318 lines
56 KiB
XML
318 lines
56 KiB
XML
<document id="2937393A0BF43581A90F5C19D7AE077B" ID-CLB-Dataset="20173" ID-DOI="10.11646/zootaxa.5116.2.3" ID-GBIF-Dataset="e4437a20-2079-4555-8e17-fa6847d621ba" ID-ISSN="1175-5326" ID-Zenodo-Dep="6367255" ID-ZooBank="EBF12EE4-4675-45AC-80C1-CFA61FD0C297" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647595347751" checkinUser="plazi" docAuthor="Hara, Hideho & Ibuki, Shinichi" docDate="2022" docId="65401F59C017FF95FF6A2EF6FB09FE10" docLanguage="en" docName="zootaxa.5116.2.3.pdf" docOrigin="Zootaxa 5116 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fagineura flavomaculata Hara 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="240" masterDocId="99796721C007FF84FFFD2E61FFD9FF99" masterDocTitle="A study of the genus Fagineura (Hymenoptera, Tenthredinidae, Nematinae)" masterLastPageNumber="252" masterPageNumber="223" pageNumber="239" updateTime="1699318279456" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="06CC80CCEE048B022DB4DADB1D5FEE82">
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<mods:title id="9B55531D2F355D34A9CCED541D754BAF">A study of the genus Fagineura (Hymenoptera, Tenthredinidae, Nematinae)</mods:title>
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<mods:name id="F9D4F08E8F977DE0563AE7779ADF1618" type="personal">
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<mods:namePart id="AA1F2238442A14ED88BD7B584B3D1A77">Hara, Hideho</mods:namePart>
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<mods:affiliation id="2919C0CA89F8F28439D2828E0BB00DC4">Nishi 4 Kita 3 4 - 29, Bibai, Hokkaido, 072 - 0033 Japan. harahideho @ bell. ocn. ne. jp; https: // orcid. org / 0000 - 0001 - 5834 - 9158 Wami 1355 - 13, Nakagawa, Tochigi, 324 - 0612 Japan. banbi-fa @ ktd. biglobe. ne. jp</mods:affiliation>
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<mods:nameIdentifier id="E469074BB3DBD82AE6C5109D174F5A23" type="email">harahideho@bell.ocn.ne.jp</mods:nameIdentifier>
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<mods:name id="B30DA2EB77DDD179BBCB7DAD741FB07A" type="personal">
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<mods:roleTerm id="F75018608111A1B064841F9A61E13130">Author</mods:roleTerm>
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<mods:namePart id="D10C34F28B09889724CF0F3FC3BABD6E">Ibuki, Shinichi</mods:namePart>
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<mods:title id="BE97271FBF9C832FE3D333A150A0D77C">Zootaxa</mods:title>
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<mods:part id="3434524B1F384E6304C8B14CC15C6D59">
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<mods:date id="709BD47DB5F6934EDF2F8A6224EC4D57">2022</mods:date>
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<mods:number id="35146EC2E23303D9BD55B2247AB4A2B8">2022-03-18</mods:number>
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<mods:number id="B8E557FBE048EB3251CB0E272311DDEE">5116</mods:number>
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<treatment id="65401F59C017FF95FF6A2EF6FB09FE10" ID-DOI="http://doi.org/10.5281/zenodo.6367279" ID-GBIF-Taxon="194098928" ID-Zenodo-Dep="6367279" LSID="urn:lsid:plazi:treatment:65401F59C017FF95FF6A2EF6FB09FE10" httpUri="http://treatment.plazi.org/id/65401F59C017FF95FF6A2EF6FB09FE10" lastPageId="17" lastPageNumber="240" pageId="16" pageNumber="239">
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<subSubSection id="A5F3FDC4C017FF94FF6A2EF6FD58FF28" box="[151,641,151,178]" pageId="16" pageNumber="239" type="nomenclature">
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<paragraph id="ED56AE4FC017FF94FF6A2EF6FD58FF28" blockId="16.[151,832,151,214]" box="[151,641,151,178]" pageId="16" pageNumber="239">
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<heading id="B61E1923C017FF94FF6A2EF6FD58FF28" bold="true" box="[151,641,151,178]" fontSize="11" level="1" pageId="16" pageNumber="239" reason="1">
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<emphasis id="DF9D725DC017FF94FF6A2EF6FD58FF28" bold="true" box="[151,641,151,178]" pageId="16" pageNumber="239">
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<taxonomicName id="2AE9D5CCC017FF94FF6A2EF6FDCFFF28" authority="Hara" authorityName="Hara" authorityYear="2022" box="[151,534,151,178]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="239" phylum="Arthropoda" rank="species" species="flavomaculata" status="sp. nov.">
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<emphasis id="DF9D725DC017FF94FF6A2EF6FE15FF2B" bold="true" box="[151,460,151,178]" italics="true" pageId="16" pageNumber="239">Fagineura flavomaculata</emphasis>
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Hara
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</taxonomicName>
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,
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<taxonomicNameLabel id="C4AECF26C017FF94FDDE2EF6FD58FF28" box="[547,641,151,177]" pageId="16" pageNumber="239" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A5F3FDC4C017FF94FF6A2EDAFCF0F978" pageId="16" pageNumber="239" type="description">
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<paragraph id="ED56AE4FC017FF94FF6A2EDAFC99FF4F" blockId="16.[151,832,151,214]" box="[151,832,187,214]" pageId="16" pageNumber="239">
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(
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<figureCitation id="75D2B2CAC017FF94FF622EDAFE87FF4C" box="[159,350,187,213]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (A–E, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (F–H, N, female; O–Q, late instar, final instar and mature final instar larva; holotype) and F. quercivora (I–K, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. L–Q, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. A–D, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="16" pageNumber="239">Figs 4F–H, N–Q</figureCitation>
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,
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<figureCitation id="75D2B2CAC017FF94FE972EDAFE5AFF4C" box="[362,387,187,213]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). A–L, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; P–S, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="16" pageNumber="239">6I</figureCitation>
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,
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<figureCitation id="75D2B2CAC017FF94FE722EDAFE02FF4F" box="[399,475,187,214]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); D–I, left mandible, outer view, left anterior; J–O, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="16" pageNumber="239">7G, M</figureCitation>
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,
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<figureCitation id="75D2B2CAC017FF94FE1B2EDAFD91FF4C" box="[486,584,187,214]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).A–E, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; G–N, dorsal part of mesepisternum, anterolateral view; O–V, claw, anterior or posterior view; W–Y, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="16" pageNumber="239">8B, K, S</figureCitation>
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,
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<figureCitation id="75D2B2CAC017FF94FDA82EDAFD44FF4F" box="[597,669,187,214]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). A–H, Dorsal view; I–P, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="16" pageNumber="239">9E, M</figureCitation>
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,
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<figureCitation id="75D2B2CAC017FF94FD542EDAFD02FF4C" box="[681,731,187,213]" captionStart="FIGURE 10" captionStartId="23.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-374@23.[151,1436,957,1917]" captionTargetPageId="23" captionText="FIGURE 10. Ovipositor or lance, lateral view: Fagineura crenativora (A, B), F. parva (C, holotype), F. togashii (D, E), F. fulvistriata (F), F. flavomaculata (G, holotype), F. quercivora (H, I; H, holotype) and F. glabella (J, holotype)." figureDoi="http://doi.org/10.5281/zenodo.6367287" httpUri="https://zenodo.org/record/6367287/files/figure.png" pageId="16" pageNumber="239">10G</figureCitation>
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,
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<figureCitation id="75D2B2CAC017FF94FD1A2EDAFCEEFF4F" box="[743,823,187,214]" captionStart="FIGURE 12" captionStartId="25.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1862]" captionTargetId="figure-19@25.[151,1436,181,1870]" captionTargetPageId="25" captionText="FIGURE 12. Lancet: Fagineura togashii (A, B), F. fulvistriata (C, holotype) and F. flavomaculata (D, E; D, holotype). A, B, C, Reversed." figureDoi="http://doi.org/10.5281/zenodo.6367293" httpUri="https://zenodo.org/record/6367293/files/figure.png" pageId="16" pageNumber="239">12D, E</figureCitation>
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)
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</paragraph>
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<paragraph id="ED56AE4FC017FF94FF6A2F62FB71FCA0" blockId="16.[151,1437,259,2014]" pageId="16" pageNumber="239">
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<emphasis id="DF9D725DC017FF94FF6A2F62FE58FE84" bold="true" box="[151,385,259,285]" pageId="16" pageNumber="239">Description: female</emphasis>
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. Length 7.0–
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(
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<typeStatus id="325210EDC017FF94FD802F62FD38FE84" box="[637,737,259,285]" pageId="16" pageNumber="239" type="holotype">holotype</typeStatus>
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<quantity id="2A1103AAC017FF94FD152F62FC9AFE87" box="[744,835,259,286]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="16" pageNumber="239" unit="mm" value="8.5">8.5 mm</quantity>
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). Black (
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<figureCitation id="75D2B2CAC017FF94FC5A2F62FBFDFE84" box="[935,1060,259,285]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (A–E, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (F–H, N, female; O–Q, late instar, final instar and mature final instar larva; holotype) and F. quercivora (I–K, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. L–Q, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. A–D, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="16" pageNumber="239">Figs 4F–H</figureCitation>
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). Head capsule brown yellow except for medial areas of frons and vertex and occiput (
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<figureCitation id="75D2B2CAC017FF94FD0A2F46FC9FFEDB" box="[759,838,295,322]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). A–L, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; P–S, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="16" pageNumber="239">Fig. 6I</figureCitation>
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); postocellar area dark brown except for wide medial area and narrow lateral margin. Mandible pale yellow, with wide apex dark red brown. Palpi pale yellow. Thorax pale yellow on pronotum and tegula; mesoscutum yellow to yellow brown on anterolateral area of median lobe and usually on narrow medial area of lateral lobe; mesoscutellum usually with pair of yellow to brown spots; mesoscutellar appendage yellow except for narrow apex; metanotum yellow except for anterolateral area and medial area of metascutellum; postspiracular sclerite pale yellow grey (
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<figureCitation id="75D2B2CAC017FF94FCEB2FBAFCA8FE6C" box="[790,881,475,501]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).A–E, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; G–N, dorsal part of mesepisternum, anterolateral view; O–V, claw, anterior or posterior view; W–Y, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="16" pageNumber="239">Fig. 8K</figureCitation>
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); epicnemium dark brown, often posteriorly pale yellow; mesepisternum dorsally yellow to yellow brown and narrowly pale yellow grey beside postspiracular sclerite; katepimeron, metepimeron and dorsal area of metepisternum yellow. Legs with trochanters, trochantelli and their adjacent areas pale yellow (
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<figureCitation id="75D2B2CAC017FF94FDF12C26FDBEFDF8" box="[524,615,583,609]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (A–E, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (F–H, N, female; O–Q, late instar, final instar and mature final instar larva; holotype) and F. quercivora (I–K, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. L–Q, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. A–D, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="16" pageNumber="239">Fig. 4H</figureCitation>
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); fore and middle femora dorsally yellow; fore and middle tibiae basally and ventrally pale yellow; hind femur dorsally yellow on basal two-thirds; hind tibia pale yellow on wide basal area; fore and middle tarsi pale yellow, sometimes dorsally darkened. Wings colorless transparent (
|
||
<figureCitation id="75D2B2CAC017FF94FB5B2CEEFAD9FD30" box="[1190,1280,655,681]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (A–E, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (F–H, N, female; O–Q, late instar, final instar and mature final instar larva; holotype) and F. quercivora (I–K, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. L–Q, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. A–D, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="16" pageNumber="239">Fig. 4G</figureCitation>
|
||
); veins brown to black, with vein C and basal part of vein A yellow; stigma yellow brown, marginally darkened. Abdomen with tergum 1 yellow on lateral and posterior areas; wide lateral areas of terga 1–8 pale yellow; narrow posteromedial parts of terga 2 or 2–4 yellow; tergum 9 brown to black, posteroventrally narrowly yellow; sterna 4–6 or 5–6 centrally yellow brown; sternum 7 with pair of yellow brown spots; valvifer 2 often dark brown.
|
||
</paragraph>
|
||
<paragraph id="ED56AE4FC017FF94FF3A2D22FD85FBC0" blockId="16.[151,1437,259,2014]" pageId="16" pageNumber="239">
|
||
Head in dorsal view with length behind eye 0.4–0.5 × eye length (
|
||
<figureCitation id="75D2B2CAC017FF94FC592D22FC2AFCC7" box="[932,1011,835,862]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). A–L, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; P–S, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="16" pageNumber="239">Fig. 6I</figureCitation>
|
||
); length behind lateral ocellus 2.6–3.5 × length of lateral ocellus. OOL:POL:OOCL 0.7–0.8:1.0:1.1–1.3. Frontal area with lateral ridge distinct, sometimes very slight; frontal field with distinct or indistinct long lateral convexity. Distance between eyes at anterior tentorial pit 1.2–1.3 × major axis of eye (
|
||
<figureCitation id="75D2B2CAC017FF94FDD32DCEFD5CFC50" box="[558,645,943,969]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (A–E, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (F–H, N, female; O–Q, late instar, final instar and mature final instar larva; holotype) and F. quercivora (I–K, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. L–Q, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. A–D, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="16" pageNumber="239">Fig. 4F</figureCitation>
|
||
). Inner edges of eyes nearly parallel. Malar space 0.3–0.4 × as long as median ocellus width. Antenna 2.4 × as long as head width (
|
||
<figureCitation id="75D2B2CAC017FF94FCB12DB2FC71FC74" box="[844,936,979,1005]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (A–E, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (F–H, N, female; O–Q, late instar, final instar and mature final instar larva; holotype) and F. quercivora (I–K, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. L–Q, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. A–D, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="16" pageNumber="239">Fig. 4G</figureCitation>
|
||
); flagellum tapered; flagellomere 1 0.7 × as long as major axis of eye; flagellomere 2 1.1 × as long as flagellomere 1. Mesepisternum distinctly expanded beside postspiracular sclerite (
|
||
<figureCitation id="75D2B2CAC017FF94FE662A7AFE2CFBAC" box="[411,501,1051,1077]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).A–E, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; G–N, dorsal part of mesepisternum, anterolateral view; O–V, claw, anterior or posterior view; W–Y, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="16" pageNumber="239">Fig. 8K</figureCitation>
|
||
). Hind tibia with posterior spur 0.8–1.0 × as long as apical breadth of tibia in lateral view. Fore wing without crossvein 2r-rs.
|
||
</paragraph>
|
||
<paragraph id="ED56AE4FC017FF94FF3A2A02FD46FACC" blockId="16.[151,1437,259,2014]" pageId="16" pageNumber="239">
|
||
Valvula
|
||
<quantity id="2A1103AAC017FF94FED92A02FE96FBE4" box="[292,335,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="16" pageNumber="239" unit="in" value="3.0">3 in</quantity>
|
||
dorsal view about twice as wide as cercus, slightly or hardly concave near pointed apex (
|
||
<figureCitation id="75D2B2CAC017FF94FAC52A02FA56FBE4" box="[1336,1423,1123,1149]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). A–H, Dorsal view; I–P, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="16" pageNumber="239">Fig. 9E</figureCitation>
|
||
), in lateral view rounded or nearly angulated apically, with dorsal edge straight and ventral edge rounded (
|
||
<figureCitation id="75D2B2CAC017FF94FACD2AE6FA49FB38" box="[1328,1424,1159,1185]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). A–H, Dorsal view; I–P, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="16" pageNumber="239">Fig. 9M</figureCitation>
|
||
). Lance with dorsal edge slightly rounded (
|
||
<figureCitation id="75D2B2CAC017FF94FD9B2ACAFD17FB5C" box="[614,718,1195,1221]" captionStart="FIGURE 10" captionStartId="23.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-374@23.[151,1436,957,1917]" captionTargetPageId="23" captionText="FIGURE 10. Ovipositor or lance, lateral view: Fagineura crenativora (A, B), F. parva (C, holotype), F. togashii (D, E), F. fulvistriata (F), F. flavomaculata (G, holotype), F. quercivora (H, I; H, holotype) and F. glabella (J, holotype)." figureDoi="http://doi.org/10.5281/zenodo.6367287" httpUri="https://zenodo.org/record/6367287/files/figure.png" pageId="16" pageNumber="239">Fig. 10G</figureCitation>
|
||
). Lancet with radix 0.5–0.6 × as long as lamnium (
|
||
<figureCitation id="75D2B2CAC017FF94FAF92ACAFA57FB5F" box="[1284,1422,1195,1222]" captionStart="FIGURE 12" captionStartId="25.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1862]" captionTargetId="figure-19@25.[151,1436,181,1870]" captionTargetPageId="25" captionText="FIGURE 12. Lancet: Fagineura togashii (A, B), F. fulvistriata (C, holotype) and F. flavomaculata (D, E; D, holotype). A, B, C, Reversed." figureDoi="http://doi.org/10.5281/zenodo.6367293" httpUri="https://zenodo.org/record/6367293/files/figure.png" pageId="16" pageNumber="239">Figs 12D, E</figureCitation>
|
||
); lamnium with 19–20 annuli; basal annuli slightly sinuous; middle annuli straight and erect; apical annuli straight and slightly oblique; annulus 1 and some apical annuli without ctenidium; ctenidia widely separated from each other, each consisting of some irregular transverse rows of setae and slightly expanded dorsally; basal serrulae with anterior slope about as long as posterior slope.
|
||
</paragraph>
|
||
<paragraph id="ED56AE4FC017FF94FF3A2B3EFAAAF990" blockId="16.[151,1437,259,2014]" pageId="16" pageNumber="239">
|
||
Head and thorax with punctures minute; interspaces between punctures generally smooth. Mesoscutum mostly covered with setiferous punctures (
|
||
<figureCitation id="75D2B2CAC017FF94FDDD2BE2FDAEFA04" box="[544,631,1411,1437]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).A–E, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; G–N, dorsal part of mesepisternum, anterolateral view; O–V, claw, anterior or posterior view; W–Y, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="16" pageNumber="239">Fig. 8B</figureCitation>
|
||
); posterolateral hollow with rugose and granular microsculpture. Mesopostnotum with rugose and granular microsculpture, medially smooth. Metapostnotum mostly smooth. Postspiracular sclerite with many setae. Mesepisternum widely or narrowly glabrous beside postspiracular sclerite (
|
||
<figureCitation id="75D2B2CAC017FF94FAC92BAAFA56FA7C" box="[1332,1423,1483,1509]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).A–E, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; G–N, dorsal part of mesepisternum, anterolateral view; O–V, claw, anterior or posterior view; W–Y, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="16" pageNumber="239">Fig. 8K</figureCitation>
|
||
). Katepimeron widely covered with setae. Abdomen microsculptured imbricately, with punctures inconspicuous.
|
||
</paragraph>
|
||
<paragraph id="ED56AE4FC017FF94FF3A2872FE51F9B4" blockId="16.[151,1437,259,2014]" box="[199,392,1555,1582]" pageId="16" pageNumber="239">
|
||
<emphasis id="DF9D725DC017FF94FF3A2872FEDDF9B4" bold="true" box="[199,260,1555,1581]" pageId="16" pageNumber="239">Male</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph id="ED56AE4FC017FF94FF3A2856FCF0F978" blockId="16.[151,1437,259,2014]" pageId="16" pageNumber="239">
|
||
<emphasis id="DF9D725DC017FF94FF3A2856FE48F9C8" bold="true" box="[199,401,1591,1617]" pageId="16" pageNumber="239">Immature stages</emphasis>
|
||
. Late instar larva: head black (
|
||
<figureCitation id="75D2B2CAC017FF94FD082856FC88F9C8" box="[757,849,1591,1617]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (A–E, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (F–H, N, female; O–Q, late instar, final instar and mature final instar larva; holotype) and F. quercivora (I–K, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. L–Q, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. A–D, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="16" pageNumber="239">Fig. 4O</figureCitation>
|
||
); thoracic legs pale grey, apically and basally darkened; trunk pale grey, yellow on thoracic segment 1, with black supraspiracular and pleural stripes; supraspiracular stripe broken posteriorly; pleural stripe broken throughout; setae short. Final instar larva:
|
||
<quantity id="2A1103AAC017FF94FB70281EFB04F903" box="[1165,1245,1663,1690]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="239" unit="mm" value="20.0">20 mm</quantity>
|
||
long; color as in late instar, but thoracic legs mostly pale grey and abdominal segment 10 yellow; abdominal tergum 10 not darkened; setae inconspicuous; when mature, trunk yellow (
|
||
<figureCitation id="75D2B2CAC017FF94FD3F28A6FCC4F978" box="[706,797,1735,1761]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (A–E, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (F–H, N, female; O–Q, late instar, final instar and mature final instar larva; holotype) and F. quercivora (I–K, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. L–Q, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. A–D, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="16" pageNumber="239">Fig. 4Q</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5F3FDC4C017FF94FF3A288AFC8EF80C" pageId="16" pageNumber="239" type="materials_examined">
|
||
<paragraph id="ED56AE4FC017FF94FF3A288AFC8EF80C" blockId="16.[151,1437,259,2014]" pageId="16" pageNumber="239">
|
||
<emphasis id="DF9D725DC017FF94FF3A288AFE75F89C" bold="true" box="[199,428,1771,1797]" pageId="16" pageNumber="239">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="5D81A412C017FF94FE41288AFEA7F8D4" ID-GBIF-Occurrence="3710521375" collectingDate="2018-05-02" collectingDateMax="2019-04-17" collectingDateMin="2018-05-02" collectionCode="V" collectorName="S. Ibuki" country="Japan" latitude="36.766666" location="Koisago" longLatPrecision="1185" longitude="140.13333" municipality="Nakagawa" pageId="16" pageNumber="239" specimenCount="6" specimenCount-female="5" specimenCount-larva="1" stateProvince="Tochigi Pref." typeStatus="holotype">
|
||
<typeStatus id="325210EDC017FF94FE41288AFDF0F89C" box="[444,553,1771,1797]" pageId="16" pageNumber="239" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<specimenCount id="FBEF65C6C017FF94FDCB288DFD92F89F" box="[566,587,1772,1798]" pageId="16" pageNumber="239" type="female">♀</specimenCount>
|
||
, “
|
||
<collectingCountry id="95FEEEDFC017FF94FD98288AFD18F89C" box="[613,705,1771,1797]" name="Japan" pageId="16" pageNumber="239">JAPAN</collectingCountry>
|
||
: Honshu,
|
||
<collectingRegion id="2F2D60ADC017FF94FCC9288AFC15F89C" box="[820,972,1771,1797]" country="Japan" name="Tochigi" pageId="16" pageNumber="239">Tochigi Pref.</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="0D323435C017FF94FC24288AFB8BF89C" box="[985,1106,1771,1797]" pageId="16" pageNumber="239">Nakagawa</collectingMunicipality>
|
||
,
|
||
<location id="E836F894C017FF94FB9D288AFB19F89C" LSID="urn:lsid:plazi:treatment:65401F59C017FF95FF6A2EF6FB09FE10:E836F894C017FF94FB9D288AFB19F89C" box="[1120,1216,1771,1797]" country="Japan" latitude="36.766666" longLatPrecision="1185" longitude="140.13333" municipality="Nakagawa" name="Koisago" pageId="16" pageNumber="239" stateProvince="Tochigi Pref.">Koisago</location>
|
||
,
|
||
<geoCoordinate id="88DDC888C017FF94FB30288AFAF7F89C" box="[1229,1326,1771,1797]" degrees="36" direction="north" minutes="46" orientation="latitude" pageId="16" pageNumber="239" precision="925" value="36.766666">36°46’N</geoCoordinate>
|
||
<geoCoordinate id="88DDC888C017FF94FAC5288AFA4EF89C" box="[1336,1431,1771,1797]" degrees="140" direction="east" minutes="8" orientation="longitude" pageId="16" pageNumber="239" precision="925" value="140.13333">140°8’E</geoCoordinate>
|
||
, coll.
|
||
<specimenCount id="FBEF65C6C017FF94FF33296EFEDFF8B0" box="[206,262,1807,1833]" pageId="16" pageNumber="239" type="larva">larva</specimenCount>
|
||
on
|
||
<taxonomicName id="2AE9D5CCC017FF94FED02971FDD0F8B0" box="[301,521,1808,1834]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="239" phylum="Tracheophyta" rank="species" species="acutissima">
|
||
<emphasis id="DF9D725DC017FF94FED02971FDD0F8B0" box="[301,521,1808,1834]" italics="true" pageId="16" pageNumber="239">
|
||
<location id="E836F894C017FF94FED02971FE52F8B3" LSID="urn:lsid:plazi:treatment:65401F59C017FF95FF6A2EF6FB09FE10:E836F894C017FF94FED02971FE52F8B3" box="[301,395,1808,1834]" country="Japan" latitude="36.766666" longLatPrecision="1185" longitude="140.13333" municipality="Nakagawa" name="Quercus" pageId="16" pageNumber="239" stateProvince="Tochigi Pref.">Quercus</location>
|
||
acutissima
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<date id="9957888FC017FF94FDF3296EFD58F8B0" box="[526,641,1807,1834]" pageId="16" pageNumber="239" value="2018-05-02">
|
||
<collectingDate id="89137167C017FF94FDF3296EFD58F8B0" box="[526,641,1807,1834]" pageId="16" pageNumber="239" value="2018-05-02">2. V. 2018</collectingDate>
|
||
</date>
|
||
, mat. 9.
|
||
<collectionCode id="8BF8368AC017FF94FD212971FD2BF8B3" box="[732,754,1808,1834]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="239" type="Museum">V</collectionCode>
|
||
., em.
|
||
<date id="9957888FC017FF94FCD2296EFC60F8B0" box="[815,953,1807,1834]" pageId="16" pageNumber="239" value="2019-04-17">
|
||
<collectingDate id="89137167C017FF94FCD2296EFC60F8B0" box="[815,953,1807,1834]" pageId="16" pageNumber="239" value="2019-04-17">17. IV. 2019</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="401CCB99C017FF94FC3E296EFBC7F8B0" box="[963,1054,1807,1833]" pageId="16" pageNumber="239">S. Ibuki</collectorName>
|
||
” (
|
||
<figureCitation id="75D2B2CAC017FF94FBC8296EFB28F8B0" box="[1077,1265,1807,1833]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (A–E, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (F–H, N, female; O–Q, late instar, final instar and mature final instar larva; holotype) and F. quercivora (I–K, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. L–Q, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. A–D, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="16" pageNumber="239">
|
||
<location id="E836F894C017FF94FBC8296EFBBEF8B0" LSID="urn:lsid:plazi:treatment:65401F59C017FF95FF6A2EF6FB09FE10:E836F894C017FF94FBC8296EFBBEF8B0" box="[1077,1127,1807,1833]" country="Japan" latitude="36.766666" longLatPrecision="1185" longitude="140.13333" municipality="Nakagawa" name="Figs" pageId="16" pageNumber="239" stateProvince="Tochigi Pref.">Figs</location>
|
||
<specimenCount id="FBEF65C6C017FF94FB91296EFB53F8B0" box="[1132,1162,1807,1833]" pageId="16" pageNumber="239" type="female">4F</specimenCount>
|
||
–H, N–Q
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="75D2B2CAC017FF94FB01296EFACCF8B0" box="[1276,1301,1807,1833]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). A–L, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; P–S, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="16" pageNumber="239">6I</figureCitation>
|
||
,
|
||
<figureCitation id="75D2B2CAC017FF94FADD296EFAB3F8B3" box="[1312,1386,1807,1834]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); D–I, left mandible, outer view, left anterior; J–O, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="16" pageNumber="239">7G, M</figureCitation>
|
||
,
|
||
<figureCitation id="75D2B2CAC017FF94FA89296EFF13F8D4" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).A–E, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; G–N, dorsal part of mesepisternum, anterolateral view; O–V, claw, anterior or posterior view; W–Y, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="16" pageNumber="239">8B, K, S</figureCitation>
|
||
,
|
||
<figureCitation id="75D2B2CAC017FF94FF2B2952FF2EF8D4" box="[214,247,1843,1869]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). A–H, Dorsal view; I–P, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="16" pageNumber="239">9E</figureCitation>
|
||
,
|
||
<figureCitation id="75D2B2CAC017FF94FEFE2952FEECF8D4" box="[259,309,1843,1869]" captionStart="FIGURE 10" captionStartId="23.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-374@23.[151,1436,957,1917]" captionTargetPageId="23" captionText="FIGURE 10. Ovipositor or lance, lateral view: Fagineura crenativora (A, B), F. parva (C, holotype), F. togashii (D, E), F. fulvistriata (F), F. flavomaculata (G, holotype), F. quercivora (H, I; H, holotype) and F. glabella (J, holotype)." figureDoi="http://doi.org/10.5281/zenodo.6367287" httpUri="https://zenodo.org/record/6367287/files/figure.png" pageId="16" pageNumber="239">10G</figureCitation>
|
||
,
|
||
<figureCitation id="75D2B2CAC017FF94FEBD2952FEACF8D4" box="[320,373,1843,1869]" captionStart="FIGURE 12" captionStartId="25.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1862]" captionTargetId="figure-19@25.[151,1436,181,1870]" captionTargetPageId="25" captionText="FIGURE 12. Lancet: Fagineura togashii (A, B), F. fulvistriata (C, holotype) and F. flavomaculata (D, E; D, holotype). A, B, C, Reversed." figureDoi="http://doi.org/10.5281/zenodo.6367293" httpUri="https://zenodo.org/record/6367293/files/figure.png" pageId="16" pageNumber="239">12D</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="325210EDC017FF94FE752955FE20F8D7" box="[392,505,1844,1870]" pageId="16" pageNumber="239" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<collectingCountry id="95FEEEDFC017FF94FDFE2952FD85F8D4" box="[515,604,1843,1869]" name="Japan" pageId="16" pageNumber="239">JAPAN</collectingCountry>
|
||
:
|
||
<collectionCode id="8BF8368AC017FF94FD9B2952FD06F8D4" box="[614,735,1843,1869]" pageId="16" pageNumber="239">HONSHU</collectionCode>
|
||
:
|
||
<specimenCount id="FBEF65C6C017FF94FD142952FCD4F8D4" box="[745,781,1843,1869]" pageId="16" pageNumber="239" type="female">4♀</specimenCount>
|
||
,
|
||
<collectingRegion id="2F2D60ADC017FF94FCE42952FC1DF8D4" box="[793,964,1843,1869]" country="Japan" name="Okayama" pageId="16" pageNumber="239">Okayama Pref.</collectingRegion>
|
||
, Soja, Minagi, coll. larvae on
|
||
<taxonomicName id="2AE9D5CCC017FF94FAEC2955FA4EF8D7" box="[1297,1431,1844,1870]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="239" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="DF9D725DC017FF94FAEC2955FAB6F8D7" box="[1297,1391,1844,1870]" italics="true" pageId="16" pageNumber="239">Quercus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2AE9D5CCC017FF94FF6A2939FED2F8E8" box="[151,267,1880,1906]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="239" phylum="Tracheophyta" rank="species" species="serrata">
|
||
<emphasis id="DF9D725DC017FF94FF6A2939FED2F8E8" box="[151,267,1880,1906]" italics="true" pageId="16" pageNumber="239">Q. serrata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2AE9D5CCC017FF94FEE42939FE6DF8E8" box="[281,436,1880,1906]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="239" phylum="Tracheophyta" rank="species" species="acutissima">
|
||
<emphasis id="DF9D725DC017FF94FEE42939FE6DF8E8" box="[281,436,1880,1906]" italics="true" pageId="16" pageNumber="239">Q. acutissima</emphasis>
|
||
</taxonomicName>
|
||
<date id="9957888FC017FF94FE412936FDEBF8E8" box="[444,562,1879,1906]" pageId="16" pageNumber="239" value="1973-05-03">
|
||
3.
|
||
<collectionCode id="8BF8368AC017FF94FE252939FE35F8EB" box="[472,492,1880,1906]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="239" type="Museum">V</collectionCode>
|
||
. 1973
|
||
</date>
|
||
, em.
|
||
<date id="9957888FC017FF94FD932936FD37F8E8" box="[622,750,1879,1906]" pageId="16" pageNumber="239" value="1974-04-04">4. IV. 1974</date>
|
||
,
|
||
<collectionCode id="8BF8368AC017FF94FD072939FCD5F8EB" box="[762,780,1880,1906]" name="Tavera, Department of Geology and Geophysics" pageId="16" pageNumber="239">T</collectionCode>
|
||
. Kondo (
|
||
<figureCitation id="75D2B2CAC017FF94FC8E2936FC08F8E8" box="[883,977,1879,1905]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). A–H, Dorsal view; I–P, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="16" pageNumber="239">Fig. 9M</figureCitation>
|
||
) ―
|
||
<collectingCountry id="95FEEEDFC017FF94FBF92936FBB0F8E8" box="[1028,1129,1879,1905]" name="South Korea" pageId="16" pageNumber="239">KOREA</collectingCountry>
|
||
:
|
||
<specimenCount id="FBEF65C6C017FF94FB8E2936FB4FF8E8" box="[1139,1174,1879,1905]" pageId="16" pageNumber="239" type="female">1♀</specimenCount>
|
||
,
|
||
<collectingRegion id="2F2D60ADC017FF94FB582936FAE5F8E8" box="[1189,1340,1879,1905]" country="South Korea" name="Gyeonggi" pageId="16" pageNumber="239">Gyeonggi-do</collectingRegion>
|
||
, Suwon,
|
||
<date id="9957888FC017FF94FF6A291AFEFDF80C" box="[151,292,1915,1942]" pageId="16" pageNumber="239" value="1927-04-25">25. IV. 1927</date>
|
||
, K, Sato;
|
||
<specimenCount id="FBEF65C6C017FF94FE6F291AFE6CF80C" box="[402,437,1915,1941]" pageId="16" pageNumber="239" type="female">1♀</specimenCount>
|
||
, same data but
|
||
<date id="9957888FC017FF94FD99291AFD0FF80C" box="[612,726,1915,1942]" pageId="16" pageNumber="239" value="1931-05-05">
|
||
5.
|
||
<collectionCode id="8BF8368AC017FF94FD82291DFD4AF80F" box="[639,659,1916,1942]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="239" type="Museum">V</collectionCode>
|
||
. 1931
|
||
</date>
|
||
(
|
||
<figureCitation id="75D2B2CAC017FF94FD18291AFC92F80C" box="[741,843,1915,1941]" captionStart="FIGURE 12" captionStartId="25.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1862]" captionTargetId="figure-19@25.[151,1436,181,1870]" captionTargetPageId="25" captionText="FIGURE 12. Lancet: Fagineura togashii (A, B), F. fulvistriata (C, holotype) and F. flavomaculata (D, E; D, holotype). A, B, C, Reversed." figureDoi="http://doi.org/10.5281/zenodo.6367293" httpUri="https://zenodo.org/record/6367293/files/figure.png" pageId="16" pageNumber="239">Fig. 12E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5F3FDC4C017FF94FF3A29FEFEF8F844" pageId="16" pageNumber="239" type="etymology">
|
||
<paragraph id="ED56AE4FC017FF94FF3A29FEFEF8F844" blockId="16.[151,1437,259,2014]" pageId="16" pageNumber="239">
|
||
<emphasis id="DF9D725DC017FF94FF3A29FEFE91F820" bold="true" box="[199,328,1951,1977]" pageId="16" pageNumber="239">Etymology</emphasis>
|
||
. The specific name is an adjective, meaning yellow spotted, and comes from the yellow markings on the body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5F3FDC4C016FF95FF3A2EF7FD53FED8" pageId="17" pageNumber="240" type="distribution">
|
||
<paragraph id="ED56AE4FC016FF95FF3A2EF7FDABFF28" blockId="17.[151,1437,150,393]" box="[199,626,150,178]" pageId="17" pageNumber="240">
|
||
<emphasis id="DF9D725DC016FF95FF3A2EF7FE80FF29" bold="true" box="[199,345,150,176]" pageId="17" pageNumber="240">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="95FEEEDFC016FF95FE9A2EF6FE7EFF28" box="[359,423,151,177]" name="Japan" pageId="17" pageNumber="240">Japan</collectingCountry>
|
||
(Honshu),
|
||
<collectingCountry id="95FEEEDFC016FF95FDDB2EF6FDB7FF28" box="[550,622,151,177]" name="South Korea" pageId="17" pageNumber="240">Korea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="ED56AE4FC016FF95FF3A2EDBFB5EFF4C" blockId="17.[151,1437,150,393]" box="[199,1159,186,213]" pageId="17" pageNumber="240">
|
||
<emphasis id="DF9D725DC016FF95FF3A2EDBFE8BFF4D" bold="true" box="[199,338,186,212]" pageId="17" pageNumber="240">Life history</emphasis>
|
||
. Host plants:
|
||
<taxonomicName id="2AE9D5CCC016FF95FE132EDAFD82FF4C" authorityName="Dumortier" authorityYear="1829" box="[494,603,187,213]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="17" pageNumber="240" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
|
||
:
|
||
<taxonomicName id="2AE9D5CCC016FF95FD982EDAFC71FF4C" authority="Carruth." authorityName="Carruth." box="[613,936,187,213]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="17" pageNumber="240" phylum="Tracheophyta" rank="species" species="acutissima">
|
||
<emphasis id="DF9D725DC016FF95FD982EDAFC9BFF4D" box="[613,834,187,213]" italics="true" pageId="17" pageNumber="240">Quercus acutissima</emphasis>
|
||
Carruth.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2AE9D5CCC016FF95FC4E2EDAFB5EFF4C" authority="Murray." box="[947,1159,187,213]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="17" pageNumber="240" phylum="Tracheophyta" rank="species" species="serrata">
|
||
<emphasis id="DF9D725DC016FF95FC4E2EDAFBFFFF4D" box="[947,1062,187,213]" italics="true" pageId="17" pageNumber="240">Q. serrata</emphasis>
|
||
Murray.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="ED56AE4FC016FF95FF3A2EBEFD53FED8" blockId="17.[151,1437,150,393]" pageId="17" pageNumber="240">
|
||
This species has one generation a year. S. Ibuki collected a solitary larva on early May. It matured a week later and made a cocoon in the soil under rearing condition. A female emerged on middle April in the next year and oviposited in a young shoot in a cage (
|
||
<figureCitation id="75D2B2CAC016FF95FDDE2F46FDA4FED8" box="[547,637,295,321]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (A–E, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (F–H, N, female; O–Q, late instar, final instar and mature final instar larva; holotype) and F. quercivora (I–K, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. L–Q, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. A–D, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="17" pageNumber="240">Fig. 4N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5F3FDC4C016FF95FF3A2F2AFB09FE10" pageId="17" pageNumber="240" type="discussion">
|
||
<paragraph id="ED56AE4FC016FF95FF3A2F2AFB09FE10" blockId="17.[151,1437,150,393]" pageId="17" pageNumber="240">
|
||
<emphasis id="DF9D725DC016FF95FF3A2F2AFEEDFEFC" bold="true" box="[199,308,331,357]" pageId="17" pageNumber="240">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="2AE9D5CCC016FF95FEB82F2AFDBCFEFC" authorityName="Hara" authorityYear="2022" box="[325,613,331,357]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="240" phylum="Arthropoda" rank="species" species="flavomaculata">
|
||
<emphasis id="DF9D725DC016FF95FEB82F2AFDBCFEFC" box="[325,613,331,357]" italics="true" pageId="17" pageNumber="240">Fagineura flavomaculata</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="2AE9D5CCC016FF95FCFF2F2AFC42FEFC" authorityName="Hara" authorityYear="2022" box="[770,923,331,357]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="240" phylum="Arthropoda" rank="species" species="fulvistriata">
|
||
<emphasis id="DF9D725DC016FF95FCFF2F2AFC42FEFC" box="[770,923,331,357]" italics="true" pageId="17" pageNumber="240">F. fulvistriata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2AE9D5CCC016FF95FC2A2F2AFB96FEFC" authorityName="Hara" authorityYear="2022" box="[983,1103,331,357]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="240" phylum="Arthropoda" rank="species" species="togashii">
|
||
<emphasis id="DF9D725DC016FF95FC2A2F2AFB96FEFC" box="[983,1103,331,357]" italics="true" pageId="17" pageNumber="240">F. togashii</emphasis>
|
||
</taxonomicName>
|
||
in color, but they are distinguished as stated in the key. For more their differences, see the remarks of the latter two species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |