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<mods:title id="76CE9CE0B1D752DFF0C48281EEC6C651">The digger wasps of the genus Alysson Panzer (Hymenoptera: Crabronidae: Bembicinae) of Russia and adjacent territories, with a key to species and new synonymies</mods:title>
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<emphasis id="C783EA82A53FFF8A2787F9C7FEB0F9FC" bold="true" box="[151,362,1601,1627]" italics="true" pageId="4" pageNumber="280">Alysson pertheesi</emphasis>
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<emphasis id="C783EA82A53FFF8A2787F90EFE93F939" box="[151,329,1672,1694]" italics="true" pageId="4" pageNumber="280">Alysson pertheesi</emphasis>
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: 178
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, ♀.
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: ♀♀,
<collectingCountry id="8DE07600A53FFF8A25D9F90EFCF7F939" box="[713,813,1672,1694]" name="Lithuania" pageId="4" pageNumber="280">Lithuania</collectingCountry>
, Vilnius (ZMHU).
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<emphasis id="C783EA82A53FFF8A2787F920FEE9F91B" box="[151,307,1702,1724]" italics="true" pageId="4" pageNumber="280">Alyson festivus</emphasis>
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: 129
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, ♀ (as
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, incorrect original termination).
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or
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: ♀,
<collectingCountry id="8DE07600A53FFF8A222FF921FA42F91B" box="[1343,1432,1703,1724]" name="Hungary" pageId="4" pageNumber="280">Hungary</collectingCountry>
, Budapest (TMB). Synonymized with
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<emphasis id="C783EA82A53FFF8A2554F943FD2DF97C" box="[580,759,1733,1755]" italics="true" pageId="4" pageNumber="280">Alysson pertheesi</emphasis>
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by
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: 250
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.
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<emphasis id="C783EA82A53FFF8A2787F962FE93F95D" box="[151,329,1764,1786]" italics="true" pageId="4" pageNumber="280">Alysson pertheesi</emphasis>
:
<treatmentCitation id="74561081A53FFF8A2646F962FDEEF95D" author="Handlirsch" box="[342,564,1764,1786]" page="251" pageId="4" pageNumber="280" year="1888">
<bibRefCitation id="91664B61A53FFF8A2646F962FDDBF95D" author="Handlirsch" box="[342,513,1764,1786]" pageId="4" pageNumber="280" refString="Handlirsch, A. (1888) Monographie der mit Nysson und Bembex verwandten Grabwespen VI. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe. Abtheilung I, 96, 219 - 311, pl. I - II." type="journal article" year="1888">Handlirsch 1888</bibRefCitation>
: 251
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; de
<treatmentCitation id="74561081A53FFF8A254CF962FCE5F95D" author="Dalla" box="[604,831,1764,1786]" page="564" pageId="4" pageNumber="280" year="1897">
<bibRefCitation id="91664B61A53FFF8A254CF962FCD6F95D" author="Dalla" box="[604,780,1764,1786]" pageId="4" pageNumber="280" refString="de Dalla Torre, C. G. (1897) Catalogus Hymenopterorum hucusque Descriptorum Systematicus et Synonimicus. Vol. VIII. Fossores (Sphegidae). Guilelmi Engelmann, Lipsiae, viii + 750 pp." type="book" year="1897">Dalla Torre 1897</bibRefCitation>
: 564
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;
<treatmentCitation id="74561081A53FFF8A2459F962FBCAF95D" author="Kokujev" box="[841,1040,1764,1786]" page="216" pageId="4" pageNumber="280" year="1906">
<bibRefCitation id="91664B61A53FFF8A2459F962FC04F95D" author="Kokujev" box="[841,990,1764,1786]" pageId="4" pageNumber="280" refString="Kokujev, N. R. (1906) O russkich predstavitelakh podsemeystva Alysonini Dalla Torre (Hymenoptera, Crabronidae). Horae Societatis Entomologicae Rossicae, 37, 209 - 219." type="journal article" year="1906">Kokujev 1906</bibRefCitation>
: 216
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;
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: 25
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, 1968: 5, 1974: 361;
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: 458
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<bibRefCitation id="91664B61A53FFF8A26E8F884FD71F8BF" author="Kolesnikov" box="[504,683,1794,1816]" pageId="4" pageNumber="280" refString="Kolesnikov, V. A. (1977) Royushchiye osy (Hymenoptera, Sphecidae) Bryanskoy Oblasti i ikh znachenie kak entomofagov. Entomologicheskoye Obozreniye, 56, 315 - 325." type="journal article" year="1977">Kolesnikov 1977</bibRefCitation>
: 318
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;
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<bibRefCitation id="91664B61A53FFF8A25FDF884FC52F8BF" author="Pulawski" box="[749,904,1794,1816]" pageId="4" pageNumber="280" refString="Pulawski, W. J. (1978) Nadsem. Sphecoidea. In: Medvedev, G. S. (Ed.), Opredelitel nasekomykh evropeyskoi chasti SSSR. Tom III. Pereponchatokrylye. Pervaya chast. Nauka, Leningrad, pp. 173 - 279." type="book chapter" year="1978">Pulawski 1978</bibRefCitation>
: 212
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: 85
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;
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: 119
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<treatmentCitation id="74561081A53FFF8A27D7F8A7FEA0F890" author="Tsuneki" box="[199,378,1825,1847]" page="18" pageId="4" pageNumber="280" year="1982">
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: 18
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;
<treatmentCitation id="74561081A53FFF8A2695F8A7FD9FF890" author="Nemkov" box="[389,581,1825,1847]" page="106" pageId="4" pageNumber="280" year="1986">
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:106
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: 199
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;
<treatmentCitation id="74561081A53FFF8A2300F8A7FEE9F8F1" author="Nemkov" page="455" pageId="4" pageNumber="280" year="1995">
<bibRefCitation id="91664B61A53FFF8A2300F8A7FF27F8F1" author="Nemkov" pageId="4" pageNumber="280" refString="Nemkov, P. G., Kazenas, V. L., Budrys, E. R. &amp; Antropov, A. V. (1995) Nadsem. Sphecoidea. 67. Sem. Sphecidae - Royushchiye osy. In: Lehr, P. A. (Ed.), Opredelitel nasekomykh Dalnego Vostoka Rossii v shesti tomakh. Tom IV. Setchatokrylyie, skorpionnitsy, pereponchatokrylyie. Chast 1. Nauka, Sankt-Peterburg, pp. 368 - 480." type="journal article" year="1995">Nemkov Kazenas, Budrys &amp; Antropov 1995</bibRefCitation>
: 455
</treatmentCitation>
;
<treatmentCitation id="74561081A53FFF8A2651F8C6FD1BF8F1" author="Shlyakhtenok" box="[321,705,1856,1878]" page="34" pageId="4" pageNumber="280" year="2002">
<bibRefCitation id="91664B61A53FFF8A2651F8C6FD43F8F1" author="Shlyakhtenok" box="[321,665,1856,1878]" pageId="4" pageNumber="280" refString="Shlyakhtenok, A. S. &amp; Skibinska, E. (2002) Zhalonosnye pereponchatokrylye (Hymenoptera, Aculeata) Belarusi: semeystvo royushchiye osy (Sphecidae). Vestnik Zoologii, 36, 31 - 40." type="journal article" year="2002">Shlyakhtenok &amp; Skibinska 2002</bibRefCitation>
: 34
</treatmentCitation>
;
<treatmentCitation id="74561081A53FFF8A25DFF8C6FC47F8F1" author="Nemkov" box="[719,925,1856,1878]" page="158" pageId="4" pageNumber="280" year="2005">
<bibRefCitation id="91664B61A53FFF8A25DFF8C6FCBDF8F1" author="Nemkov" box="[719,871,1856,1878]" pageId="4" pageNumber="280" refString="Nemkov, P. G. (2005) Fauna royushchikh os (Hymenoptera, Sphecidae, Crabronidae) ostrova Sakhalin. In: Storozhenko, S. Yu., Bogatov, V. V., Barkalov, V. Yu., Lelej, A. S. &amp; Makarchenko, E. A. (Eds.), Rastitelnyi i zhivotnyi mir ostrova Sakhalin (Materialy mezhdunarodnogo sakhalinskogo projecta). Part 2. Dalnauka, Vladivostok, pp. 141 - 167." type="journal article" year="2005">Nemkov 2005</bibRefCitation>
: 158
</treatmentCitation>
;
<treatmentCitation id="74561081A53FFF8A24BBF8C7FB96F8F1" author="Hua" box="[939,1100,1856,1878]" page="277" pageId="4" pageNumber="280" year="2006">
<bibRefCitation id="91664B61A53FFF8A24BBF8C7FBCFF8F1" author="Hua" box="[939,1045,1856,1878]" pageId="4" pageNumber="280" refString="Hua, L. (2006) Superfamily Apoidea (Sphecoidea). In: Hua, L. (Ed.), List of Chinese insects. Vol. IV. Sun-Yat-sen University Press, Guangzhou, pp. 274 - 299." type="book chapter" year="2006">Hua 2006</bibRefCitation>
: 277
</treatmentCitation>
;
<treatmentCitation id="74561081A53FFF8A2349F8C7FAC2F8F1" author="Terayama" box="[1113,1304,1856,1878]" page="2" pageId="4" pageNumber="280" year="2006">
<bibRefCitation id="91664B61A53FFF8A2349F8C7FB24F8F1" author="Terayama" box="[1113,1278,1856,1878]" pageId="4" pageNumber="280" refString="Terayama, M. (2006) Taxonomic guide to the Japanese Aculeate wasps. 6. Family Nyssonidae. Tsunekibachi, 10, 1 - 27." type="journal article" year="2006">Terayama 2006</bibRefCitation>
: 2
</treatmentCitation>
, 4;
<treatmentCitation id="74561081A53FFF8A2255F8C6FEF4F8D3" author="Nemkov" page="74" pageId="4" pageNumber="280" year="2007">
<bibRefCitation id="91664B61A53FFF8A2255F8C6FED3F8D3" author="Nemkov" pageId="4" pageNumber="280" refString="Nemkov, P. G. (2007 a) Fauna royushchikh os (Hymenoptera: Sphecidae, Crabronidae) Kurilskikh ostrovov. Evrasiatskiy Entomologicheskij Zhurnal, 6, 67 - 76." type="journal article" year="2007" yearSuffix="a">Nemkov 2007a</bibRefCitation>
: 74
</treatmentCitation>
, 2007b: 997, 2008: 24, 2009: 127;
<treatmentCitation id="74561081A53FFF8A2581F8D8FC86F8D3" author="Baghirov" box="[657,860,1886,1908]" page="141" pageId="4" pageNumber="280" year="2011">
<bibRefCitation id="91664B61A53FFF8A2581F8D8FCF0F8D3" author="Baghirov" box="[657,810,1886,1908]" pageId="4" pageNumber="280" refString="Baghirov, R. T. (2011) Fauna royushchikh os (Hymenoptera: Sphecidae, Crabronidae) Altayskogo Kraya i Respubliki Altay. Entomologicheskoye Obozreniye, 90, 138 - 158." type="journal article" year="2011">Baghirov 2011</bibRefCitation>
: 141
</treatmentCitation>
;
<treatmentCitation id="74561081A53FFF8A2477F8D8FBE1F8D3" author="Nemkov" box="[871,1083,1886,1908]" page="445" pageId="4" pageNumber="280" year="2012">
<bibRefCitation id="91664B61A53FFF8A2477F8D8FBD2F8D3" author="Nemkov" box="[871,1032,1886,1908]" pageId="4" pageNumber="280" refString="Nemkov, P. G. (2012 b) Sektsiya Spheciformes - Royushchiye osy. In: Lelej, A. S. (Ed.), Annotated Catalogue of the Insects of Russian Far East. Volume I. Hymenoptera. Dalnauka, Vladivostok, pp. 433 - 447." type="book chapter" year="2012" yearSuffix="b">Nemkov 2012b</bibRefCitation>
: 445
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F5483690A53FFF8A2787F8FBFE04F815" blockId="4.[151,1436,1672,2031]" pageId="4" pageNumber="280">
<treatmentCitationGroup id="D5E711BEA53FFF8A2787F8FBFE04F815" pageId="4" pageNumber="280">
<taxonomicName id="32F74D13A53FFF8A2787F8FBFE9EF834" ID-CoL="7J99Q" box="[151,324,1917,1939]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis id="C783EA82A53FFF8A2787F8FBFE9EF834" box="[151,324,1917,1939]" italics="true" pageId="4" pageNumber="280">Alysson fuscatus</emphasis>
</taxonomicName>
<taxonomicName id="32F74D13A53FFF8A265FF8F8FD3AF834" ID-CoL="CCFK" authority="Kokujev 1906: 216" authorityName="Kokujev" authorityPageNumber="216" authorityYear="1906" box="[335,736,1917,1939]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="variety" species="fuscatus" variety="jaroslavensis">
var.
<emphasis id="C783EA82A53FFF8A2690F8FBFDDFF834" box="[384,517,1917,1939]" italics="true" pageId="4" pageNumber="280">jaroslavensis</emphasis>
<treatmentCitation id="74561081A53FFF8A2500F8FBFD3AF834" author="Kokujev" box="[528,736,1917,1939]" page="216" pageId="4" pageNumber="280" year="1906">
<bibRefCitation id="91664B61A53FFF8A2500F8FBFD72F834" author="Kokujev" box="[528,680,1917,1939]" pageId="4" pageNumber="280" refString="Kokujev, N. R. (1906) O russkich predstavitelakh podsemeystva Alysonini Dalla Torre (Hymenoptera, Crabronidae). Horae Societatis Entomologicae Rossicae, 37, 209 - 219." type="journal article" year="1906">Kokujev 1906</bibRefCitation>
: 216
</treatmentCitation>
</taxonomicName>
, ♀.
<typeStatus id="2A4C8832A53FFF8A2404F8FBFCACF834" box="[788,886,1917,1939]" pageId="4" pageNumber="280" type="holotype">Holotype</typeStatus>
: ♀,
<collectingCountry id="8DE07600A53FFF8A24B9F8FBFC35F834" box="[937,1007,1917,1939]" name="Russia" pageId="4" pageNumber="280">Russia</collectingCountry>
, Yaroslavskaya Oblast, Yaroslavl (ZIN), examined.
<emphasis id="C783EA82A53FFF8A2626F81AFE02F816" bold="true" box="[310,472,1948,1969]" pageId="4" pageNumber="280">New synonymy</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F5483690A53FFF8A2787F83CFB23F877" blockId="4.[151,1436,1672,2031]" box="[151,1273,1978,2000]" pageId="4" pageNumber="280">
<treatmentCitationGroup id="D5E711BEA53FFF8A2787F83CFB23F877" box="[151,1273,1978,2000]" pageId="4" pageNumber="280">
<taxonomicName id="32F74D13A53FFF8A2787F83CFDD6F877" ID-CoL="CCFG" authority="Tsuneki 1967: 7" authorityName="Tsuneki" authorityPageNumber="7" authorityYear="1967" box="[151,524,1978,2000]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="harbinensis">
<emphasis id="C783EA82A53FFF8A2787F83CFEB8F877" box="[151,354,1978,2000]" italics="true" pageId="4" pageNumber="280">Alysson harbinensis</emphasis>
<treatmentCitation id="74561081A53FFF8A2678F83CFDD6F877" author="Tsuneki" box="[360,524,1978,2000]" page="7" pageId="4" pageNumber="280" year="1967">
<bibRefCitation id="91664B61A53FFF8A2678F83CFE2EF877" author="Tsuneki" box="[360,500,1978,2000]" pageId="4" pageNumber="280" refString="Tsuneki, K. (1967) Further studies on the fossorial Hymenoptera from Manchuria. Etizenia, 23,1 - 17." type="book" year="1967">Tsuneki 1967</bibRefCitation>
: 7
</treatmentCitation>
</taxonomicName>
, ♀.
<typeStatus id="2A4C8832A53FFF8A2527F83CFD43F877" box="[567,665,1978,2000]" pageId="4" pageNumber="280" type="holotype">Holotype</typeStatus>
: ♀,
<collectingCountry id="8DE07600A53FFF8A25D2F83CFCDBF877" box="[706,769,1978,2000]" name="China" pageId="4" pageNumber="280">China</collectingCountry>
, Heilongjiang, Harbin (NNML).
<emphasis id="C783EA82A53FFF8A2341F83DFB29F877" bold="true" box="[1105,1267,1979,2000]" pageId="4" pageNumber="280">New synonymy</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F5483690A53FFF8A2787F85FFA92F848" blockId="4.[151,1436,1672,2031]" box="[151,1352,2009,2031]" pageId="4" pageNumber="280">
<treatmentCitationGroup id="D5E711BEA53FFF8A2787F85FFA92F848" box="[151,1352,2009,2031]" pageId="4" pageNumber="280">
<taxonomicName id="32F74D13A53FFF8A2787F85FFDCEF848" ID-CoL="CCFG" authority="Tsuneki 1968: 5" authorityName="Tsuneki" authorityPageNumber="5" authorityYear="1968" box="[151,532,2009,2031]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="harbinensis">
<emphasis id="C783EA82A53FFF8A2787F85FFEB8F848" box="[151,354,2009,2031]" italics="true" pageId="4" pageNumber="280">Alysson harbinensis</emphasis>
:
<treatmentCitation id="74561081A53FFF8A267FF85FFDCEF848" author="Tsuneki" box="[367,532,2009,2031]" page="5" pageId="4" pageNumber="280" year="1968">
<bibRefCitation id="91664B61A53FFF8A267FF85FFE21F848" author="Tsuneki" box="[367,507,2009,2031]" pageId="4" pageNumber="280" refString="Tsuneki, K. (1968) Studies on the Formosan Sphecidae (VI.). The subfamily Nyssoninae (Hymenoptera) with notes on Bembecinus of the Ryukyus and Korea. Etizenia, 31, 1 - 26, pls. I - II." type="journal article" year="1968">Tsuneki 1968</bibRefCitation>
: 5
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="74561081A53FFF8A250FF85FFCE0F848" author="Bohart" box="[543,826,2009,2031]" page="458" pageId="4" pageNumber="280" year="1976">
<bibRefCitation id="91664B61A53FFF8A250FF85FFCDDF848" author="Bohart" box="[543,775,2009,2031]" pageId="4" pageNumber="280" refString="Bohart, R. M. &amp; Menke, A. S. (1976) Sphecid Wasps of the World. A generic revision. University of California Press, Berkeley, Los Angeles, London, ix + 695 pp." type="book" year="1976">Bohart &amp; Menke 1976</bibRefCitation>
: 458
</treatmentCitation>
;
<treatmentCitation id="74561081A53FFF8A2454F85CFBF7F848" author="Wu" box="[836,1069,2009,2031]" page="214" pageId="4" pageNumber="280" year="1987">
<bibRefCitation id="91664B61A53FFF8A2454F85CFC20F848" author="Wu" box="[836,1018,2009,2031]" pageId="4" pageNumber="280" refString="Wu, Y. &amp; Zhou, Q. (1987) A study of the genus Alysson and descriptions of new species from China. Acta Entomologica Sinica, 30, 214 - 222." type="journal article" year="1987">Wu &amp; Zhou 1987</bibRefCitation>
: 214
</treatmentCitation>
; 1996: 136;
<treatmentCitation id="74561081A53FFF8A23BBF85CFA9EF848" author="Hua" box="[1195,1348,2009,2031]" page="277" pageId="4" pageNumber="280" year="2006">
<bibRefCitation id="91664B61A53FFF8A23BBF85CFACBF848" author="Hua" box="[1195,1297,2009,2031]" pageId="4" pageNumber="280" refString="Hua, L. (2006) Superfamily Apoidea (Sphecoidea). In: Hua, L. (Ed.), List of Chinese insects. Vol. IV. Sun-Yat-sen University Press, Guangzhou, pp. 274 - 299." type="book chapter" year="2006">Hua 2006</bibRefCitation>
: 277
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F5483690A53EFF8B2787FF1EFD18FF4C" blockId="5.[151,1436,152,327]" pageId="5" pageNumber="281">
<treatmentCitationGroup id="D5E711BEA53EFF8B2787FF1EFD18FF4C" bridged="true" pageId="5" pageNumber="281">
<taxonomicName id="32F74D13A53EFF8B2787FF1EFE93FF09" ID-CoL="CCFY" box="[151,329,152,174]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="pertheesi">
<emphasis id="C783EA82A53EFF8B2787FF1EFE93FF09" box="[151,329,152,174]" italics="true" pageId="5" pageNumber="281">Alysson pertheesi</emphasis>
</taxonomicName>
forma
<emphasis id="C783EA82A53EFF8B2682FF1FFE0AFF09" box="[402,464,153,174]" italics="true" pageId="5" pageNumber="281">typica</emphasis>
<treatmentCitation id="74561081A53EFF8B26C6FF1EFD2FFF09" author="Tsuneki" box="[470,757,152,174]" page="13" pageId="5" pageNumber="281" year="1973">
<bibRefCitation id="91664B61A53EFF8B26C6FF1EFD15FF09" author="Tsuneki" box="[470,719,152,174]" pageId="5" pageNumber="281" refString="Tsuneki, K. &amp; Itami, I. (1973) Variation in colour of Alysson pertheesi Gorski with notes on other characters. The Life Study (Fukui), 17, 13." type="journal article" year="1973">Tsuneki and Itami, 1973</bibRefCitation>
: 13
</treatmentCitation>
. Not available (Article 16).
<taxonomicName id="32F74D13A53EFF8B2787FF30FE90FF6B" ID-CoL="CCFY" box="[151,330,182,204]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="pertheesi">
<emphasis id="C783EA82A53EFF8B2787FF30FE90FF6B" box="[151,330,182,204]" italics="true" pageId="5" pageNumber="281">Alysson pertheesi</emphasis>
</taxonomicName>
forma
<emphasis id="C783EA82A53EFF8B2686FF3EFE17FF6A" box="[406,461,184,205]" italics="true" pageId="5" pageNumber="281">nigra</emphasis>
<treatmentCitation id="74561081A53EFF8B26C5FF30FD23FF6B" author="Tsuneki" box="[469,761,182,204]" page="13" pageId="5" pageNumber="281" year="1973">
<bibRefCitation id="91664B61A53EFF8B26C5FF30FD08FF6B" author="Tsuneki" box="[469,722,182,204]" pageId="5" pageNumber="281" refString="Tsuneki, K. &amp; Itami, I. (1973) Variation in colour of Alysson pertheesi Gorski with notes on other characters. The Life Study (Fukui), 17, 13." type="journal article" year="1973">Tsuneki and Itami, 1973</bibRefCitation>
: 13
</treatmentCitation>
, ♀.
<typeStatus id="2A4C8832A53EFF8B2436FF30FC5DFF6B" box="[806,903,182,204]" pageId="5" pageNumber="281" type="syntype">Syntypes</typeStatus>
: ♀ ♀,
<collectingCountry id="8DE07600A53EFF8B24DEFF31FBD0FF6B" box="[974,1034,183,204]" name="Japan" pageId="5" pageNumber="281">Japan</collectingCountry>
, Niigata Prefecture, Shidaihama (Itami coll., K. Tsuneki coll.). Not available (Article 16).
</treatmentCitationGroup>
</paragraph>
<paragraph id="F5483690A53EFF8B2787FF72FEFDFEE0" blockId="5.[151,1436,152,327]" pageId="5" pageNumber="281">
<treatmentCitationGroup id="D5E711BEA53EFF8B2787FF72FEFDFEE0" pageId="5" pageNumber="281">
<taxonomicName id="32F74D13A53EFF8B2787FF72FD2CFEAD" ID-CoL="CCFK" authority="Bohart &amp; Menke 1976: 458" authorityName="Bohart &amp; Menke" authorityPageNumber="458" authorityYear="1976" box="[151,758,244,266]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="subSpecies" species="spinosus" subSpecies="jaroslavensis">
<emphasis id="C783EA82A53EFF8B2787FF72FE14FEAD" box="[151,462,244,266]" italics="true" pageId="5" pageNumber="281">Alysson spinosus jaroslavensis</emphasis>
:
<treatmentCitation id="74561081A53EFF8B26CCFF72FD2CFEAD" author="Bohart" box="[476,758,244,266]" page="458" pageId="5" pageNumber="281" year="1976">
<bibRefCitation id="91664B61A53EFF8B26CCFF72FD1EFEAD" author="Bohart" box="[476,708,244,266]" pageId="5" pageNumber="281" refString="Bohart, R. M. &amp; Menke, A. S. (1976) Sphecid Wasps of the World. A generic revision. University of California Press, Berkeley, Los Angeles, London, ix + 695 pp." type="book" year="1976">Bohart &amp; Menke 1976</bibRefCitation>
: 458
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="74561081A53EFF8B2411FF72FBA8FEAD" author="Shlyakhtenok" box="[769,1138,244,266]" page="34" pageId="5" pageNumber="281" year="2002">
<bibRefCitation id="91664B61A53EFF8B2411FF72FB97FEAD" author="Shlyakhtenok" box="[769,1101,244,266]" pageId="5" pageNumber="281" refString="Shlyakhtenok, A. S. &amp; Skibinska, E. (2002) Zhalonosnye pereponchatokrylye (Hymenoptera, Aculeata) Belarusi: semeystvo royushchiye osy (Sphecidae). Vestnik Zoologii, 36, 31 - 40." type="journal article" year="2002">Shlyakhtenok &amp; Skibinska 2002</bibRefCitation>
: 34
</treatmentCitation>
.
<taxonomicName id="32F74D13A53EFF8B2787FE94FD8EFE8F" ID-CoL="CCFK" authority="Pulawski 1978: 212" authorityName="Pulawski" authorityPageNumber="212" authorityYear="1978" box="[151,596,274,296]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="jaroslavensis">
<emphasis id="C783EA82A53EFF8B2787FE94FEA9FE8F" box="[151,371,274,296]" italics="true" pageId="5" pageNumber="281">Alysson jaroslavensis</emphasis>
:
<treatmentCitation id="74561081A53EFF8B2694FE94FD8EFE8F" author="Pulawski" box="[388,596,274,296]" page="212" pageId="5" pageNumber="281" year="1978">
<bibRefCitation id="91664B61A53EFF8B2694FE94FDFAFE8F" author="Pulawski" box="[388,544,274,296]" pageId="5" pageNumber="281" refString="Pulawski, W. J. (1978) Nadsem. Sphecoidea. In: Medvedev, G. S. (Ed.), Opredelitel nasekomykh evropeyskoi chasti SSSR. Tom III. Pereponchatokrylye. Pervaya chast. Nauka, Leningrad, pp. 173 - 279." type="book chapter" year="1978">Pulawski 1978</bibRefCitation>
: 212
</treatmentCitation>
</taxonomicName>
, 213;
<treatmentCitation id="74561081A53EFF8B2588FE94FC7DFE8F" author="Pankevich" box="[664,935,274,296]" page="95" pageId="5" pageNumber="281" year="1989">
<bibRefCitation id="91664B61A53EFF8B2588FE94FCA4FE8F" author="Pankevich" box="[664,894,274,296]" pageId="5" pageNumber="281" refString="Pankevich, T. P., Belavskaya, V. I., Molchanova, P. V., Shlyakhtenok, A. S. &amp; Blinov, V. V. (1989) Otryad pereponchatokrylyie - Hymenoptera, In: Kryzhanovskiy, O. L. (Ed.), Flora i Fauna Zapovednikov SSSR. Nasekomyye Berezinskogo zapovednika. Operativnoinformatsionnyi material. Moskva, pp. 81 - 101." type="book chapter" year="1989">Pankevich et al. 1989</bibRefCitation>
: 95
</treatmentCitation>
;
<treatmentCitation id="74561081A53EFF8B24A4FE94FB2AFE8F" author="Blagoveshchenskaya" box="[948,1264,274,296]" page="91" pageId="5" pageNumber="281" year="1994">
<bibRefCitation id="91664B61A53EFF8B24A4FE94FB12FE8F" author="Blagoveshchenskaya" box="[948,1224,274,296]" pageId="5" pageNumber="281" refString="Blagoveshchenskaya, N. N. (1994) Katalog fauny zhalyashchikh pereponchatokrylykh (podotryad Aculeata) Ulyanovskoy Oblasti. Priroda Ulyanovskoy Oblasti, 5, 82 - 93." type="journal article" year="1994">Blagoveshchenskaya 1994</bibRefCitation>
: 91
</treatmentCitation>
;
<treatmentCitation id="74561081A53EFF8B23EEFE94FEF9FEE0" author="Voblenko" page="15" pageId="5" pageNumber="281" year="1996">
<bibRefCitation id="91664B61A53EFF8B23EEFE94FF27FEE0" author="Voblenko" pageId="5" pageNumber="281" refString="Voblenko, A. S., Gorobchishin, V. A. &amp; Nesterov, M. A. (1996) Digger wasps (Hymenoptera, Sphecidae) of Ukrainian Polesye. Sphecos, 30, 14 - 15." type="journal article" year="1996">Voblenko et al. 1996</bibRefCitation>
: 15
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BDED651BA53EFF8B2787FEF2FE96FC57" pageId="5" pageNumber="281" type="materials_examined">
<paragraph id="F5483690A53EFF8B2787FEF2FE96FC57" blockId="5.[151,1436,372,1773]" pageId="5" pageNumber="281">
<emphasis id="C783EA82A53EFF8B2787FEF2FEA0FE2A" bold="true" box="[151,378,372,397]" pageId="5" pageNumber="281">Material examined</emphasis>
.
<typeStatus id="2A4C8832A53EFF8B2699FEF2FE2BFE2A" box="[393,497,372,397]" pageId="5" pageNumber="281" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="32F74D13A53EFF8B2508FEF2FD09FE2B" box="[536,723,372,396]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis id="C783EA82A53EFF8B2508FEF2FD09FE2B" box="[536,723,372,396]" italics="true" pageId="5" pageNumber="281">Alysson fuscatus</emphasis>
</taxonomicName>
<taxonomicName id="32F74D13A53EFF8B25CBFEF3FC45FE2B" box="[731,927,372,396]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="variety" species="spinosus" variety="jaroslavensis">
var.
<emphasis id="C783EA82A53EFF8B241CFEF2FC45FE2B" box="[780,927,372,396]" italics="true" pageId="5" pageNumber="281">jaroslavensis</emphasis>
</taxonomicName>
Kokujev—♀,
<collectingCountry id="8DE07600A53EFF8B2357FEF2FB4CFE2A" box="[1095,1174,372,397]" name="Russia" pageId="5" pageNumber="281">Russia</collectingCountry>
: Yaroslavskaya Oblast, Yaroslavl,
<date id="81491050A53EFF8B260AFE1EFE7FFE17" box="[282,421,407,432]" pageId="5" pageNumber="281" value="1896-09-14">14.IX 1896</date>
(Kokujev) (
<collectionCode id="93E6AE55A53EFF8B2555FE1EFDA3FE17" box="[581,633,408,432]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
).
<collectingCountry id="8DE07600A53EFF8B2587FE11FD3CFE17" box="[663,742,407,432]" name="Russia" pageId="5" pageNumber="281">
<emphasis id="C783EA82A53EFF8B2587FE11FD3CFE17" bold="true" box="[663,742,407,432]" pageId="5" pageNumber="281">Russia</emphasis>
</collectingCountry>
:
<specimenCount id="E3F1FD19A53EFF8B2411FE1EFCFFFE17" box="[769,805,408,432]" pageId="5" pageNumber="281" type="generic">1♂</specimenCount>
, Leningradskaya Oblast, Tolmachevo,
<date id="81491050A53EFF8B2218FE1EFA46FE17" box="[1288,1436,407,432]" pageId="5" pageNumber="281" value="1936-07-13">13.VII 1936</date>
(Shtakelberg) (
<collectionCode id="93E6AE55A53EFF8B2658FE3AFEA6FE73" box="[328,380,444,468]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
); 2♀,
<specimenCount id="E3F1FD19A53EFF8B26C5FE3AFE23FE73" box="[469,505,444,468]" pageId="5" pageNumber="281" type="generic">1♂</specimenCount>
, Kostromskaya Oblast, Kostroma,
<date id="81491050A53EFF8B24BAFE3AFB9AFE73" box="[938,1088,444,468]" pageId="5" pageNumber="281" value="1924-07-24">24.VII 1924</date>
,
<date id="81491050A53EFF8B2345FE3AFB31FE72" box="[1109,1259,444,469]" pageId="5" pageNumber="281" value="1926-07-22">22.VII 1926</date>
,
<date id="81491050A53EFF8B2210FE3AFA46FE73" box="[1280,1436,444,468]" pageId="5" pageNumber="281" value="1933-08-10">10.VIII 1933</date>
(Gussakovskij) (
<collectionCode id="93E6AE55A53EFF8B2642FE66FE5CFE5F" box="[338,390,480,504]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
);
<specimenCount id="E3F1FD19A53EFF8B268CFE66FE1AFE5F" box="[412,448,480,504]" pageId="5" pageNumber="281" type="generic">1♂</specimenCount>
, Kostromskaya Oblast, Troitsk,
<date id="81491050A53EFF8B2424FE59FC65FE5F" box="[820,959,479,504]" pageId="5" pageNumber="281" value="1924-07-25">25.VII 1924</date>
,
<date id="81491050A53EFF8B24C8FE66FBBEFE5F" box="[984,1124,480,504]" pageId="5" pageNumber="281" value="1929-07-22">22.VII 1929</date>
(Gussakovskij) (
<collectionCode id="93E6AE55A53EFF8B2238FE66FA86FE5F" box="[1320,1372,480,504]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
); 1♀, Kostromskaya Oblast, Vasilevskoe,
<date id="81491050A53EFF8B2521FD82FD66FDBB" box="[561,700,516,540]" pageId="5" pageNumber="281" value="1933-07-21">21.VII 1933</date>
(Gussakovskij) (
<collectionCode id="93E6AE55A53EFF8B2490FD82FC6EFDBB" box="[896,948,516,540]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
); 1♀,
<specimenCount id="E3F1FD19A53EFF8B24EDFD82FBF8FDBB" box="[1021,1058,516,540]" pageId="5" pageNumber="281" type="generic">3♂</specimenCount>
, Kostromskaya Oblast, Baidarki,
<date id="81491050A53EFF8B2787FDAEFF39FDE7" box="[151,227,552,576]" pageId="5" pageNumber="281" value="1933-07">19.VII</date>
,
<date id="81491050A53EFF8B27E0FDAEFEAFFDE7" box="[240,373,552,576]" pageId="5" pageNumber="281" value="1933-08-01">1.VIII 1933</date>
(Gussakovskij) (
<collectionCode id="93E6AE55A53EFF8B2527FDAEFDB1FDE7" box="[567,619,552,576]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
); 1♀, Volgogradskaya Oblast, Volgograd (coll. F. Morawitz) (
<collectionCode id="93E6AE55A53EFF8B2239FDAEFA87FDE7" box="[1321,1373,552,576]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
); 2♀, Krasnoyarskii Krai,
<collectingCountry id="8DE07600A53EFF8B266AFDCAFE14FDC2" box="[378,462,588,613]" name="Malaysia" pageId="5" pageNumber="281">Malaya</collectingCountry>
Minusa, 4,
<date id="81491050A53EFF8B2542FDCBFDBAFDC3" box="[594,608,589,612]" pageId="5" pageNumber="281" value="2012-07-07">7</date>
.VII 2012 (Proshchalykin, Loktionov) (
<collectionCode id="93E6AE55A53EFF8B2308FDCAFB82FDC3" box="[1048,1112,588,612]" pageId="5" pageNumber="281">IBSS</collectionCode>
); 2♀, Krasnoyarskii Krai,
<quantity id="320F9B75A53EFF8B2290FDCAFF61FD2F" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="281" unit="km" value="40.0">40 km</quantity>
NE Minusinsk, Tes, Tuba River,
<date id="81491050A53EFF8B2540FDF6FD0AFD2F" box="[592,720,624,648]" pageId="5" pageNumber="281" value="2012-07-07">7.VII 2012</date>
(Proshchalykin, Loktionov) (
<collectionCode id="93E6AE55A53EFF8B233BFDF6FBB1FD2F" box="[1067,1131,624,648]" pageId="5" pageNumber="281">IBSS</collectionCode>
); 1♀, Krasnoyarskii Krai, Minusinsk,
<date id="81491050A53EFF8B260BFD12FE4FFD0B" box="[283,405,660,684]" pageId="5" pageNumber="281" value="2012-07-09">9.VII 2012</date>
(Proshchalykin, Loktionov) (
<collectionCode id="93E6AE55A53EFF8B25F0FD12FCFAFD0B" box="[736,800,660,684]" pageId="5" pageNumber="281">IBSS</collectionCode>
); 4♀,
<specimenCount id="E3F1FD19A53EFF8B2474FD12FC53FD0B" box="[868,905,660,684]" pageId="5" pageNumber="281" type="generic">2♂</specimenCount>
, Irkutskaya Oblast, Angarsk,
<date id="81491050A53EFF8B23C4FD12FA46FD0B" box="[1236,1436,660,684]" pageId="5" pageNumber="281" value="1983-07-11" valueMax="1983-09-09" valueMin="1983-07-11">11.VII9.IX 1983</date>
(Nemkov) (
<collectionCode id="93E6AE55A53EFF8B260EFD3EFE84FD77" box="[286,350,696,720]" pageId="5" pageNumber="281">IBSS</collectionCode>
); 4♀, Irkutskaya Oblast,
<quantity id="320F9B75A53EFF8B2599FD31FD0EFD77" box="[649,724,695,720]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="281" unit="km" value="15.0">15 km</quantity>
E Ust-Orda, Ordinsk,
<date id="81491050A53EFF8B24F3FD31FB5DFD77" box="[995,1159,695,720]" pageId="5" pageNumber="281" value="1994-08-02" valueMax="1994-08-06" valueMin="1994-08-02">26.VIII 1994</date>
(Nemkov) (
<collectionCode id="93E6AE55A53EFF8B220AFD3EFA80FD77" box="[1306,1370,696,720]" pageId="5" pageNumber="281">IBSS</collectionCode>
); 1♀, Khabarovskii Krai, Khabarovsk,
<date id="81491050A53EFF8B251FFD5AFD79FD53" box="[527,675,732,756]" pageId="5" pageNumber="281" value="1979-08-22">22.VIII 1979</date>
(Ermolenko) (
<collectionCode id="93E6AE55A53EFF8B245BFD5AFCA5FD53" box="[843,895,732,756]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
);
<specimenCount id="E3F1FD19A53EFF8B2485FD5AFC63FD53" box="[917,953,732,756]" pageId="5" pageNumber="281" type="generic">1♂</specimenCount>
, Khabarovskii Krai, Amurzet,
<date id="81491050A53EFF8B220BFD5AFA46FD52" box="[1307,1436,732,757]" pageId="5" pageNumber="281" value="1985-06-17">17.VI 1985</date>
(Belokobylskij) (
<collectionCode id="93E6AE55A53EFF8B2646FC86FE50FCBF" box="[342,394,768,792]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
); 1♀, Primorskii Krai, Yakovlevka,
<date id="81491050A53EFF8B2430FC86FC68FCBF" box="[800,946,767,792]" pageId="5" pageNumber="281" value="1926-08-23">23.VIII 1926</date>
(Dyakonov, Filipev) (
<collectionCode id="93E6AE55A53EFF8B23BCFC86FB3AFCBF" box="[1196,1248,768,792]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
); 1♀, Primorskii Krai, Olga,
<date id="81491050A53EFF8B2630FCA2FE76FC9B" box="[288,428,804,828]" pageId="5" pageNumber="281" value="1979-07-27">27.VII 1979</date>
(Belokobylskij) (
<collectionCode id="93E6AE55A53EFF8B2567FCA2FD71FC9B" box="[631,683,804,828]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
); 1♀, Primorskii Krai, Ussuriiskii Reserve, Kamenushka,
<date id="81491050A53EFF8B2259FCA2FF15FCC7" pageId="5" pageNumber="281" value="1982-08-12">12.VIII 1982</date>
(Nemkov) (
<collectionCode id="93E6AE55A53EFF8B264CFCCEFE46FCC7" box="[348,412,840,864]" pageId="5" pageNumber="281">IBSS</collectionCode>
); 1♀, Primorskii Krai, Ussuriiskii Reserve, Staraya Baza,
<date id="81491050A53EFF8B2325FCCEFB6CFCC7" box="[1077,1206,840,864]" pageId="5" pageNumber="281" value="1991-06-29">29.VI 1991</date>
(Lelej) (
<collectionCode id="93E6AE55A53EFF8B220CFCCEFA86FCC7" box="[1308,1372,840,864]" pageId="5" pageNumber="281">IBSS</collectionCode>
); 1♀, Primorskii Krai, Ryazanovka,
<date id="81491050A53EFF8B251FFCEAFD7EFC23" box="[527,676,876,901]" pageId="5" pageNumber="281" value="1992-07-15">15.VII 1992</date>
(Lelej) (
<collectionCode id="93E6AE55A53EFF8B240EFCEAFC84FC23" box="[798,862,876,900]" pageId="5" pageNumber="281">IBSS</collectionCode>
);
<specimenCount id="E3F1FD19A53EFF8B246EFCEAFC7EFC23" box="[894,932,876,900]" pageId="5" pageNumber="281" type="generic">2♂</specimenCount>
, Primorskii Krai, Khasan,
<date id="81491050A53EFF8B23EEFCEAFA46FC23" box="[1278,1436,876,900]" pageId="5" pageNumber="281" value="1998-08-12">12.VIII 1998</date>
(Belokobylskij) (
<collectionCode id="93E6AE55A53EFF8B2649FC16FE57FC0F" box="[345,397,912,936]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
); 3♀, Sakhalinskaya Oblast, Sakhalin Island,
<quantity id="320F9B75A53EFF8B24B1FC16FC31FC0F" box="[929,1003,911,936]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="281" unit="km" value="10.0">10 km</quantity>
E Boshnyakovo,
<date id="81491050A53EFF8B23ACFC16FA92FC0F" box="[1212,1352,912,936]" pageId="5" pageNumber="281" value="2003-07-24">24.VII 2003</date>
(Lelej, Storozhenko) (
<collectionCode id="93E6AE55A53EFF8B2651FC32FE5BFC6B" box="[321,385,948,972]" pageId="5" pageNumber="281">IBSS</collectionCode>
).
<collectingCountry id="8DE07600A53EFF8B2688FC32FDD0FC6A" box="[408,522,948,973]" name="Mongolia" pageId="5" pageNumber="281">
<emphasis id="C783EA82A53EFF8B2688FC32FDD0FC6A" bold="true" box="[408,522,948,973]" pageId="5" pageNumber="281">Mongolia</emphasis>
</collectingCountry>
:
<specimenCount id="E3F1FD19A53EFF8B250DFC32FD9BFC6B" box="[541,577,948,972]" pageId="5" pageNumber="281" type="generic">1♂</specimenCount>
, Khövsgöl aimag, Delger-Muren River near Buren-Khan,
<date id="81491050A53EFF8B23FEFC32FA46FC6A" box="[1262,1436,948,973]" pageId="5" pageNumber="281" value="1968-06-28" valueMax="1968-06-29" valueMin="1968-06-28">2829.VI 1968</date>
(Kozlov) (
<collectionCode id="93E6AE55A53EFF8B261BFC5EFEE5FC57" box="[267,319,984,1008]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="281">ZIN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="BDED651BA53EFF8B27D7FC7AFDD9F94A" pageId="5" pageNumber="281" type="distribution">
<paragraph id="F5483690A53EFF8B27D7FC7AFC52FB02" blockId="5.[151,1436,372,1773]" pageId="5" pageNumber="281">
<emphasis id="C783EA82A53EFF8B27D7FC7AFE83FBB2" bold="true" box="[199,345,1020,1045]" pageId="5" pageNumber="281">Distribution</emphasis>
.
<collectingCountry id="8DE07600A53EFF8B2662FC7AFE26FBB2" box="[370,508,1020,1045]" name="Netherlands" pageId="5" pageNumber="281">Netherlands</collectingCountry>
,
<collectingCountry id="8DE07600A53EFF8B2501FC7BFDBAFBB3" box="[529,608,1021,1044]" name="France" pageId="5" pageNumber="281">France</collectingCountry>
,
<collectingCountry id="8DE07600A53EFF8B2566FC7AFD3BFBB2" box="[630,737,1020,1045]" name="Denmark" pageId="5" pageNumber="281">Denmark</collectingCountry>
,
<collectingCountry id="8DE07600A53EFF8B25E7FC7AFC96FBB2" box="[759,844,1020,1045]" name="Austria" pageId="5" pageNumber="281">Austria</collectingCountry>
,
<collectingCountry id="8DE07600A53EFF8B2472FC7AFC69FBB2" box="[866,947,1020,1045]" name="Poland" pageId="5" pageNumber="281">Poland</collectingCountry>
,
<collectingCountry id="8DE07600A53EFF8B24D8FC7AFBF7FBB2" box="[968,1069,1020,1045]" name="Slovakia" pageId="5" pageNumber="281">Slovakia</collectingCountry>
,
<collectingCountry id="8DE07600A53EFF8B2353FC7BFB7CFBB2" box="[1091,1190,1021,1045]" name="Hungary" pageId="5" pageNumber="281">Hungary</collectingCountry>
,
<collectingCountry id="8DE07600A53EFF8B23ACFC7AFAF0FBB2" box="[1212,1322,1020,1045]" name="Lithuania" pageId="5" pageNumber="281">Lithuania</collectingCountry>
,
<collectingCountry id="8DE07600A53EFF8B2250FC7AFA42FBB2" box="[1344,1432,1020,1045]" name="Belarus" pageId="5" pageNumber="281">Belarus</collectingCountry>
,
<collectingCountry id="8DE07600A53EFF8B2787FB99FF2EFB9F" box="[151,244,1055,1080]" name="Ukraine" pageId="5" pageNumber="281">Ukraine</collectingCountry>
,
<collectingCountry id="8DE07600A53EFF8B2616FB99FE8BFB9F" box="[262,337,1055,1080]" name="Russia" pageId="5" pageNumber="281">Russia</collectingCountry>
(Leningradskaya Oblast, Yaroslavskaya Oblast, Kostromskaya Oblast, Moskovskaya Oblast, Bryanskaya Oblast, Volgogradskaya Oblast, Ulyanovskaya Oblast, Altai, Krasnoyarskii Krai, Irkutskaya Oblast, Buryatiya, Amurskaya Oblast, Khabarovskii Krai, Primorskii Krai, Sakhalin, Kuril Islands: Kunashir),
<collectingCountry id="8DE07600A53EFF8B2238FBE1FA42FB27" box="[1320,1432,1127,1152]" name="Mongolia" pageId="5" pageNumber="281">Mongolia</collectingCountry>
,
<collectingCountry id="8DE07600A53EFF8B2787FB0AFF00FB02" box="[151,218,1164,1189]" name="China" pageId="5" pageNumber="281">China</collectingCountry>
(Heilongjiang,
<collectingCountry id="8DE07600A53EFF8B269AFB0AFE3AFB02" box="[394,480,1164,1189]" name="Taiwan" pageId="5" pageNumber="281">Taiwan</collectingCountry>
),
<collectingCountry id="8DE07600A53EFF8B26E4FB0AFD1FFB03" box="[500,709,1164,1189]" name="South Korea" pageId="5" pageNumber="281">Republic of Korea</collectingCountry>
,
<collectingCountry id="8DE07600A53EFF8B25C0FB0BFCCAFB03" box="[720,784,1165,1188]" name="Japan" pageId="5" pageNumber="281">Japan</collectingCountry>
(Honshu).
</paragraph>
<paragraph id="F5483690A53EFF8B27D7FB36FE66FA22" blockId="5.[151,1436,372,1773]" pageId="5" pageNumber="281">
<emphasis id="C783EA82A53EFF8B27D7FB36FD8BFB6D" bold="true" box="[199,593,1200,1226]" pageId="5" pageNumber="281">Justification of New Synonymy</emphasis>
.
<bibRefCitation id="91664B61A53EFF8B2572FB36FCC4FB6D" author="Kokujev" box="[610,798,1200,1226]" pageId="5" pageNumber="281" refString="Kokujev, N. R. (1906) O russkich predstavitelakh podsemeystva Alysonini Dalla Torre (Hymenoptera, Crabronidae). Horae Societatis Entomologicae Rossicae, 37, 209 - 219." type="journal article" year="1906">Kokujev (1906)</bibRefCitation>
described
<taxonomicName id="32F74D13A53EFF8B24B3FB36FBAAFB6E" box="[931,1136,1199,1226]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="variety" species="spinosus" variety="jaroslavensis">
var.
<emphasis id="C783EA82A53EFF8B24C7FB29FBAAFB6E" box="[983,1136,1199,1225]" italics="true" pageId="5" pageNumber="281">jaroslavensis</emphasis>
</taxonomicName>
as a form of
<taxonomicName id="32F74D13A53EFF8B2209FB29FA46FB6E" box="[1305,1436,1199,1225]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis id="C783EA82A53EFF8B2209FB29FA46FB6E" box="[1305,1436,1199,1225]" italics="true" pageId="5" pageNumber="281">A. fuscatus</emphasis>
</taxonomicName>
(correctly
<taxonomicName id="32F74D13A53EFF8B2604FB52FE45FB49" box="[276,415,1236,1262]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="C783EA82A53EFF8B2604FB52FE45FB49" box="[276,415,1236,1262]" italics="true" pageId="5" pageNumber="281">A. spinosus</emphasis>
</taxonomicName>
).
<bibRefCitation id="91664B61A53EFF8B26ACFB53FD23FB48" author="Bohart" box="[444,761,1237,1263]" pageId="5" pageNumber="281" refString="Bohart, R. M. &amp; Menke, A. S. (1976) Sphecid Wasps of the World. A generic revision. University of California Press, Berkeley, Los Angeles, London, ix + 695 pp." type="book" year="1976">Bohart and Menke (1976)</bibRefCitation>
treated
<taxonomicName id="32F74D13A53EFF8B2471FB52FB51FB49" box="[865,1163,1236,1262]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="subSpecies" species="fuscatus" subSpecies="jaroslavensis">
<emphasis id="C783EA82A53EFF8B2471FB52FB51FB49" box="[865,1163,1236,1262]" italics="true" pageId="5" pageNumber="281">A. fuscatus jaroslavensis</emphasis>
</taxonomicName>
as a subspecies of
<taxonomicName id="32F74D13A53EFF8B2293FB52FF26FAB3" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="C783EA82A53EFF8B2293FB52FF26FAB3" italics="true" pageId="5" pageNumber="281">A. spinosus</emphasis>
</taxonomicName>
. However, it differs from the latter in having a red propodeum (black in
<taxonomicName id="32F74D13A53EFF8B235BFB7CFB0AFAB3" box="[1099,1232,1274,1300]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="C783EA82A53EFF8B235BFB7CFB0AFAB3" box="[1099,1232,1274,1300]" italics="true" pageId="5" pageNumber="281">A. spinosus</emphasis>
</taxonomicName>
) and is similar in this and other characters to
<taxonomicName id="32F74D13A53EFF8B26F7FA99FDAFFA9E" box="[487,629,1311,1337]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="pertheesi">
<emphasis id="C783EA82A53EFF8B26F7FA99FDAFFA9E" box="[487,629,1311,1337]" italics="true" pageId="5" pageNumber="281">A. pertheesi</emphasis>
</taxonomicName>
.
<bibRefCitation id="91664B61A53EFF8B2597FAA6FC90FA9D" author="Pulawski" box="[647,842,1312,1338]" pageId="5" pageNumber="281" refString="Pulawski, W. J. (1978) Nadsem. Sphecoidea. In: Medvedev, G. S. (Ed.), Opredelitel nasekomykh evropeyskoi chasti SSSR. Tom III. Pereponchatokrylye. Pervaya chast. Nauka, Leningrad, pp. 173 - 279." type="book chapter" year="1978">Pulawski (1978)</bibRefCitation>
regarded
<taxonomicName id="32F74D13A53EFF8B24D4FA99FB5BFA9E" box="[964,1153,1311,1337]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="jaroslavensis">
<emphasis id="C783EA82A53EFF8B24D4FA99FB5BFA9E" box="[964,1153,1311,1337]" italics="true" pageId="5" pageNumber="281">A. jaroslavensis</emphasis>
</taxonomicName>
as a valid species. The examination of the
<typeStatus id="2A4C8832A53EFF8B266EFAC3FE3FFAF8" box="[382,485,1349,1375]" pageId="5" pageNumber="281" type="holotype">holotype</typeStatus>
of
<taxonomicName id="32F74D13A53EFF8B251EFAC2FD08FAF9" box="[526,722,1348,1374]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis id="C783EA82A53EFF8B251EFAC2FD08FAF9" box="[526,722,1348,1374]" italics="true" pageId="5" pageNumber="281">Alysson fuscatus</emphasis>
</taxonomicName>
<taxonomicName id="32F74D13A53EFF8B25CBFAC3FC7DFAF9" box="[731,935,1348,1375]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="variety" species="fuscatus" variety="jaroslavensis">
var.
<emphasis id="C783EA82A53EFF8B241EFAC2FC7DFAF9" box="[782,935,1348,1374]" italics="true" pageId="5" pageNumber="281">jaroslavensis</emphasis>
</taxonomicName>
demonstrated that it is a junior subjective synonym of
<taxonomicName id="32F74D13A53EFF8B2639FAECFE6FFA23" box="[297,437,1386,1412]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="pertheesi">
<emphasis id="C783EA82A53EFF8B2639FAECFE6FFA23" box="[297,437,1386,1412]" italics="true" pageId="5" pageNumber="281">A. pertheesi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F5483690A53EFF8B27D7FA16FDD9F94A" blockId="5.[151,1436,372,1773]" pageId="5" pageNumber="281">
In the original description of
<taxonomicName id="32F74D13A53EFF8B2536FA16FC5DFA0F" authority="Tsuneki (1967)" authorityName="Tsuneki" authorityYear="1967" box="[550,903,1423,1448]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="harbinensis">
<emphasis id="C783EA82A53EFF8B2536FA16FD17FA0F" box="[550,717,1424,1448]" italics="true" pageId="5" pageNumber="281">A. harbinensis</emphasis>
<bibRefCitation id="91664B61A53EFF8B25C8FA09FC5DFA0F" author="Tsuneki" box="[728,903,1423,1448]" pageId="5" pageNumber="281" refString="Tsuneki, K. (1967) Further studies on the fossorial Hymenoptera from Manchuria. Etizenia, 23,1 - 17." type="book" year="1967">Tsuneki (1967)</bibRefCitation>
</taxonomicName>
indicated that the female (the male was still unknown) differs from
<taxonomicName id="32F74D13A53EFF8B268DFA32FDF9FA6B" box="[413,547,1460,1484]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="pertheesi">
<emphasis id="C783EA82A53EFF8B268DFA32FDF9FA6B" box="[413,547,1460,1484]" italics="true" pageId="5" pageNumber="281">A. pertheesi</emphasis>
</taxonomicName>
mainly in having a red metapleuron and the propodeal enclosure (enclosure all black in
<taxonomicName id="32F74D13A53EFF8B27ECFA5EFEB9FA57" box="[252,355,1496,1520]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="pertheesi">
<emphasis id="C783EA82A53EFF8B27ECFA5EFEB9FA57" box="[252,355,1496,1520]" italics="true" pageId="5" pageNumber="281">pertheesi</emphasis>
</taxonomicName>
). The considerable color variation in the Eastern Asian populations of
<taxonomicName id="32F74D13A53EFF8B2387FA5EFAFAFA57" box="[1175,1312,1496,1520]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="pertheesi">
<emphasis id="C783EA82A53EFF8B2387FA5EFAFAFA57" box="[1175,1312,1496,1520]" italics="true" pageId="5" pageNumber="281">A. pertheesi</emphasis>
</taxonomicName>
was noted later by
<bibRefCitation id="91664B61A53EFF8B27EDFA7AFDF1F9B3" author="Tsuneki" box="[253,555,1532,1557]" pageId="5" pageNumber="281" refString="Tsuneki, K. &amp; Itami, I. (1973) Variation in colour of Alysson pertheesi Gorski with notes on other characters. The Life Study (Fukui), 17, 13." type="journal article" year="1973">Tsuneki and Itami (1973)</bibRefCitation>
. I was not able to examine the
<typeStatus id="2A4C8832A53EFF8B24BDFA7AFBD5F9B2" box="[941,1039,1532,1557]" pageId="5" pageNumber="281" type="holotype">holotype</typeStatus>
of
<taxonomicName id="32F74D13A53EFF8B232FFA7AFB3CF9B3" box="[1087,1254,1532,1556]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="harbinensis">
<emphasis id="C783EA82A53EFF8B232FFA7AFB3CF9B3" box="[1087,1254,1532,1556]" italics="true" pageId="5" pageNumber="281">A. harbinensis</emphasis>
</taxonomicName>
because of the temporary closure of the collection at the Leiden Museum, but the diagnostic characters of this species are clearly within the limits of the individual variation of
<taxonomicName id="32F74D13A53EFF8B25D2F9C2FC96F9FB" box="[706,844,1604,1628]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="pertheesi">
<emphasis id="C783EA82A53EFF8B25D2F9C2FC96F9FB" box="[706,844,1604,1628]" italics="true" pageId="5" pageNumber="281">A. pertheesi</emphasis>
</taxonomicName>
. Specimens with a red propodeal enclosure (as in
<taxonomicName id="32F74D13A53EFF8B2787F9EEFEC0F927" box="[151,282,1640,1664]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="harbinensis">
<emphasis id="C783EA82A53EFF8B2787F9EEFEC0F927" box="[151,282,1640,1664]" italics="true" pageId="5" pageNumber="281">harbinensis</emphasis>
</taxonomicName>
) occur in all parts of
<collectingCountry id="8DE07600A53EFF8B2534F9E1FDB5F927" box="[548,623,1639,1664]" name="Russia" pageId="5" pageNumber="281">Russia</collectingCountry>
along with the typically colored specimens (with a black enclosure), although in the western
<collectingCountry id="8DE07600A53EFF8B26B5F90AFE2AF902" box="[421,496,1676,1701]" name="Russia" pageId="5" pageNumber="281">Russia</collectingCountry>
the typical form prevails over the
<taxonomicName id="32F74D13A53EFF8B2465F90AFC22F903" box="[885,1016,1676,1700]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="harbinensis">
<emphasis id="C783EA82A53EFF8B2465F90AFC22F903" box="[885,1016,1676,1700]" italics="true" pageId="5" pageNumber="281">harbinensis</emphasis>
</taxonomicName>
form, while in the eastern
<collectingCountry id="8DE07600A53EFF8B2237F90AFAA8F902" box="[1319,1394,1676,1701]" name="Russia" pageId="5" pageNumber="281">Russia</collectingCountry>
the specimens with black enclosure are found very rarely. Therefore I regard
<taxonomicName id="32F74D13A53EFF8B24E4F936FB40F96F" box="[1012,1178,1712,1736]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="harbinensis">
<emphasis id="C783EA82A53EFF8B24E4F936FB40F96F" box="[1012,1178,1712,1736]" italics="true" pageId="5" pageNumber="281">A. harbinensis</emphasis>
</taxonomicName>
as a junior subjective synonym of
<taxonomicName id="32F74D13A53EFF8B2633F952FDD9F94A" authority="Gorski." authorityName="Gorski." box="[291,515,1748,1773]" class="Insecta" family="Crabronidae" genus="Alysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="pertheesi">
<emphasis id="C783EA82A53EFF8B2633F952FE72F94B" box="[291,424,1748,1772]" italics="true" pageId="5" pageNumber="281">A. pertheesi</emphasis>
Gorski.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>