282 lines
27 KiB
XML
282 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/nl.44.70359" ID-GBIF-Dataset="e68244ef-cd68-488f-9afb-c23ca55ae447" ID-GBIF-Taxon="189469089" ID-Pensoft-Pub="2367-5365-44-193" ID-Pensoft-UUID="18B5BF4D68665E9EB5D8A8BF6B64641D" ID-ZooBank="EA6D65A049FE420FB0E55908EBF72D02" LSID="urn:lsid:zoobank.org:act:F744DF74-AE2D-4236-8089-076C74D3EB81" ModsDocID="2367-5365-44-193" checkinTime="1633780208711" checkinUser="pensoft" docAuthor="Zilli, Alberto" docDate="2021" docId="7BF5F85461F2507F95175E720F7A12AD" docLanguage="en" docName="NotaLepidopt 44: 193-211" docOrigin="Nota Lepidopterologica 44" docPubDate="2021-10-08" docSource="http://dx.doi.org/10.3897/nl.44.70359" docTitle="Tabwecala Zilli 2021, gen. nov." docType="treatment" docUuid="F744DF74-AE2D-4236-8089-076C74D3EB81" docUuidSource="ZooBank" docVersion="3" id="18B5BF4D68665E9EB5D8A8BF6B64641D" lastPageNumber="193" masterDocId="18B5BF4D68665E9EB5D8A8BF6B64641D" masterDocTitle="Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu and its systematic position in the ' Ophiusini - Poaphilini ' clade (Lepidoptera, Erebidae)" masterLastPageNumber="211" masterPageNumber="193" pageId="0" pageNumber="193" updateTime="1635284740426" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu and its systematic position in the ' Ophiusini - Poaphilini ' clade (Lepidoptera, Erebidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Zilli, Alberto</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3416-8069</mods:nameIdentifier>
|
||
<mods:affiliation>Natural History Museum, Life Sciences, Cromwell Road, SW 7 5 BD London, UK; a. zilli @ nhm. ac. uk; https: // orcid. org / 0000 - 0002 - 3416 - 8069</mods:affiliation>
|
||
<mods:nameIdentifier type="email">a.zilli@nhm.ac.uk</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Nota Lepidopterologica</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2021</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2021-10-08</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>44</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>193</mods:start>
|
||
<mods:end>211</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/nl.44.70359</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/nl.44.70359</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">2367-5365-44-193</mods:identifier>
|
||
<mods:identifier type="ZooBank">EA6D65A049FE420FB0E55908EBF72D02</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">18B5BF4D68665E9EB5D8A8BF6B64641D</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="189469089" LSID="urn:lsid:zoobank.org:act:F744DF74-AE2D-4236-8089-076C74D3EB81" httpUri="http://treatment.plazi.org/id/7BF5F85461F2507F95175E720F7A12AD" lastPageNumber="193" pageId="0" pageNumber="193">
|
||
<subSubSection pageId="0" pageNumber="193" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<taxonomicName LSID="http://zoobank.org/F744DF74-AE2D-4236-8089-076C74D3EB81" authority="Zilli, 2021" authorityName="Zilli" authorityYear="2021" class="Insecta" family="Erebidae" genus="Tabwecala" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Tabwecala" order="Lepidoptera" pageId="0" pageNumber="193" rank="genus" status="gen. nov.">Tabwecala</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="193">gen. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="193" type="type species">
|
||
<paragraph pageId="0" pageNumber="193">Type species.</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<taxonomicName authorityName="Zilli" authorityYear="2021" class="Insecta" family="Erebidae" genus="Tabwecala" kingdom="Animalia" lsidName="Tabwecala robinsoni" order="Lepidoptera" pageId="0" pageNumber="193" rank="species" species="robinsoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Tabwecala robinsoni</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (described below), by present designation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="193" type="etymology">
|
||
<paragraph pageId="0" pageNumber="193">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
The generic epithet derives from the combination of
|
||
<normalizedToken originalValue="“Tabwe-”">"Tabwe-"</normalizedToken>
|
||
(from Mt Tabwemasana in Espiritu Santo Island) and
|
||
<normalizedToken originalValue="“-cala”">"-cala"</normalizedToken>
|
||
(from the Greek
|
||
<normalizedToken originalValue="“κᾰλή”">"κᾰλή"</normalizedToken>
|
||
, meaning
|
||
<normalizedToken originalValue="“nice”">"nice"</normalizedToken>
|
||
); it is feminine in gender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="193" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="193">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
The absence of sexual dimorphism in the labial palpi, the presence of two anterior spines on the male profemur, the absence of spines on male tibiae, the presence of an androteca on male mesotibia, the strongly modified male metatibia bearing a dorsal hair pencil, the tuba analis without scaphium and with a deep, hood-like scaphial pocket, and the sterigma projected posteriorly into a free end, represent altogether a combination of characters not seen, at the
|
||
<normalizedToken originalValue="author’s">author's</normalizedToken>
|
||
best knowledge, in other genera of the
|
||
<taxonomicName authorityName="Leach" authorityYear="1815" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="family">Erebidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="193" type="description">
|
||
<paragraph pageId="0" pageNumber="193">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<emphasis bold="true" pageId="0" pageNumber="193">Male</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Habitus of Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu, Espiritu Santo, Mt Tabwemasana. 1. Holotype ♂; 2. Paratype ♀. Scale bar: 1 cm (1, 2.)." figureDoi="10.3897/nl.44.70359.figures1-2" httpUri="https://binary.pensoft.net/fig/597051" pageId="0" pageNumber="193">1</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="193">
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
Large, feebly sunk into thorax, frons slightly bulged, unscaled at middle (scales evidently brushed off by labial palps after emergence as in numerous other
|
||
<taxonomicName authorityName="Leach" authorityYear="1815" lsidName="" pageId="0" pageNumber="193" rank="subFamily" subFamily="Erebinae">Erebinae</taxonomicName>
|
||
), vertex roughly scaled, eye large, globular, antenna filiform, scape stout, barrel-like, covered by scales projected into small dorsal hood, pedicel small globular, flagellum consisting of short cylindrical, ventrally very shortly ciliate antennomeres, pilifer conspicuous, with long bristles, haustellum well developed, with tiny flat apical sensilla, labial palpus slender, upcurved, first joint roughly scaled, second joint compactly scaled, third joint long and thin, rod-like, two thirds as long as second, with no apical club.
|
||
</paragraph>
|
||
<caption doi="10.3897/nl.44.70359.figures1-2" httpUri="https://binary.pensoft.net/fig/597051" pageId="0" pageNumber="193" start="Figures 1, 2" startId="F1">
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<emphasis bold="true" pageId="0" pageNumber="193">Figures 1, 2.</emphasis>
|
||
Habitus of
|
||
<taxonomicName authorityName="Zilli" authorityYear="2021" class="Insecta" family="Erebidae" genus="Tabwecala" kingdom="Animalia" lsidName="Tabwecala robinsoni" order="Lepidoptera" pageId="0" pageNumber="193" rank="species" species="robinsoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Tabwecala robinsoni</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov., from Vanuatu, Espiritu Santo, Mt Tabwemasana.
|
||
<emphasis bold="true" pageId="0" pageNumber="193">1.</emphasis>
|
||
Holotype ♂;
|
||
<emphasis bold="true" pageId="0" pageNumber="193">2.</emphasis>
|
||
Paratype ♀. Scale bar: 1 cm (
|
||
<emphasis bold="true" pageId="0" pageNumber="193">1, 2.</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<emphasis bold="true" pageId="0" pageNumber="193">
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Thorax</emphasis>
|
||
.
|
||
</emphasis>
|
||
Stout, both patagium and tegula broad and long, roughly scaled, meso-metanotum with hair-like scales, without crests; pectus conspicuously hairy. Wings broad; forewing vein R1 from middle of cell, areole present, splitting between Rs2 and Rs3 close to termen in apical area of wing, but topology of Rs branches from cell very variable (see dedicated paragraph below), M visible in outer half of cell, other veins as in other
|
||
<taxonomicName authorityName="Leach" authorityYear="1815" lsidName="" pageId="0" pageNumber="193" rank="subFamily" subFamily="Erebinae">Erebinae</taxonomicName>
|
||
, with CuA2 from before middle of cell; hindwing veins as in other
|
||
<taxonomicName authorityName="Leach" authorityYear="1815" lsidName="" pageId="0" pageNumber="193" rank="subFamily" subFamily="Erebinae">Erebinae</taxonomicName>
|
||
. Legs slender (Fig.
|
||
<figureCitation captionStart="Figures 3, 4" captionStartId="F2" captionText="Figures 3, 4. Legs of Tabwecala robinsoni gen. nov., sp. nov. 3. Male; 4. Female (this also with procoxa). Scale bar: 2 mm. (3, 4.)" figureDoi="10.3897/nl.44.70359.figures3-4" httpUri="https://binary.pensoft.net/fig/597052" pageId="0" pageNumber="193">3</figureCitation>
|
||
); foreleg: profemur elongated, slightly wider at base and gradually tapered distally, with paired preapical spines, protibia not spined, barely shorter than femur, and feebly incrassate at middle, with epiphysis half length of tibia, protarsus conspicuously spined ventrally on tarsomeres I-IV, spines arranged into 3 rows, terminal spines of each tarsomere longest, spines replaced by bristles on tarsomere V, pretarsus with semi-circular arolium, onychia sharply bent, with midventral tooth, and pulvilli short; midleg: longest of the three legs, mesofemur long and slender, of almost uniform width, mesotibia greatly developed, longer and wider than mesofemur, its edges wrapping internally so as to originate a groove (androteca, cf. Berio 1955,
|
||
<bibRefCitation author="Berio, E" journalOrPublisher="Annali del Museo civico di Storia naturale " Giacomo Doria " (Genova)" pageId="0" pageNumber="193" pagination="276 - 327" publicationUrl="https://www.biodiversitylibrary.org/page/35318765" refId="B8" refString="Berio, E, 1959. Studi sulla sistematica delle cosiddette " Catocalinae " e " Othreinae " (Lepidoptera, Noctuidae). Annali del Museo civico di Storia naturale "Giacomo Doria" (Genova) 71 (1960): 276 - 327, https://www.biodiversitylibrary.org/page/35318765" title="Studi sulla sistematica delle cosiddette " Catocalinae " e " Othreinae " (Lepidoptera, Noctuidae)." url="https://www.biodiversitylibrary.org/page/35318765" volume="71" year="1959">1959</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Berio, E" journalOrPublisher="Annali del Museo civico di Storia naturale " Giacomo Doria " (Genova)" pageId="0" pageNumber="193" pagination="181 - 332" publicationUrl="https://www.biodiversitylibrary.org/page/35589416" refId="B9" refString="Berio, E, 1965. Le catocaline africane a tibie spinose del Museo di Tervuren. Annali del Museo civico di Storia naturale "Giacomo Doria" (Genova) 75 (1964-1965): 181 - 332, https://www.biodiversitylibrary.org/page/35589416" title="Le catocaline africane a tibie spinose del Museo di Tervuren." url="https://www.biodiversitylibrary.org/page/35589416" volume="75" year="1965">1965</bibRefCitation>
|
||
) enclosing a long hair-pencil kept in place by flat scales, mesotibial spurs one terminal pair, mesotarsus slender, spines and bristles on I-V as in protarsus, pretarsus as in protarsus; hindleg: metafemur slender and of almost uniform width, 7/10 as long as mesofemur, metatibia not spined and greatly modified, very short, about 2/3 length of metafemur, and swollen distally and dorsally so as to show a teardrop outline with dorso-apical bulge (respecting tibio-tarsal axis) for insertion of conspicuous pencil of long, hair-like androconial scales, this leaning middorsally along basal tarsomere, metatibial spurs two pairs, long and flimsy, first pair from middle of metatibia, second pair from well before apex, mesotarsus with basal tarsomere greatly developed, thick and longer than tarsomeres II-V altogether, widest at base and gradually tapered apically, clothed externally with thin long setae and showing midventrally in its distal two thirds comb of long bristles, tarsomeres II-V and pretarsus as in fore- and midleg.
|
||
</paragraph>
|
||
<caption doi="10.3897/nl.44.70359.figures3-4" httpUri="https://binary.pensoft.net/fig/597052" pageId="0" pageNumber="193" start="Figures 3, 4" startId="F2">
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<emphasis bold="true" pageId="0" pageNumber="193">Figures 3, 4.</emphasis>
|
||
Legs of
|
||
<taxonomicName authorityName="Zilli" authorityYear="2021" class="Insecta" family="Erebidae" genus="Tabwecala" kingdom="Animalia" lsidName="Tabwecala robinsoni" order="Lepidoptera" pageId="0" pageNumber="193" rank="species" species="robinsoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Tabwecala robinsoni</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="193">3.</emphasis>
|
||
Male;
|
||
<emphasis bold="true" pageId="0" pageNumber="193">4.</emphasis>
|
||
Female (this also with procoxa). Scale bar: 2 mm. (
|
||
<emphasis bold="true" pageId="0" pageNumber="193">3, 4.</emphasis>
|
||
)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<emphasis bold="true" pageId="0" pageNumber="193">
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Abdomen</emphasis>
|
||
.
|
||
</emphasis>
|
||
Sternum A2 broad, subrectangular, with short, broad-based anterior apodemes, other segments of pregenital abdomen with subrectangular urotergites and urosternites, much wider than high, tergum A8 trapezoid, sternum A8 shallowly cleft on its posterior margin.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="193">Male genitalia</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figures 5, 6" captionStartId="F3" captionText="Figures 5, 6. Genitalia of Tabwecala robinsoni gen. nov., sp. nov. 5. Male, paratype; 6. Female, paratype (sternum A 7 on left). Scale bar: 1 mm (5, 6.)." figureDoi="10.3897/nl.44.70359.figures5-6" httpUri="https://binary.pensoft.net/fig/597053" pageId="0" pageNumber="193">5</figureCitation>
|
||
). Tegumen and vinculum robust, the latter shorter than tegumen and without inwardly produced saccus, valva oblong, with strongly sclerotised proximo-ventral portion, deriving from fusion between costal process and clasper, narrowly joined to sacculus, this produced into short saccular process, and dorso-distal membranous portion, which is broadly but shallowly corematous dorsally at base, where a conspicuous scent tuft is inserted. Uncus with dorsal process far from base. Tuba analis of complex configuration, dorsally membranous, showing distinct hood-like scaphial pocket (best seen in side view) above anus to accommodate tip of uncus. Juxta long, symmetrical, of the
|
||
<normalizedToken originalValue="‘X’">'X'</normalizedToken>
|
||
-type, articulated at very base of valvae. Phallus arched and compressed, with recurved foot-shaped coecum, showing small sclerotised platelet at base of vesica in distal excision of ventral edge of its shaft, vesica multi-lobed.
|
||
</paragraph>
|
||
<caption doi="10.3897/nl.44.70359.figures5-6" httpUri="https://binary.pensoft.net/fig/597053" pageId="0" pageNumber="193" start="Figures 5, 6" startId="F3">
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<emphasis bold="true" pageId="0" pageNumber="193">Figures 5, 6.</emphasis>
|
||
Genitalia of
|
||
<taxonomicName authorityName="Zilli" authorityYear="2021" class="Insecta" family="Erebidae" genus="Tabwecala" kingdom="Animalia" lsidName="Tabwecala robinsoni" order="Lepidoptera" pageId="0" pageNumber="193" rank="species" species="robinsoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Tabwecala robinsoni</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="193">5.</emphasis>
|
||
Male, paratype;
|
||
<emphasis bold="true" pageId="0" pageNumber="193">6.</emphasis>
|
||
Female, paratype (sternum A7 on left). Scale bar: 1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="193">5, 6.</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<emphasis bold="true" pageId="0" pageNumber="193">Female</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Habitus of Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu, Espiritu Santo, Mt Tabwemasana. 1. Holotype ♂; 2. Paratype ♀. Scale bar: 1 cm (1, 2.)." figureDoi="10.3897/nl.44.70359.figures1-2" httpUri="https://binary.pensoft.net/fig/597051" pageId="0" pageNumber="193">2</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="193">
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
As in male, no sexual dimorphism detectable, including labial palpus, except for base of flagellum, slightly thinner, and periscapular scales, shorter and not forming hood.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<emphasis bold="true" pageId="0" pageNumber="193">
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Thorax</emphasis>
|
||
.
|
||
</emphasis>
|
||
Pro-metathorax and wings as in male, legs without the sexual secondary modifications seen in male (namely, profemural spines, mesotibial androteca, shortened and swollen metatibia bearing dorsal scent pencil and weak spurs, strongly developed basal segment of metatarsus with setae and comb), mesotibia irregularly spined externally, metatibia not spined, basal segment of metatarsus as long as tarsomeres II+III, with three rows of spines ventrally (Fig.
|
||
<figureCitation captionStart="Figures 3, 4" captionStartId="F2" captionText="Figures 3, 4. Legs of Tabwecala robinsoni gen. nov., sp. nov. 3. Male; 4. Female (this also with procoxa). Scale bar: 2 mm. (3, 4.)" figureDoi="10.3897/nl.44.70359.figures3-4" httpUri="https://binary.pensoft.net/fig/597052" pageId="0" pageNumber="193">4</figureCitation>
|
||
); other features as in male.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<emphasis bold="true" pageId="0" pageNumber="193">
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Abdomen</emphasis>
|
||
.
|
||
</emphasis>
|
||
As in male but tergum A7 broad and high, approximately square-shaped, and sternum A7 greatly reduced to narrow and long subrectangular lodix plate (Fig.
|
||
<figureCitation captionStart="Figures 5, 6" captionStartId="F3" captionText="Figures 5, 6. Genitalia of Tabwecala robinsoni gen. nov., sp. nov. 5. Male, paratype; 6. Female, paratype (sternum A 7 on left). Scale bar: 1 mm (5, 6.)." figureDoi="10.3897/nl.44.70359.figures5-6" httpUri="https://binary.pensoft.net/fig/597053" pageId="0" pageNumber="193">6</figureCitation>
|
||
) overlapping ostium bursae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="193">Female genitalia</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figures 5, 6" captionStartId="F3" captionText="Figures 5, 6. Genitalia of Tabwecala robinsoni gen. nov., sp. nov. 5. Male, paratype; 6. Female, paratype (sternum A 7 on left). Scale bar: 1 mm (5, 6.)." figureDoi="10.3897/nl.44.70359.figures5-6" httpUri="https://binary.pensoft.net/fig/597053" pageId="0" pageNumber="193">6</figureCitation>
|
||
). Complete sterigma present and of complex configuration, corresponding to broadly dilated midventral section of intersegmental membrane A7-A8 produced midventrally into stout, strongly sclerotised longitudinal outgrowth that terminates posteriorly into free end overlapping midventral sclerotisation which extends between from post-sterigmatal area to sternum A8; ostium bursae opening at approximately one third from anterior margin of sterigma. Bursa copulatrix with broad cervix and appendix anteriorly oriented originating from left of this. Ovipositor short, papillae anales soft, elongated, densely setose. Gonapophyses rod-like.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="193" type="venational variation">
|
||
<paragraph pageId="0" pageNumber="193">Venational variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
Two different configurations regarding topology of forewing Rs veins, not linked to the sexes of specimens examined, have been found. As noted in the 'materials and
|
||
<normalizedToken originalValue="methods’">methods'</normalizedToken>
|
||
, these are reported here after a descriptive approach (italics between brackets) and an interpretative one (normal text and Figs
|
||
<figureCitation captionStart="Figures 7, 8" captionStartId="F4" captionText="Figures 7, 8. Tabwecala robinsoni gen. nov., sp. nov., different branching patterns of veins from forewing areole (shaded light red) with interpretation of homologies (see criteria in Material and methods). 7. Long-areoled configuration; 8. Short-areoled configuration; M - medial; R - radial; Rs - radial sector; Sc - subcostal. Scale bar: 1 mm (7, 8.)." figureDoi="10.3897/nl.44.70359.figures7-8" httpUri="https://binary.pensoft.net/fig/597054" pageId="0" pageNumber="193">7</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 7, 8" captionStartId="F4" captionText="Figures 7, 8. Tabwecala robinsoni gen. nov., sp. nov., different branching patterns of veins from forewing areole (shaded light red) with interpretation of homologies (see criteria in Material and methods). 7. Long-areoled configuration; 8. Short-areoled configuration; M - medial; R - radial; Rs - radial sector; Sc - subcostal. Scale bar: 1 mm (7, 8.)." figureDoi="10.3897/nl.44.70359.figures7-8" httpUri="https://binary.pensoft.net/fig/597054" pageId="0" pageNumber="193">8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
(1) [
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Areole long; Rs1 from just beyond middle of areole; Rs2 and Rs3 long stalked, from upper distal corner of areole; Rs4 from lower distal corner of cell</emphasis>
|
||
] Rs1 and stalk (Rs2+Rs3) long stalked, from before end of cell, areole very long, bound superiorly by stalk (Rs1+(Rs2+Rs3)) and stalk (Rs2+Rs3), inferiorly by Rs (viz. cell) and Rs4, and closed by short anastomosis that from Rs4 reaches stalk (Rs2+Rs3) (Fig.
|
||
<figureCitation captionStart="Figures 7, 8" captionStartId="F4" captionText="Figures 7, 8. Tabwecala robinsoni gen. nov., sp. nov., different branching patterns of veins from forewing areole (shaded light red) with interpretation of homologies (see criteria in Material and methods). 7. Long-areoled configuration; 8. Short-areoled configuration; M - medial; R - radial; Rs - radial sector; Sc - subcostal. Scale bar: 1 mm (7, 8.)." figureDoi="10.3897/nl.44.70359.figures7-8" httpUri="https://binary.pensoft.net/fig/597054" pageId="0" pageNumber="193">7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
(2) [
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Areole short; Rs1 from 4/5 length of areole; stalk of Rs2 and Rs3 shortly stalked with Rs4, from end of areole</emphasis>
|
||
] Rs1 and Rs2' stalked, from before end of cell, Rs2' free from bifurcation with Rs1 for short tract, then anastomosed and cojoined into short stalk ((Rs2
|
||
<normalizedToken originalValue="”+Rs3)+Rs4”">"+Rs3)+Rs4"</normalizedToken>
|
||
) which then separates into long stalk (Rs2"+Rs3) and Rs4", areole thus short, bound superiorly by stalk (Rs1+Rs2') and Rs2', and inferiorly by Rs (viz. cell) and Rs4' (Fig.
|
||
<figureCitation captionStart="Figures 7, 8" captionStartId="F4" captionText="Figures 7, 8. Tabwecala robinsoni gen. nov., sp. nov., different branching patterns of veins from forewing areole (shaded light red) with interpretation of homologies (see criteria in Material and methods). 7. Long-areoled configuration; 8. Short-areoled configuration; M - medial; R - radial; Rs - radial sector; Sc - subcostal. Scale bar: 1 mm (7, 8.)." figureDoi="10.3897/nl.44.70359.figures7-8" httpUri="https://binary.pensoft.net/fig/597054" pageId="0" pageNumber="193">8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/nl.44.70359.figures7-8" httpUri="https://binary.pensoft.net/fig/597054" pageId="0" pageNumber="193" start="Figures 7, 8" startId="F4">
|
||
<paragraph pageId="0" pageNumber="193">
|
||
<emphasis bold="true" pageId="0" pageNumber="193">Figures 7, 8.</emphasis>
|
||
<taxonomicName authorityName="Zilli" authorityYear="2021" class="Insecta" family="Erebidae" genus="Tabwecala" kingdom="Animalia" lsidName="Tabwecala robinsoni" order="Lepidoptera" pageId="0" pageNumber="193" rank="species" species="robinsoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="193">Tabwecala robinsoni</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov., different branching patterns of veins from forewing areole (shaded light red) with interpretation of homologies (see criteria in Material and methods).
|
||
<emphasis bold="true" pageId="0" pageNumber="193">7.</emphasis>
|
||
Long-areoled configuration;
|
||
<emphasis bold="true" pageId="0" pageNumber="193">8.</emphasis>
|
||
Short-areoled configuration;
|
||
<emphasis bold="true" pageId="0" pageNumber="193">M</emphasis>
|
||
- medial;
|
||
<emphasis bold="true" pageId="0" pageNumber="193">R</emphasis>
|
||
- radial;
|
||
<emphasis bold="true" pageId="0" pageNumber="193">Rs</emphasis>
|
||
- radial sector;
|
||
<emphasis bold="true" pageId="0" pageNumber="193">Sc</emphasis>
|
||
- subcostal. Scale bar: 1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="193">7, 8.</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="193" type="distribution">
|
||
<paragraph pageId="0" pageNumber="193">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">The genus is currently known only from a restricted area of the Pacific (Vanuatu: Espiritu Santo).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="193" type="systematic position">
|
||
<paragraph pageId="0" pageNumber="193">Systematic position.</paragraph>
|
||
<paragraph pageId="0" pageNumber="193">
|
||
Features of the new genus indicate its position in tribe
|
||
<taxonomicName authorityName="Guenee" authorityYear="1852" lsidName="" pageId="0" pageNumber="193" rank="tribe" tribe="Poaphilini">Poaphilini</taxonomicName>
|
||
of the
|
||
<taxonomicName authorityName="Leach" authorityYear="1815" lsidName="" pageId="0" pageNumber="193" rank="subFamily" subFamily="Erebinae">Erebinae</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Leach" authorityYear="1815" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="family">Erebidae</taxonomicName>
|
||
), a group which is however morphologically ill-defined, as it will be reviewed in the discussion below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |