A0802DB9E91A29E1E1AE67E60E9FB1C" pageId="4" pageNumber="95">
Table 1. Scalation and morphological characteristics for the species of the
<taxonomicName id="7AFF1B4D09E4D44F860309D73F8A83D5" lsidName="nigroviridis" pageId="4" pageNumber="95" rank="species" species="nigroviridis">nigroviridis</taxonomicName>
group. Juvenile specimens examined are excluded. M = males; F = females. (*) Taken from
<bibRefCitation id="826783095406FD0C2FC7FCC901BFCD89" author="Navarro, J" journalOrPublisher="Boletin del Museo Nacional de Historia Natural de Chile" pageId="16" pageNumber="107" pagination="99 - 113" title="Liolaemuspatriciaturrae y Liolaemusisabelae, dos nuevas especies de lagartijas para el norte de Chile. Aspectos biogeograficos y citotaxonomicos (Squamata, Tropiduridae)." volume="44" year="1993">
Navarro and
<normalizedToken id="F1BDAF85264FF9AB6D7F4C3A2F17368E" originalValue="Núñez">Nunez</normalizedToken>
(1993)
</bibRefCitation>
. (**) Examined specimen plus Young-Downey and Moreno (1991) data. (***) Taken from Young-Downey and Moreno (1991). (****) Counted only for one specimen.
</paragraph>
</caption>
<paragraph id="A7D28DE2B5EB7DE6E880D159950BFA90" pageId="4" pageNumber="95">
<table id="B102285C33CB15256204948D28276887" pageId="4" pageNumber="95">
<tr id="4833175ABB9D2A968A1DC14AE4C613D7" pageId="4" pageNumber="95">
<th id="6C2EAD54B641B1239879DB11C2C3CF40" colspan="1" pageId="4" pageNumber="95" rowspan="1">
<taxonomicName id="DE3B74A33F5252C600207725361705E7" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus constanzae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="constanzae">Liolaemus constanzae</taxonomicName>
(M = 14, F = 13)
</th>
<th id="93721C838D86A117AFF51FB614F8E278" colspan="1" pageId="4" pageNumber="95" rowspan="1">
<taxonomicName id="181D4DDC103AC5D4FF09737A22838C69" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus isabelae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="isabelae">Liolaemus isabelae</taxonomicName>
(M = 4)
</th>
<th id="ED0ED82621A6E1F22ADD93670E19F89C" colspan="1" pageId="4" pageNumber="95" rowspan="1">
<taxonomicName id="C6E9C9F2938F763BAAB45F73C413075B" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus juanortizi" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="juanortizi">Liolaemus juanortizi</taxonomicName>
(M = 1)
</th>
<th id="AB9627B49EB21237A8AED091BF1EBEB4" colspan="1" pageId="4" pageNumber="95" rowspan="1">
<taxonomicName id="0187CE0769BCC7E6DFD6B216CBFABD87" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus lorenzmuelleri" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="lorenzmuelleri">Liolaemus lorenzmuelleri</taxonomicName>
(M = 3, F = 5)
</th>
<th id="996A5D0CD0E119C385023ECD77626B62" colspan="1" pageId="4" pageNumber="95" rowspan="1">
<taxonomicName id="F1A1463644EA346B20234D3E749B69F4" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus maldonadae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="maldonadae">Liolaemus maldonadae</taxonomicName>
(M = 3)
</th>
<th id="3F654F9A624071B2F3D2A17A70109F53" colspan="1" pageId="4" pageNumber="95" rowspan="1">
<taxonomicName id="DE567809434396B40BA4847F5289D11F" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus melanopleurus" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="melanopleurus">Liolaemus melanopleurus</taxonomicName>
(M = 2)
</th>
<th id="D915F13BAB38ED2989DEC29210F57120" colspan="1" pageId="4" pageNumber="95" rowspan="1">
<taxonomicName id="5E7F589ED5FA8196E0206A3FF8D74BA5" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
(M = 9, F = 4)
</th>
<th id="5791A5CF66E83C20921902AF3F92FDA6" colspan="1" pageId="4" pageNumber="95" rowspan="1">
<taxonomicName id="73870F86062F5A61DF92AC0861B91F63" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
sp. n. (M = 2, F = 3)
</th>
</tr>
<tr id="F3DB4684DC59A118D137D743699415D1" pageId="4" pageNumber="95">
<td id="8A7B801ECFD490D3ECA962BEBE0B4589" colspan="1" pageId="4" pageNumber="95" rowspan="1">SVL</td>
</tr>
</table>
</paragraph>
<paragraph id="8DDE0F5228A8305DBEBC87034C84C007" pageId="4" pageNumber="95">
<taxonomicName id="6AFEF8B1D7D1B537D696BC6853034AB0" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
differs from
<taxonomicName id="CE83E77AC9986B93AB86498E3CDB3C17" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus isabelae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="isabelae">Liolaemus isabelae</taxonomicName>
(Fig. 3C), because in the latter the nasal and the rostral scales are in contact only in 25% of specimens, whereas in
<taxonomicName id="E01B62EA94A0907B40A8BD9BFBD20910" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
, these scales are always in contact. Males of
<taxonomicName id="84CC5E3217C4A29D4515B7F02FC9FB75" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus isabelae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="isabelae">Liolaemus isabelae</taxonomicName>
have black ventral coloration, a yellow dorsal color with a black vertebral line, black bars in the paravertebral fields, and a black lateral band, or some males have a completely black dorsal color; all traits that are absent in
<taxonomicName id="07071D2E5BD94218A1548B868BA5988F" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
. Additionally, the southern distributional limit of
<taxonomicName id="1A81A9D74CCD72CB9AAFE47F76C23755" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus isabelae" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="isabelae">Liolaemus isabelae</taxonomicName>
in Salar de Pedernales, Atacama Region, Chile (
<bibRefCitation id="B8F380AD84DE10D695D80F316EF35B71" author="Pincheira-Donoso, D" journalOrPublisher="Publicacion Ocasional del Museo Nacional de Historia Natural de Chile" pageId="16" pageNumber="107" pagination="7 - 486" title="Las especies chilenas del genero Liolaemus (IguanidaeTropiduridae, Liolaeminae). Taxonomia, sistematica y evolucion." volume="59" year="2005">
Pincheira-Donoso and
<normalizedToken id="A37407DF5C6D7DE8687BFF0E9D6A96B3" originalValue="Núñez">Nunez</normalizedToken>
2005
</bibRefCitation>
) is more than 650 km north of the type locality recorded for
<taxonomicName id="767D129A0597F75557073F4436089B34" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
.
</paragraph>
<paragraph id="15C894EDC53704DC45B354BAE90F368F" lastPageId="5" lastPageNumber="96" pageId="4" pageNumber="95">
<taxonomicName id="0C6FB6988BD0B5E8B68D8530FC36C024" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
resembles
<taxonomicName id="E7A15444767D722CD3E116F70294E996" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus lorenzmuelleri" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="lorenzmuelleri">Liolaemus lorenzmuelleri</taxonomicName>
(Fig. 3E) and
<taxonomicName id="92BAF426DF7235859DDE441CE9AC8A91" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus juanortizi" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="juanortizi">Liolaemus juanortizi</taxonomicName>
(Fig. 3D), species suggested as conspecific (
<bibRefCitation id="12BA37AD26FE84CC176C2D67EC588D5F" author="Pincheira-Donoso, D" journalOrPublisher="Publicacion Ocasional del Museo Nacional de Historia Natural de Chile" pageId="16" pageNumber="107" pagination="7 - 486" title="Las especies chilenas del genero Liolaemus (IguanidaeTropiduridae, Liolaeminae). Taxonomia, sistematica y evolucion." volume="59" year="2005">
Pincheira-Donoso and
<normalizedToken id="CA098E3F15288144876B9C67F41F569A" originalValue="Núñez">Nunez</normalizedToken>
2005
</bibRefCitation>
). However, the dorsal scales in
<taxonomicName id="23F62722274D83F7993567BF22A932D9" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus lorenzmuelleri" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="lorenzmuelleri">Liolaemus lorenzmuelleri</taxonomicName>
and
<taxonomicName id="6527446744EF424226263FC457E92664" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus juanortizi" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="juanortizi">Liolaemus juanortizi</taxonomicName>
are noticeably larger than those of
<taxonomicName id="7953926D1641F416380BD67285020504" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
, and have a distinct
<normalizedToken id="2E7637AFAC0310308D93A15D24A6EADD" originalValue="“ovoid”">&quot;ovoid&quot;</normalizedToken>
shape.
<taxonomicName id="DD5138F5BAF997D5A35FFAEDAF82EDE7" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
has more dorsal scales (60.0
<normalizedToken id="C046CB151FA410B7D6A794EF27E916F5" originalValue="±">+/-</normalizedToken>
2.9) than
<taxonomicName id="C6A9D5D41686EAC12897A82962954E7B" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus lorenzmuelleri" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="lorenzmuelleri">Liolaemus lorenzmuelleri</taxonomicName>
(48.4
<normalizedToken id="FB6BD12B4163A64107732F650C525509" originalValue="±">+/-</normalizedToken>
4.2) (t = -5.4, P &lt;0.01). On the other hand, while only one specimen of
<taxonomicName id="699427E8510D3C7D810A9376B7D57571" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus juanortizi" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="juanortizi">Liolaemus juanortizi</taxonomicName>
was examined, this one has 52 dorsal scales, which is below of the range for
<taxonomicName id="94D135F7B77F09D0283541840DA91F76" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
(Table 1).
<taxonomicName id="BA4C608F81BBBD91D84FF0DCD2373C9E" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
has more midbody scales (60.4
<normalizedToken id="A500B693371A54AC7FFD6248E8941398" originalValue="±">+/-</normalizedToken>
1.7) than
<taxonomicName id="BB950DC845C936C8F133E8C409EF6D8B" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus lorenzmuelleri" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="lorenzmuelleri">Liolaemus lorenzmuelleri</taxonomicName>
(54.9
<normalizedToken id="8406CCEF44225FBFD4168DF24825E5ED" originalValue="±">+/-</normalizedToken>
4.5) (t = 2.6, P &lt;0.05) and
<taxonomicName id="1132DE0F1F2AF30C409DFBA378A4F73F" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus juanortizi" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="juanortizi">Liolaemus juanortizi</taxonomicName>
(56.7
<normalizedToken id="FB0C18200E8423EACF6371A8E6670470" originalValue="±">+/-</normalizedToken>
2.1) (t = 3.2, P &lt;0.05).
<taxonomicName id="23C398D3D135FC07E2B94BD294402CA5" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus lorenzmuelleri" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="lorenzmuelleri">Liolaemus lorenzmuelleri</taxonomicName>
has a dark vertebral line and dark transversal lines running from the paravertebral fields to the flanks, whereas
<taxonomicName id="EDF640326B94F73C3208A1E1A8018CD0" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
has no dorsal pattern. The dorsal pattern of
<taxonomicName id="3E55FA9237C7F6F5FD9CE353CF5A14EB" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus juanortizi" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="juanortizi">Liolaemus juanortizi</taxonomicName>
is very similar to
<taxonomicName id="2FE2C133F6C6EB6D7CA278CCAF5784E1" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus lorenzmuelleri" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="lorenzmuelleri">Liolaemus lorenzmuelleri</taxonomicName>
, but some specimens have a black ventral coloration, a black lateral band, and the lack of a dark vertebral line, whereas
<taxonomicName id="B08A02B5B5B3D32DB645B2B215A195B3" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
has no
<pageBreakToken id="2C71A88DC09E099954E07B48E4116088" pageId="5" pageNumber="96" start="start">black</pageBreakToken>
ventral color or black lateral band. Additionally, the southern distributional limit of
<taxonomicName id="2DD1C14827CB25C95992E100EA77F476" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus lorenzmuelleri" pageId="5" pageNumber="96" phylum="Chordata" rank="species" species="lorenzmuelleri">Liolaemus lorenzmuelleri</taxonomicName>
(Embalse La Laguna, Coquimbo Region, Chile) is more than 240 km north of the type locality recorded for
<taxonomicName id="B96EE3EF3CF9D7454C62F53F5D05E027" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="5" pageNumber="96" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
; and the southern distributional limit of
<taxonomicName id="F126144C23B85EEFFFB41CF9BCF7EFE9" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus juanortizi" pageId="5" pageNumber="96" phylum="Chordata" rank="species" species="juanortizi">Liolaemus juanortizi</taxonomicName>
in Quebrada Contrabando, Atacama Region, Chile (MNHNCL collection catalog, unpublished) is more than 520 km north of the type locality recorded for
<taxonomicName id="C1CE096298EDC5AABEBAA8B42D3178E4" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="5" pageNumber="96" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
.
</paragraph>
<paragraph id="BC6976E3263CB1BE798714A8176B3F13" pageId="5" pageNumber="96">
<taxonomicName id="8B92306256152DC7D0D59AEB52221893" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="5" pageNumber="96" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
differs from
<taxonomicName id="27BEF9166AF65904995A1B9087660C35" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus melanopleurus" pageId="5" pageNumber="96" phylum="Chordata" rank="species" species="melanopleurus">Liolaemus melanopleurus</taxonomicName>
(a species with only three known specimens from an undetermined location, Fig. 3B) in that the latter has a blue-gray dorsal coloration (
<bibRefCitation id="05883837A63C7E4891E9475706520992" author="Philippi, RA" journalOrPublisher="Halle, Eduard Anton" pageId="16" pageNumber="107" title="Reise durch die wueste Atacama, auf befehl der chilenischen regierung im sommer 1853 - 1854." year="1860">Philippi 1860</bibRefCitation>
) and a black lateral band running from the axilla to the midbody, features absent in
<taxonomicName id="FEC45DEA4F65A0C67414E3A94F1733B3" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="5" pageNumber="96" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
. Although the type locality of
<taxonomicName id="96A6CBE5D97F2F54A8DDECBC9A5639A5" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus melanopleurus" pageId="5" pageNumber="96" phylum="Chordata" rank="species" species="melanopleurus">Liolaemus melanopleurus</taxonomicName>
is undetermined, the syntypes were collected by Philippi in his journey through the Atacama Desert, between the vicinities of
<normalizedToken id="A7D6148AD55E655FC9D6A352CD89B9C1" originalValue="Copiapó">Copiapo</normalizedToken>
(27°23'S) and San Pedro de Atacama (22°54'S), more than 530 km north of the type locality recorded for
<taxonomicName id="2199108C7F65F7AE192740D04454D331" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="5" pageNumber="96" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
.
</paragraph>
<paragraph id="292D9D4D9131A04CC8D1C42C91C34E83" lastPageId="6" lastPageNumber="97" pageId="5" pageNumber="96">
<taxonomicName id="251D7925A7AEE79CBF59BC42D7AB610E" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="5" pageNumber="96" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
differs from
<taxonomicName id="747973842572A236028E87CE7F0770AE" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus maldonadae" pageId="5" pageNumber="96" phylum="Chordata" rank="species" species="maldonadae">Liolaemus maldonadae</taxonomicName>
(Fig. 3F), because males of the latter have a yellowish or reddish dorsal color with black transverse dorsal and ventral bars and black lateral band, whereas
<taxonomicName id="0006DC858052253071AAD1C9D641893C" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="5" pageNumber="96" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
has no dorsal pattern or black trans
<pageBreakToken id="721D8529BB8AE5E1051EB6AEEB8AACAB" pageId="6" pageNumber="97" start="start">verse</pageBreakToken>
ventral bars. Dorsal scales in
<taxonomicName id="A50162CABCA9762217CF845189FA9D4A" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus maldonadae" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="maldonadae">Liolaemus maldonadae</taxonomicName>
are noticeably larger than found in
<taxonomicName id="A542EF453916F800C45FFCF6D1A7BC6F" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
, and they have an
<normalizedToken id="6DF1E3764D82D27951A29BC02E1D99B4" originalValue="“ovoid”">&quot;ovoid&quot;</normalizedToken>
shape. Dorsal and ventral scale counts in
<taxonomicName id="36C8D1F1EBF65AB28CC22D46869863A1" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus maldonadae" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="maldonadae">Liolaemus maldonadae</taxonomicName>
do not overlap with the same scale counts in
<taxonomicName id="197E6ED6ED2B06F0BC01F55CA6B94C9B" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
(Table 1). Finally, the southern distributional limit of
<taxonomicName id="C029E8681E455026C9C26A516DCE1DC3" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus maldonadae" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="maldonadae">Liolaemus maldonadae</taxonomicName>
in Los Molles (
<bibRefCitation id="61D7481EC572F82037A0EF78CD238A5B" author="Nunez, H" journalOrPublisher="Boletin del Museo Nacional de Historia Natural de Chile" pageId="16" pageNumber="107" pagination="79 - 88" title="Liolaemusmaldonadae y Liolaemuscristiani, dos especies nuevas de lagartijas para Chile (Reptilia, Squamata)." volume="42" year="1991">
<normalizedToken id="91368A801940367F6F99A488845F9F14" originalValue="Núñez">Nunez</normalizedToken>
et al. 1991
</bibRefCitation>
) is more than 150 km north of the type locality of
<taxonomicName id="2F8DD6A5C805F81D2B537FD3EE833AB5" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
.
</paragraph>
<paragraph id="912DC724B634A67C6758B7379FBF9228" lastPageId="7" lastPageNumber="98" pageId="6" pageNumber="97">
<taxonomicName id="E780ACA4A9494AA67536D34F12EA5ACC" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
is found in sympatry with
<taxonomicName id="B13F689AE80EF66FCE49EF67BFAACD2C" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
(Fig. 4), but is larger than
<taxonomicName id="1259EC133F41990489F444B9F4BDD8FC" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
(
<normalizedToken id="C552331F106B19B60B063474E0328D42" originalValue="Mann–Whitney">Mann-Whitney</normalizedToken>
U = 8.0, P &lt;0.05, Table 1).
<taxonomicName id="ADB290EC0614FD063BEA424CA3BBF10A" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
also has more dorsal scales (60.0
<normalizedToken id="572F973ECD8DCBC331F8878864CE32A7" originalValue="±">+/-</normalizedToken>
2.9) than
<taxonomicName id="4241B52D7BCAD14B263B31AE3373029D" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
(49.4
<normalizedToken id="D55C07EA3552156F4B017F98552868F6" originalValue="±">+/-</normalizedToken>
2.7) (t = 7.4, P &lt;0.01).
<taxonomicName id="D6A78153EFC6F53E46D6F3DCFAD2C453" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
has strongly mucronated dorsal scales, whereas
<taxonomicName id="7C9C3C8DAAB9BE73A366969E0B5670B2" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
has no mucrons (Fig. 5).
<taxonomicName id="432C1DEB9FDE166E4A809156321EAB6E" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
has sexual dichromatism, absent in
<taxonomicName id="2ECBC9822D4B8FA1C11DB83D56899063" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
. Males of
<taxonomicName id="4E3DE886725CD096D37635219D682B40" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
have a bluish or yellowish green dorsal color
<pageBreakToken id="434E8B570326F4570029148AF7CBD6EB" pageId="7" pageNumber="98" start="start">with</pageBreakToken>
black reticulation, and females have a brown dorsal color with a black lateral band, black vertebral line, and black paravertebral spots. In contrast,
<taxonomicName id="7CF8026DB56A45BE0B1B620829185D92" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
has a brown dorsal color without any pattern.
</paragraph>
<caption id="2D50E24A19E93046722356F4FA11DDAA" pageId="7" pageNumber="98">
<paragraph id="6D8CE8ABB2F9F74051B5991604D1F183" pageId="7" pageNumber="98">
Figure 4. Variation in
<taxonomicName id="6BDC2BA7D853C5A4D25DB0DE6A322285" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
. A Male from Farellones (picture by H.
<normalizedToken id="DC67BB15ED2A0EAB75691A744208F049" originalValue="Díaz">Diaz</normalizedToken>
) B Male from Carpa Mountain (picture by JTP) C Male from Provincia Mountain (picture by JTP) D Female from Juncal (picture by JTP).
</paragraph>
</caption>
<caption id="724D37823EF1E8133221F55B149BB693" pageId="7" pageNumber="98">
<paragraph id="7A47E0974A84C23EA450B194F6101BE8" pageId="7" pageNumber="98">
Figure 5. Dorsal scales, 8 mm width of view. A Male of
<taxonomicName id="826AC9E7B9EEA0672822634C0071432E" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
sp. n. B
<taxonomicName id="9A1D95FE94D82572BA16A0535B3BCCB5" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="D6AD8C4D9601EAB4CFAC7620D80A41C7" pageId="7" pageNumber="98">
Molecular data show that
<taxonomicName id="D3D0AA7185B8B3F69EC2C32909F4CF75" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
is not closely related to
<taxonomicName id="C7B014D420E8B366D607F2D75703A866" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus monticola" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="monticola">Liolaemus monticola</taxonomicName>
(Fig. 1). Moreover,
<taxonomicName id="DBD5BA5DD7EEFB688678BD1D3ADFD8A3" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus monticola" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="monticola">Liolaemus monticola</taxonomicName>
is smaller (maximum SVL = 65.6 mm) than
<taxonomicName id="86E3D3606CD2EC79DBA1E756ED36F01C" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
(max. SVL = 89.1 mm) (t = 3.9, P &lt;0.01) according to our samples, and although
<bibRefCitation id="90615B45DC6931A10F1A262F668D3371" author="Pincheira-Donoso, D" journalOrPublisher="Publicacion Ocasional del Museo Nacional de Historia Natural de Chile" pageId="16" pageNumber="107" pagination="7 - 486" title="Las especies chilenas del genero Liolaemus (IguanidaeTropiduridae, Liolaeminae). Taxonomia, sistematica y evolucion." volume="59" year="2005">
Pincheira-Donoso and
<normalizedToken id="EC294D69F614D5FD9CD80AA72ED58208" originalValue="Núñez">Nunez</normalizedToken>
(2005)
</bibRefCitation>
recorded a max. SVL = 67.3 mm for
<taxonomicName id="BB2966DDC8E51C9B8823CB06FC9BED27" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus monticola" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="monticola">Liolaemus monticola</taxonomicName>
, the difference between both species is marked. Moreover,
<taxonomicName id="57DA7CFCE68E07676B21F5469A2A9456" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus monticola" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="monticola">Liolaemus monticola</taxonomicName>
exhibit a characteristic black lateral band between the axilla and midbody (diffuse in females), and males have white dots dispersed on the dorsum and a series of small black spots on the dorsum (Fig. 6). All these traits are absent in
<taxonomicName id="3C35FD3647523CAAE51EE764D9ECBE9C" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
. The upper altitudinal limit of
<taxonomicName id="70EF74DB69AD288D21110D970B11EFE7" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus monticola" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="monticola">Liolaemus monticola</taxonomicName>
distributions is 2000 m a.s.l. (
<bibRefCitation id="C21F4046A5794AB532B17FE355842069" author="Espinoza, RE" journalOrPublisher="Proceedings of the National Academy of Sciences of the United States of America" pageId="15" pageNumber="106" pagination="16819 - 16824" title="Recurrent evolution of herbivory in small, cold-climate lizards: Breaking the ecophysiological rules of reptilian herbivory." url="10.1073/pnas.0401226101" volume="48" year="2004">Espinoza et al. 2004</bibRefCitation>
,
<bibRefCitation id="1CBAC0052D3F8E8568F122F9FA6B0B8F" author="Fuentes, ER" journalOrPublisher="Journal of Herpetology" pageId="15" pageNumber="106" pagination="123 - 124" title="A note on the diet of Liolaemusmonticola (Reptilia, Lacertilia, Iguanidae)." url="10.2307/1563768" volume="13" year="1979">Fuentes and Ipinza 1979</bibRefCitation>
), whereas
<taxonomicName id="4520D17E7E5C5F45EAD45613556BFDF1" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
has a lower altitudinal distribution limit of 2820 m a.s.l.
</paragraph>
<caption id="85071192C438E7B6E6954C40BB322960" pageId="7" pageNumber="98">
<paragraph id="66A694099FC0332208063BE9721CB8DA" pageId="7" pageNumber="98">
Figure 6. Variation in species probably confused with
<taxonomicName id="CAE3CC9AECBB514B7241344CA6246F46" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
sp. n. A
<taxonomicName id="4EF63D7DAB670C94A5CCAEF2AB5548D5" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus monticola" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="monticola">Liolaemus monticola</taxonomicName>
from Salto de Apoquindo (picture by JTP) B
<taxonomicName id="5A662A9659636E446A67115A925F1F07" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus monticola" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="monticola">Liolaemus monticola</taxonomicName>
from La Cruz Mountain (picture by J.
<normalizedToken id="6B09A536E0AC2630056E3FCFC217B8AF" originalValue="Abarca-Díaz">Abarca-Diaz</normalizedToken>
) C
<taxonomicName id="562FC13A32CBA11D815F21201FDB6C3F" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus monticola" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="monticola">Liolaemus monticola</taxonomicName>
from Provincia Mountain (picture by JTP) D
<taxonomicName id="78BD6A6A29AE42A57DD9C506CFA19509" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus bellii" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="bellii">Liolaemus bellii</taxonomicName>
from La Parva (JR Martini) E
<taxonomicName id="BA4067DBA2E83249F316B0A3BC6D51BC" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus bellii" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="bellii">Liolaemus bellii</taxonomicName>
from Lagunillas (picture by JTP) F
<taxonomicName id="70D68EF7040CE9B1F1087875E71A13C6" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus bellii" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="bellii">Liolaemus bellii</taxonomicName>
from San
<normalizedToken id="3C30A36A4F5E076155516B7FD6AAA960" originalValue="Ramón">Ramon</normalizedToken>
Mountain (picture by JTP).
</paragraph>
</caption>
<paragraph id="2B8910C8195F3D464644DD75C46B25B4" lastPageId="8" lastPageNumber="99" pageId="7" pageNumber="98">
Molecular data show that
<taxonomicName id="37F07C76A9DAE312C4A9F15703AF3B33" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
is not closely related to
<taxonomicName id="D0658420C2EEE6F0AEF7A478B9FD6508" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus bellii" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="bellii">Liolaemus bellii</taxonomicName>
(Fig. 1). Moreover,
<taxonomicName id="3F7206C5ED28E0316725A693215D6BF0" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus bellii" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="bellii">Liolaemus bellii</taxonomicName>
is smaller (maximum SVL = 80.8 mm) than
<taxonomicName id="FBE6F05E75003D46CD04515C45D0419B" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
(max. SVL = 89.1 mm) (t = 2.7, P &lt;0.05).
<taxonomicName id="BA433A629EB630777080E5E3FE2E7110" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
has more midbody scales (60.4
<normalizedToken id="8DB24FAABBDC185D60B7CFBF02A84C4E" originalValue="±">+/-</normalizedToken>
1.7) than
<taxonomicName id="A97F4B78CA16C42BAD4BA7DB2FAA4A71" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus bellii" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="bellii">Liolaemus bellii</taxonomicName>
(52.9
<normalizedToken id="C3AC6C71B4BA2D563D55E41AE90110F6" originalValue="±">+/-</normalizedToken>
2.6) (t = 6.1, P &lt;0.01); more dorsal scales (60.0
<normalizedToken id="A555A04E084840E862529E426AD03B36" originalValue="±">+/-</normalizedToken>
2.9) than
<taxonomicName id="9594E31C8544C66112D98F624EF41878" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus bellii" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="bellii">Liolaemus bellii</taxonomicName>
(43.3
<normalizedToken id="3106A1319198F474D9A16B490D60E62A" originalValue="±">+/-</normalizedToken>
3.1) (t = 10.2, P &lt;0.01); and more ventral scales (96.2
<normalizedToken id="BC2D0ECC9CC31CCFC7D5266A29DEC9EC" originalValue="±">+/-</normalizedToken>
4.8) than
<taxonomicName id="06EB003315A6EA620FBFE03485EE9E9D" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus bellii" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="bellii">Liolaemus bellii</taxonomicName>
(89.7
<normalizedToken id="4440E90817571845CF6B9A6CBBCC211D" originalValue="±">+/-</normalizedToken>
4.6) (
<normalizedToken id="159ECF9201FBEAF014825A94E31D4B44" originalValue="Mann–Whitney">Mann-Whitney</normalizedToken>
U = 10.5, P &lt;0.05). Dorsal scales in
<taxonomicName id="B593719F3DDBBFA51752355824F65F00" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus bellii" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="bellii">Liolaemus bellii</taxonomicName>
are strongly keeled and mucronated, whereas there are no mucrons in
<taxonomicName id="E752FFB947CB5CE3B8AAD03C46BC66F8" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
.
<pageBreakToken id="8F875DC899C7DE665DF13A49CDB8A161" pageId="8" pageNumber="99" start="start">Moreover</pageBreakToken>
,
<taxonomicName id="48FD77C9E1965EAD3C53BA7D12377C5D" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus bellii" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="bellii">Liolaemus bellii</taxonomicName>
exhibit a characteristic series of black dorsal
<normalizedToken id="2D617776D8505E60E60F002F6F0D0909" originalValue="“W”">&quot;W&quot;</normalizedToken>
o
<normalizedToken id="FB2803D1E3FAA1A0517A19A806A46F39" originalValue="“V”">&quot;V&quot;</normalizedToken>
shaped spots (Fig. 6), whereas
<taxonomicName id="6E73B824FA72D2EB5CD3F9F231971785" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
has no dorsal pattern.
</paragraph>
</subSubSection>
<subSubSection id="DA412A450D9093A9B9B50EB2B1541C69" lastPageId="9" lastPageNumber="100" pageId="8" pageNumber="99" type="description of the holotype">
<paragraph id="31D12AA91A9F0C87FCDD82BA5F8C2170" pageId="8" pageNumber="99">Description of the holotype.</paragraph>
<paragraph id="4973A5A397703CB377EB8AE6A17B222C" pageId="8" pageNumber="99">Adult male. SVL = 84.7 mm. Horizontal diameter of the eye: 4.3 mm. Subocular length: 4.5 mm. Length of the fourth supralabial: 4.1 mm. Head length (from the posterior border of the auditory meatus to the tip of the snout): 22.1 mm. Head height (distance between the two ear openings): 10.4 mm. Head width (at the level of ear openings): 15.8 mm. Neck width: 12.4 mm. Interorbital distance: 6.3 mm. Ear-eye distance: 7.5 mm. Internarine distance: 3.8 mm. Ear width: 2.5 mm. Ear height: 3.5 mm. Axillary-groin distance: 34.9 mm. Body width: 24.7 mm. Forelimb length: 25.7 mm. Hindlimb length: 46.1 mm. Length of the right hand: 10.4 mm. Length of the right foot: 22.4 mm. Tail length (not autotomized): 132.4 mm, with relation tail length/SVL = 1.56. Pentagonal rostral scale, wider (4.2 mm) than high (1.4 mm).</paragraph>
<paragraph id="46E12EC11C830CE0E6CB3E765C896323" lastPageId="9" lastPageNumber="100" pageId="8" pageNumber="99">
Two postrostrals. Four internasals. Heptagonal interparietal, with a central, small, and whitish central spot marking the position of the parietal eye. Interparietal smaller than the parietals, surrounded by seven scales. Seven scales between the interparietal and rostral. Thirteen scales between the occiput and the rostral. Orbital semicircle incomplete on the right side and complete on the left (formed by thirteen scales). Three supraoculars on the left side and four on the right. Six superciliary scales. Frontal area divided into three scales (1 posterior and 2 anterior). Preocular separated from the lorilabials by one loreal scale. Two scales between nasal and canthal. Nasal in contact with the rostral, surrounded by six scales. One row of lorilabials between the supralabials and subocular. Four lorilabials in contact with the subocular. Six supralabials, the fourth is curved upward without contacting the subocular. Four infralabials scales. Pentagonal mental scale, in contact with four scales. Four pairs of post-mental shields, the second is separated by two scales. Temporal scales smooth or slightly keeled, imbricated. Six temporal scales between the level of superciliary scales and the rictal level. Four scales on the anterior edge of the ear, which do not cover the auditory meatus. Poorly differ
<pageBreakToken id="BFBECD2371B98BDE2FCA16A243F38817" pageId="9" pageNumber="100" start="start">entiated</pageBreakToken>
auricular scale, pentagonal and located at the upper part of the meatus. Thirty gulars between the auditory meatus. Lateral neck fold is
<normalizedToken id="67D82E9AAFE2268325BA5841C3F8EDA2" originalValue="“Y”">&quot;Y&quot;</normalizedToken>
shaped. Ventrolateral fold running from the neck to the groin. Dorsolateral fold slightly developed, running from the ear to the base of the tail. Midbody scales: 60. Dorsal scales are lanceolated, imbricated, keeled (without mucrons), with few interstitial granules. Dorsal smaller than the ventrals. Dorsal scales: 58. Ventrals scales are polymorphic (rounded, rhomboidal, pentagonal or hexagonal) smooth, imbricated, without interstitial granules. Ventrals: 91. Three precloacal pores. Supra-femoral scales lanceolate, imbricated, smooth or keeled. Infra-femoral scales lanceolate or rounded, smooth and imbricated. Supra-antebrachials scales are rounded or lanceolated, imbricated and smooth or keeled. Infra-antebrachials are rounded, imbricated and smooth. Dorsal scales of tail are pentagonal or rhomboidal, imbricated and keeled. Ventral tail scales are rounded or rhomboidal, smooth and imbricated. Lamellae of the fingers: I: 9, II: 13, III: 20, IV: 20 and V: 13. Lamellae of the toes: I: 11, II: 15, III: 21, VI: 27 and V: 17.
</paragraph>
</subSubSection>
<subSubSection id="2BA1CB295D247D0E766F3BAC3C07CC87" lastPageId="10" lastPageNumber="101" pageId="9" pageNumber="100" type="color of the holotype in life">
<paragraph id="1905042FEFFBB22CDBCE7E8B4E1C9E04" pageId="9" pageNumber="100">Color of the holotype in life.</paragraph>
<paragraph id="CA325D5F46CAAD74314B4C23C239B559" lastPageId="10" lastPageNumber="101" pageId="9" pageNumber="100">
The specimen is notable for its lack of pattern and uniform color. The head is brown and darker than the body. There are several white dots dispersed over the head and cheeks. The dorsum is coppery brown and has a few white-spotted scales that did not form a pattern. The subocular is brown and crossed by three white, vertical lines. The dorsal surface of the tail is light brown and without a pattern. The limbs are a dorsal-brown, similar to the dorsal surface, with white dots
<pageBreakToken id="1F205698F1D4490E22CD271633700B88" pageId="10" pageNumber="101" start="start">dispersed</pageBreakToken>
on the forelimbs and white transversal lines on the hindlimbs. The flanks are whitish with abundant dark brown scales. Ventrally, the hands, feet, thighs, vent, and tail are yellowish. The belly is whitish with dark dispersed spots and a dark ventral stripe. The throat is whitish with a dark thick reticulation. The precloacal pores are orange.
</paragraph>
</subSubSection>
<subSubSection id="F13FE0657AAB60D69D6F7528B3EF479D" pageId="10" pageNumber="101" type="variation in the type series">
<paragraph id="4646E40DBBF6BB7B779BAD3955C68251" pageId="10" pageNumber="101">Variation in the type series.</paragraph>
<paragraph id="FEE52AB60077AFE99FA032CEAD5C7C80" pageId="10" pageNumber="101">Males are larger and more corpulent than females. In two males: SVL: 84.7-89.1 mm. Axilla-groin distance: 34.9-37.8 mm. Head length: 21.9-22.1 mm. Head width: 15.8-16.3 mm. Head height: 10.4-11.2 mm. Leg length: 45.4-46.1 mm. Arm length: 25.0-25.8 mm. Tail length: 132.4 mm in one specimen, with relation tail length/SVL = 1.56 (autotomized in the other). In three females: SVL: 67.7-73.1 mm. Axilla-groin distance: 33.1-35.7 mm. Head length: 17.8-20.0 mm. Head width: 11.8-13.3 mm. Head height: 7.5-8.3 mm. Leg length: 32.0-34.8 mm. Arm length: 19.2-21.3 mm. Tail length: 98.1 mm in one specimen, with relation tail length/SVL = 1.45 (autotomized in other).</paragraph>
<paragraph id="E67790981E35441055B86C30229FA57D" pageId="10" pageNumber="101">
The variation of the scalation in
<taxonomicName id="48A1B14802E8C1F73684E02325E38E2B" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="10" pageNumber="101" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
is as follows. Midbody scales: 58-62 (60.4
<normalizedToken id="7744E4D28FBEDC938958993803309005" originalValue="±">+/-</normalizedToken>
1.7). Dorsal scales: 56-63 (60.0
<normalizedToken id="0AD3627317FE9BD90522FB5B072124BF" originalValue="±">+/-</normalizedToken>
2.9). Ventral scales 91-102 (96.2
<normalizedToken id="A49EB9C06FF0B6504140CB6FF0CCF4B2" originalValue="±">+/-</normalizedToken>
4.8). Fourth finger lamellae: 17-20 (19.0
<normalizedToken id="B516C716EEAF4818DC9B68452ABA2BD2" originalValue="±">+/-</normalizedToken>
1.4). Fourth toe lamellae: 25-27 (26.4
<normalizedToken id="5711B9248AD62EB0FCDFC06EEEFBD4C0" originalValue="±">+/-</normalizedToken>
0.9). Supralabial scales: 6. Infralabial scales: 4-5 (4.4
<normalizedToken id="0435346B2B82AA11662CAAEFEF6FC324" originalValue="±">+/-</normalizedToken>
0.6). Interparietal scale pentagonal, hexagonal or heptagonal, bordered by 5-7 scales (6.0
<normalizedToken id="C65EBBB7466492C5AFBD24FFEB10B308" originalValue="±">+/-</normalizedToken>
0.7). Nasal and rostral always in contact. Precloacal pores in males: 3. Precloacal pores are absent in females.
</paragraph>
<paragraph id="2861B2B4A13197F0FE371E17D6FB13D1" pageId="10" pageNumber="101">In general, all specimens have the pattern and color described for the holotype, with slight variations in shade. The male paratype has a dark brown throat. Two females have inconspicuous dark rings and an inconspicuous vertebral stripe on the dorsal surface of the tail. Also, two females have an olive hue on the snout. One female has a very inconspicuous series of dark crossbars on the paravertebral fields, which, while difficult to count, approximated eight. The juvenile has a similar pattern and color as the holotype, but it has an inconspicuous and fragmented dark vertebral line and inconspicuous dark spots on the paravertebral fields.</paragraph>
</subSubSection>
<subSubSection id="5597942E1DE13B898C5A45670B7A5395" lastPageId="11" lastPageNumber="102" pageId="10" pageNumber="101" type="distribution and natural history">
<paragraph id="C7BFC064A58DFB8DA2ECB845D1706499" pageId="10" pageNumber="101">Distribution and natural history.</paragraph>
<paragraph id="95D8138791CB5DD77473BE7047582749" pageId="10" pageNumber="101">
This species is currently only known from the type locality in the surroundings of the Chepical Lagoon, approximately 30 km NE of Alicahue, in the San Felipe de Aconcagua Province,
<normalizedToken id="80C5869CD57E077FEFEB6C58BD59E014" originalValue="Valparaíso">Valparaiso</normalizedToken>
Region, Chile (Fig. 7). Specimens were collected on the west shore of the Chepical Lagoon (
<geoCoordinate id="A34839655AC12CD20EE3A00E38D42137" direction="south" orientation="latitude" precision="925" value="-32.25">32°15'S</geoCoordinate>
-
<geoCoordinate id="ABC38F956D0D109C105FA272DD3868F4" direction="west" orientation="longitude" precision="925" value="-70.5">70°30'W</geoCoordinate>
, 3050 m a.s.l.). This new species was found inhabiting rocky areas with little shrubby vegetation composed mainly of high-Andean forbs, such as
<taxonomicName id="070F3E1D8D2CFCE644E07CD3E2802E66" class="Magnoliopsida" family="Asteraceae" genus="Chuquiraga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chuquiraga oppositifolia" order="Asterales" pageId="10" pageNumber="101" phylum="Tracheophyta" rank="species" species="oppositifolia">Chuquiraga oppositifolia</taxonomicName>
and
<taxonomicName id="0C97ED153DC354E823A4108229E48401" class="Magnoliopsida" family="Apiaceae" genus="Azorella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Azorella" order="Apiales" pageId="10" pageNumber="101" phylum="Tracheophyta" rank="genus">Azorella</taxonomicName>
sp. (Fig. 8). This lizard was found in abundance and was observed to have saxicolous habits. It was active between 9:00 h and 18:00 h and took refuge under rocks. Moreover, this species was found in syntopy with
<taxonomicName id="08DC1BD7548E5A85369DBD2C2B46DC66" class="Squamata" family="Liolaemidae" genus="Phymaturus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phymaturus alicahuense" pageId="10" pageNumber="101" phylum="Chordata" rank="species" species="alicahuense">Phymaturus alicahuense</taxonomicName>
<normalizedToken id="754E7031056F5A537D34E5BD755349A0" originalValue="Núñez">Nunez</normalizedToken>
, Veloso, Espejo, Veloso,
<normalizedToken id="15CFA6D2598D60CE671530EF0C04407C" originalValue="Cortés">Cortes</normalizedToken>
&amp; Araya 2010. Specimens were also observed at lower altitudes (
<geoCoordinate id="8232580DA0A5C65B037D4A6F4DCF2CDC" direction="south" orientation="latitude" precision="925" value="-32.266666">32°16'S</geoCoordinate>
-
<geoCoordinate id="7C44A723CEC2244BE1622142A3D011B9" direction="west" orientation="longitude" precision="925" value="-70.5">70°30'W</geoCoordinate>
, 2820 m a.s.l.) in similar environments, altitudes at which this species was found in sympatry with a few specimens of
<taxonomicName id="6AEDD89F408305234E38DB36515179AF" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="10" pageNumber="101" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
.
</paragraph>
<caption id="289504D57E6DBD54110EA9445BB3FC6F" pageId="10" pageNumber="101">
<paragraph id="9F50136C9B45DB13EC9E2F621688B0AD" pageId="10" pageNumber="101">
Figure 7. Distributional map for
<taxonomicName id="CCCFDC78351FFEAD6118CDD7A0E8CA40" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="10" pageNumber="101" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
sp. n. along with geographically proximate species of the
<taxonomicName id="2078F548D27E5B85E02C858AE655AD5D" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="10" pageNumber="101" phylum="Chordata" rank="species" species="nigroviridis">nigroviridis</taxonomicName>
group. Red star:
<taxonomicName id="1BA767B8D6B78AD7734D8C6CD9CFFEE9" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="10" pageNumber="101" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
sp. n., Chepical Lagoon, type locality. Green circles:
<taxonomicName id="6239DB0AB4CB18EF95EFE111412D5872" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="10" pageNumber="101" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
(1 = Manque, 2 = El Arpa, 3 = Juncal, 4 = Riecillo, 5 = La Campana, 6 = El Roble, without number = near Chepical Lagoon). Pink squares:
<taxonomicName id="5576B54698CED08A859DDE3CE67F6C2B" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus maldonadae" pageId="10" pageNumber="101" phylum="Chordata" rank="species" species="maldonadae">Liolaemus maldonadae</taxonomicName>
(1 = near Alcohuaz, 2 = Los Molles). Yellow triangles:
<taxonomicName id="0AF9BEBB8F05252544A1677DDA154CE6" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus lorenzmuelleri" pageId="10" pageNumber="101" phylum="Chordata" rank="species" species="lorenzmuelleri">Liolaemus lorenzmuelleri</taxonomicName>
(1 =
<normalizedToken id="16343B058EF06E630D340DBE2B037AED" originalValue="Baños">Banos</normalizedToken>
del Toro, 2 = Embalse La Laguna).
</paragraph>
</caption>
<caption id="3A3DB0B91A805B555B3CC13F10E6682B" pageId="10" pageNumber="101">
<paragraph id="83089FB571747C921432561293DF3A8C" pageId="10" pageNumber="101">
Figure 8. View of the type locality of
<taxonomicName id="815D623061A08A94907EC594DB30A4FE" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="10" pageNumber="101" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
sp. n., a high Andean environment.
</paragraph>
</caption>
<paragraph id="E3F31DAF02DEADAA58E670A77F46170A" lastPageId="11" lastPageNumber="102" pageId="10" pageNumber="101">
One of the collected specimens had a yellow flower inside of its mouth. An analysis of intestinal contents showed that
<taxonomicName id="54394E89CA01754D3E016367B4222B9E" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus uniformis" pageId="10" pageNumber="101" phylum="Chordata" rank="species" species="uniformis">Liolaemus uniformis</taxonomicName>
is omnivorous; plant and
<taxonomicName id="8479A735493365141A0E44B16101A34A" lsidName="Liolaemus" order="Hymenoptera" pageId="10" pageNumber="101" rank="order">Hymenoptera</taxonomicName>
remains were found. A large quantity of nematodes from an unidentified species was
<pageBreakToken id="BB88DE1FC975A78694D02EB1A5BFDDD0" pageId="11" pageNumber="102" start="start">found</pageBreakToken>
in the intestines. While the reproductive mode is yet unknown, at the time of sampling (December) no evidence of embryos was found but one female had several small oocytes. Comparisons with the reproductive modes of other species in the
<taxonomicName id="FD05B1DA49395119324D3EBEC258CDA5" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="11" pageNumber="102" phylum="Chordata" rank="species" species="nigroviridis">nigroviridis</taxonomicName>
group would not be helpful as there is little available data. It is known that
<taxonomicName id="270F3D5E85A4B4A380C5740496A576F0" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus nigroviridis" pageId="11" pageNumber="102" phylum="Chordata" rank="species" species="nigroviridis">Liolaemus nigroviridis</taxonomicName>
is viviparous (
<bibRefCitation id="3CD04675D21A34BE1D44305CB55E347F" author="Donoso-Barros, R" journalOrPublisher="Ediciones de la Universidad de Chile, Santiago" pageId="15" pageNumber="106" title="Reptiles de Chile" year="1966">Donoso-Barros 1966</bibRefCitation>
) and
<taxonomicName id="05381845314A24A5B46370F439F7C63A" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus lorenzmuelleri" pageId="11" pageNumber="102" phylum="Chordata" rank="species" species="lorenzmuelleri">Liolaemus lorenzmuelleri</taxonomicName>
is oviparous (
<bibRefCitation id="E763166A2A140B501BFF6A0F483FE2D5" author="Cortes, A" journalOrPublisher="Ediciones Universidad de La Serena, La Serena" pageId="15" pageNumber="106" title="Fauna de vertebrados de los Andes de Coquimbo: Cordillera de Dona Ana." year="1995">
<normalizedToken id="1AA8D9F1CA5DFF0392D42DB8266AE031" originalValue="Cortés">Cortes</normalizedToken>
et al. 1995
</bibRefCitation>
).
<bibRefCitation id="844492D6B9CB072BD59DAE292DA0BF33" author="Pincheira-Donoso, D" journalOrPublisher="Publicacion Ocasional del Museo Nacional de Historia Natural de Chile" pageId="16" pageNumber="107" pagination="7 - 486" title="Las especies chilenas del genero Liolaemus (IguanidaeTropiduridae, Liolaeminae). Taxonomia, sistematica y evolucion." volume="59" year="2005">
Pincheira-Donoso and
<normalizedToken id="1EEF7DD7D5E63278126DF682E797E06F" originalValue="Núñez">Nunez</normalizedToken>
(2005)
</bibRefCitation>
reported that
<taxonomicName id="7B3377D49F73E4EB38B9C50D7AC4771A" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus maldonadae" pageId="11" pageNumber="102" phylum="Chordata" rank="species" species="maldonadae">Liolaemus maldonadae</taxonomicName>
and
<taxonomicName id="AD98DBC04A4546D3FF525D31A1144A55" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus isabelae" pageId="11" pageNumber="102" phylum="Chordata" rank="species" species="isabelae">Liolaemus isabelae</taxonomicName>
are viviparous, but the source of this information is unclear (see
<bibRefCitation id="46EDB90117A87A072F4581E0DD07B67C" author="Lobo, F" journalOrPublisher="Zootaxa" pageId="16" pageNumber="107" pagination="1 - 30" title="A critical review and systematic discussion of recent classification proposals for liolaemid lizards." volume="2549" year="2010">Lobo et al. 2010</bibRefCitation>
:4) since the reproductive mode was not mentioned in the original descriptions (
<bibRefCitation id="E9C2DAED6C79D60CADB3DB431F0D88D3" author="Navarro, J" journalOrPublisher="Boletin del Museo Nacional de Historia Natural de Chile" pageId="16" pageNumber="107" pagination="99 - 113" title="Liolaemuspatriciaturrae y Liolaemusisabelae, dos nuevas especies de lagartijas para el norte de Chile. Aspectos biogeograficos y citotaxonomicos (Squamata, Tropiduridae)." volume="44" year="1993">
Navarro and
<normalizedToken id="F17C6BA9FD84FB7F4F174E52CBB73766" originalValue="Núñez">Nunez</normalizedToken>
1993
</bibRefCitation>
,
<bibRefCitation id="41D43453A257AC95B9E1AD05F616AAD5" author="Nunez, H" journalOrPublisher="Boletin del Museo Nacional de Historia Natural de Chile" pageId="16" pageNumber="107" pagination="79 - 88" title="Liolaemusmaldonadae y Liolaemuscristiani, dos especies nuevas de lagartijas para Chile (Reptilia, Squamata)." volume="42" year="1991">
<normalizedToken id="CC6A01B374D49D46CE5DC8077E05F3F0" originalValue="Núñez">Nunez</normalizedToken>
et al. 1991
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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