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<mods:title id="3CF5C0FBFD31FC890BF9CACD6950DB13">A study of the genus Fagineura (Hymenoptera, Tenthredinidae, Nematinae)</mods:title>
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<mods:namePart id="07B64AB19AFCF7B2E3712F0269F4A6E7">Hara, Hideho</mods:namePart>
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<taxonomicName id="2AE9D5CCC006FF85FF6A2EBEFDBBFF60" authority="Vikberg &amp; Zinovjev, 2000" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[151,610,223,249]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="genus">
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<emphasis id="DF9D725DC006FF85FF6A2EBEFEC1FF60" bold="true" box="[151,280,223,249]" italics="true" pageId="1" pageNumber="224">Fagineura</emphasis>
Vikberg &amp; Zinovjev, 2000
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<taxonomicName id="2AE9D5CCC006FF85FF6A2F45FDCCFEA5" authority="Vikberg &amp; Zinovjev, 2000" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[151,533,292,316]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC006FF85FF6A2F45FEDBFEA2" box="[151,258,292,315]" italics="true" pageId="1" pageNumber="224">Fagineura</emphasis>
Vikberg &amp; Zinovjev, 2000
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(in
<bibRefCitation id="8978D3BEC006FF85FDC22F45FCC6FEA5" author="Shinohara, A. &amp; Vikberg, V. &amp; Zinovjev, A. &amp; Yamagami, A." box="[575,799,292,316]" pageId="1" pageNumber="224" pagination="113 - 124" refId="ref22313" refString="Shinohara, A., Vikberg, V., Zinovjev, A. &amp; Yamagami, A. (2000) Fagineura crenativora, a new genus and species of sawfly (Hymenoptera, Tenthredinidae, Nematinae) injurious to beech trees in Japan. Bulletin of the National Science Museum, Series A, 26 (3), 113 - 124." type="journal article" year="2000">
Shinohara
<emphasis id="DF9D725DC006FF85FD502F45FD38FEA5" box="[685,737,292,316]" italics="true" pageId="1" pageNumber="224">et al.</emphasis>
2000
</bibRefCitation>
): 114;
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<bibRefCitation id="8978D3BEC006FF85FC9E2F45FBFAFEA2" author="Togashi, I." box="[867,1059,291,316]" pageId="1" pageNumber="224" pagination="169 - 173" refId="ref22763" refString="Togashi, I. (2006) A new sawfly, Fagineura quercivora (Hymenoptera: Tenthredinidae) feeding on Quercus serrata and Q. mongolica crispula in Honshu, Japan. Proceedings of the entomological Society of Washington, 108 (1), 169 - 173." type="journal article" year="2006">Togashi 2006: 169</bibRefCitation>
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;
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<bibRefCitation id="8978D3BEC006FF85FBD32F45FAC4FEA2" author="Taeger, A. &amp; Blank, S. M. &amp; Liston, A. D." box="[1070,1309,291,316]" pageId="1" pageNumber="224" pagination="1 - 1064" refId="ref22428" refString="Taeger, A., Blank, S. M. &amp; Liston, A. D. (2010) World catalog of Symphyta (Hymenoptera). Zootaxa, 2580 (1), 1 - 1064. https: // doi. org / 10.11646 / zootaxa. 2580.1" type="journal article" year="2010">
Taeger
<emphasis id="DF9D725DC006FF85FB842F45FB74FEA5" box="[1145,1197,292,316]" italics="true" pageId="1" pageNumber="224">et al.</emphasis>
2010: 405
</bibRefCitation>
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;
<treatmentCitation id="6C48885EC006FF85FAD52F45FEFBFEC3" author="Prous, M. &amp; Blank, S. M. &amp; Goulet, H. &amp; Heibo, E. &amp; Liston, A. &amp; Malm, T. &amp; Nyman, T. &amp; Schmidt, S. &amp; Smith, D. R. &amp; Vardal, H. &amp; Viitasaari, M. &amp; Vikberg, V. &amp; Taeger, A." page="18" pageId="1" pageNumber="224" year="2014">
<bibRefCitation id="8978D3BEC006FF85FAD52F45FEFBFEC3" author="Prous, M. &amp; Blank, S. M. &amp; Goulet, H. &amp; Heibo, E. &amp; Liston, A. &amp; Malm, T. &amp; Nyman, T. &amp; Schmidt, S. &amp; Smith, D. R. &amp; Vardal, H. &amp; Viitasaari, M. &amp; Vikberg, V. &amp; Taeger, A." pageId="1" pageNumber="224" pagination="1 - 69" refId="ref21989" refString="Prous, M., Blank, S. M., Goulet, H., Heibo, E., Liston, A., Malm, T., Nyman, T., Schmidt, S., Smith, D. R., Vardal, H., Viitasaari, M., Vikberg, V. &amp; Taeger, A. (2014) The genera of Nematinae (Hymenoptera, Tenthredinidae). Journal of Hymenoptera Research, 40, 1 - 69. https: // doi. org / 10.3897 / JHR. 40.7442" type="journal article" year="2014">
Prous
<emphasis id="DF9D725DC006FF85FA942F45FA44FEA5" box="[1385,1437,292,316]" italics="true" pageId="1" pageNumber="224">et al.</emphasis>
2014: 18
</bibRefCitation>
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, 26, 47;
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<bibRefCitation id="8978D3BEC006FF85FE872F23FDE1FEC3" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[378,568,322,346]" pageId="1" pageNumber="224" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC006FF85FE5E2F22FE0EFEC3" box="[419,471,322,346]" italics="true" pageId="1" pageNumber="224">et al.</emphasis>
2019: 32
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;
<treatmentCitation id="6C48885EC006FF85FDBE2F22FD0EFEC3" author="Hara, H." box="[579,727,322,346]" page="77" pageId="1" pageNumber="224" year="2019">
<bibRefCitation id="8978D3BEC006FF85FDBE2F22FD0EFEC3" author="Hara, H." box="[579,727,322,346]" pageId="1" pageNumber="224" pagination="69 - 87" refId="ref21558" refString="Hara, H. (2019) Subfamily Nematinae. In: Editorial Committee of Catalogue of the Insects of Japan (Ed.), Catalogue of the Insects of Japan. Vol. 9. Hymenoptera. Part 1. Symphyta. Entomological Society of Japan, Kyoto, pp. 69 - 87. [in Japanese]" type="book chapter" year="2019">Hara 2019: 77</bibRefCitation>
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;
<treatmentCitation id="6C48885EC006FF85FD1C2F22FC5BFEC3" author="Hara, H." box="[737,898,322,346]" page="338" pageId="1" pageNumber="224" year="2020">
<bibRefCitation id="8978D3BEC006FF85FD1C2F22FC5BFEC3" author="Hara, H." box="[737,898,322,346]" pageId="1" pageNumber="224" pagination="323 - 350" refId="ref21619" refString="Hara, H. (2020) Subfamily Nematinae. In: Naito, T., Shinohara, A., Hara, H. &amp; Ito, F., Sawflies and Woodwasps of Japan. Hokkaido University Press, Sapporo, pp. 75 - 89 + 323 - 350. [in Japanese]" type="book chapter" year="2020">Hara 2020: 338</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6C48885EC006FF85FC712F23FB81FEC3" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[908,1112,322,346]" httpUri="http://treatment.plazi.org/id/C1CB47A79E3559FD884FE85C1821CE11" page="126" pageId="1" pageNumber="224" year="2021">
<bibRefCitation id="8978D3BEC006FF85FC712F23FB81FEC3" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[908,1112,322,346]" pageId="1" pageNumber="224" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">
Liu
<emphasis id="DF9D725DC006FF85FC4B2F22FC30FEC3" box="[950,1001,322,346]" italics="true" pageId="1" pageNumber="224">et al.</emphasis>
2021: 126
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.
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</paragraph>
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<paragraph id="ED56AE4FC006FF85FF3A2F00FB92FEE0" blockId="1.[151,1437,291,377]" box="[199,1099,353,377]" pageId="1" pageNumber="224">
<typeStatus id="325210EDC006FF85FF3A2F00FF20FEE0" box="[199,249,353,377]" pageId="1" pageNumber="224">Type</typeStatus>
species:
<taxonomicName id="2AE9D5CCC006FF85FEAB2F03FC89FEE0" authority="Vikberg &amp; Zinovjev, 2000" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[342,848,353,377]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC006FF85FEAB2F03FDE4FEE0" box="[342,573,354,377]" italics="true" pageId="1" pageNumber="224">Fagineura crenativora</emphasis>
Vikberg &amp; Zinovjev, 2000
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, by original designation.
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<subSubSection id="A5F3FDC4C006FF85FF6A2FC6FB4BFDEC" pageId="1" pageNumber="224" type="diagnosis">
<paragraph id="ED56AE4FC006FF85FF6A2FC6FB4BFDEC" blockId="1.[151,1437,423,2034]" pageId="1" pageNumber="224">
<emphasis id="DF9D725DC006FF85FF6A2FC6FE44FE58" bold="true" box="[151,413,423,449]" pageId="1" pageNumber="224">Diagnostic characters</emphasis>
. Outer margin of eye without distinct furrow (
<figureCitation id="75D2B2CAC006FF85FC452FC6FBE5FE5B" box="[952,1084,423,450]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="1" pageNumber="224">Figs 6AN</figureCitation>
). Inter-antennal area protruded anteriorly beyond anterior ridge of frontal area (
<figureCitation id="75D2B2CAC006FF85FD4F2FAAFCEDFE7F" box="[690,820,459,486]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="1" pageNumber="224">Figs 6AN</figureCitation>
). Torulus completely surrounded by narrow ridge (
<figureCitation id="75D2B2CAC006FF85FA922FAAFF64FD93" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="1" pageNumber="224">Fig. 6O</figureCitation>
). Paraantennal field covered with setae except for narrow medial part (
<figureCitation id="75D2B2CAC006FF85FC202F8EFB8FFD93" box="[989,1110,495,522]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="1" pageNumber="224">Figs 6PS</figureCitation>
). Malar space length 0.20.8 × median ocellus width. Labrum apically rounded (
<figureCitation id="75D2B2CAC006FF85FD162C72FCB7FDB4" box="[747,878,531,558]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="1" pageNumber="224">Figs 7B, C</figureCitation>
). Left and right mandibles equal in length (
<figureCitation id="75D2B2CAC006FF85FA972C72FF07FDC8" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="1" pageNumber="224">Figs 7AC</figureCitation>
); basal outer surface not widely flattened or concave (
<figureCitation id="75D2B2CAC006FF85FCCA2C56FC61FDC8" box="[823,952,567,593]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="1" pageNumber="224">Figs 7DO</figureCitation>
). Maxillary palpus not shortened (
<figureCitation id="75D2B2CAC006FF85FAC82C56FA57FDC8" box="[1333,1422,567,593]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="1" pageNumber="224">Fig. 7A</figureCitation>
); palpomere 2 length 0.7 × palpomere 3 length; palpomere 6 length 1.11.2 × torulus height.
</paragraph>
</subSubSection>
<subSubSection id="A5F3FDC4C006FF87FF3A2C1EFEB8FE48" lastPageId="3" lastPageNumber="226" pageId="1" pageNumber="224" type="description">
<paragraph id="ED56AE4FC006FF85FF3A2C1EFCFFFB9B" blockId="1.[151,1437,423,2034]" pageId="1" pageNumber="224">
Notaulus distinct (
<figureCitation id="75D2B2CAC006FF85FE6B2C1EFDCCFD00" box="[406,533,639,665]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="1" pageNumber="224">Figs 8AE</figureCitation>
). Mesoscutellar appendage 0.91.5 × as long as minor axis of cenchrus. Median membranous part of mesopostnotum not or slightly widened anteriorly. Mesepisternum smooth between punctures, with distinct groove along anterior edge (
<figureCitation id="75D2B2CAC006FF85FD932CA6FD1CFD78" box="[622,709,711,737]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="1" pageNumber="224">Fig. 8F</figureCitation>
); this groove dorsally extending into epicnemium. Anterior fore tibial spur with distinct velum; hind leg normal; hind tarsomere 1 not grooved on anterior and posterior surfaces; tarsal claws without basal lobe, with inner tooth long and slightly curved (
<figureCitation id="75D2B2CAC006FF85FC112D6EFBB6FCB0" box="[1004,1135,783,809]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="1" pageNumber="224">Figs 8OV</figureCitation>
); inner tooth as long as or somewhat shorter than apical tooth; depth of concavity between apical and inner teeth longer than distance between their apices. In fore wing, posterior end of transverse section of vein Sc situated before or at joint of vein M and vein R (
<figureCitation id="75D2B2CAC006FF85FF0C2D1AFEA0FC0C" box="[241,377,891,917]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="1" pageNumber="224">Figs 8WY</figureCitation>
); vein R+M longer than first section of vein Rs (
<figureCitation id="75D2B2CAC006FF85FC612D1AFC2FFC0C" box="[924,1014,891,917]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="1" pageNumber="224">Fig. 8X</figureCitation>
); joint of crossvein 2r-m with vein M located apical to joint of crossvein 2m-cu with vein M (
<figureCitation id="75D2B2CAC006FF85FD022DFEFC83FC20" box="[767,858,927,953]" captionStart="FIGURE 1" captionStartId="7.[151,250,1832,1856]" captionTargetBox="[158,1435,276,1794]" captionTargetId="figure-57@7.[151,1435,270,1806]" captionTargetPageId="7" captionText="FIGURE 1. Fagineura crenativora (AK, female; LN, male). A, E, I, L, Head, frontal view; B, F, J, M, dorsal view; C, G, K, N, lateral or ventrolateral view; D, H, ventral view. AH, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367257" httpUri="https://zenodo.org/record/6367257/files/figure.png" pageId="1" pageNumber="224">Figs 1B</figureCitation>
, etc.); basal section of vein 2A+3A straight and apically separated from vein 1A. In hind wing, vein 2A complete and cell 1A closed; section of vein 1A between cell 1A and crossvein cu-a 0.71.7 × as long as crossvein cu-a.
</paragraph>
<paragraph id="ED56AE4FC006FF85FF3A2A6DFBFEFBF4" blockId="1.[151,1437,423,2034]" pageId="1" pageNumber="224">
In female abdomen, tergum 9 normal size (
<figureCitation id="75D2B2CAC006FF85FD5B2A6AFCDBFBBC" box="[678,770,1035,1061]" captionStart="FIGURE 1" captionStartId="7.[151,250,1832,1856]" captionTargetBox="[158,1435,276,1794]" captionTargetId="figure-57@7.[151,1435,270,1806]" captionTargetPageId="7" captionText="FIGURE 1. Fagineura crenativora (AK, female; LN, male). A, E, I, L, Head, frontal view; B, F, J, M, dorsal view; C, G, K, N, lateral or ventrolateral view; D, H, ventral view. AH, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367257" httpUri="https://zenodo.org/record/6367257/files/figure.png" pageId="1" pageNumber="224">Figs 1D</figureCitation>
, etc.); its visible length 0.72.0 × visible length of tergum 8 at level of spiracle 8. In male abdomen, tergum 8 with tergal hollow with distinct anterior edge (
<figureCitation id="75D2B2CAC006FF85FB3D2A4EFA94FBD0" box="[1216,1357,1071,1097]" captionStart="FIGURE 14" captionStartId="27.[151,250,1869,1893]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-19@27.[151,1436,181,1845]" captionTargetPageId="27" captionText="FIGURE 14. Apex of male abdomen and genitalia: Fagineura crenativora (A, DF, K, L), F. parva (B, G, H, M, N) and F. togashii (C, I, J, O, P). AC, Apex of abdomen, dorsal view; D, G, I, genitalia, dorsal view; E, F, H, J, genitalia, ventral view (E, H, J, penis valve removed); KP, penis valve, left dorsal." figureDoi="http://doi.org/10.5281/zenodo.6367297" httpUri="https://zenodo.org/record/6367297/files/figure.png" pageId="1" pageNumber="224">Figs 14AC</figureCitation>
). Penis valve with pseudoceps and paravalva divided (
<figureCitation id="75D2B2CAC006FF85FD5F2A32FCF4FBF4" box="[674,813,1107,1133]" captionStart="FIGURE 14" captionStartId="27.[151,250,1869,1893]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-19@27.[151,1436,181,1845]" captionTargetPageId="27" captionText="FIGURE 14. Apex of male abdomen and genitalia: Fagineura crenativora (A, DF, K, L), F. parva (B, G, H, M, N) and F. togashii (C, I, J, O, P). AC, Apex of abdomen, dorsal view; D, G, I, genitalia, dorsal view; E, F, H, J, genitalia, ventral view (E, H, J, penis valve removed); KP, penis valve, left dorsal." figureDoi="http://doi.org/10.5281/zenodo.6367297" httpUri="https://zenodo.org/record/6367297/files/figure.png" pageId="1" pageNumber="224">Figs 14KP</figureCitation>
); paravalva expanded.
</paragraph>
<paragraph id="ED56AE4FC006FF85FF3A2A16FA5CFB0B" blockId="1.[151,1437,423,2034]" box="[199,1413,1143,1170]" pageId="1" pageNumber="224">
<emphasis id="DF9D725DC006FF85FF3A2A16FE15FB08" bold="true" box="[199,460,1143,1169]" pageId="1" pageNumber="224">Character assessment</emphasis>
. We here comment on several characters adopted by previous authors for
<taxonomicName id="2AE9D5CCC006FF85FAF52A19FAA7FB08" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[1288,1406,1144,1169]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC006FF85FAF52A19FAA7FB08" box="[1288,1406,1144,1169]" italics="true" pageId="1" pageNumber="224">Fagineura</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="ED56AE4FC006FF85FF3A2AFDFF35FABB" blockId="1.[151,1437,423,2034]" pageId="1" pageNumber="224">
- Postocellar area length.
<bibRefCitation id="8978D3BEC006FF85FE1C2AFAFD49FB2C" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[481,656,1179,1206]" pageId="1" pageNumber="224" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC006FF85FDF32AFDFDE6FB2C" box="[526,575,1179,1205]" italics="true" pageId="1" pageNumber="224">et al</emphasis>
. (2019
</bibRefCitation>
,
<bibRefCitation id="8978D3BEC006FF85FD672AFAFD0AFB2C" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[666,723,1179,1205]" pageId="1" pageNumber="224" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">2021</bibRefCitation>
) stated “postocellar area more than 2.0 × as wide as long” under the diagnosis of
<taxonomicName id="2AE9D5CCC006FF85FEA92AA1FE13FB40" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[340,458,1216,1241]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC006FF85FEA92AA1FE13FB40" box="[340,458,1216,1241]" italics="true" pageId="1" pageNumber="224">Fagineura</emphasis>
</taxonomicName>
. In our material, the ratio is more than 2.0 × in the
<taxonomicName id="2AE9D5CCC006FF85FBE32AA1FB64FB40" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[1054,1213,1216,1241]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC006FF85FBE32AA1FB64FB40" box="[1054,1213,1216,1241]" italics="true" pageId="1" pageNumber="224">F. crenativora</emphasis>
</taxonomicName>
group (
<figureCitation id="75D2B2CAC006FF85FAEA2ADEFA4CFB43" box="[1303,1429,1215,1242]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="1" pageNumber="224">Figs 6AE</figureCitation>
) but sometimes 2.0 × or less in the
<taxonomicName id="2AE9D5CCC006FF85FDD42A85FD18FB64" authorityName="Togashi" authorityYear="2006" box="[553,705,1252,1277]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC006FF85FDD42A85FD18FB64" box="[553,705,1252,1277]" italics="true" pageId="1" pageNumber="224">F. quercivora</emphasis>
</taxonomicName>
group (
<figureCitation id="75D2B2CAC006FF85FCE12A82FC45FB67" box="[796,924,1251,1278]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="1" pageNumber="224">Figs 6H, L</figureCitation>
) (for the species groups, see under Remarks below).
</paragraph>
<paragraph id="ED56AE4FC006FF85FF3A2B4DFDF1F9D8" blockId="1.[151,1437,423,2034]" pageId="1" pageNumber="224">
- Malar space length.
<bibRefCitation id="8978D3BEC006FF85FE402B4AFDB6FADC" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[445,623,1323,1350]" pageId="1" pageNumber="224" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC006FF85FE162B4DFDC4FADC" box="[491,541,1323,1349]" italics="true" pageId="1" pageNumber="224">et al</emphasis>
. (2019
</bibRefCitation>
,
<bibRefCitation id="8978D3BEC006FF85FD862B4AFD6DFADC" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[635,692,1323,1349]" pageId="1" pageNumber="224" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">2021</bibRefCitation>
) stated “malar space shorter than diameter of median ocellus, and in most species not exceeding 0.5 × of diameter of median ocellus” under the diagnosis of
<taxonomicName id="2AE9D5CCC006FF85FB5F2B31FAC1FAF0" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[1186,1304,1360,1385]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC006FF85FB5F2B31FAC1FAF0" box="[1186,1304,1360,1385]" italics="true" pageId="1" pageNumber="224">Fagineura</emphasis>
</taxonomicName>
. According to them, the ratios of four Chinese species are 0.30.8 ×. In the other seven species of the genus, the malar space is 0.20.8 × as long as the median ocellus width (= horizontal diameter). We also consider the short malar space to be a good generic character of
<taxonomicName id="2AE9D5CCC006FF85FE332BDDFD9DFA4C" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[462,580,1468,1493]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC006FF85FE332BDDFD9DFA4C" box="[462,580,1468,1493]" italics="true" pageId="1" pageNumber="224">Fagineura</emphasis>
</taxonomicName>
.
<bibRefCitation id="8978D3BEC006FF85FDAF2BDDFCFBFA4C" author="Prous, M. &amp; Blank, S. M. &amp; Goulet, H. &amp; Heibo, E. &amp; Liston, A. &amp; Malm, T. &amp; Nyman, T. &amp; Schmidt, S. &amp; Smith, D. R. &amp; Vardal, H. &amp; Viitasaari, M. &amp; Vikberg, V. &amp; Taeger, A." box="[594,802,1467,1494]" pageId="1" pageNumber="224" pagination="1 - 69" refId="ref21989" refString="Prous, M., Blank, S. M., Goulet, H., Heibo, E., Liston, A., Malm, T., Nyman, T., Schmidt, S., Smith, D. R., Vardal, H., Viitasaari, M., Vikberg, V. &amp; Taeger, A. (2014) The genera of Nematinae (Hymenoptera, Tenthredinidae). Journal of Hymenoptera Research, 40, 1 - 69. https: // doi. org / 10.3897 / JHR. 40.7442" type="journal article" year="2014">
Prous
<emphasis id="DF9D725DC006FF85FD642BDDFD13FA4C" box="[665,714,1467,1493]" italics="true" pageId="1" pageNumber="224">et al</emphasis>
. (2014)
</bibRefCitation>
erroneously treated
<taxonomicName id="2AE9D5CCC006FF85FBFA2BDDFBA4FA4C" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[1031,1149,1468,1493]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC006FF85FBFA2BDDFBA4FA4C" box="[1031,1149,1468,1493]" italics="true" pageId="1" pageNumber="224">Fagineura</emphasis>
</taxonomicName>
as one of the genera with “Malar space 1.02.0 times as long as diameter of front ocellus” in the key to
<taxonomicName id="2AE9D5CCC006FF85FBF72BBEFB5AFA60" authorityName="Thomson" authorityYear="1871" box="[1034,1155,1503,1529]" class="Insecta" family="Tenthredinidae" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="subFamily" subFamily="Nematinae">Nematinae</taxonomicName>
genera (pp. 2226, from the lower half of the couplet 18 to the couplet 22). We suggest some modification of the key under Remarks below so that
<taxonomicName id="2AE9D5CCC006FF85FF152849FE87F9D8" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[232,350,1576,1601]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC006FF85FF152849FE87F9D8" box="[232,350,1576,1601]" italics="true" pageId="1" pageNumber="224">Fagineura</emphasis>
</taxonomicName>
can be identified.
</paragraph>
<paragraph id="ED56AE4FC006FF86FF3A282DFB20FED8" blockId="1.[151,1437,423,2034]" lastBlockId="2.[151,1437,151,1977]" lastPageId="2" lastPageNumber="225" pageId="1" pageNumber="224">
- Shape of mandibles. Vikberg &amp; Zinovjev (in
<bibRefCitation id="8978D3BEC006FF85FD34282AFC6EF9FC" author="Shinohara, A. &amp; Vikberg, V. &amp; Zinovjev, A. &amp; Yamagami, A." box="[713,951,1611,1638]" pageId="1" pageNumber="224" pagination="113 - 124" refId="ref22313" refString="Shinohara, A., Vikberg, V., Zinovjev, A. &amp; Yamagami, A. (2000) Fagineura crenativora, a new genus and species of sawfly (Hymenoptera, Tenthredinidae, Nematinae) injurious to beech trees in Japan. Bulletin of the National Science Museum, Series A, 26 (3), 113 - 124." type="journal article" year="2000">
Shinohara
<emphasis id="DF9D725DC006FF85FCBD282DFCA9F9FC" box="[832,880,1611,1637]" italics="true" pageId="1" pageNumber="224">et al</emphasis>
. 2000
</bibRefCitation>
) established
<taxonomicName id="2AE9D5CCC006FF85FBBA282DFB64F9FC" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[1095,1213,1612,1637]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC006FF85FBBA282DFB64F9FC" box="[1095,1213,1612,1637]" italics="true" pageId="1" pageNumber="224">Fagineura</emphasis>
</taxonomicName>
for a single species,
<taxonomicName id="2AE9D5CCC006FF85FF6A2811FEEFF910" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[151,310,1648,1673]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC006FF85FF6A2811FEEFF910" box="[151,310,1648,1673]" italics="true" pageId="1" pageNumber="224">F. crenativora</emphasis>
</taxonomicName>
. One of the generic characters they stated is “Mandibles symmetric, rather narrow, gradually tapering from base to apex, without carina in apical half”. More precisely, each of the mandibles of
<taxonomicName id="2AE9D5CCC006FF85FB4B28F5FA8AF934" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[1206,1363,1684,1709]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC006FF85FB4B28F5FA8AF934" box="[1206,1363,1684,1709]" italics="true" pageId="1" pageNumber="224">F. crenativora</emphasis>
</taxonomicName>
tapers rather sharply on the basal half and gradually on the apical half in the outer view (
<figureCitation id="75D2B2CAC006FF85FBD028D6FB1DF94B" box="[1069,1220,1719,1746]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="1" pageNumber="224">Figs 7A, D, J</figureCitation>
) and has a low and dull apical carina on both anterior and posterior surfaces (
<figureCitation id="75D2B2CAC006FF85FCD528BAFC7AF96C" box="[808,931,1755,1782]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="1" pageNumber="224">Figs 7P, Q</figureCitation>
). This character state is common to
<taxonomicName id="2AE9D5CCC006FF85FAC328BDFA45F96C" authorityName="Hara" authorityYear="2022" box="[1342,1436,1756,1781]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="DF9D725DC006FF85FAC328BDFA45F96C" box="[1342,1436,1756,1781]" italics="true" pageId="1" pageNumber="224">F. parva</emphasis>
</taxonomicName>
described in this paper (
<figureCitation id="75D2B2CAC006FF85FE59289EFDFAF883" box="[420,547,1791,1818]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="1" pageNumber="224">Figs 7E, K</figureCitation>
).
<bibRefCitation id="8978D3BEC006FF85FDCA289EFD30F880" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[567,745,1791,1818]" pageId="1" pageNumber="224" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC006FF85FD982961FD4EF880" box="[613,663,1791,1817]" italics="true" pageId="1" pageNumber="224">et al</emphasis>
. (2019
</bibRefCitation>
,
<bibRefCitation id="8978D3BEC006FF85FD08289EFCF7F880" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[757,814,1791,1817]" pageId="1" pageNumber="224" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">2021</bibRefCitation>
) who described four Chinese species,
<taxonomicName id="2AE9D5CCC006FF85FB222961FAA0F880" authorityName="Liu, Li &amp; Wei" authorityYear="2021" box="[1247,1401,1791,1817]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="DF9D725DC006FF85FB222961FAA0F880" box="[1247,1401,1791,1817]" italics="true" pageId="1" pageNumber="224">F. brevicornis</emphasis>
</taxonomicName>
,
<taxonomicName id="2AE9D5CCC006FF85FA752961FEF1F8A4" authorityName="Liu, Li &amp; Wei" authorityYear="2019" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="flactoserrula">
<emphasis id="DF9D725DC006FF85FA752961FEF1F8A4" italics="true" pageId="1" pageNumber="224">F. flactoserrula</emphasis>
</taxonomicName>
,
<taxonomicName id="2AE9D5CCC006FF85FEC82945FE09F8A4" authorityName="Liu, Li &amp; Wei" authorityYear="2021" box="[309,464,1827,1853]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="longitangia">
<emphasis id="DF9D725DC006FF85FEC82945FE09F8A4" box="[309,464,1827,1853]" italics="true" pageId="1" pageNumber="224">F. longitangia</emphasis>
</taxonomicName>
and
<taxonomicName id="2AE9D5CCC006FF85FDF82945FD7FF8A4" authorityName="Liu, Li &amp; Wei" authorityYear="2019" box="[517,678,1827,1853]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="xanthosoma">
<emphasis id="DF9D725DC006FF85FDF82945FD7FF8A4" box="[517,678,1827,1853]" italics="true" pageId="1" pageNumber="224">F. xanthosoma</emphasis>
</taxonomicName>
, stated only “mandibles symmetrical” under the diagnosis of the genus, and details of mandibles are not given. These six species belong to the
<taxonomicName id="2AE9D5CCC006FF85FC1C2929FBA7F8F8" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[993,1150,1864,1889]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC006FF85FC1C2929FBA7F8F8" box="[993,1150,1864,1889]" italics="true" pageId="1" pageNumber="224">F. crenativora</emphasis>
</taxonomicName>
group. Other five species belong to the
<taxonomicName id="2AE9D5CCC006FF85FED2290DFE1DF81C" authorityName="Togashi" authorityYear="2006" box="[303,452,1900,1925]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC006FF85FED2290DFE1DF81C" box="[303,452,1900,1925]" italics="true" pageId="1" pageNumber="224">F. quercivora</emphasis>
</taxonomicName>
group and their mandibles differ from those of
<taxonomicName id="2AE9D5CCC006FF85FC28290DFBABF81C" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[981,1138,1900,1925]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC006FF85FC28290DFBABF81C" box="[981,1138,1900,1925]" italics="true" pageId="1" pageNumber="224">F. crenativora</emphasis>
</taxonomicName>
and
<taxonomicName id="2AE9D5CCC006FF85FB5A290DFADAF81C" authorityName="Hara" authorityYear="2022" box="[1191,1283,1900,1925]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="DF9D725DC006FF85FB5A290DFADAF81C" box="[1191,1283,1900,1925]" italics="true" pageId="1" pageNumber="224">F. parva</emphasis>
</taxonomicName>
and probably from four Chinese species. The mandibles of the
<taxonomicName id="2AE9D5CCC006FF85FD3D29F1FC8FF830" authorityName="Togashi" authorityYear="2006" box="[704,854,1936,1961]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC006FF85FD3D29F1FC8FF830" box="[704,854,1936,1961]" italics="true" pageId="1" pageNumber="224">F. quercivora</emphasis>
</taxonomicName>
group are slightly asymmetric. The left mandible is constricted in the middle with the middle part very gradually tapered or being the same thickness in the outer view (
<figureCitation id="75D2B2CAC006FF85FF6229B6FED7F86B" box="[159,270,2007,2034]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="1" pageNumber="224">Figs 7FI</figureCitation>
) and with a sharp apical carina on the anterior surface (
<figureCitation id="75D2B2CAC006FF85FC7F29B6FC01F86B" box="[898,984,2007,2034]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="1" pageNumber="224">Fig. 7R</figureCitation>
) and a low and dull apical carina on the posterior surface (
<figureCitation id="75D2B2CAC005FF86FE9A2EF6FE67FF28" box="[359,446,151,177]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="2" pageNumber="225">Fig. 7S</figureCitation>
). The right mandible is constricted in the middle with the middle part being the same thickness or slightly thickened toward the apex in the outer view (
<figureCitation id="75D2B2CAC005FF86FC8A2EDAFC2BFF4C" box="[887,1010,187,213]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="2" pageNumber="225">Figs 7LO</figureCitation>
) and with a low and dull apical carina on the anterior surface (
<figureCitation id="75D2B2CAC005FF86FE572EBEFDDBFF60" box="[426,514,223,249]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="2" pageNumber="225">Fig. 7R</figureCitation>
) and a sharp apical carina on the posterior surface (
<figureCitation id="75D2B2CAC005FF86FBA72EBEFB6BFF60" box="[1114,1202,223,249]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="2" pageNumber="225">Fig. 7S</figureCitation>
). For “
<taxonomicName id="2AE9D5CCC005FF86FAF82EBEFEE0FE84" authority="Togashi, 2006" authorityName="Togashi" authorityYear="2006" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC005FF86FAF82EBEFA45FF61" box="[1285,1436,223,248]" italics="true" pageId="2" pageNumber="225">F. quercivora</emphasis>
<bibRefCitation id="8978D3BEC005FF86FF6A2F62FEE0FE84" author="Togashi, I." box="[151,313,259,285]" pageId="2" pageNumber="225" pagination="169 - 173" refId="ref22763" refString="Togashi, I. (2006) A new sawfly, Fagineura quercivora (Hymenoptera: Tenthredinidae) feeding on Quercus serrata and Q. mongolica crispula in Honshu, Japan. Proceedings of the entomological Society of Washington, 108 (1), 169 - 173." type="journal article" year="2006">Togashi, 2006</bibRefCitation>
</taxonomicName>
” (=
<taxonomicName id="2AE9D5CCC005FF86FE962F62FE38FE84" authorityName="Hara" authorityYear="2022" box="[363,481,259,285]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="togashii">
<emphasis id="DF9D725DC005FF86FE962F62FE38FE84" box="[363,481,259,285]" italics="true" pageId="2" pageNumber="225">F. togashii</emphasis>
</taxonomicName>
),
<bibRefCitation id="8978D3BEC005FF86FE042F62FD7DFE87" author="Togashi, I." box="[505,676,259,286]" pageId="2" pageNumber="225" pagination="169 - 173" refId="ref22763" refString="Togashi, I. (2006) A new sawfly, Fagineura quercivora (Hymenoptera: Tenthredinidae) feeding on Quercus serrata and Q. mongolica crispula in Honshu, Japan. Proceedings of the entomological Society of Washington, 108 (1), 169 - 173." type="journal article" year="2006">Togashi (2006)</bibRefCitation>
wrote “left mandible narrow, rather gradually tapered to apex” and gave the figure (fig.
<quantity id="2A1103AAC005FF86FE802F46FE71FED8" box="[381,424,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="225" unit="in" value="4.0">4 in</quantity>
<bibRefCitation id="8978D3BEC005FF86FE532F46FD90FEDB" author="Togashi, I." box="[430,585,295,322]" pageId="2" pageNumber="225" pagination="169 - 173" refId="ref22763" refString="Togashi, I. (2006) A new sawfly, Fagineura quercivora (Hymenoptera: Tenthredinidae) feeding on Quercus serrata and Q. mongolica crispula in Honshu, Japan. Proceedings of the entomological Society of Washington, 108 (1), 169 - 173." type="journal article" year="2006">Togashi 2006</bibRefCitation>
), but his description and figure are not accurate (see
<figureCitation id="75D2B2CAC005FF86FB652F46FB35FED8" box="[1176,1260,295,321]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="2" pageNumber="225">Fig. 7F</figureCitation>
).
</paragraph>
<paragraph id="ED56AE4FC005FF86FF3A2F2AFDAFFDA4" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="225">
<bibRefCitation id="8978D3BEC005FF86FF3A2F2AFE41FEFC" author="Prous, M. &amp; Blank, S. M. &amp; Goulet, H. &amp; Heibo, E. &amp; Liston, A. &amp; Malm, T. &amp; Nyman, T. &amp; Schmidt, S. &amp; Smith, D. R. &amp; Vardal, H. &amp; Viitasaari, M. &amp; Vikberg, V. &amp; Taeger, A." box="[199,408,331,357]" pageId="2" pageNumber="225" pagination="1 - 69" refId="ref21989" refString="Prous, M., Blank, S. M., Goulet, H., Heibo, E., Liston, A., Malm, T., Nyman, T., Schmidt, S., Smith, D. R., Vardal, H., Viitasaari, M., Vikberg, V. &amp; Taeger, A. (2014) The genera of Nematinae (Hymenoptera, Tenthredinidae). Journal of Hymenoptera Research, 40, 1 - 69. https: // doi. org / 10.3897 / JHR. 40.7442" type="journal article" year="2014">
Prous
<emphasis id="DF9D725DC005FF86FEF32F2AFE9FFEFC" box="[270,326,331,357]" italics="true" pageId="2" pageNumber="225">et al.</emphasis>
(2014)
</bibRefCitation>
divided the character of mandibles into two states in the key to genera of world
<taxonomicName id="2AE9D5CCC005FF86FAE12F2AFA41FEFC" authorityName="Thomson" authorityYear="1871" box="[1308,1432,331,357]" class="Insecta" family="Tenthredinidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="subFamily" subFamily="Nematinae">Nematinae</taxonomicName>
, the couplet 6 “
<emphasis id="DF9D725DC005FF86FEB82F0EFE8AFE10" bold="true" box="[325,339,367,393]" pageId="2" pageNumber="225">a</emphasis>
Left mandible markedly constricted near middle and right mandible tapered regularly towards apex” in the first line and “
<emphasis id="DF9D725DC005FF86FE352FF2FE3DFE34" bold="true" box="[456,484,403,429]" pageId="2" pageNumber="225">aa</emphasis>
Left and right mandible both tapered regularly towards apex” in the second line (see also the couplets 26 and 28), and treated
<taxonomicName id="2AE9D5CCC005FF86FDA62FD6FD08FE49" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[603,721,439,464]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FDA62FD6FD08FE49" box="[603,721,439,464]" italics="true" pageId="2" pageNumber="225">Fagineura</emphasis>
</taxonomicName>
as having both states. However, the state a does not agree with any
<taxonomicName id="2AE9D5CCC005FF86FF342FBAFE40FE6C" box="[201,409,475,501]" pageId="2" pageNumber="225">
<emphasis id="DF9D725DC005FF86FF342FBAFEE6FE6D" box="[201,319,475,500]" italics="true" pageId="2" pageNumber="225">Fagineura</emphasis>
species
</taxonomicName>
(
<figureCitation id="75D2B2CAC005FF86FE572FBAFDF4FE6C" box="[426,557,475,501]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="2" pageNumber="225">Figs 7DO</figureCitation>
), and the state aa almost agrees with
<taxonomicName id="2AE9D5CCC005FF86FC132FBAFB57FE6D" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[1006,1166,475,500]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC005FF86FC132FBAFB57FE6D" box="[1006,1166,475,500]" italics="true" pageId="2" pageNumber="225">F. crenativora</emphasis>
</taxonomicName>
and
<taxonomicName id="2AE9D5CCC005FF86FB352FBAFAFEFE6D" authorityName="Hara" authorityYear="2022" box="[1224,1319,475,500]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="DF9D725DC005FF86FB352FBAFAFEFE6D" box="[1224,1319,475,500]" italics="true" pageId="2" pageNumber="225">F. parva</emphasis>
</taxonomicName>
(
<figureCitation id="75D2B2CAC005FF86FAC52FBAFF31FD80" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="2" pageNumber="225">Figs 7D, E, J, K</figureCitation>
) (and possibly also with the four Chinese species) but not with the
<taxonomicName id="2AE9D5CCC005FF86FC142F9EFB59FD81" authorityName="Togashi" authorityYear="2006" box="[1001,1152,511,536]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC005FF86FC142F9EFB59FD81" box="[1001,1152,511,536]" italics="true" pageId="2" pageNumber="225">F. quercivora</emphasis>
</taxonomicName>
group. We suggest some modification of the key in Remarks below.
</paragraph>
<paragraph id="ED56AE4FC005FF86FF3A2C26FC85FCA0" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="225">
- Setae on katepimeron. Vikberg &amp; Zinovjev (in
<bibRefCitation id="8978D3BEC005FF86FD092C26FC33FDF8" author="Shinohara, A. &amp; Vikberg, V. &amp; Zinovjev, A. &amp; Yamagami, A." box="[756,1002,583,609]" pageId="2" pageNumber="225" pagination="113 - 124" refId="ref22313" refString="Shinohara, A., Vikberg, V., Zinovjev, A. &amp; Yamagami, A. (2000) Fagineura crenativora, a new genus and species of sawfly (Hymenoptera, Tenthredinidae, Nematinae) injurious to beech trees in Japan. Bulletin of the National Science Museum, Series A, 26 (3), 113 - 124." type="journal article" year="2000">
Shinohara
<emphasis id="DF9D725DC005FF86FC932C26FC78FDF8" box="[878,929,583,609]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. 2000
</bibRefCitation>
) wrote “Anterior third of mesopleural katepimeron glabrous, posterior part pilose” for
<taxonomicName id="2AE9D5CCC005FF86FD3B2C0AFCE5FD1D" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[710,828,619,644]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FD3B2C0AFCE5FD1D" box="[710,828,619,644]" italics="true" pageId="2" pageNumber="225">Fagineura</emphasis>
</taxonomicName>
.
<bibRefCitation id="8978D3BEC005FF86FCB02C0AFBFEFD1C" author="Prous, M. &amp; Blank, S. M. &amp; Goulet, H. &amp; Heibo, E. &amp; Liston, A. &amp; Malm, T. &amp; Nyman, T. &amp; Schmidt, S. &amp; Smith, D. R. &amp; Vardal, H. &amp; Viitasaari, M. &amp; Vikberg, V. &amp; Taeger, A." box="[845,1063,619,645]" pageId="2" pageNumber="225" pagination="1 - 69" refId="ref21989" refString="Prous, M., Blank, S. M., Goulet, H., Heibo, E., Liston, A., Malm, T., Nyman, T., Schmidt, S., Smith, D. R., Vardal, H., Viitasaari, M., Vikberg, V. &amp; Taeger, A. (2014) The genera of Nematinae (Hymenoptera, Tenthredinidae). Journal of Hymenoptera Research, 40, 1 - 69. https: // doi. org / 10.3897 / JHR. 40.7442" type="journal article" year="2014">
Prous
<emphasis id="DF9D725DC005FF86FC6A2C0AFC0AFD1C" box="[919,979,619,645]" italics="true" pageId="2" pageNumber="225">et al.</emphasis>
(2014)
</bibRefCitation>
adopted the katepimeron extensively covered with setae as the generic character. In our specimens, the katepimeron is usually covered with setae extensively or moderately, but in
<taxonomicName id="2AE9D5CCC005FF86FDEE2CD2FDA9FD55" authorityName="Hara" authorityYear="2022" box="[531,624,691,716]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="DF9D725DC005FF86FDEE2CD2FDA9FD55" box="[531,624,691,716]" italics="true" pageId="2" pageNumber="225">F. parva</emphasis>
</taxonomicName>
and a few specimens of
<taxonomicName id="2AE9D5CCC005FF86FC762CD2FBF3FD55" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[907,1066,691,716]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC005FF86FC762CD2FBF3FD55" box="[907,1066,691,716]" italics="true" pageId="2" pageNumber="225">F. crenativora</emphasis>
</taxonomicName>
, the katepimeron has only a few or several setae. According to
<bibRefCitation id="8978D3BEC005FF86FE162CB6FD7FFD68" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[491,678,727,753]" pageId="2" pageNumber="225" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC005FF86FDE72CB6FD92FD68" box="[538,587,727,753]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. (2019)
</bibRefCitation>
,
<taxonomicName id="2AE9D5CCC005FF86FD4F2CB6FC86FD68" authorityName="Liu, Li &amp; Wei" authorityYear="2019" box="[690,863,727,753]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="flactoserrula">
<emphasis id="DF9D725DC005FF86FD4F2CB6FC86FD68" box="[690,863,727,753]" italics="true" pageId="2" pageNumber="225">F. flactoserrula</emphasis>
</taxonomicName>
also has only a few setae on the katepimeron. This character is widely variable in the
<taxonomicName id="2AE9D5CCC005FF86FDDD2C9AFD66FC8D" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[544,703,763,788]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC005FF86FDDD2C9AFD66FC8D" box="[544,703,763,788]" italics="true" pageId="2" pageNumber="225">F. crenativora</emphasis>
</taxonomicName>
group, although the katepimeron is never entirely glabrous as in some species of the related genera such as
<taxonomicName id="2AE9D5CCC005FF86FD8B2D7EFD65FCA1" authorityName="Newman" authorityYear="1837" box="[630,700,799,824]" class="Insecta" family="Tenthredinidae" genus="Euura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FD8B2D7EFD65FCA1" box="[630,700,799,824]" italics="true" pageId="2" pageNumber="225">Euura</emphasis>
</taxonomicName>
and
<taxonomicName id="2AE9D5CCC005FF86FD0E2D7EFC8CFCA1" box="[755,853,799,824]" class="Insecta" family="Tenthredinidae" genus="Nematus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FD0E2D7EFC8CFCA1" box="[755,853,799,824]" italics="true" pageId="2" pageNumber="225">Nematus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="ED56AE4FC005FF86FF3A2D22FBA8FC74" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="225">
- Distance between cenchri.
<bibRefCitation id="8978D3BEC005FF86FDFB2D22FD61FCC4" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[518,696,835,861]" pageId="2" pageNumber="225" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC005FF86FDC92D22FDBFFCC4" box="[564,614,835,861]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. (2019
</bibRefCitation>
,
<bibRefCitation id="8978D3BEC005FF86FD3E2D22FD25FCC4" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[707,764,835,861]" pageId="2" pageNumber="225" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">2021</bibRefCitation>
) wrote “distance between cenchri almost as long as breadth of a cenchrus” or “distance between cenchri as long as or shorter than breadth of a cenchrus” under the diagnosis of
<taxonomicName id="2AE9D5CCC005FF86FF482DEAFEF2FC3D" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[181,299,907,932]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FF482DEAFEF2FC3D" box="[181,299,907,932]" italics="true" pageId="2" pageNumber="225">Fagineura</emphasis>
</taxonomicName>
. In our material, the ratio of the distance between cenchri to the major axis of a cenchrus is 0.81.5 × in the genus, 0.91.4 × in the
<taxonomicName id="2AE9D5CCC005FF86FE222DCEFDA5FC51" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[479,636,943,968]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC005FF86FE222DCEFDA5FC51" box="[479,636,943,968]" italics="true" pageId="2" pageNumber="225">F. crenativora</emphasis>
</taxonomicName>
group and 0.81.5 × in the
<taxonomicName id="2AE9D5CCC005FF86FC532DCEFB9BFC51" authorityName="Togashi" authorityYear="2006" box="[942,1090,943,968]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC005FF86FC532DCEFB9BFC51" box="[942,1090,943,968]" italics="true" pageId="2" pageNumber="225">F. quercivora</emphasis>
</taxonomicName>
group. This character is hardly useful to distinguish
<taxonomicName id="2AE9D5CCC005FF86FE7C2DB2FE2EFC75" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[385,503,979,1004]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FE7C2DB2FE2EFC75" box="[385,503,979,1004]" italics="true" pageId="2" pageNumber="225">Fagineura</emphasis>
</taxonomicName>
and the two species groups from other nematine genera.
</paragraph>
<paragraph id="ED56AE4FC005FF86FF3A2D96FBA0FB70" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="225">
- Apical dilation of vein C of fore wing. Vikberg &amp; Zinovjev (in
<bibRefCitation id="8978D3BEC005FF86FC502D96FB7BFB88" author="Shinohara, A. &amp; Vikberg, V. &amp; Zinovjev, A. &amp; Yamagami, A." box="[941,1186,1015,1041]" pageId="2" pageNumber="225" pagination="113 - 124" refId="ref22313" refString="Shinohara, A., Vikberg, V., Zinovjev, A. &amp; Yamagami, A. (2000) Fagineura crenativora, a new genus and species of sawfly (Hymenoptera, Tenthredinidae, Nematinae) injurious to beech trees in Japan. Bulletin of the National Science Museum, Series A, 26 (3), 113 - 124." type="journal article" year="2000">
Shinohara
<emphasis id="DF9D725DC005FF86FBDA2D96FB80FB88" box="[1063,1113,1015,1041]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. 2000
</bibRefCitation>
) and
<bibRefCitation id="8978D3BEC005FF86FB1E2D96FA41FB88" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[1251,1432,1015,1041]" pageId="2" pageNumber="225" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC005FF86FAEF2D96FA9CFB88" box="[1298,1349,1015,1041]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. (2019
</bibRefCitation>
,
<bibRefCitation id="8978D3BEC005FF86FF6A2A7AFF09FBAC" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[151,208,1051,1077]" pageId="2" pageNumber="225" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">2021</bibRefCitation>
) separated
<taxonomicName id="2AE9D5CCC005FF86FEAD2A7AFE1FFBAD" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[336,454,1051,1076]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FEAD2A7AFE1FFBAD" box="[336,454,1051,1076]" italics="true" pageId="2" pageNumber="225">Fagineura</emphasis>
</taxonomicName>
from
<taxonomicName id="2AE9D5CCC005FF86FDF62A7AFD55FBAC" box="[523,652,1051,1077]" class="Insecta" family="Tenthredinidae" genus="Pristiphora" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FDF62A7AFD55FBAC" box="[523,652,1051,1077]" italics="true" pageId="2" pageNumber="225">Pristiphora</emphasis>
</taxonomicName>
and its related genera by the less dilated vein C of the fore wing. This character is not very valuable to distinguish these genera. The apex of vein C is sometimes greatly dilated in the
<taxonomicName id="2AE9D5CCC005FF86FA752A5EFEC0FBE5" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC005FF86FA752A5EFEC0FBE5" italics="true" pageId="2" pageNumber="225">F. crenativora</emphasis>
</taxonomicName>
group (
<figureCitation id="75D2B2CAC005FF86FE8C2A02FE12FBE4" box="[369,459,1123,1149]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="2" pageNumber="225">Fig. 8X</figureCitation>
), usually so in the
<taxonomicName id="2AE9D5CCC005FF86FD622A02FCEDFBE5" authorityName="Togashi" authorityYear="2006" box="[671,820,1123,1148]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC005FF86FD622A02FCEDFBE5" box="[671,820,1123,1148]" italics="true" pageId="2" pageNumber="225">F. quercivora</emphasis>
</taxonomicName>
group (
<figureCitation id="75D2B2CAC005FF86FC712A02FC3DFBE4" box="[908,996,1123,1149]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="2" pageNumber="225">Fig. 8Y</figureCitation>
) and sometimes less noticeably dilated in
<taxonomicName id="2AE9D5CCC005FF86FF492AE6FEECFB38" box="[180,309,1159,1185]" class="Insecta" family="Tenthredinidae" genus="Pristiphora" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FF492AE6FEECFB38" box="[180,309,1159,1185]" italics="true" pageId="2" pageNumber="225">Pristiphora</emphasis>
</taxonomicName>
so that the ratio of the width of cell Sc to the width of vein C at the middle point of the base of vein Rs+M widely overlaps between the two genera. The ratio is 0.11.0 × in
<taxonomicName id="2AE9D5CCC005FF86FC382ACAFBE0FB5D" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[965,1081,1195,1220]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FC382ACAFBE0FB5D" box="[965,1081,1195,1220]" italics="true" pageId="2" pageNumber="225">Fagineura</emphasis>
</taxonomicName>
(0.31.0 × in the
<taxonomicName id="2AE9D5CCC005FF86FAFF2ACAFA45FB5D" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[1282,1436,1195,1220]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC005FF86FAFF2ACAFA45FB5D" box="[1282,1436,1195,1220]" italics="true" pageId="2" pageNumber="225">F. crenativora</emphasis>
</taxonomicName>
group and 0.10.5 × in the
<taxonomicName id="2AE9D5CCC005FF86FE422AAEFD88FB71" authorityName="Togashi" authorityYear="2006" box="[447,593,1231,1256]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC005FF86FE422AAEFD88FB71" box="[447,593,1231,1256]" italics="true" pageId="2" pageNumber="225">F. quercivora</emphasis>
</taxonomicName>
group) and 00.4 × in
<taxonomicName id="2AE9D5CCC005FF86FCB12AAEFC13FB70" box="[844,970,1231,1257]" class="Insecta" family="Tenthredinidae" genus="Pristiphora" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FCB12AAEFC13FB70" box="[844,970,1231,1257]" italics="true" pageId="2" pageNumber="225">Pristiphora</emphasis>
</taxonomicName>
in our material.
</paragraph>
<paragraph id="ED56AE4FC005FF86FF3A2A92FEB5FA04" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="225">
- Presence or absence of crossvein 2r-rs of fore wing. Previously known species of
<taxonomicName id="2AE9D5CCC005FF86FB872A92FB29FA95" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[1146,1264,1267,1292]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FB872A92FB29FA95" box="[1146,1264,1267,1292]" italics="true" pageId="2" pageNumber="225">Fagineura</emphasis>
</taxonomicName>
have no crossvein 2r-rs. Therefore, the absence of crossvein 2r-rs has been adopted as the generic character by previous authors (Vikberg &amp; Zinovjev in
<bibRefCitation id="8978D3BEC005FF86FE562B5AFD79FACC" author="Shinohara, A. &amp; Vikberg, V. &amp; Zinovjev, A. &amp; Yamagami, A." box="[427,672,1339,1365]" pageId="2" pageNumber="225" pagination="113 - 124" refId="ref22313" refString="Shinohara, A., Vikberg, V., Zinovjev, A. &amp; Yamagami, A. (2000) Fagineura crenativora, a new genus and species of sawfly (Hymenoptera, Tenthredinidae, Nematinae) injurious to beech trees in Japan. Bulletin of the National Science Museum, Series A, 26 (3), 113 - 124." type="journal article" year="2000">
Shinohara
<emphasis id="DF9D725DC005FF86FDD92B5AFD8FFACC" box="[548,598,1339,1365]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. 2000
</bibRefCitation>
,
<bibRefCitation id="8978D3BEC005FF86FD512B5AFC8EFACC" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[684,855,1339,1365]" pageId="2" pageNumber="225" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC005FF86FD262B5AFCD4FACC" box="[731,781,1339,1365]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. 2019
</bibRefCitation>
,
<bibRefCitation id="8978D3BEC005FF86FC9F2B5AFC47FACC" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[866,926,1339,1365]" pageId="2" pageNumber="225" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">2021</bibRefCitation>
). In this paper, we move
<taxonomicName id="2AE9D5CCC005FF86FB402B5AFEEEFAE0" authority="Togashi, 1997" authorityName="Togashi" authorityYear="1997" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC005FF86FB402B5AFA45FACD" box="[1213,1436,1339,1364]" italics="true" pageId="2" pageNumber="225">Dineura quercivora</emphasis>
<bibRefCitation id="8978D3BEC005FF86FF6A2B3EFEEEFAE0" author="Togashi, I." box="[151,311,1375,1401]" pageId="2" pageNumber="225" pagination="195 - 198" refId="ref22718" refString="Togashi, I. (1997) A new species of Dineura Dahlbom (Hymenoptera: Tenthredinidae) feeding on Quercus dentata Thunberg (Dicotyledoneae: Fagaceae) from Japan. Japanese Journal of Entomology, 65 (1), 195 - 198." type="journal article" year="1997">Togashi, 1997</bibRefCitation>
</taxonomicName>
of which the
<typeStatus id="325210EDC005FF86FE2B2B3EFDE1FAE0" box="[470,568,1375,1401]" pageId="2" pageNumber="225" type="holotype">holotype</typeStatus>
has crossvein 2r-rs to
<taxonomicName id="2AE9D5CCC005FF86FCC62B3EFC68FAE1" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[827,945,1375,1400]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FCC62B3EFC68FAE1" box="[827,945,1375,1400]" italics="true" pageId="2" pageNumber="225">Fagineura</emphasis>
</taxonomicName>
(for the reason, see Remarks of
<taxonomicName id="2AE9D5CCC005FF86FADB2B3EFA45FAE1" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[1318,1436,1375,1400]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FADB2B3EFA45FAE1" box="[1318,1436,1375,1400]" italics="true" pageId="2" pageNumber="225">Fagineura</emphasis>
</taxonomicName>
and
<taxonomicName id="2AE9D5CCC005FF86FF3A2BE2FE85FA05" authorityName="Togashi" authorityYear="1997" box="[199,348,1411,1436]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC005FF86FF3A2BE2FE85FA05" box="[199,348,1411,1436]" italics="true" pageId="2" pageNumber="225">F. quercivora</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="ED56AE4FC005FF86FF3A2BC6FE7FF9B4" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="225">
- Apical emargination of valvula 3. Vikberg &amp; Zinovjev (in
<bibRefCitation id="8978D3BEC005FF86FC7B2BC6FB5BFA58" author="Shinohara, A. &amp; Vikberg, V. &amp; Zinovjev, A. &amp; Yamagami, A." box="[902,1154,1447,1473]" pageId="2" pageNumber="225" pagination="113 - 124" refId="ref22313" refString="Shinohara, A., Vikberg, V., Zinovjev, A. &amp; Yamagami, A. (2000) Fagineura crenativora, a new genus and species of sawfly (Hymenoptera, Tenthredinidae, Nematinae) injurious to beech trees in Japan. Bulletin of the National Science Museum, Series A, 26 (3), 113 - 124." type="journal article" year="2000">
Shinohara
<emphasis id="DF9D725DC005FF86FBFF2BC6FBEEFA58" box="[1026,1079,1447,1473]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. 2000
</bibRefCitation>
) considered the apically emarginate valvula 3 to be a character of the genus. However,
<taxonomicName id="2AE9D5CCC005FF86FCBA2BAAFC30FA7C" authorityName="Liu, Li &amp; Wei" authorityYear="2019" box="[839,1001,1483,1509]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="xanthosoma">
<emphasis id="DF9D725DC005FF86FCBA2BAAFC30FA7C" box="[839,1001,1483,1509]" italics="true" pageId="2" pageNumber="225">F. xanthosoma</emphasis>
</taxonomicName>
of the
<taxonomicName id="2AE9D5CCC005FF86FBC82BAAFB08FA7D" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[1077,1233,1483,1508]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC005FF86FBC82BAAFB08FA7D" box="[1077,1233,1483,1508]" italics="true" pageId="2" pageNumber="225">F. crenativora</emphasis>
</taxonomicName>
group and several species of the
<taxonomicName id="2AE9D5CCC005FF86FEC12B8EFE0DF991" authorityName="Togashi" authorityYear="1997" box="[316,468,1519,1544]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC005FF86FEC12B8EFE0DF991" box="[316,468,1519,1544]" italics="true" pageId="2" pageNumber="225">F. quercivora</emphasis>
</taxonomicName>
group have the apically not or indistinctly emarginate valvula 3 (fig. 2f in
<bibRefCitation id="8978D3BEC005FF86FACE2B8EFF0BF9B4" author="Liu, M. &amp; Li, Z. &amp; Wei, M." pageId="2" pageNumber="225" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC005FF86FA9F2B8EFA4CF990" box="[1378,1429,1519,1545]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. 2019
</bibRefCitation>
;
<figureCitation id="75D2B2CAC005FF86FF232872FE40F9B4" box="[222,409,1555,1581]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). AH, Dorsal view; IP, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="2" pageNumber="225">Figs 9C, D, F, G</figureCitation>
).
</paragraph>
<paragraph id="ED56AE4FC005FF86FF3A2856FD65F924" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="225">
- Presence or absence of pores in tangium of lancet. The pored tangium was considered the generic character of
<taxonomicName id="2AE9D5CCC005FF86FF4A283AFEF4F9ED" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[183,301,1627,1652]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC005FF86FF4A283AFEF4F9ED" box="[183,301,1627,1652]" italics="true" pageId="2" pageNumber="225">Fagineura</emphasis>
</taxonomicName>
by Vikberg &amp; Zinovjev (in
<bibRefCitation id="8978D3BEC005FF86FD91283AFCB9F9EC" author="Shinohara, A. &amp; Vikberg, V. &amp; Zinovjev, A. &amp; Yamagami, A." box="[620,864,1627,1653]" pageId="2" pageNumber="225" pagination="113 - 124" refId="ref22313" refString="Shinohara, A., Vikberg, V., Zinovjev, A. &amp; Yamagami, A. (2000) Fagineura crenativora, a new genus and species of sawfly (Hymenoptera, Tenthredinidae, Nematinae) injurious to beech trees in Japan. Bulletin of the National Science Museum, Series A, 26 (3), 113 - 124." type="journal article" year="2000">
Shinohara
<emphasis id="DF9D725DC005FF86FD18283AFCCEF9EC" box="[741,791,1627,1653]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. 2000
</bibRefCitation>
) and
<bibRefCitation id="8978D3BEC005FF86FC5D283AFBAEF9EC" author="Prous, M. &amp; Blank, S. M. &amp; Goulet, H. &amp; Heibo, E. &amp; Liston, A. &amp; Malm, T. &amp; Nyman, T. &amp; Schmidt, S. &amp; Smith, D. R. &amp; Vardal, H. &amp; Viitasaari, M. &amp; Vikberg, V. &amp; Taeger, A." box="[928,1143,1627,1653]" pageId="2" pageNumber="225" pagination="1 - 69" refId="ref21989" refString="Prous, M., Blank, S. M., Goulet, H., Heibo, E., Liston, A., Malm, T., Nyman, T., Schmidt, S., Smith, D. R., Vardal, H., Viitasaari, M., Vikberg, V. &amp; Taeger, A. (2014) The genera of Nematinae (Hymenoptera, Tenthredinidae). Journal of Hymenoptera Research, 40, 1 - 69. https: // doi. org / 10.3897 / JHR. 40.7442" type="journal article" year="2014">
Prous
<emphasis id="DF9D725DC005FF86FC15283AFBF8F9EC" box="[1000,1057,1627,1653]" italics="true" pageId="2" pageNumber="225">et al.</emphasis>
(2014)
</bibRefCitation>
. However, the tangium of
<taxonomicName id="2AE9D5CCC005FF86FF6A281EFE7FF900" authorityName="Liu, Li &amp; Wei" authorityYear="2019" box="[151,422,1663,1689]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="flactoserrula">
<emphasis id="DF9D725DC005FF86FF6A281EFE7FF900" box="[151,422,1663,1689]" italics="true" pageId="2" pageNumber="225">Fagineura flactoserrula</emphasis>
</taxonomicName>
has only one pore (fig. 1j in
<bibRefCitation id="8978D3BEC005FF86FD0F281EFC47F900" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[754,926,1663,1689]" pageId="2" pageNumber="225" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC005FF86FCDC281EFC8AF900" box="[801,851,1663,1689]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. 2019
</bibRefCitation>
), and those of
<taxonomicName id="2AE9D5CCC005FF86FBB8281EFB7AF901" authorityName="Hara" authorityYear="2022" box="[1093,1187,1663,1688]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="DF9D725DC005FF86FBB8281EFB7AF901" box="[1093,1187,1663,1688]" italics="true" pageId="2" pageNumber="225">F. parva</emphasis>
</taxonomicName>
and the
<taxonomicName id="2AE9D5CCC005FF86FAFB281EFA45F901" authorityName="Togashi" authorityYear="1997" box="[1286,1436,1663,1688]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC005FF86FAFB281EFA45F901" box="[1286,1436,1663,1688]" italics="true" pageId="2" pageNumber="225">F. quercivora</emphasis>
</taxonomicName>
group have no pores (
<figureCitation id="75D2B2CAC005FF86FE7728C2FE2AF924" box="[394,499,1699,1725]" captionStart="FIGURE 11" captionStartId="24.[151,250,1126,1150]" captionTargetBox="[151,1436,181,1094]" captionTargetId="figure-26@24.[151,1436,181,1103]" captionTargetPageId="24" captionText="FIGURE 11. Lancet: Fagineura crenativora (A, B), F. parva (C, holotype)." figureDoi="http://doi.org/10.5281/zenodo.6367291" httpUri="https://zenodo.org/record/6367291/files/figure.png" pageId="2" pageNumber="225">Figs 11C</figureCitation>
,
<figureCitation id="75D2B2CAC005FF86FE0328C2FD89F924" box="[510,592,1699,1725]" captionStart="FIGURE 12" captionStartId="25.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1862]" captionTargetId="figure-19@25.[151,1436,181,1870]" captionTargetPageId="25" captionText="FIGURE 12. Lancet: Fagineura togashii (A, B), F. fulvistriata (C, holotype) and F. flavomaculata (D, E; D, holotype). A, B, C, Reversed." figureDoi="http://doi.org/10.5281/zenodo.6367293" httpUri="https://zenodo.org/record/6367293/files/figure.png" pageId="2" pageNumber="225">12AE</figureCitation>
,
<figureCitation id="75D2B2CAC005FF86FDA128C2FD69F924" box="[604,688,1699,1725]" captionStart="FIGURE 13" captionStartId="26.[151,250,1847,1871]" captionTargetBox="[151,1436,181,1823]" captionTargetId="figure-26@26.[151,1436,181,1823]" captionTargetPageId="26" captionText="FIGURE 13. Lancet: Fagineura quercivora (AC; A, holotype) and F. glabella (D, E; D, holotype). A, Reversed." figureDoi="http://doi.org/10.5281/zenodo.6367295" httpUri="https://zenodo.org/record/6367295/files/figure.png" pageId="2" pageNumber="225">13AE</figureCitation>
).
</paragraph>
<paragraph id="ED56AE4FC005FF86FF3A28A6FDE0F820" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="225">
- Presence or absence of ctenidium in most basal annulus of lancet.
<bibRefCitation id="8978D3BEC005FF86FC3028A6FB5BF978" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[973,1154,1735,1761]" pageId="2" pageNumber="225" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC005FF86FC0028A6FBF6F978" box="[1021,1071,1735,1761]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. (2019
</bibRefCitation>
,
<bibRefCitation id="8978D3BEC005FF86FB7228A6FB11F978" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[1167,1224,1735,1761]" pageId="2" pageNumber="225" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">2021</bibRefCitation>
) adopted the presence of a ctenidium in the annulus 1 of a lancet as the generic character. In all the members of the
<taxonomicName id="2AE9D5CCC005FF86FB03288AFA44F89D" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[1278,1437,1771,1796]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC005FF86FB03288AFA44F89D" box="[1278,1437,1771,1796]" italics="true" pageId="2" pageNumber="225">F. crenativora</emphasis>
</taxonomicName>
group, a ctenidium is present in the annulus 1 and it occurs in a transverse colorless area (
<figureCitation id="75D2B2CAC005FF86FB70296EFAC3F8B0" box="[1165,1306,1807,1833]" captionStart="FIGURE 11" captionStartId="24.[151,250,1126,1150]" captionTargetBox="[151,1436,181,1094]" captionTargetId="figure-26@24.[151,1436,181,1103]" captionTargetPageId="24" captionText="FIGURE 11. Lancet: Fagineura crenativora (A, B), F. parva (C, holotype)." figureDoi="http://doi.org/10.5281/zenodo.6367291" httpUri="https://zenodo.org/record/6367291/files/figure.png" pageId="2" pageNumber="225">Figs 11AC</figureCitation>
; figs 2AD in
<bibRefCitation id="8978D3BEC005FF86FF492952FE87F8D4" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[180,350,1843,1869]" pageId="2" pageNumber="225" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">
Liu
<emphasis id="DF9D725DC005FF86FF1F2952FECDF8D4" box="[226,276,1843,1869]" italics="true" pageId="2" pageNumber="225">et al</emphasis>
. 2021
</bibRefCitation>
). In
<taxonomicName id="2AE9D5CCC005FF86FE6D2952FE34F8D5" authorityName="Hara" authorityYear="2022" box="[400,493,1843,1868]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="DF9D725DC005FF86FE6D2952FE34F8D5" box="[400,493,1843,1868]" italics="true" pageId="2" pageNumber="225">F. parva</emphasis>
</taxonomicName>
, the ctenidium in the annulus 1 is small, but the colorless area with the ctenidium is well developed (
<figureCitation id="75D2B2CAC005FF86FEAE2936FE60F8E8" box="[339,441,1879,1905]" captionStart="FIGURE 11" captionStartId="24.[151,250,1126,1150]" captionTargetBox="[151,1436,181,1094]" captionTargetId="figure-26@24.[151,1436,181,1103]" captionTargetPageId="24" captionText="FIGURE 11. Lancet: Fagineura crenativora (A, B), F. parva (C, holotype)." figureDoi="http://doi.org/10.5281/zenodo.6367291" httpUri="https://zenodo.org/record/6367291/files/figure.png" pageId="2" pageNumber="225">Fig. 11C</figureCitation>
). In the
<taxonomicName id="2AE9D5CCC005FF86FDE42936FD76F8E9" authorityName="Togashi" authorityYear="1997" box="[537,687,1879,1904]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC005FF86FDE42936FD76F8E9" box="[537,687,1879,1904]" italics="true" pageId="2" pageNumber="225">F. quercivora</emphasis>
</taxonomicName>
group, the ctenidium and related colorless area are inconspicuous or absent (
<figureCitation id="75D2B2CAC005FF86FEF1291AFE4EF80C" box="[268,407,1915,1941]" captionStart="FIGURE 12" captionStartId="25.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1862]" captionTargetId="figure-19@25.[151,1436,181,1870]" captionTargetPageId="25" captionText="FIGURE 12. Lancet: Fagineura togashii (A, B), F. fulvistriata (C, holotype) and F. flavomaculata (D, E; D, holotype). A, B, C, Reversed." figureDoi="http://doi.org/10.5281/zenodo.6367293" httpUri="https://zenodo.org/record/6367293/files/figure.png" pageId="2" pageNumber="225">Figs 12AE</figureCitation>
,
<figureCitation id="75D2B2CAC005FF86FE5F291AFE2FF80C" box="[418,502,1915,1941]" captionStart="FIGURE 13" captionStartId="26.[151,250,1847,1871]" captionTargetBox="[151,1436,181,1823]" captionTargetId="figure-26@26.[151,1436,181,1823]" captionTargetPageId="26" captionText="FIGURE 13. Lancet: Fagineura quercivora (AC; A, holotype) and F. glabella (D, E; D, holotype). A, Reversed." figureDoi="http://doi.org/10.5281/zenodo.6367295" httpUri="https://zenodo.org/record/6367295/files/figure.png" pageId="2" pageNumber="225">13AE</figureCitation>
). The development of the colorless area with a ctenidium in the annulus 1 is a good character of the
<taxonomicName id="2AE9D5CCC005FF86FEB129FEFE33F821" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[332,490,1951,1976]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC005FF86FEB129FEFE33F821" box="[332,490,1951,1976]" italics="true" pageId="2" pageNumber="225">F. crenativora</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="ED56AE4FC004FF87FF3A2EF6FDCCFE84" blockId="3.[151,1437,151,2009]" pageId="3" pageNumber="226">
- Tergum 8 of male.
<bibRefCitation id="8978D3BEC004FF87FE562EF6FDBBFF28" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[427,610,151,177]" pageId="3" pageNumber="226" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC004FF87FE252EF6FDD0FF28" box="[472,521,151,177]" italics="true" pageId="3" pageNumber="226">et al</emphasis>
. (2019)
</bibRefCitation>
wrote “in males, the posterior end of tergum 8 with distinct apical projection” as one of the character to distinguish
<taxonomicName id="2AE9D5CCC004FF87FD7D2EDAFD2FFF4D" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[640,758,187,212]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FD7D2EDAFD2FFF4D" box="[640,758,187,212]" italics="true" pageId="3" pageNumber="226">Fagineura</emphasis>
</taxonomicName>
from
<taxonomicName id="2AE9D5CCC004FF87FCC02EDAFC67FF4C" box="[829,958,187,213]" class="Insecta" family="Tenthredinidae" genus="Pristiphora" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FCC02EDAFC67FF4C" box="[829,958,187,213]" italics="true" pageId="3" pageNumber="226">Pristiphora</emphasis>
</taxonomicName>
. Actually, certain members of
<taxonomicName id="2AE9D5CCC004FF87FAE62EDAFA45FF4C" box="[1307,1436,187,213]" class="Insecta" family="Tenthredinidae" genus="Pristiphora" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FAE62EDAFA45FF4C" box="[1307,1436,187,213]" italics="true" pageId="3" pageNumber="226">Pristiphora</emphasis>
</taxonomicName>
also have a distinct procidentia (compare
<figureCitation id="75D2B2CAC004FF87FD9A2EBEFD37FF60" box="[615,750,223,249]" captionStart="FIGURE 14" captionStartId="27.[151,250,1869,1893]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-19@27.[151,1436,181,1845]" captionTargetPageId="27" captionText="FIGURE 14. Apex of male abdomen and genitalia: Fagineura crenativora (A, DF, K, L), F. parva (B, G, H, M, N) and F. togashii (C, I, J, O, P). AC, Apex of abdomen, dorsal view; D, G, I, genitalia, dorsal view; E, F, H, J, genitalia, ventral view (E, H, J, penis valve removed); KP, penis valve, left dorsal." figureDoi="http://doi.org/10.5281/zenodo.6367297" httpUri="https://zenodo.org/record/6367297/files/figure.png" pageId="3" pageNumber="226">Figs 14AC</figureCitation>
with figs
<quantity id="2A1103AAC004FF87FCA32EBFFC18FF60" box="[862,961,222,249]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" metricValueMax="1.4478" metricValueMin="1.3969999999999998" pageId="3" pageNumber="226" unit="in" value="56.0" valueMax="57.0" valueMin="55.0">5557 in</quantity>
<bibRefCitation id="8978D3BEC004FF87FC3A2EBEFB5EFF60" author="Prous, M. &amp; Kramp, K. &amp; Vikberg, V. &amp; Liston, A." box="[967,1159,223,249]" pageId="3" pageNumber="226" pagination="1 - 190" refId="ref22093" refString="Prous, M., Kramp, K., Vikberg, V. &amp; Liston, A. (2017) North-Western Palaearctic species of Pristiphora (Hymenoptera, Tenthredinidae). Journal of Hymenoptera Research, 59, 1 - 190. https: // doi. org / 10.3897 / jhr. 59.12656" type="journal article" year="2017">
Prous
<emphasis id="DF9D725DC004FF87FBF32EBEFBE7FF60" box="[1038,1086,223,249]" italics="true" pageId="3" pageNumber="226">et al</emphasis>
. 2017
</bibRefCitation>
, figs 3Q, V in Hara
<emphasis id="DF9D725DC004FF87FA982EBEFA4CFF60" box="[1381,1429,223,249]" italics="true" pageId="3" pageNumber="226">et al</emphasis>
. 2018, fig. 5H in
<bibRefCitation id="8978D3BEC004FF87FEAD2F62FDD0FE84" author="Hara, H. &amp; Iwasaki, A. &amp; Shinohara, A." box="[336,521,259,285]" pageId="3" pageNumber="226" pagination="393 - 404" refId="ref21677" refString="Hara, H., Iwasaki, A. &amp; Shinohara, A. (2021) Taxonomic and biological notes on Pristiphora ribisi Togashi, 1990, (Hymenoptera, Tenthredinidae) injurious to gooseberries and currants. Zootaxa, 4903 (3), 393 - 404. https: // doi. org / 10.11646 / zootaxa. 4903.3.5" type="journal article" year="2021">
Hara
<emphasis id="DF9D725DC004FF87FE732F62FE66FE84" box="[398,447,259,285]" italics="true" pageId="3" pageNumber="226">et al</emphasis>
. 2021
</bibRefCitation>
).
</paragraph>
<paragraph id="ED56AE4FC004FF87FF3A2F46FEB8FE48" blockId="3.[151,1437,151,2009]" pageId="3" pageNumber="226">
- Extra larval molt. The larva of
<taxonomicName id="2AE9D5CCC004FF87FDD72F46FD1EFED9" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[554,711,295,320]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC004FF87FDD72F46FD1EFED9" box="[554,711,295,320]" italics="true" pageId="3" pageNumber="226">F. crenativora</emphasis>
</taxonomicName>
has an extra molt after the feeding period (
<bibRefCitation id="8978D3BEC004FF87FB632F46FA48FED8" author="Shinohara, A. &amp; Vikberg, V. &amp; Zinovjev, A. &amp; Yamagami, A." box="[1182,1425,295,321]" pageId="3" pageNumber="226" pagination="113 - 124" refId="ref22313" refString="Shinohara, A., Vikberg, V., Zinovjev, A. &amp; Yamagami, A. (2000) Fagineura crenativora, a new genus and species of sawfly (Hymenoptera, Tenthredinidae, Nematinae) injurious to beech trees in Japan. Bulletin of the National Science Museum, Series A, 26 (3), 113 - 124." type="journal article" year="2000">
Shinohara
<emphasis id="DF9D725DC004FF87FAEA2F46FA9EFED8" box="[1303,1351,295,321]" italics="true" pageId="3" pageNumber="226">et al</emphasis>
. 2000
</bibRefCitation>
). Vikberg &amp; Zinovjev (in
<bibRefCitation id="8978D3BEC004FF87FE5A2F2AFD41FEFC" author="Shinohara, A. &amp; Vikberg, V. &amp; Zinovjev, A. &amp; Yamagami, A." box="[423,664,331,357]" pageId="3" pageNumber="226" pagination="113 - 124" refId="ref22313" refString="Shinohara, A., Vikberg, V., Zinovjev, A. &amp; Yamagami, A. (2000) Fagineura crenativora, a new genus and species of sawfly (Hymenoptera, Tenthredinidae, Nematinae) injurious to beech trees in Japan. Bulletin of the National Science Museum, Series A, 26 (3), 113 - 124." type="journal article" year="2000">
Shinohara
<emphasis id="DF9D725DC004FF87FDE22F2AFD89FEFC" box="[543,592,331,357]" italics="true" pageId="3" pageNumber="226">et al</emphasis>
. 2000
</bibRefCitation>
) mentioned the presence of an extra larval molt as one of the generic characters. We observed that the mature larva of
<taxonomicName id="2AE9D5CCC004FF87FD472F0EFCF6FE10" authorityName="Hara" authorityYear="2022" box="[698,815,367,393]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="togashii">
<emphasis id="DF9D725DC004FF87FD472F0EFCF6FE10" box="[698,815,367,393]" italics="true" pageId="3" pageNumber="226">F. togashii</emphasis>
</taxonomicName>
of the
<taxonomicName id="2AE9D5CCC004FF87FC832F0EFBCAFE11" authorityName="Togashi" authorityYear="1997" box="[894,1043,367,392]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC004FF87FC832F0EFBCAFE11" box="[894,1043,367,392]" italics="true" pageId="3" pageNumber="226">F. quercivora</emphasis>
</taxonomicName>
group entered into the soil without an extra molt. The presence or absence of an extra molt may differ between the
<taxonomicName id="2AE9D5CCC004FF87FBCA2FF2FB0EFE35" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[1079,1239,403,428]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC004FF87FBCA2FF2FB0EFE35" box="[1079,1239,403,428]" italics="true" pageId="3" pageNumber="226">F. crenativora</emphasis>
</taxonomicName>
group and the
<taxonomicName id="2AE9D5CCC004FF87FA752FF2FEC8FE49" authorityName="Togashi" authorityYear="1997" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC004FF87FA752FF2FEC8FE49" italics="true" pageId="3" pageNumber="226">F. quercivora</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection id="A5F3FDC4C004FF80FF3A2FBAFC40FD80" lastPageId="4" lastPageNumber="227" pageId="3" pageNumber="226" type="discussion">
<paragraph id="ED56AE4FC004FF87FF3A2FBAFA45FB7D" blockId="3.[151,1437,151,2009]" pageId="3" pageNumber="226">
<emphasis id="DF9D725DC004FF87FF3A2FBAFEEDFE6C" bold="true" box="[199,308,475,501]" pageId="3" pageNumber="226">Remarks</emphasis>
. The key to the genera of world
<taxonomicName id="2AE9D5CCC004FF87FD642FBAFCCBFE6C" authorityName="Thomson" authorityYear="1871" box="[665,786,475,501]" class="Insecta" family="Tenthredinidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="subFamily" subFamily="Nematinae">Nematinae</taxonomicName>
by
<bibRefCitation id="8978D3BEC004FF87FCC72FBAFBD1FE6C" author="Prous, M. &amp; Blank, S. M. &amp; Goulet, H. &amp; Heibo, E. &amp; Liston, A. &amp; Malm, T. &amp; Nyman, T. &amp; Schmidt, S. &amp; Smith, D. R. &amp; Vardal, H. &amp; Viitasaari, M. &amp; Vikberg, V. &amp; Taeger, A." box="[826,1032,475,501]" pageId="3" pageNumber="226" pagination="1 - 69" refId="ref21989" refString="Prous, M., Blank, S. M., Goulet, H., Heibo, E., Liston, A., Malm, T., Nyman, T., Schmidt, S., Smith, D. R., Vardal, H., Viitasaari, M., Vikberg, V. &amp; Taeger, A. (2014) The genera of Nematinae (Hymenoptera, Tenthredinidae). Journal of Hymenoptera Research, 40, 1 - 69. https: // doi. org / 10.3897 / JHR. 40.7442" type="journal article" year="2014">
Prous
<emphasis id="DF9D725DC004FF87FC7D2FBAFC6EFE6C" box="[896,951,475,501]" italics="true" pageId="3" pageNumber="226">et al.</emphasis>
(2014)
</bibRefCitation>
needs some modification to identify
<taxonomicName id="2AE9D5CCC004FF87FF6A2F9EFED4FD81" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[151,269,511,536]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FF6A2F9EFED4FD81" box="[151,269,511,536]" italics="true" pageId="3" pageNumber="226">Fagineura</emphasis>
</taxonomicName>
. The
<taxonomicName id="2AE9D5CCC004FF87FEAD2F9EFDF9FD80" box="[336,544,511,537]" pageId="3" pageNumber="226">
<emphasis id="DF9D725DC004FF87FEAD2F9EFE1FFD81" box="[336,454,511,536]" italics="true" pageId="3" pageNumber="226">Fagineura</emphasis>
species
</taxonomicName>
without crossvein 2r-rs go to the couplet 6 but part of them do not agree with either line of the couplet as stated in Character assessment (Shape of mandibles). To fit the part of
<taxonomicName id="2AE9D5CCC004FF87FB0B2C42FAB5FDA5" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[1270,1388,547,572]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FB0B2C42FAB5FDA5" box="[1270,1388,547,572]" italics="true" pageId="3" pageNumber="226">Fagineura</emphasis>
</taxonomicName>
and some
<taxonomicName id="2AE9D5CCC004FF87FF272C26FE8BFDF9" authorityName="Rohwer" authorityYear="1911" box="[218,338,583,608]" class="Insecta" family="Tenthredinidae" genus="Nematinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FF272C26FE8BFDF9" box="[218,338,583,608]" italics="true" pageId="3" pageNumber="226">Nematinus</emphasis>
</taxonomicName>
(see the last paragraph), the state aa in the second line of the couplet 6 needs to be modified as follows: Left and right mandible both tapered regularly towards apex
<emphasis id="DF9D725DC004FF87FC512C0AFB18FD30" italics="true" pageId="3" pageNumber="226">or markedly constricted near middle, or left mandible tapered regularly towards apex and right mandible markedly constricted near middle</emphasis>
(modified parts are shown in italics; the former modified part is for the part of
<taxonomicName id="2AE9D5CCC004FF87FCE22CD2FC4CFD55" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[799,917,691,716]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FCE22CD2FC4CFD55" box="[799,917,691,716]" italics="true" pageId="3" pageNumber="226">Fagineura</emphasis>
</taxonomicName>
and the latter for some
<taxonomicName id="2AE9D5CCC004FF87FB652CD2FAC9FD55" authorityName="Rohwer" authorityYear="1911" box="[1176,1296,691,716]" class="Insecta" family="Tenthredinidae" genus="Nematinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FB652CD2FAC9FD55" box="[1176,1296,691,716]" italics="true" pageId="3" pageNumber="226">Nematinus</emphasis>
</taxonomicName>
).
<taxonomicName id="2AE9D5CCC004FF87FADB2CD2FA45FD55" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[1318,1436,691,716]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FADB2CD2FA45FD55" box="[1318,1436,691,716]" italics="true" pageId="3" pageNumber="226">Fagineura</emphasis>
</taxonomicName>
goes through the modified second half of the couplet 6 to the couplet 18, but disagrees with either half of the couplet 18, because it has the subapical tooth of a tarsal claw not larger than the apical tooth* (aa in the second half), a short malar space (b in the first half) and a widely emarginate clypeus (cc in the second half) [*
<bibRefCitation id="8978D3BEC004FF87FB182D7EFA45FCA0" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[1253,1436,799,825]" pageId="3" pageNumber="226" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">
Liu
<emphasis id="DF9D725DC004FF87FAEE2D7EFA9DFCA0" box="[1299,1348,799,825]" italics="true" pageId="3" pageNumber="226">et al</emphasis>
. (2021)
</bibRefCitation>
stated “inner tooth of tarsal claw longer than outer tooth” for “
<taxonomicName id="2AE9D5CCC004FF87FCAE2D22FC31FCC5" authorityName="Togashi" authorityYear="1997" box="[851,1000,835,860]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC004FF87FCAE2D22FC31FCC5" box="[851,1000,835,860]" italics="true" pageId="3" pageNumber="226">F. quercivora</emphasis>
</taxonomicName>
” (=
<taxonomicName id="2AE9D5CCC004FF87FBE12D22FB48FCC4" authorityName="Hara" authorityYear="2022" box="[1052,1169,835,861]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="togashii">
<emphasis id="DF9D725DC004FF87FBE12D22FB48FCC4" box="[1052,1169,835,861]" italics="true" pageId="3" pageNumber="226">F. togashii</emphasis>
</taxonomicName>
) probably based on fig. 11 by
<bibRefCitation id="8978D3BEC004FF87FF262D06FE5EFC1B" author="Togashi, I." box="[219,391,871,898]" pageId="3" pageNumber="226" pagination="169 - 173" refId="ref22763" refString="Togashi, I. (2006) A new sawfly, Fagineura quercivora (Hymenoptera: Tenthredinidae) feeding on Quercus serrata and Q. mongolica crispula in Honshu, Japan. Proceedings of the entomological Society of Washington, 108 (1), 169 - 173." type="journal article" year="2006">Togashi (2006)</bibRefCitation>
, but fig. 11 is inaccurate (compare it with
<figureCitation id="75D2B2CAC004FF87FCA02D06FC1DFC18" box="[861,964,870,897]" captionStart="FIGURE 11" captionStartId="24.[151,250,1126,1150]" captionTargetBox="[151,1436,181,1094]" captionTargetId="figure-26@24.[151,1436,181,1103]" captionTargetPageId="24" captionText="FIGURE 11. Lancet: Fagineura crenativora (A, B), F. parva (C, holotype)." figureDoi="http://doi.org/10.5281/zenodo.6367291" httpUri="https://zenodo.org/record/6367291/files/figure.png" pageId="3" pageNumber="226">Fig. 11Q</figureCitation>
)]. By modifying the state “bb Malar space 1.02.0 times as long as diameter of front ocellus” in the second half to “bb Malar space
<emphasis id="DF9D725DC004FF87FB762DEAFB77FC3C" box="[1163,1198,907,933]" italics="true" pageId="3" pageNumber="226">0.2</emphasis>
2.0 times as long as diameter of front ocellus” (see Malar space length in Character assessment),
<taxonomicName id="2AE9D5CCC004FF87FC0A2DCEFBB4FC51" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[1015,1133,943,968]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FC0A2DCEFBB4FC51" box="[1015,1133,943,968]" italics="true" pageId="3" pageNumber="226">Fagineura</emphasis>
</taxonomicName>
goes through the modified second half of the couplet 18 to the couplet 22 consisting of
<taxonomicName id="2AE9D5CCC004FF87FCB02DB2FC1AFC75" authorityName="Vikberg &amp; Zinovjev" authorityYear="2000" box="[845,963,979,1004]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FCB02DB2FC1AFC75" box="[845,963,979,1004]" italics="true" pageId="3" pageNumber="226">Fagineura</emphasis>
</taxonomicName>
and
<taxonomicName id="2AE9D5CCC004FF87FC012DB2FB9BFC75" authorityName="Newman" authorityYear="1837" box="[1020,1090,979,1004]" class="Insecta" family="Tenthredinidae" genus="Euura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FC012DB2FB9BFC75" box="[1020,1090,979,1004]" italics="true" pageId="3" pageNumber="226">Euura</emphasis>
</taxonomicName>
. For the reasons mentioned in Character assessment (Setae on katepimeron and Presence or absence of pores in tangium of lancet), the couplet 22 needs to be modified as follows:
<key id="9978F97EC004FF87FF3A2A5AFA45FB7D" pageId="3" pageNumber="226">
<keyStep id="561DB6EAC004FF87FF3A2A5AFA45FB7D" pageId="3" pageNumber="226">
<keyLead id="56180D7AC004FF87FF3A2A5AFA45FB7D" pageId="3" pageNumber="226">
<emphasis id="DF9D725DC004FF87FF3A2A5AFEF2FBC8" bold="true" box="[199,299,1083,1105]" pageId="3" pageNumber="226">22(21) a</emphasis>
Katepimeron extensively covered with hairs,
<emphasis id="DF9D725DC004FF87FD162A5DFBD5FBC8" box="[747,1036,1084,1105]" italics="true" pageId="3" pageNumber="226">sometimes with only few hairs</emphasis>
.
<emphasis id="DF9D725DC004FF87FEE02A38FEF3FBF7" bold="true" box="[285,298,1113,1134]" pageId="3" pageNumber="226">b</emphasis>
Sawsheath slightly emarginate or not emarginate in dorsal view.
<emphasis id="DF9D725DC004FF87FEE02A17FEF1FB12" bold="true" box="[285,296,1142,1163]" pageId="3" pageNumber="226">c</emphasis>
Tangium of lancet with
<emphasis id="DF9D725DC004FF87FDE42A16FDA5FB12" box="[537,636,1142,1164]" italics="true" pageId="3" pageNumber="226">or without</emphasis>
campaniform sensilla............
<taxonomicName id="2AE9D5CCC004FF87FC202A17FA81FB12" authority="Vikberg and Zinovjev, 2000" authorityName="Vikberg and Zinovjev" authorityYear="2000" box="[989,1368,1142,1164]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FC202A17FB9FFB12" bold="true" box="[989,1094,1142,1163]" italics="true" pageId="3" pageNumber="226">Fagineura</emphasis>
Vikberg and Zinovjev,
<quantity id="2A1103AAC004FF87FAD52A17FA81FB12" box="[1320,1368,1142,1163]" metricMagnitude="1" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="226" unit="in" value="2000.0">2000</quantity>
</taxonomicName>
in part
<emphasis id="DF9D725DC004FF87FF3A2AF5FEECFB30" bold="true" box="[199,309,1172,1193]" pageId="3" pageNumber="226">- aa</emphasis>
Katepimeron without hairs,
<emphasis id="DF9D725DC004FF87FDB52AF5FCE3FB30" box="[584,826,1172,1193]" italics="true" pageId="3" pageNumber="226">sometimes with few hairs</emphasis>
*.
<emphasis id="DF9D725DC004FF87FEE02AD0FEE1FB5F" bold="true" box="[285,312,1201,1222]" pageId="3" pageNumber="226">bb</emphasis>
Sawsheath not emarginate in dorsal view.
<emphasis id="DF9D725DC004FF87FEE02AAFFEEBFB7A" bold="true" box="[285,306,1230,1251]" pageId="3" pageNumber="226">cc</emphasis>
Tangium of lancet without campaniform sensilla...............................
<taxonomicName id="2AE9D5CCC004FF87FB7D2AAEFA81FB7A" authority="Newman, 1837" authorityName="Newman" authorityYear="1837" box="[1152,1368,1230,1252]" class="Insecta" family="Tenthredinidae" genus="Euura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FB7D2AAEFB19FB7D" bold="true" box="[1152,1216,1231,1252]" italics="true" pageId="3" pageNumber="226">Euura</emphasis>
Newman,
<quantity id="2A1103AAC004FF87FAD52AAFFA81FB7A" box="[1320,1368,1230,1251]" metricMagnitude="1" metricUnit="m" metricValue="4.66598" pageId="3" pageNumber="226" unit="in" value="1837.0">1837</quantity>
</taxonomicName>
in part
</keyLead>
</keyStep>
</key>
</paragraph>
<paragraph id="ED56AE4FC004FF87FEDD2A93FC63FA91" blockId="3.[151,1437,151,2009]" box="[288,954,1266,1288]" pageId="3" pageNumber="226">
[* This modification is based on our material of gall-inducing
<taxonomicName id="2AE9D5CCC004FF87FC8B2A92FC6BFA91" authorityName="Newman" authorityYear="1837" box="[886,946,1267,1288]" class="Insecta" family="Tenthredinidae" genus="Euura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FC8B2A92FC6BFA91" box="[886,946,1267,1288]" italics="true" pageId="3" pageNumber="226">Euura</emphasis>
</taxonomicName>
]
</paragraph>
<paragraph id="ED56AE4FC004FF87FF3A2B72FDFDFA78" blockId="3.[151,1437,151,2009]" pageId="3" pageNumber="226">
Even with this modified couplet, most of the
<taxonomicName id="2AE9D5CCC004FF87FD2C2B72FC9EFAB5" authorityName="Vikberg and Zinovjev" authorityYear="2000" box="[721,839,1299,1324]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FD2C2B72FC9EFAB5" box="[721,839,1299,1324]" italics="true" pageId="3" pageNumber="226">Fagineura</emphasis>
</taxonomicName>
and
<taxonomicName id="2AE9D5CCC004FF87FC7F2B72FC11FAB5" authorityName="Newman" authorityYear="1837" box="[898,968,1299,1324]" class="Insecta" family="Tenthredinidae" genus="Euura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FC7F2B72FC11FAB5" box="[898,968,1299,1324]" italics="true" pageId="3" pageNumber="226">Euura</emphasis>
</taxonomicName>
coming here will be distinguishable, but the male of
<taxonomicName id="2AE9D5CCC004FF87FEE32B56FEA3FAC9" authorityName="Hara" authorityYear="2022" box="[286,378,1335,1360]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="DF9D725DC004FF87FEE32B56FEA3FAC9" box="[286,378,1335,1360]" italics="true" pageId="3" pageNumber="226">F. parva</emphasis>
</taxonomicName>
and part of the male of
<taxonomicName id="2AE9D5CCC004FF87FD772B56FCF1FAC9" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[650,808,1335,1360]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC004FF87FD772B56FCF1FAC9" box="[650,808,1335,1360]" italics="true" pageId="3" pageNumber="226">F. crenativora</emphasis>
</taxonomicName>
are not separable from the males of some gall-inducing
<taxonomicName id="2AE9D5CCC004FF87FF6A2B3AFF04FAED" authorityName="Newman" authorityYear="1837" box="[151,221,1371,1396]" class="Insecta" family="Tenthredinidae" genus="Euura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FF6A2B3AFF04FAED" box="[151,221,1371,1396]" italics="true" pageId="3" pageNumber="226">Euura</emphasis>
</taxonomicName>
. Their males are separated as follows: the tergal hollow of abdominal tergum 8 is well defined in the former (
<figureCitation id="75D2B2CAC004FF87FF622B1EFEF5FA00" box="[159,300,1407,1433]" captionStart="FIGURE 14" captionStartId="27.[151,250,1869,1893]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-19@27.[151,1436,181,1845]" captionTargetPageId="27" captionText="FIGURE 14. Apex of male abdomen and genitalia: Fagineura crenativora (A, DF, K, L), F. parva (B, G, H, M, N) and F. togashii (C, I, J, O, P). AC, Apex of abdomen, dorsal view; D, G, I, genitalia, dorsal view; E, F, H, J, genitalia, ventral view (E, H, J, penis valve removed); KP, penis valve, left dorsal." figureDoi="http://doi.org/10.5281/zenodo.6367297" httpUri="https://zenodo.org/record/6367297/files/figure.png" pageId="3" pageNumber="226">Figs 14AC</figureCitation>
) but not clearly defined in the latter; a paraantennal field is mostly covered with setae in the former (
<figureCitation id="75D2B2CAC004FF87FF622BC2FECAFA24" box="[159,275,1443,1469]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="3" pageNumber="226">Figs 6P, Q</figureCitation>
) but often widely glabrous in the latter; a median groove is present above a frontal pit in the former (
<figureCitation id="75D2B2CAC004FF87FA962BC2FF0CFA78" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="3" pageNumber="226">Figs 6P, Q</figureCitation>
) but often absent in the latter.
</paragraph>
<paragraph id="ED56AE4FC004FF87FF3A2B8AFEFDF8D0" blockId="3.[151,1437,151,2009]" pageId="3" pageNumber="226">
In the key by
<bibRefCitation id="8978D3BEC004FF87FE912B8AFD91F99C" author="Prous, M. &amp; Blank, S. M. &amp; Goulet, H. &amp; Heibo, E. &amp; Liston, A. &amp; Malm, T. &amp; Nyman, T. &amp; Schmidt, S. &amp; Smith, D. R. &amp; Vardal, H. &amp; Viitasaari, M. &amp; Vikberg, V. &amp; Taeger, A." box="[364,584,1515,1541]" pageId="3" pageNumber="226" pagination="1 - 69" refId="ref21989" refString="Prous, M., Blank, S. M., Goulet, H., Heibo, E., Liston, A., Malm, T., Nyman, T., Schmidt, S., Smith, D. R., Vardal, H., Viitasaari, M., Vikberg, V. &amp; Taeger, A. (2014) The genera of Nematinae (Hymenoptera, Tenthredinidae). Journal of Hymenoptera Research, 40, 1 - 69. https: // doi. org / 10.3897 / JHR. 40.7442" type="journal article" year="2014">
Prous
<emphasis id="DF9D725DC004FF87FE4B2B8AFE28F99C" box="[438,497,1515,1541]" italics="true" pageId="3" pageNumber="226">et al.</emphasis>
(2014)
</bibRefCitation>
, some
<taxonomicName id="2AE9D5CCC004FF87FD662B8AFCEAF99D" authorityName="Togashi" authorityYear="1997" box="[667,819,1515,1540]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC004FF87FD662B8AFCEAF99D" box="[667,819,1515,1540]" italics="true" pageId="3" pageNumber="226">F. quercivora</emphasis>
</taxonomicName>
with crossvein 2r-rs run through the couplet 5 to the couplet 26 but their mandibles do not fit both the states d and dd in the couplet 26. The state dd needs the following modification: Left and right mandible both tapered regularly towards apex
<emphasis id="DF9D725DC004FF87FBB22852FF33F9E8" italics="true" pageId="3" pageNumber="226">or markedly constricted near middle,</emphasis>
or left mandible markedly constricted near middle and right mandible tapered regularly towards apex. The same modification is needed for the state aa of the couplet 28. With these modifications, some
<taxonomicName id="2AE9D5CCC004FF87FB33281AFABDF90D" authorityName="Togashi" authorityYear="1997" box="[1230,1380,1659,1684]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC004FF87FB33281AFABDF90D" box="[1230,1380,1659,1684]" italics="true" pageId="3" pageNumber="226">F. quercivora</emphasis>
</taxonomicName>
with crossvein 2r-rs run through the modified second half of the couplet 26 and the modified second half of the couplet 28 to the couplet 32 consisting of
<taxonomicName id="2AE9D5CCC004FF87FDED28A2FCFBF944" authority="Rohwer, 1916" authorityName="Rohwer" authorityYear="1916" box="[528,802,1731,1757]" class="Insecta" family="Tenthredinidae" genus="Moricella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FDED28A2FDA7F944" box="[528,638,1731,1757]" italics="true" pageId="3" pageNumber="226">Moricella</emphasis>
Rohwer, 1916
</taxonomicName>
and
<taxonomicName id="2AE9D5CCC004FF87FCA528A2FBB5F944" authority="Dahlbom, 1835" authorityName="Dahlbom" authorityYear="1835" box="[856,1132,1730,1757]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FCA528A2FC6CF945" box="[856,949,1731,1756]" italics="true" pageId="3" pageNumber="226">Dineura</emphasis>
Dahlbom, 1835
</taxonomicName>
.
<taxonomicName id="2AE9D5CCC004FF87FB8A28A2FAB4F945" authorityName="Togashi" authorityYear="2006" box="[1143,1389,1731,1756]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC004FF87FB8A28A2FAB4F945" box="[1143,1389,1731,1756]" italics="true" pageId="3" pageNumber="226">Fagineura quercivora</emphasis>
</taxonomicName>
differs from those genera by the anterior fore tibial spur with a velum (without a velum in those two genera) and the left and right mandible both markedly constricted near the middle (both tapered regularly towards the apex in those two genera).
</paragraph>
<paragraph id="ED56AE4FC004FF80FF3A2932FC40FD80" blockId="3.[151,1437,151,2009]" lastBlockId="4.[151,1436,151,537]" lastPageId="4" lastPageNumber="227" pageId="3" pageNumber="226">
We divide
<taxonomicName id="2AE9D5CCC004FF87FEC22932FE6CF8F5" authorityName="Vikberg and Zinovjev" authorityYear="2000" box="[319,437,1875,1900]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC004FF87FEC22932FE6CF8F5" box="[319,437,1875,1900]" italics="true" pageId="3" pageNumber="226">Fagineura</emphasis>
</taxonomicName>
into two species groups, the
<taxonomicName id="2AE9D5CCC004FF87FD072932FC4EF8F5" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[762,919,1875,1900]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC004FF87FD072932FC4EF8F5" box="[762,919,1875,1900]" italics="true" pageId="3" pageNumber="226">F. crenativora</emphasis>
</taxonomicName>
group and the
<taxonomicName id="2AE9D5CCC004FF87FBC32932FB0AF8F5" authorityName="Togashi" authorityYear="2006" box="[1086,1235,1875,1900]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC004FF87FBC32932FB0AF8F5" box="[1086,1235,1875,1900]" italics="true" pageId="3" pageNumber="226">F. quercivora</emphasis>
</taxonomicName>
group. These two groups have different mandibles, frontal pits, female cerci, lances, lancets and penis valves as stated in the couplet 1 of the key below. As far as we know about the ecology of
<taxonomicName id="2AE9D5CCC004FF87FCEF29FAFC3BF82C" box="[786,994,1947,1973]" pageId="3" pageNumber="226">
<emphasis id="DF9D725DC004FF87FCEF29FAFC51F82D" box="[786,904,1947,1972]" italics="true" pageId="3" pageNumber="226">Fagineura</emphasis>
species
</taxonomicName>
, the larva of
<taxonomicName id="2AE9D5CCC004FF87FB9229FAFAD2F82D" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[1135,1291,1947,1972]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC004FF87FB9229FAFAD2F82D" box="[1135,1291,1947,1972]" italics="true" pageId="3" pageNumber="226">F. crenativora</emphasis>
</taxonomicName>
of the
<taxonomicName id="2AE9D5CCC004FF87FAAB29FAFF2FF841" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC004FF87FAAB29FAFF2FF841" italics="true" pageId="3" pageNumber="226">F. crenativora</emphasis>
</taxonomicName>
group feeds on
<taxonomicName id="2AE9D5CCC004FF87FE5729DEFE29F841" box="[426,496,1983,2008]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="226" phylum="Tracheophyta" rank="genus">
<emphasis id="DF9D725DC004FF87FE5729DEFE29F841" box="[426,496,1983,2008]" italics="true" pageId="3" pageNumber="226">Fagus</emphasis>
</taxonomicName>
and has an extra molt, but the larvae of all members of the
<taxonomicName id="2AE9D5CCC004FF87FB7529DEFAC4F841" authorityName="Togashi" authorityYear="2006" box="[1160,1309,1983,2008]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC004FF87FB7529DEFAC4F841" box="[1160,1309,1983,2008]" italics="true" pageId="3" pageNumber="226">F. quercivora</emphasis>
</taxonomicName>
group feed on
<taxonomicName id="2AE9D5CCC003FF80FF462EF6FEC0FF28" box="[187,281,151,177]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="227" phylum="Tracheophyta" rank="genus">
<emphasis id="DF9D725DC003FF80FF462EF6FEC0FF28" box="[187,281,151,177]" italics="true" pageId="4" pageNumber="227">Quercus</emphasis>
</taxonomicName>
and the larva of
<taxonomicName id="2AE9D5CCC003FF80FE2A2EF6FD94FF28" authorityName="Hara" authorityYear="2022" box="[471,589,151,177]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="togashii">
<emphasis id="DF9D725DC003FF80FE2A2EF6FD94FF28" box="[471,589,151,177]" italics="true" pageId="4" pageNumber="227">F. togashii</emphasis>
</taxonomicName>
of the
<taxonomicName id="2AE9D5CCC003FF80FD602EF6FCEAFF29" authorityName="Togashi" authorityYear="2006" box="[669,819,151,176]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC003FF80FD602EF6FCEAFF29" box="[669,819,151,176]" italics="true" pageId="4" pageNumber="227">F. quercivora</emphasis>
</taxonomicName>
group does not have an extra molt. We have not found any apomorphic character that suggests a close relationship between the two groups. The markedly constricted right mandible of the
<taxonomicName id="2AE9D5CCC003FF80FEAE2EBEFE33FF61" authorityName="Togashi" authorityYear="2006" box="[339,490,223,248]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC003FF80FEAE2EBEFE33FF61" box="[339,490,223,248]" italics="true" pageId="4" pageNumber="227">F. quercivora</emphasis>
</taxonomicName>
group (
<figureCitation id="75D2B2CAC003FF80FDB92EBEFD18FF60" box="[580,705,223,249]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="4" pageNumber="227">Figs 7LO</figureCitation>
) appears to be an unusual feature in the
<taxonomicName id="2AE9D5CCC003FF80FB692EBEFAC9FF60" authorityName="Thomson" authorityYear="1871" box="[1172,1296,223,249]" class="Insecta" family="Tenthredinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="subFamily" subFamily="Nematinae">Nematinae</taxonomicName>
, though this character is also found in some
<taxonomicName id="2AE9D5CCC003FF80FDFE2F62FD0CFE84" box="[515,725,259,285]" pageId="4" pageNumber="227">
<emphasis id="DF9D725DC003FF80FDFE2F62FDA2FE85" box="[515,635,259,284]" italics="true" pageId="4" pageNumber="227">Nematinus</emphasis>
species
</taxonomicName>
(fig. 2I in
<bibRefCitation id="8978D3BEC003FF80FCA92F62FC09FE84" author="Hara, H." box="[852,976,259,285]" pageId="4" pageNumber="227" pagination="61 - 66" refId="ref21517" refString="Hara, H. (2017) A new sawfly, Nematus viburnum sp. nov., related to N. lucidus (Hymenoptera, Tenthredinidae). Japanese Journal of Systematic Entomology, 23, 61 - 66." type="journal article" year="2017">Hara 2017</bibRefCitation>
). This fact may suggest that the
<taxonomicName id="2AE9D5CCC003FF80FABE2F62FF06FED9" authorityName="Togashi" authorityYear="2006" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC003FF80FABE2F62FF06FED9" italics="true" pageId="4" pageNumber="227">F. quercivora</emphasis>
</taxonomicName>
group is closely related to
<taxonomicName id="2AE9D5CCC003FF80FDF62F46FD5AFED9" authorityName="Rohwer" authorityYear="1911" box="[523,643,295,320]" class="Insecta" family="Tenthredinidae" genus="Nematinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC003FF80FDF62F46FD5AFED9" box="[523,643,295,320]" italics="true" pageId="4" pageNumber="227">Nematinus</emphasis>
</taxonomicName>
. On the other hand, the
<taxonomicName id="2AE9D5CCC003FF80FC722F46FBFDFED9" authorityName="Togashi" authorityYear="2006" box="[911,1060,295,320]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC003FF80FC722F46FBFDFED9" box="[911,1060,295,320]" italics="true" pageId="4" pageNumber="227">F. quercivora</emphasis>
</taxonomicName>
group has a markedly constricted left mandible (
<figureCitation id="75D2B2CAC003FF80FEC32F2AFE60FEFC" box="[318,441,331,357]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="4" pageNumber="227">Figs 7FH</figureCitation>
) like
<taxonomicName id="2AE9D5CCC003FF80FE012F2AFD9BFEFD" authorityName="Newman" authorityYear="1837" box="[508,578,331,356]" class="Insecta" family="Tenthredinidae" genus="Euura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC003FF80FE012F2AFD9BFEFD" box="[508,578,331,356]" italics="true" pageId="4" pageNumber="227">Euura</emphasis>
</taxonomicName>
,
<taxonomicName id="2AE9D5CCC003FF80FDAF2F2AFD6DFEFD" box="[594,692,331,356]" class="Insecta" family="Tenthredinidae" genus="Nematus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC003FF80FDAF2F2AFD6DFEFD" box="[594,692,331,356]" italics="true" pageId="4" pageNumber="227">Nematus</emphasis>
</taxonomicName>
,
<taxonomicName id="2AE9D5CCC003FF80FD3E2F2AFC9DFEFC" box="[707,836,331,357]" class="Insecta" family="Tenthredinidae" genus="Pristiphora" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC003FF80FD3E2F2AFC9DFEFC" box="[707,836,331,357]" italics="true" pageId="4" pageNumber="227">Pristiphora</emphasis>
</taxonomicName>
and their related genera, and the larva of
<taxonomicName id="2AE9D5CCC003FF80FAD82F2AFA45FEFC" authorityName="Hara" authorityYear="2022" box="[1317,1436,331,357]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="togashii">
<emphasis id="DF9D725DC003FF80FAD82F2AFA45FEFC" box="[1317,1436,331,357]" italics="true" pageId="4" pageNumber="227">F. togashii</emphasis>
</taxonomicName>
of the
<taxonomicName id="2AE9D5CCC003FF80FF1D2F0EFEAFFE11" authorityName="Togashi" authorityYear="2006" box="[224,374,367,392]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC003FF80FF1D2F0EFEAFFE11" box="[224,374,367,392]" italics="true" pageId="4" pageNumber="227">F. quercivora</emphasis>
</taxonomicName>
group goes to the couplet 30 leading to part of
<taxonomicName id="2AE9D5CCC003FF80FC722F0EFC0CFE11" authorityName="Newman" authorityYear="1837" box="[911,981,367,392]" class="Insecta" family="Tenthredinidae" genus="Euura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC003FF80FC722F0EFC0CFE11" box="[911,981,367,392]" italics="true" pageId="4" pageNumber="227">Euura</emphasis>
</taxonomicName>
in the larval key of the West Palaearctic genera by
<bibRefCitation id="8978D3BEC003FF80FEED2FF2FE30FE34" author="Prous, M. &amp; Liston, A. &amp; Kramp, K. &amp; Savina, H. &amp; Vardal, H. &amp; Taeger, A." box="[272,489,403,429]" pageId="4" pageNumber="227" pagination="63 - 127" refId="ref22149" refString="Prous, M., Liston, A., Kramp, K., Savina, H., Vardal, H. &amp; Taeger, A. (2019) The West Palaearctic genera of Nematinae (Hymenoptera, Tenthredinidae). ZooKeys, 875, 63 - 127. https: // doi. org / 10.3897 / zookeys. 875.35748" type="journal article" year="2019">
Prous
<emphasis id="DF9D725DC003FF80FEA52FF2FE4BFE34" box="[344,402,403,429]" italics="true" pageId="4" pageNumber="227">et al.</emphasis>
(2019)
</bibRefCitation>
. Genetic study will be needed to clarify the relationship between the
<taxonomicName id="2AE9D5CCC003FF80FAF82FF2FA45FE35" authorityName="Togashi" authorityYear="2006" box="[1285,1436,403,428]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC003FF80FAF82FF2FA45FE35" box="[1285,1436,403,428]" italics="true" pageId="4" pageNumber="227">F. quercivora</emphasis>
</taxonomicName>
group and other nematines. We believe that it is best to place the
<taxonomicName id="2AE9D5CCC003FF80FC892FD6FBD3FE49" authorityName="Togashi" authorityYear="2006" box="[884,1034,439,464]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC003FF80FC892FD6FBD3FE49" box="[884,1034,439,464]" italics="true" pageId="4" pageNumber="227">F. quercivora</emphasis>
</taxonomicName>
group in
<taxonomicName id="2AE9D5CCC003FF80FB8A2FD6FB34FE49" authorityName="Vikberg and Zinovjev" authorityYear="2000" box="[1143,1261,439,464]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC003FF80FB8A2FD6FB34FE49" box="[1143,1261,439,464]" italics="true" pageId="4" pageNumber="227">Fagineura</emphasis>
</taxonomicName>
at the moment, because the group is more similar to the
<taxonomicName id="2AE9D5CCC003FF80FDA12FBAFD23FE6D" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[604,762,475,500]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC003FF80FDA12FBAFD23FE6D" box="[604,762,475,500]" italics="true" pageId="4" pageNumber="227">F. crenativora</emphasis>
</taxonomicName>
group than any other nematines and the genus classification of the
<taxonomicName id="2AE9D5CCC003FF80FF222F9EFE81FD80" authorityName="Thomson" authorityYear="1871" box="[223,344,511,537]" class="Insecta" family="Tenthredinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="subFamily" subFamily="Nematinae">Nematinae</taxonomicName>
is still not stable, especially for East Asian species.
</paragraph>
</subSubSection>
<subSection id="8466B7A4C003FF80FF6A2C0BFC3BFB38" pageId="4" pageNumber="227" type="multiple">
<subSubSection id="A5F3FDC4C003FF80FF6A2C0BFD06FD1C" box="[151,735,618,645]" pageId="4" pageNumber="227" type="multiple">
<paragraph id="ED56AE4FC003FF80FF6A2C0BFD06FD1C" blockId="4.[151,735,618,645]" box="[151,735,618,645]" pageId="4" pageNumber="227">
<heading id="B61E1923C003FF80FF6A2C0BFD06FD1C" bold="true" box="[151,735,618,645]" fontSize="11" level="1" pageId="4" pageNumber="227" reason="1">
<emphasis id="DF9D725DC003FF80FF6A2C0BFD06FD1C" bold="true" box="[151,735,618,645]" pageId="4" pageNumber="227">
List of species groups and species of
<taxonomicName id="2AE9D5CCC003FF80FDA32C0AFD06FD1C" authorityName="Vikberg and Zinovjev" authorityYear="2000" box="[606,735,619,645]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC003FF80FDA32C0AFD06FD1C" bold="true" box="[606,735,619,645]" italics="true" pageId="4" pageNumber="227">Fagineura</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A5F3FDC4C003FF80FF6A2CD2FC3BFB38" pageId="4" pageNumber="227" type="synonymic_list">
<paragraph id="ED56AE4FC003FF80FF6A2CD2FE5CFD54" blockId="4.[151,1443,691,1186]" box="[151,389,691,717]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF6A2CD2FEECFD55" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[151,309,691,716]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC003FF80FF6A2CD2FEECFD55" box="[151,309,691,716]" italics="true" pageId="4" pageNumber="227">F. crenativora</emphasis>
</taxonomicName>
group:
</paragraph>
<paragraph id="ED56AE4FC003FF80FF3A2CB6FCDBFD6B" blockId="4.[151,1443,691,1186]" box="[199,770,727,754]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF3A2CB6FD96FD68" authority="Liu, Li &amp; Wei, 2021" authorityName="Liu, Li &amp; Wei" authorityYear="2021" box="[199,591,727,753]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="DF9D725DC003FF80FF3A2CB6FEBBFD68" box="[199,354,727,753]" italics="true" pageId="4" pageNumber="227">F. brevicornis</emphasis>
<bibRefCitation id="8978D3BEC003FF80FE952CB6FD96FD68" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[360,591,727,753]" pageId="4" pageNumber="227" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">Liu, Li &amp; Wei, 2021</bibRefCitation>
</taxonomicName>
.
<collectingCountry id="95FEEEDFC003FF80FDA72CB6FD44FD68" box="[602,669,727,753]" name="China" pageId="4" pageNumber="227">China</collectingCountry>
(
<collectingRegion id="2F2D60ADC003FF80FD512CB6FD2EFD6B" box="[684,759,727,754]" country="China" name="Hubei" pageId="4" pageNumber="227">Hubei</collectingRegion>
).
</paragraph>
<paragraph id="ED56AE4FC003FF80FF3A2C9AFB7DFC8F" blockId="4.[151,1443,691,1186]" box="[199,1188,763,790]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF3A2C9AFD4CFC8C" authority="Vikberg &amp; Zinovjev, 2000" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[199,661,763,789]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC003FF80FF3A2C9AFEBCFC8D" box="[199,357,763,788]" italics="true" pageId="4" pageNumber="227">F. crenativora</emphasis>
Vikberg &amp; Zinovjev, 2000
</taxonomicName>
.
<collectingCountry id="95FEEEDFC003FF80FD622C9AFD06FC8C" box="[671,735,763,789]" name="Japan" pageId="4" pageNumber="227">Japan</collectingCountry>
(
<collectingRegion id="2F2D60ADC003FF80FD132C9AFCB8FC8C" box="[750,865,763,789]" country="Japan" name="Hokkaido" pageId="4" pageNumber="227">Hokkaido</collectingRegion>
, Honshu, Shikoku, Kyushu).
</paragraph>
<paragraph id="ED56AE4FC003FF80FF3A2D7EFCCDFCA3" blockId="4.[151,1443,691,1186]" box="[199,788,799,826]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF3A2D7EFDBBFCA0" authority="Liu, Li &amp; Wei, 2019" authorityName="Liu, Li &amp; Wei" authorityYear="2019" box="[199,610,799,825]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="flactoserrula">
<emphasis id="DF9D725DC003FF80FF3A2D7EFEADFCA0" box="[199,372,799,825]" italics="true" pageId="4" pageNumber="227">F. flactoserrula</emphasis>
<bibRefCitation id="8978D3BEC003FF80FE872D7EFDBBFCA0" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[378,610,799,825]" pageId="4" pageNumber="227" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">Liu, Li &amp; Wei, 2019</bibRefCitation>
</taxonomicName>
.
<collectingCountry id="95FEEEDFC003FF80FD912D7EFD76FCA0" box="[620,687,799,825]" name="China" pageId="4" pageNumber="227">China</collectingCountry>
(
<collectingRegion id="2F2D60ADC003FF80FD432D7EFCD0FCA3" box="[702,777,799,826]" country="China" name="Hubei" pageId="4" pageNumber="227">Hubei</collectingRegion>
).
</paragraph>
<paragraph id="ED56AE4FC003FF80FF3A2D22FCD2FCC4" blockId="4.[151,1443,691,1186]" box="[199,779,835,861]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF3A2D22FD88FCC4" authority="Liu, Li &amp; Wei, 2021" authorityName="Liu, Li &amp; Wei" authorityYear="2021" box="[199,593,835,861]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="longitangia">
<emphasis id="DF9D725DC003FF80FF3A2D22FEBDFCC4" box="[199,356,835,861]" italics="true" pageId="4" pageNumber="227">F. longitangia</emphasis>
<bibRefCitation id="8978D3BEC003FF80FE962D22FD88FCC4" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[363,593,835,861]" pageId="4" pageNumber="227" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">Liu, Li &amp; Wei, 2021</bibRefCitation>
</taxonomicName>
.
<collectingCountry id="95FEEEDFC003FF80FDA12D22FD46FCC4" box="[604,671,835,861]" name="China" pageId="4" pageNumber="227">China</collectingCountry>
(
<collectingRegion id="2F2D60ADC003FF80FD532D22FD26FCC4" box="[686,767,835,861]" country="China" name="Hunan" pageId="4" pageNumber="227">Hunan</collectingRegion>
).
</paragraph>
<paragraph id="ED56AE4FC003FF80FF3A2D06FD29FC1B" blockId="4.[151,1443,691,1186]" box="[199,752,871,898]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF3A2D06FEBDFC18" authority="Hara" authorityName="Hara" authorityYear="2022" box="[199,356,871,897]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
<emphasis id="DF9D725DC003FF80FF3A2D06FEFDFC19" box="[199,292,871,896]" italics="true" pageId="4" pageNumber="227">F. parva</emphasis>
Hara
</taxonomicName>
,
<taxonomicNameLabel id="C4AECF26C003FF80FE922D06FE1FFC18" box="[367,454,871,897]" pageId="4" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
<collectingCountry id="95FEEEDFC003FF80FE312D06FDD5FC18" box="[460,524,871,897]" name="Japan" pageId="4" pageNumber="227">Japan</collectingCountry>
(Honshu, Shikoku).
</paragraph>
<paragraph id="ED56AE4FC003FF80FF3A2DEAFCBDFC3C" blockId="4.[151,1443,691,1186]" box="[199,868,907,933]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF3A2DEAFD81FC3C" authority="Liu, Li &amp; Wei, 2019" authorityName="Liu, Li &amp; Wei" authorityYear="2019" box="[199,600,907,933]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="xanthosoma">
<emphasis id="DF9D725DC003FF80FF3A2DEAFEB3FC3C" box="[199,362,907,933]" italics="true" pageId="4" pageNumber="227">F. xanthosoma</emphasis>
<bibRefCitation id="8978D3BEC003FF80FE8C2DEAFD81FC3C" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[369,600,907,933]" pageId="4" pageNumber="227" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">Liu, Li &amp; Wei, 2019</bibRefCitation>
</taxonomicName>
.
<collectingCountry id="95FEEEDFC003FF80FD9F2DEAFD7CFC3C" box="[610,677,907,933]" name="China" pageId="4" pageNumber="227">China</collectingCountry>
(
<collectingRegion id="2F2D60ADC003FF80FD492DEAFD24FC3C" box="[692,765,907,933]" country="China" name="Hubei" pageId="4" pageNumber="227">Hubei</collectingRegion>
,
<collectingRegion id="2F2D60ADC003FF80FCF52DEAFC81FC3C" box="[776,856,907,933]" country="China" name="Hunan" pageId="4" pageNumber="227">Hunan</collectingRegion>
).
</paragraph>
<paragraph id="ED56AE4FC003FF80FF6A2DCEFEA4FC50" blockId="4.[151,1443,691,1186]" box="[151,381,943,969]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF6A2DCEFEF4FC51" authorityName="Togashi" authorityYear="2006" box="[151,301,943,968]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC003FF80FF6A2DCEFEF4FC51" box="[151,301,943,968]" italics="true" pageId="4" pageNumber="227">F. quercivora</emphasis>
</taxonomicName>
group:
</paragraph>
<paragraph id="ED56AE4FC003FF80FF3A2DB2FC87FC74" blockId="4.[151,1443,691,1186]" box="[199,862,979,1006]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF3A2DB2FE1AFC74" authority="Hara" authorityName="Hara" authorityYear="2022" box="[199,451,979,1005]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="flavomaculata" status="sp. nov.">
<emphasis id="DF9D725DC003FF80FF3A2DB2FE5AFC74" box="[199,387,979,1005]" italics="true" pageId="4" pageNumber="227">F. flavomaculata</emphasis>
Hara
</taxonomicName>
,
<taxonomicNameLabel id="C4AECF26C003FF80FE332DB2FDFCFC74" box="[462,549,979,1005]" pageId="4" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
<collectingCountry id="95FEEEDFC003FF80FDD62DB2FDB2FC74" box="[555,619,979,1005]" name="Japan" pageId="4" pageNumber="227">Japan</collectingCountry>
(Honshu) and
<collectingCountry id="95FEEEDFC003FF80FCEF2DB2FC83FC74" box="[786,858,979,1005]" name="South Korea" pageId="4" pageNumber="227">Korea</collectingCountry>
.
</paragraph>
<paragraph id="ED56AE4FC003FF80FF3A2D96FD67FB8B" blockId="4.[151,1443,691,1186]" box="[199,702,1015,1042]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF3A2D96FE47FB88" authority="Hara" authorityName="Hara" authorityYear="2022" box="[199,414,1015,1041]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="fulvistriata" status="sp. nov.">
<emphasis id="DF9D725DC003FF80FF3A2D96FE87FB88" box="[199,350,1015,1041]" italics="true" pageId="4" pageNumber="227">F. fulvistriata</emphasis>
Hara
</taxonomicName>
,
<taxonomicNameLabel id="C4AECF26C003FF80FE542D96FDD9FB88" box="[425,512,1015,1041]" pageId="4" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
<collectingCountry id="95FEEEDFC003FF80FDFB2D96FD9FFB88" box="[518,582,1015,1041]" name="Japan" pageId="4" pageNumber="227">Japan</collectingCountry>
(Honshu).
</paragraph>
<paragraph id="ED56AE4FC003FF80FF3A2A7AFCC3FBAC" blockId="4.[151,1443,691,1186]" box="[199,794,1051,1078]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF3A2A7AFEA7FBAC" authority="Hara" authorityName="Hara" authorityYear="2022" box="[199,382,1051,1077]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="glabella" status="sp. nov.">
<emphasis id="DF9D725DC003FF80FF3A2A7AFEE6FBAC" box="[199,319,1051,1077]" italics="true" pageId="4" pageNumber="227">F. glabella</emphasis>
Hara
</taxonomicName>
,
<taxonomicNameLabel id="C4AECF26C003FF80FE772A7AFE38FBAC" box="[394,481,1051,1077]" pageId="4" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
<collectingCountry id="95FEEEDFC003FF80FE1A2A7AFDFEFBAC" box="[487,551,1051,1077]" name="Japan" pageId="4" pageNumber="227">Japan</collectingCountry>
(Honshu) and
<collectingCountry id="95FEEEDFC003FF80FD332A7AFCCFFBAC" box="[718,790,1051,1077]" name="South Korea" pageId="4" pageNumber="227">Korea</collectingCountry>
.
</paragraph>
<paragraph id="ED56AE4FC003FF80FF3A2A5EFA4BFBC0" blockId="4.[151,1443,691,1186]" box="[199,1426,1086,1114]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF3A2A5EFDCEFBC0" authority="(Togashi, 1997)" authorityName="Hara &amp; Ibuki" authorityYear="2022" baseAuthorityName="Togashi" baseAuthorityYear="1997" box="[199,535,1087,1113]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora" status="comb. nov.">
<emphasis id="DF9D725DC003FF80FF3A2A5EFE84FBC1" box="[199,349,1087,1112]" italics="true" pageId="4" pageNumber="227">F. quercivora</emphasis>
(
<bibRefCitation id="8978D3BEC003FF80FE962A5EFDD6FBC0" author="Togashi, I." box="[363,527,1087,1113]" pageId="4" pageNumber="227" pagination="195 - 198" refId="ref22718" refString="Togashi, I. (1997) A new species of Dineura Dahlbom (Hymenoptera: Tenthredinidae) feeding on Quercus dentata Thunberg (Dicotyledoneae: Fagaceae) from Japan. Japanese Journal of Entomology, 65 (1), 195 - 198." type="journal article" year="1997">Togashi, 1997</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="C4AECF26C003FF80FDDF2A5EFD47FBC0" box="[546,670,1087,1113]" pageId="4" pageNumber="227" rank="species">comb. nov.</taxonomicNameLabel>
[=
<taxonomicName id="2AE9D5CCC003FF80FD382A5EFB97FBC0" authority="Togashi, 1997" authorityName="Togashi" authorityYear="1997" box="[709,1102,1086,1113]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC003FF80FD382A5EFC7AFBC1" box="[709,931,1087,1112]" italics="true" pageId="4" pageNumber="227">Dineura quercivora</emphasis>
<bibRefCitation id="8978D3BEC003FF80FC572A5EFB97FBC0" author="Togashi, I." box="[938,1102,1086,1113]" pageId="4" pageNumber="227" pagination="195 - 198" refId="ref22718" refString="Togashi, I. (1997) A new species of Dineura Dahlbom (Hymenoptera: Tenthredinidae) feeding on Quercus dentata Thunberg (Dicotyledoneae: Fagaceae) from Japan. Japanese Journal of Entomology, 65 (1), 195 - 198." type="journal article" year="1997">Togashi, 1997</bibRefCitation>
</taxonomicName>
].
<collectingCountry id="95FEEEDFC003FF80FBA22A5EFB46FBC0" box="[1119,1183,1087,1113]" name="Japan" pageId="4" pageNumber="227">Japan</collectingCountry>
(Honshu) and
<collectingCountry id="95FEEEDFC003FF80FABB2A5EFA57FBC0" box="[1350,1422,1087,1113]" name="South Korea" pageId="4" pageNumber="227">Korea</collectingCountry>
.
</paragraph>
<paragraph id="ED56AE4FC003FF80FF3A2A02FC3BFB38" blockId="4.[151,1443,691,1186]" pageId="4" pageNumber="227">
<taxonomicName id="2AE9D5CCC003FF80FF3A2A02FEA4FBE4" authority="Hara" authorityName="Hara" authorityYear="2022" box="[199,381,1123,1149]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="togashii" status="nom. nov.">
<emphasis id="DF9D725DC003FF80FF3A2A02FEE4FBE4" box="[199,317,1123,1149]" italics="true" pageId="4" pageNumber="227">F. togashii</emphasis>
Hara
</taxonomicName>
,
<taxonomicNameLabel id="C4AECF26C003FF80FE752A02FE2EFBE4" box="[392,503,1123,1149]" pageId="4" pageNumber="227">nom. nov.</taxonomicNameLabel>
[replacement name for
<taxonomicName id="2AE9D5CCC003FF80FCFF2A02FB99FBE4" authority="Togashi, 2006" authorityName="Togashi" authorityYear="2006" box="[770,1088,1123,1149]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC003FF80FCFF2A02FC4EFBE5" box="[770,919,1123,1148]" italics="true" pageId="4" pageNumber="227">F. quercivora</emphasis>
<bibRefCitation id="8978D3BEC003FF80FC602A02FB99FBE4" author="Togashi, I." box="[925,1088,1123,1149]" pageId="4" pageNumber="227" pagination="169 - 173" refId="ref22763" refString="Togashi, I. (2006) A new sawfly, Fagineura quercivora (Hymenoptera: Tenthredinidae) feeding on Quercus serrata and Q. mongolica crispula in Honshu, Japan. Proceedings of the entomological Society of Washington, 108 (1), 169 - 173." type="journal article" year="2006">Togashi, 2006</bibRefCitation>
</taxonomicName>
; junior secondary homonym of
<taxonomicName id="2AE9D5CCC003FF80FF182AE6FDE1FB38" authority="(Togashi, 1997)" baseAuthorityName="Togashi" baseAuthorityYear="1997" box="[229,568,1158,1185]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC003FF80FF182AE6FEA2FB39" box="[229,379,1159,1184]" italics="true" pageId="4" pageNumber="227">F. quercivora</emphasis>
(
<bibRefCitation id="8978D3BEC003FF80FE742AE6FDF6FB38" author="Togashi, I." box="[393,559,1158,1185]" pageId="4" pageNumber="227" pagination="195 - 198" refId="ref22718" refString="Togashi, I. (1997) A new species of Dineura Dahlbom (Hymenoptera: Tenthredinidae) feeding on Quercus dentata Thunberg (Dicotyledoneae: Fagaceae) from Japan. Japanese Journal of Entomology, 65 (1), 195 - 198." type="journal article" year="1997">Togashi, 1997</bibRefCitation>
)
</taxonomicName>
].
<collectingCountry id="95FEEEDFC003FF80FDB72AE6FD53FB38" box="[586,650,1159,1185]" name="Japan" pageId="4" pageNumber="227">Japan</collectingCountry>
(Honshu, Kyushu) and
<collectingCountry id="95FEEEDFC003FF80FC6B2AE6FC07FB38" box="[918,990,1159,1185]" name="South Korea" pageId="4" pageNumber="227">Korea</collectingCountry>
.
</paragraph>
</subSubSection>
</subSection>
<subSection id="8466B7A4C003FF81FF6A2A92FA45FB54" lastPageId="5" lastPageNumber="228" pageId="4" pageNumber="227" type="key">
<subSubSection id="A5F3FDC4C003FF80FF6A2A92FD38FA94" box="[151,737,1266,1293]" pageId="4" pageNumber="227" type="multiple">
<paragraph id="ED56AE4FC003FF80FF6A2A92FD38FA94" blockId="4.[151,737,1266,1293]" box="[151,737,1266,1293]" pageId="4" pageNumber="227">
<heading id="B61E1923C003FF80FF6A2A92FD38FA94" bold="true" box="[151,737,1266,1293]" fontSize="11" level="1" pageId="4" pageNumber="227" reason="1">
<emphasis id="DF9D725DC003FF80FF6A2A92FD38FA94" bold="true" box="[151,737,1266,1293]" pageId="4" pageNumber="227">
Key to species groups and species of
<taxonomicName id="2AE9D5CCC003FF80FD9D2A92FD38FA94" authorityName="Vikberg and Zinovjev" authorityYear="2000" box="[608,737,1267,1293]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="genus">
<emphasis id="DF9D725DC003FF80FD9D2A92FD38FA94" bold="true" box="[608,737,1267,1293]" italics="true" pageId="4" pageNumber="227">Fagineura</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A5F3FDC4C003FF81FF6A2B59FA45FB54" lastPageId="5" lastPageNumber="228" pageId="4" pageNumber="227" type="key">
<key id="9978F97EC003FF81FF6A2B59FA45FB54" lastPageId="5" lastPageNumber="228" pageId="4" pageNumber="227">
<keyStep id="561DB6EAC003FF80FF6A2B59FA45F934" pageId="4" pageNumber="227">
<paragraph id="ED56AE4FC003FF80FF6A2B59FA45FA5B" blockId="4.[151,1437,1336,1475]" pageId="4" pageNumber="227">
<keyLead id="56180D7AC003FF80FF6A2B59FA45FA5B" pageId="4" pageNumber="227">
1 Mandibles symmetric; each gradually tapered from middle to apex in outer view, with low and dull apical carina on each of anterior and posterior surfaces* (
<figureCitation id="75D2B2CAC003FF80FDE72B34FCDBFAF2" box="[538,770,1365,1387]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="4" pageNumber="227">Figs 7A, D, E, J, K, P, Q</figureCitation>
). Frontal pit dorsally with median groove (
<figureCitation id="75D2B2CAC003FF80FB6A2B34FB25FAF2" box="[1175,1276,1365,1387]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="4" pageNumber="227">Figs 6P, Q</figureCitation>
). Female: cercus about 46 × as long as wide (
<figureCitation id="75D2B2CAC003FF80FE012B13FD4BFA11" box="[508,658,1394,1416]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). AH, Dorsal view; IP, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="4" pageNumber="227">Figs 9A, B, I, J</figureCitation>
); lance without subdorsal carina on radix* (
<figureCitation id="75D2B2CAC003FF80FBC42B13FB6AFA11" box="[1081,1203,1394,1416]" captionStart="FIGURE 10" captionStartId="23.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-374@23.[151,1436,957,1917]" captionTargetPageId="23" captionText="FIGURE 10. Ovipositor or lance, lateral view: Fagineura crenativora (A, B), F. parva (C, holotype), F. togashii (D, E), F. fulvistriata (F), F. flavomaculata (G, holotype), F. quercivora (H, I; H, holotype) and F. glabella (J, holotype)." figureDoi="http://doi.org/10.5281/zenodo.6367287" httpUri="https://zenodo.org/record/6367287/files/figure.png" pageId="4" pageNumber="227">Figs 10AC</figureCitation>
); lancet with sclerotized part before lamnium being narrow transverse band (
<figureCitation id="75D2B2CAC003FF80FD192BF1FCBEFA3F" box="[740,871,1424,1446]" captionStart="FIGURE 11" captionStartId="24.[151,250,1126,1150]" captionTargetBox="[151,1436,181,1094]" captionTargetId="figure-26@24.[151,1436,181,1103]" captionTargetPageId="24" captionText="FIGURE 11. Lancet: Fagineura crenativora (A, B), F. parva (C, holotype)." figureDoi="http://doi.org/10.5281/zenodo.6367291" httpUri="https://zenodo.org/record/6367291/files/figure.png" pageId="4" pageNumber="227">Figs 11 AC</figureCitation>
); annulus 1 anteriorly with transverse colorless area with ctenidium. Male: valvispina directed posteroventrally* (
<figureCitation id="75D2B2CAC003FF80FD012BCCFCA1FA5A" box="[764,888,1453,1475]" captionStart="FIGURE 14" captionStartId="27.[151,250,1869,1893]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-19@27.[151,1436,181,1845]" captionTargetPageId="27" captionText="FIGURE 14. Apex of male abdomen and genitalia: Fagineura crenativora (A, DF, K, L), F. parva (B, G, H, M, N) and F. togashii (C, I, J, O, P). AC, Apex of abdomen, dorsal view; D, G, I, genitalia, dorsal view; E, F, H, J, genitalia, ventral view (E, H, J, penis valve removed); KP, penis valve, left dorsal." figureDoi="http://doi.org/10.5281/zenodo.6367297" httpUri="https://zenodo.org/record/6367297/files/figure.png" pageId="4" pageNumber="227">Figs 14KN</figureCitation>
).........................
<taxonomicName id="2AE9D5CCC003FF80FB532BCFFAEDFA5A" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[1198,1332,1454,1475]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC003FF80FB532BCFFAEDFA5A" box="[1198,1332,1454,1475]" italics="true" pageId="4" pageNumber="227">F. crenativora</emphasis>
</taxonomicName>
group ... 2
</keyLead>
</paragraph>
<footnote id="8EF2B241C003FF80FF1F2BABFB3BFA79" box="[226,1250,1482,1504]" pageId="4" pageNumber="227">
<paragraph id="ED56AE4FC003FF80FF1F2BABFB3BFA79" blockId="4.[226,1250,1482,1504]" box="[226,1250,1482,1504]" pageId="4" pageNumber="227">
[* The character states are unknown for
<taxonomicName id="2AE9D5CCC003FF80FD982BAAFD31FA46" authorityName="Liu, Li &amp; Wei" authorityYear="2021" box="[613,744,1482,1504]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="DF9D725DC003FF80FD982BAAFD31FA46" box="[613,744,1482,1504]" italics="true" pageId="4" pageNumber="227">F. brevicornis</emphasis>
</taxonomicName>
,
<taxonomicName id="2AE9D5CCC003FF80FD082BAAFC51FA46" authorityName="Liu, Li &amp; Wei" authorityYear="2019" box="[757,904,1482,1504]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="flactoserrula">
<emphasis id="DF9D725DC003FF80FD082BAAFC51FA46" box="[757,904,1482,1504]" italics="true" pageId="4" pageNumber="227">F. flactoserrula</emphasis>
</taxonomicName>
,
<taxonomicName id="2AE9D5CCC003FF80FC692BAAFBC2FA46" authorityName="Liu, Li &amp; Wei" authorityYear="2021" box="[916,1051,1482,1504]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="longitangia">
<emphasis id="DF9D725DC003FF80FC692BAAFBC2FA46" box="[916,1051,1482,1504]" italics="true" pageId="4" pageNumber="227">F. longitangia</emphasis>
</taxonomicName>
and
<taxonomicName id="2AE9D5CCC003FF80FBB42BAAFB0CFA46" authorityName="Liu, Li &amp; Wei" authorityYear="2019" box="[1097,1237,1482,1504]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="xanthosoma">
<emphasis id="DF9D725DC003FF80FBB42BAAFB0CFA46" box="[1097,1237,1482,1504]" italics="true" pageId="4" pageNumber="227">F. xanthosoma</emphasis>
</taxonomicName>
.]
</paragraph>
</footnote>
<paragraph id="ED56AE4FC003FF80FF6A2B89FA45F934" blockId="4.[151,1437,1512,1710]" pageId="4" pageNumber="227">
<keyLead id="56180D7AC003FF80FF6A2B89FA45F934" pageId="4" pageNumber="227">
- Mandibles slightly asymmetric; left one with middle part very gradually tapered or being same thickness in outer view, with sharp apical carina on anterior surface (
<figureCitation id="75D2B2CAC003FF80FD992864FD3DF983" box="[612,740,1541,1563]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="4" pageNumber="227">Figs 7FI, R</figureCitation>
); right one with middle part being same thickness or slightly thickened toward apex in outer view, with sharp apical carina on posterior surface (
<figureCitation id="75D2B2CAC003FF80FC572843FBE9F9A1" box="[938,1072,1570,1592]" captionStart="FIGURE 7" captionStartId="20.[151,250,1845,1869]" captionTargetBox="[151,1436,189,1821]" captionTargetId="figure-26@20.[151,1436,189,1821]" captionTargetPageId="20" captionText="FIGURE 7. Mandible: Fagineura crenativora (A, B, D, J, P, Q, female; A, holotype), F. parva (E, K, female, holotype), F. togashii (C, F, L, R, S, female), F. flavomaculata (G, M, female, holotype), F. quercivora (H, N, female, holotype) and F. glabella (I, O, female, holotype). A, Ventral view; B, C, anterior view (C, max. h. = maximum height); DI, left mandible, outer view, left anterior; JO, right mandible, outer view, left posterior; P, R, anterior view; Q, S, posterior view." figureDoi="http://doi.org/10.5281/zenodo.6367277" httpUri="https://zenodo.org/record/6367277/files/figure.png" pageId="4" pageNumber="227">Figs 7LO, S</figureCitation>
). Frontal pit dorsally without median groove (
<figureCitation id="75D2B2CAC003FF80FECC2821FE41F9CF" box="[305,408,1600,1622]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="4" pageNumber="227">Figs 6R, S</figureCitation>
). Female: cercus about 89 × as long as wide (
<figureCitation id="75D2B2CAC003FF80FCAA2821FC22F9CC" box="[855,1019,1600,1622]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). AH, Dorsal view; IP, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="4" pageNumber="227">Figs 9CH, KP</figureCitation>
); lance with subdorsal carina on radix (
<figureCitation id="75D2B2CAC003FF80FA8F2821FEFDF9EA" captionStart="FIGURE 10" captionStartId="23.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-374@23.[151,1436,957,1917]" captionTargetPageId="23" captionText="FIGURE 10. Ovipositor or lance, lateral view: Fagineura crenativora (A, B), F. parva (C, holotype), F. togashii (D, E), F. fulvistriata (F), F. flavomaculata (G, holotype), F. quercivora (H, I; H, holotype) and F. glabella (J, holotype)." figureDoi="http://doi.org/10.5281/zenodo.6367287" httpUri="https://zenodo.org/record/6367287/files/figure.png" pageId="4" pageNumber="227">Figs 10DJ</figureCitation>
); lancet with sclerotized part before lamnium large, extended anteriorly along olistheter (
<figureCitation id="75D2B2CAC003FF80FB84283CFB29F9EB" box="[1145,1264,1629,1651]" captionStart="FIGURE 12" captionStartId="25.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1862]" captionTargetId="figure-19@25.[151,1436,181,1870]" captionTargetPageId="25" captionText="FIGURE 12. Lancet: Fagineura togashii (A, B), F. fulvistriata (C, holotype) and F. flavomaculata (D, E; D, holotype). A, B, C, Reversed." figureDoi="http://doi.org/10.5281/zenodo.6367293" httpUri="https://zenodo.org/record/6367293/files/figure.png" pageId="4" pageNumber="227">Figs 12AE</figureCitation>
,
<figureCitation id="75D2B2CAC003FF80FB04283CFA98F9EA" box="[1273,1345,1629,1651]" captionStart="FIGURE 13" captionStartId="26.[151,250,1847,1871]" captionTargetBox="[151,1436,181,1823]" captionTargetId="figure-26@26.[151,1436,181,1823]" captionTargetPageId="26" captionText="FIGURE 13. Lancet: Fagineura quercivora (AC; A, holotype) and F. glabella (D, E; D, holotype). A, Reversed." figureDoi="http://doi.org/10.5281/zenodo.6367295" httpUri="https://zenodo.org/record/6367295/files/figure.png" pageId="4" pageNumber="227">13AE</figureCitation>
); annulus 1 anteriorly without colorless area and ctenidium, sometimes with colorless line or indistinct ctenidium. Male: valvispina directed posterodorsally* (
<figureCitation id="75D2B2CAC003FF80FE3228F9FD9FF934" box="[463,582,1688,1710]" captionStart="FIGURE 14" captionStartId="27.[151,250,1869,1893]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-19@27.[151,1436,181,1845]" captionTargetPageId="27" captionText="FIGURE 14. Apex of male abdomen and genitalia: Fagineura crenativora (A, DF, K, L), F. parva (B, G, H, M, N) and F. togashii (C, I, J, O, P). AC, Apex of abdomen, dorsal view; D, G, I, genitalia, dorsal view; E, F, H, J, genitalia, ventral view (E, H, J, penis valve removed); KP, penis valve, left dorsal." figureDoi="http://doi.org/10.5281/zenodo.6367297" httpUri="https://zenodo.org/record/6367297/files/figure.png" pageId="4" pageNumber="227">Figs 14O, P</figureCitation>
)....................................................
<taxonomicName id="2AE9D5CCC003FF80FB4928F9FAEDF934" baseAuthorityName="Togashi" baseAuthorityYear="1997" box="[1204,1332,1688,1709]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC003FF80FB4928F9FAEDF934" box="[1204,1332,1688,1709]" italics="true" pageId="4" pageNumber="227">F. quercivora</emphasis>
</taxonomicName>
group ... 9
</keyLead>
</paragraph>
</keyStep>
<footnote id="8EF2B241C003FF80FF1F28D4FD00F953" box="[226,729,1716,1739]" pageId="4" pageNumber="227">
<paragraph id="ED56AE4FC003FF80FF1F28D4FD00F953" blockId="4.[226,729,1716,1739]" box="[226,729,1716,1739]" pageId="4" pageNumber="227">
[* The character state is known only for
<taxonomicName id="2AE9D5CCC003FF80FD9A28D7FD12F953" authorityName="Hara" authorityYear="2022" box="[615,715,1717,1739]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="togashii">
<emphasis id="DF9D725DC003FF80FD9A28D7FD12F953" box="[615,715,1717,1739]" italics="true" pageId="4" pageNumber="227">F. togashii</emphasis>
</taxonomicName>
.]
</paragraph>
</footnote>
<keyStep id="561DB6EAC003FF80FF6A28B3FA45F89C" pageId="4" pageNumber="227">
<paragraph id="ED56AE4FC003FF80FF6A28B3FA45F97E" blockId="4.[151,1436,1746,1798]" box="[151,1436,1746,1768]" pageId="4" pageNumber="227">
<keyLead id="56180D7AC003FF80FF6A28B3FA45F97E" box="[151,1436,1746,1768]" pageId="4" pageNumber="227">2(1) Female.............................................................................................. 3</keyLead>
</paragraph>
<paragraph id="ED56AE4FC003FF80FF6A2891FA45F89C" blockId="4.[151,1436,1746,1798]" box="[151,1436,1776,1798]" pageId="4" pageNumber="227">
<keyLead id="56180D7AC003FF80FF6A2891FA45F89C" box="[151,1436,1776,1798]" pageId="4" pageNumber="227">- Male*............................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<footnote id="8EF2B241C003FF80FF1F296CFEEBF8D9" pageId="4" pageNumber="227">
<paragraph id="ED56AE4FC003FF80FF1F296CFEEBF8D9" blockId="4.[226,1436,1805,1856]" pageId="4" pageNumber="227">
[* Only
<taxonomicName id="2AE9D5CCC003FF80FEC8296FFE67F8BA" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[309,446,1806,1827]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC003FF80FEC8296FFE67F8BA" box="[309,446,1806,1827]" italics="true" pageId="4" pageNumber="227">F. crenativora</emphasis>
</taxonomicName>
and
<taxonomicName id="2AE9D5CCC003FF80FE0D296FFD9BF8BA" authorityName="Hara" authorityYear="2022" box="[496,578,1806,1827]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="DF9D725DC003FF80FE0D296FFD9BF8BA" box="[496,578,1806,1827]" italics="true" pageId="4" pageNumber="227">F. parva</emphasis>
</taxonomicName>
; the males of
<taxonomicName id="2AE9D5CCC003FF80FD2C296FFC8FF8BB" authorityName="Liu, Li &amp; Wei" authorityYear="2021" box="[721,854,1805,1827]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="DF9D725DC003FF80FD2C296FFC8FF8BB" box="[721,854,1805,1827]" italics="true" pageId="4" pageNumber="227">F. brevicornis</emphasis>
</taxonomicName>
,
<taxonomicName id="2AE9D5CCC003FF80FC99296FFC23F8BB" authorityName="Liu, Li &amp; Wei" authorityYear="2019" box="[868,1018,1805,1827]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="flactoserrula">
<emphasis id="DF9D725DC003FF80FC99296FFC23F8BB" box="[868,1018,1805,1827]" italics="true" pageId="4" pageNumber="227">F. flactoserrula</emphasis>
</taxonomicName>
,
<taxonomicName id="2AE9D5CCC003FF80FBF5296FFB49F8BB" authorityName="Liu, Li &amp; Wei" authorityYear="2021" box="[1032,1168,1805,1827]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="longitangia">
<emphasis id="DF9D725DC003FF80FBF5296FFB49F8BB" box="[1032,1168,1805,1827]" italics="true" pageId="4" pageNumber="227">F. longitangia</emphasis>
</taxonomicName>
and
<taxonomicName id="2AE9D5CCC003FF80FB3F296FFA96F8BB" authorityName="Liu, Li &amp; Wei" authorityYear="2019" box="[1218,1359,1805,1827]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="xanthosoma">
<emphasis id="DF9D725DC003FF80FB3F296FFA96F8BB" box="[1218,1359,1805,1827]" italics="true" pageId="4" pageNumber="227">F. xanthosoma</emphasis>
</taxonomicName>
are unknown.]
</paragraph>
</footnote>
<keyStep id="561DB6EAC003FF80FF6A2929FA45F8E3" pageId="4" pageNumber="227">
<paragraph id="ED56AE4FC003FF80FF6A2929FA45F8C4" blockId="4.[151,1436,1864,2032]" box="[151,1436,1864,1886]" pageId="4" pageNumber="227">
<keyLead id="56180D7AC003FF80FF6A2929FA45F8C4" box="[151,1436,1864,1886]" pageId="4" pageNumber="227">
3(2) Valvula 3 with apex concave in dorsal view (
<figureCitation id="75D2B2CAC003FF80FD6D2929FD22F8C4" box="[656,763,1864,1886]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). AH, Dorsal view; IP, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="4" pageNumber="227">Figs 9A, B</figureCitation>
)...................................................... 4
</keyLead>
</paragraph>
<paragraph id="ED56AE4FC003FF80FF6A2904FA45F8E3" blockId="4.[151,1436,1864,2032]" box="[151,1436,1893,1915]" pageId="4" pageNumber="227">
<keyLead id="56180D7AC003FF80FF6A2904FA45F8E3" box="[151,1436,1893,1915]" pageId="4" pageNumber="227">
- Valvula 3 with apex pointed in dorsal view (fig. 2f in
<bibRefCitation id="8978D3BEC003FF80FD192904FCAFF8E2" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[740,886,1893,1915]" pageId="4" pageNumber="227" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC003FF80FCF12907FCEFF8E3" box="[780,822,1893,1915]" italics="true" pageId="4" pageNumber="227">et al</emphasis>
. 2019
</bibRefCitation>
)...............................
<taxonomicName id="2AE9D5CCC003FF80FB0E2907FAA7F8E3" authorityName="Liu, Li &amp; Wei" authorityYear="2019" box="[1267,1406,1893,1915]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="xanthosoma">
<emphasis id="DF9D725DC003FF80FB0E2907FAA7F8E3" box="[1267,1406,1893,1915]" italics="true" pageId="4" pageNumber="227">F. xanthosoma</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C003FF80FA772904FA45F8E3" box="[1418,1436,1893,1914]" pageId="4" pageNumber="227" type="female"></specimenCount>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="561DB6EAC003FF80FF6A29E3FA45F82C" pageId="4" pageNumber="227">
<paragraph id="ED56AE4FC003FF80FF6A29E3FA45F801" blockId="4.[151,1436,1864,2032]" box="[151,1436,1922,1944]" pageId="4" pageNumber="227">
<keyLead id="56180D7AC003FF80FF6A29E3FA45F801" box="[151,1436,1922,1944]" pageId="4" pageNumber="227">
4(3) Valvula 3 narrower than long in dorsal view (
<figureCitation id="75D2B2CAC003FF80FD6829E3FD38F801" box="[661,737,1922,1944]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). AH, Dorsal view; IP, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="4" pageNumber="227">Fig. 9B</figureCitation>
). Cercus posteriorly not beyond valvula 3 or slightly beyond it....... 5
</keyLead>
</paragraph>
<paragraph id="ED56AE4FC003FF80FF6A29C1FA45F82C" blockId="4.[151,1436,1864,2032]" box="[151,1436,1952,1974]" pageId="4" pageNumber="227">
<keyLead id="56180D7AC003FF80FF6A29C1FA45F82C" box="[151,1436,1952,1974]" pageId="4" pageNumber="227">
- Valvula 3 wider than long in dorsal view (
<figureCitation id="75D2B2CAC003FF80FD8829C1FD1BF82F" box="[629,706,1952,1974]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). AH, Dorsal view; IP, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="4" pageNumber="227">Fig. 9A</figureCitation>
). Cercus posteriorly far beyond valvula 3.............
<taxonomicName id="2AE9D5CCC003FF80FB0529C1FAA7F82C" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[1272,1406,1952,1973]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC003FF80FB0529C1FAA7F82C" box="[1272,1406,1952,1973]" italics="true" pageId="4" pageNumber="227">F. crenativora</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C003FF80FA7729C1FA45F82C" box="[1418,1436,1952,1973]" pageId="4" pageNumber="227" type="female"></specimenCount>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="561DB6EAC003FF81FF6A29DCFA45FF34" lastPageId="5" lastPageNumber="228" pageId="4" pageNumber="227">
<paragraph id="ED56AE4FC003FF80FF6A29DCFA45F84A" blockId="4.[151,1436,1864,2032]" box="[151,1436,1981,2003]" pageId="4" pageNumber="227">
<keyLead id="56180D7AC003FF80FF6A29DCFA45F84A" box="[151,1436,1981,2003]" pageId="4" pageNumber="227">
5(4) Each flagellomere except for apical one slightly widened toward apex (
<figureCitation id="75D2B2CAC003FF80FC7329DCFC2EF84A" box="[910,1015,1981,2003]" captionStart="FIGURE 2" captionStartId="9.[151,250,1741,1765]" captionTargetBox="[158,1435,189,1704]" captionTargetId="figure-19@9.[151,1435,181,1717]" captionTargetPageId="9" captionText="FIGURE 2. Fagineura parva (AC, female, holotype; DF, male) and F. togashii (GJ, female; KN, male). A, D, G, K, Head, frontal view; B, E, H, L, dorsal view; C, F, I, M, ventrolateral or lateral view; J, N, ventral view. GN, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367259" httpUri="https://zenodo.org/record/6367259/files/figure.png" pageId="4" pageNumber="227">Figs 2B, C</figureCitation>
)................................. 6
</keyLead>
</paragraph>
<paragraph id="ED56AE4FC003FF81FF6A29BAFA45FF34" blockId="4.[151,1436,1864,2032]" lastBlockId="5.[151,1437,151,1230]" lastPageId="5" lastPageNumber="228" pageId="4" pageNumber="227">
<keyLead id="56180D7AC003FF81FF6A29BAFA45FF34" lastPageId="5" lastPageNumber="228" pageId="4" pageNumber="227">
- Each flagellomere except for apical one markedly widened toward apex (figs 1A, 3A, E in
<bibRefCitation id="8978D3BEC003FF80FBB129BBFB03F869" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[1100,1242,2010,2032]" pageId="4" pageNumber="227" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">
Liu
<emphasis id="DF9D725DC003FF80FB8E29BAFB44F876" box="[1139,1181,2010,2032]" italics="true" pageId="4" pageNumber="227">et al</emphasis>
. 2021
</bibRefCitation>
)......................................................................................................
<taxonomicName id="2AE9D5CCC002FF81FB072EF9FAA7FF35" authorityName="Liu, Li &amp; Wei" authorityYear="2021" box="[1274,1406,151,173]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="228" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="DF9D725DC002FF81FB072EF9FAA7FF35" box="[1274,1406,151,173]" italics="true" pageId="5" pageNumber="228">F. brevicornis</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C002FF81FA772EF9FA45FF34" box="[1418,1436,152,173]" pageId="5" pageNumber="228" type="female"></specimenCount>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="561DB6EAC002FF81FF6A2ED4FA45FEBB" pageId="5" pageNumber="228">
<paragraph id="ED56AE4FC002FF81FF6A2ED4FA45FF71" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2ED4FA45FF71" pageId="5" pageNumber="228">
6(5) Head black between eye and anterior tentorial pit (
<figureCitation id="75D2B2CAC002FF81FD362ED4FCC0FF52" box="[715,793,181,203]" captionStart="FIGURE 2" captionStartId="9.[151,250,1741,1765]" captionTargetBox="[158,1435,189,1704]" captionTargetId="figure-19@9.[151,1435,181,1717]" captionTargetPageId="9" captionText="FIGURE 2. Fagineura parva (AC, female, holotype; DF, male) and F. togashii (GJ, female; KN, male). A, D, G, K, Head, frontal view; B, E, H, L, dorsal view; C, F, I, M, ventrolateral or lateral view; J, N, ventral view. GN, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367259" httpUri="https://zenodo.org/record/6367259/files/figure.png" pageId="5" pageNumber="228">Fig. 2A</figureCitation>
). Katepimeron with only a few or several setae. Tangium of lancet with no or one pore (
<figureCitation id="75D2B2CAC002FF81FE562EB3FDD8FF7E" box="[427,513,210,232]" captionStart="FIGURE 11" captionStartId="24.[151,250,1126,1150]" captionTargetBox="[151,1436,181,1094]" captionTargetId="figure-26@24.[151,1436,181,1103]" captionTargetPageId="24" captionText="FIGURE 11. Lancet: Fagineura crenativora (A, B), F. parva (C, holotype)." figureDoi="http://doi.org/10.5281/zenodo.6367291" httpUri="https://zenodo.org/record/6367291/files/figure.png" pageId="5" pageNumber="228">Fig. 11C</figureCitation>
; fig. 1j in
<bibRefCitation id="8978D3BEC002FF81FD992EB3FD2BFF71" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[612,754,210,232]" pageId="5" pageNumber="228" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC002FF81FD762EB2FD6CFF7E" box="[651,693,210,232]" italics="true" pageId="5" pageNumber="228">et al</emphasis>
. 2019
</bibRefCitation>
)....................................................... 7
</keyLead>
</paragraph>
<paragraph id="ED56AE4FC002FF81FF6A2E91FA45FEBB" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2E91FA45FEBB" pageId="5" pageNumber="228">
- Head with pale area between eye and anterior tentorial pit (figs 3C, 4C in
<bibRefCitation id="8978D3BEC002FF81FC4A2E8EFB95FE9F" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[951,1100,239,262]" pageId="5" pageNumber="228" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">
Liu
<emphasis id="DF9D725DC002FF81FC222E91FBD2FE9D" box="[991,1035,239,261]" italics="true" pageId="5" pageNumber="228">et al</emphasis>
. 2021
</bibRefCitation>
). Katepimeron posteriorly covered with setae. Tangium of lancet with six pores (fig. 4I in
<bibRefCitation id="8978D3BEC002FF81FD0D2F6CFC5BFEBA" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[752,898,269,291]" pageId="5" pageNumber="228" pagination="125 - 137" refId="ref21924" refString="Liu, M., Li, Z. &amp; Wei, M. (2021) A key to all species of Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) worldwide with the descriptions of two new Chinese species. Journal of Hymenoptera Research, 83, 125 - 137. https: // doi. org / 10.3897 / jhr. 83.64380" type="journal article" year="2021">
Liu
<emphasis id="DF9D725DC002FF81FCE52F6CFC9BFEBB" box="[792,834,269,290]" italics="true" pageId="5" pageNumber="228">et al</emphasis>
. 2021
</bibRefCitation>
)..............................
<taxonomicName id="2AE9D5CCC002FF81FB052F6CFAA6FEBB" authorityName="Liu, Li &amp; Wei" authorityYear="2021" box="[1272,1407,269,290]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="228" phylum="Arthropoda" rank="species" species="longitangia">
<emphasis id="DF9D725DC002FF81FB052F6CFAA6FEBB" box="[1272,1407,269,290]" italics="true" pageId="5" pageNumber="228">F. longitangia</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C002FF81FA772F6CFA45FEBB" box="[1418,1436,269,290]" pageId="5" pageNumber="228" type="female"></specimenCount>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="561DB6EAC002FF81FF6A2F4BFA45FE4B" pageId="5" pageNumber="228">
<paragraph id="ED56AE4FC002FF81FF6A2F4BFA45FEE3" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2F4BFA45FEE3" pageId="5" pageNumber="228">
7(6) Mesepisternum entirely dark brown to black (
<figureCitation id="75D2B2CAC002FF81FD622F4BFD34FED9" box="[671,749,298,320]" captionStart="FIGURE 2" captionStartId="9.[151,250,1741,1765]" captionTargetBox="[158,1435,189,1704]" captionTargetId="figure-19@9.[151,1435,181,1717]" captionTargetPageId="9" captionText="FIGURE 2. Fagineura parva (AC, female, holotype; DF, male) and F. togashii (GJ, female; KN, male). A, D, G, K, Head, frontal view; B, E, H, L, dorsal view; C, F, I, M, ventrolateral or lateral view; J, N, ventral view. GN, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367259" httpUri="https://zenodo.org/record/6367259/files/figure.png" pageId="5" pageNumber="228">Fig. 2C</figureCitation>
). Lancet with transverse sclerotized band before lamnium very narrow and short, dorsally widely separated from olistheter (
<figureCitation id="75D2B2CAC002FF81FD262F26FCE8FEC7" box="[731,817,327,350]" captionStart="FIGURE 11" captionStartId="24.[151,250,1126,1150]" captionTargetBox="[151,1436,181,1094]" captionTargetId="figure-26@24.[151,1436,181,1103]" captionTargetPageId="24" captionText="FIGURE 11. Lancet: Fagineura crenativora (A, B), F. parva (C, holotype)." figureDoi="http://doi.org/10.5281/zenodo.6367291" httpUri="https://zenodo.org/record/6367291/files/figure.png" pageId="5" pageNumber="228">Fig. 11C</figureCitation>
); middle ctenidia long, dorsally almost in contact with neighboring ctenidia..................................................................................
<taxonomicName id="2AE9D5CCC002FF81FAD22F04FAA7FEE3" authorityName="Hara" authorityYear="2022" box="[1327,1406,357,378]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="228" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="DF9D725DC002FF81FAD22F04FAA7FEE3" box="[1327,1406,357,378]" italics="true" pageId="5" pageNumber="228">F. parva</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C002FF81FA772F04FA45FEE3" box="[1418,1436,357,378]" pageId="5" pageNumber="228" type="female"></specimenCount>
</keyLead>
</paragraph>
<paragraph id="ED56AE4FC002FF81FF6A2FE2FA45FE4B" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2FE2FA45FE4B" pageId="5" pageNumber="228">
- Mesepisternum widely yellow brown (fig. 1e in
<bibRefCitation id="8978D3BEC002FF81FD562FE3FCE6FE01" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[683,831,386,408]" pageId="5" pageNumber="228" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC002FF81FD2C2FE2FD23FE0E" box="[721,762,386,408]" italics="true" pageId="5" pageNumber="228">et al</emphasis>
., 2019
</bibRefCitation>
). Lancet with transverse sclerotized band before lamnium wide, dorsally extending to olistheter (fig. 1i in
<bibRefCitation id="8978D3BEC002FF81FD922FFEFCDCFE2F" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[623,773,415,438]" pageId="5" pageNumber="228" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC002FF81FD6B2FC1FD19FE2D" box="[662,704,415,437]" italics="true" pageId="5" pageNumber="228">et al</emphasis>
., 2019
</bibRefCitation>
); middle ctenidia widely separated from neighboring ctenidia (fig. 1h in
<bibRefCitation id="8978D3BEC002FF81FF062FDCFE56FE4A" author="Liu, M. &amp; Li, Z. &amp; Wei, M." box="[251,399,445,467]" pageId="5" pageNumber="228" pagination="29 - 42" refId="ref21864" refString="Liu, M., Li, Z. &amp; Wei, M. (2019) First record of the genus Fagineura Vikberg &amp; Zinovjev (Hymenoptera, Tenthredinidae) with descriptions of two new species from China. ZooKeys, 829, 29 - 42. https: // doi. org / 10.3897 / zookeys. 829.30086" type="journal article" year="2019">
Liu
<emphasis id="DF9D725DC002FF81FEDF2FDCFE95FE4B" box="[290,332,445,466]" italics="true" pageId="5" pageNumber="228">et al</emphasis>
., 2019
</bibRefCitation>
)......................................................................
<taxonomicName id="2AE9D5CCC002FF81FB162FDCFAA7FE4B" authorityName="Liu, Li &amp; Wei" authorityYear="2019" box="[1259,1406,445,466]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="228" phylum="Arthropoda" rank="species" species="flactoserrula">
<emphasis id="DF9D725DC002FF81FB162FDCFAA7FE4B" box="[1259,1406,445,466]" italics="true" pageId="5" pageNumber="228">F. flactoserrula</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C002FF81FA772FDCFA45FE4B" box="[1418,1436,445,466]" pageId="5" pageNumber="228" type="female"></specimenCount>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="561DB6EAC002FF81FF6A2FBBFA45FDFC" pageId="5" pageNumber="228">
<paragraph id="ED56AE4FC002FF81FF6A2FBBFA45FDB3" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2FBBFA45FDB3" pageId="5" pageNumber="228">
8(2) Head pale between eye and anterior tentorial pit (
<figureCitation id="75D2B2CAC002FF81FD482FBBFD27FE69" box="[693,766,474,496]" captionStart="FIGURE 1" captionStartId="7.[151,250,1832,1856]" captionTargetBox="[158,1435,276,1794]" captionTargetId="figure-57@7.[151,1435,270,1806]" captionTargetPageId="7" captionText="FIGURE 1. Fagineura crenativora (AK, female; LN, male). A, E, I, L, Head, frontal view; B, F, J, M, dorsal view; C, G, K, N, lateral or ventrolateral view; D, H, ventral view. AH, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367257" httpUri="https://zenodo.org/record/6367257/files/figure.png" pageId="5" pageNumber="228">Fig. 1L</figureCitation>
). Thorax ventrally pale mostly or widely (
<figureCitation id="75D2B2CAC002FF81FB732FBBFB26FE69" box="[1166,1279,474,496]" captionStart="FIGURE 1" captionStartId="7.[151,250,1832,1856]" captionTargetBox="[158,1435,276,1794]" captionTargetId="figure-57@7.[151,1435,270,1806]" captionTargetPageId="7" captionText="FIGURE 1. Fagineura crenativora (AK, female; LN, male). A, E, I, L, Head, frontal view; B, F, J, M, dorsal view; C, G, K, N, lateral or ventrolateral view; D, H, ventral view. AH, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367257" httpUri="https://zenodo.org/record/6367257/files/figure.png" pageId="5" pageNumber="228">Figs 1M, N</figureCitation>
). Abdomen with dorsum black basally and brown apically or mostly black and venter yellow. Paravalva with spinules ventrally (
<figureCitation id="75D2B2CAC002FF81FAED2F96FABEFD97" box="[1296,1383,503,526]" captionStart="FIGURE 14" captionStartId="27.[151,250,1869,1893]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-19@27.[151,1436,181,1845]" captionTargetPageId="27" captionText="FIGURE 14. Apex of male abdomen and genitalia: Fagineura crenativora (A, DF, K, L), F. parva (B, G, H, M, N) and F. togashii (C, I, J, O, P). AC, Apex of abdomen, dorsal view; D, G, I, genitalia, dorsal view; E, F, H, J, genitalia, ventral view (E, H, J, penis valve removed); KP, penis valve, left dorsal." figureDoi="http://doi.org/10.5281/zenodo.6367297" httpUri="https://zenodo.org/record/6367297/files/figure.png" pageId="5" pageNumber="228">Fig. 14L</figureCitation>
)..........................................................................................
<taxonomicName id="2AE9D5CCC002FF81FB052C74FAA7FDB3" authorityName="♀ Vikberg &amp; Zinovjev" authorityYear="2000" box="[1272,1406,533,554]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="228" phylum="Arthropoda" rank="species" species="crenativora">
<emphasis id="DF9D725DC002FF81FB052C74FAA7FDB3" box="[1272,1406,533,554]" italics="true" pageId="5" pageNumber="228">F. crenativora</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C002FF81FA772C74FA45FDB3" box="[1418,1436,533,554]" pageId="5" pageNumber="228" type="male"></specimenCount>
</keyLead>
</paragraph>
<paragraph id="ED56AE4FC002FF81FF6A2C52FA45FDFC" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2C52FA45FDFC" pageId="5" pageNumber="228">
- Head black between eye and anterior tentorial pit (
<figureCitation id="75D2B2CAC002FF81FD252C53FCF1FDD1" box="[728,808,562,584]" captionStart="FIGURE 2" captionStartId="9.[151,250,1741,1765]" captionTargetBox="[158,1435,189,1704]" captionTargetId="figure-19@9.[151,1435,181,1717]" captionTargetPageId="9" captionText="FIGURE 2. Fagineura parva (AC, female, holotype; DF, male) and F. togashii (GJ, female; KN, male). A, D, G, K, Head, frontal view; B, E, H, L, dorsal view; C, F, I, M, ventrolateral or lateral view; J, N, ventral view. GN, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367259" httpUri="https://zenodo.org/record/6367259/files/figure.png" pageId="5" pageNumber="228">Fig. 2D</figureCitation>
). Thorax ventrally darkened (
<figureCitation id="75D2B2CAC002FF81FBB62C53FB6EFDD1" box="[1099,1207,562,584]" captionStart="FIGURE 2" captionStartId="9.[151,250,1741,1765]" captionTargetBox="[158,1435,189,1704]" captionTargetId="figure-19@9.[151,1435,181,1717]" captionTargetPageId="9" captionText="FIGURE 2. Fagineura parva (AC, female, holotype; DF, male) and F. togashii (GJ, female; KN, male). A, D, G, K, Head, frontal view; B, E, H, L, dorsal view; C, F, I, M, ventrolateral or lateral view; J, N, ventral view. GN, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367259" httpUri="https://zenodo.org/record/6367259/files/figure.png" pageId="5" pageNumber="228">Figs 2E, F</figureCitation>
). Abdomen black, with subgenital plate brownish. Paravalva without spinules (
<figureCitation id="75D2B2CAC002FF81FD092C2EFC96FDFF" box="[756,847,591,614]" captionStart="FIGURE 14" captionStartId="27.[151,250,1869,1893]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-19@27.[151,1436,181,1845]" captionTargetPageId="27" captionText="FIGURE 14. Apex of male abdomen and genitalia: Fagineura crenativora (A, DF, K, L), F. parva (B, G, H, M, N) and F. togashii (C, I, J, O, P). AC, Apex of abdomen, dorsal view; D, G, I, genitalia, dorsal view; E, F, H, J, genitalia, ventral view (E, H, J, penis valve removed); KP, penis valve, left dorsal." figureDoi="http://doi.org/10.5281/zenodo.6367297" httpUri="https://zenodo.org/record/6367297/files/figure.png" pageId="5" pageNumber="228">Fig. 14N</figureCitation>
)........................................
<taxonomicName id="2AE9D5CCC002FF81FAD22C31FAA7FDFC" authorityName="Hara" authorityYear="2022" box="[1327,1406,592,613]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="228" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="DF9D725DC002FF81FAD22C31FAA7FDFC" box="[1327,1406,592,613]" italics="true" pageId="5" pageNumber="228">F. parva</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C002FF81FA772C31FA45FDFC" box="[1418,1436,592,613]" pageId="5" pageNumber="228" type="male"></specimenCount>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="561DB6EAC002FF81FF6A2C0CFA45FD43" pageId="5" pageNumber="228">
<paragraph id="ED56AE4FC002FF81FF6A2C0CFA45FD06" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2C0CFA45FD06" pageId="5" pageNumber="228">
9(1) Mesepisternum not or very narrowly glabrous and barely or slightly expanded beside postspiracular sclerite (
<figureCitation id="75D2B2CAC002FF81FAFF2C0CFAB8FD1B" box="[1282,1377,621,643]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="5" pageNumber="228">Figs 8I, J</figureCitation>
). Pale areas of body yellowish in color (
<figureCitation id="75D2B2CAC002FF81FDDE2CEBFD48FD06" box="[547,657,650,672]" captionStart="FIGURE 2" captionStartId="9.[151,250,1741,1765]" captionTargetBox="[158,1435,189,1704]" captionTargetId="figure-19@9.[151,1435,181,1717]" captionTargetPageId="9" captionText="FIGURE 2. Fagineura parva (AC, female, holotype; DF, male) and F. togashii (GJ, female; KN, male). A, D, G, K, Head, frontal view; B, E, H, L, dorsal view; C, F, I, M, ventrolateral or lateral view; J, N, ventral view. GN, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367259" httpUri="https://zenodo.org/record/6367259/files/figure.png" pageId="5" pageNumber="228">Figs 2GN</figureCitation>
,
<figureCitation id="75D2B2CAC002FF81FD672CEBFD01FD39" box="[666,728,650,672]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (AE, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (FH, N, female; OQ, late instar, final instar and mature final instar larva; holotype) and F. quercivora (IK, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. LQ, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. AD, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="5" pageNumber="228">4AE</figureCitation>
)......................................................... 10
</keyLead>
</paragraph>
<paragraph id="ED56AE4FC002FF81FF6A2CC9FA45FD43" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2CC9FA45FD43" pageId="5" pageNumber="228">
- Mesepisternum moderately or widely glabrous and distinctly expanded beside postspiracular sclerite (
<figureCitation id="75D2B2CAC002FF81FB472CC6FAF3FD27" box="[1210,1322,679,702]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="5" pageNumber="228">Figs 8KN</figureCitation>
). Pale areas of body yellowish or reddish in color (
<figureCitation id="75D2B2CAC002FF81FDAE2CA4FD65FD43" box="[595,700,709,731]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (AE, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (FH, N, female; OQ, late instar, final instar and mature final instar larva; holotype) and F. quercivora (IK, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. LQ, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. AD, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="5" pageNumber="228">Figs 4FK</figureCitation>
,
<figureCitation id="75D2B2CAC002FF81FD3B2CA4FCDBFD42" box="[710,770,709,731]" captionStart="FIGURE 5" captionStartId="18.[151,250,1940,1964]" captionTargetBox="[151,1435,394,1917]" captionTargetId="figure-147@18.[151,1435,381,1917]" captionTargetPageId="18" captionText="FIGURE 5. Fagineura quercivora (AC, female; I, early instar larvae; J, final instar larvae) and F. glabella (DH, female, holotype). A, D, H, Dorsal view; B, E, lateral view; C, F, ventral view; G, head, frontal view. I, Nakagawa, 5 May 2016; J, Nakagwa, 14 May 2012. AF, Photographed by F. Ito, just after killing; I, J, photographed by S. Ibuki." figureDoi="http://doi.org/10.5281/zenodo.6367273" httpUri="https://zenodo.org/record/6367273/files/figure.png" pageId="5" pageNumber="228">5AH</figureCitation>
).................................................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="561DB6EAC002FF81FF6A2C83FA45FC12" pageId="5" pageNumber="228">
<paragraph id="ED56AE4FC002FF81FF6A2C83FA45FCAB" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2C83FA45FCAB" pageId="5" pageNumber="228">
10(9) Mesepisternum not glabrous and barely expanded beside postspiracular sclerite (
<figureCitation id="75D2B2CAC002FF81FC1C2C83FBFAFD61" box="[993,1059,738,760]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="5" pageNumber="228">Fig. 8I</figureCitation>
). In female, postocellar area black (
<figureCitation id="75D2B2CAC002FF81FA882C83FF24FC8C" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="5" pageNumber="228">Fig. 6F</figureCitation>
), mesepimeron darkened with dorsal margin and posterior part yellow (
<figureCitation id="75D2B2CAC002FF81FC472C9EFC26FC8F" box="[954,1023,767,790]" captionStart="FIGURE 2" captionStartId="9.[151,250,1741,1765]" captionTargetBox="[158,1435,189,1704]" captionTargetId="figure-19@9.[151,1435,181,1717]" captionTargetPageId="9" captionText="FIGURE 2. Fagineura parva (AC, female, holotype; DF, male) and F. togashii (GJ, female; KN, male). A, D, G, K, Head, frontal view; B, E, H, L, dorsal view; C, F, I, M, ventrolateral or lateral view; J, N, ventral view. GN, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367259" httpUri="https://zenodo.org/record/6367259/files/figure.png" pageId="5" pageNumber="228">Fig. 2I</figureCitation>
), and abdomen with terga 1, 2 yellow and laterotergites 16 mostly yellow and laterotergites 7, 8 mostly black (
<figureCitation id="75D2B2CAC002FF81FC862D7CFC07FCAB" box="[891,990,797,819]" captionStart="FIGURE 2" captionStartId="9.[151,250,1741,1765]" captionTargetBox="[158,1435,189,1704]" captionTargetId="figure-19@9.[151,1435,181,1717]" captionTargetPageId="9" captionText="FIGURE 2. Fagineura parva (AC, female, holotype; DF, male) and F. togashii (GJ, female; KN, male). A, D, G, K, Head, frontal view; B, E, H, L, dorsal view; C, F, I, M, ventrolateral or lateral view; J, N, ventral view. GN, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367259" httpUri="https://zenodo.org/record/6367259/files/figure.png" pageId="5" pageNumber="228">Figs 2H, I</figureCitation>
).........................
<taxonomicName id="2AE9D5CCC002FF81FAFA2D7CFAB5FCAB" authorityName="Hara" authorityYear="2022" box="[1287,1388,797,818]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="228" phylum="Arthropoda" rank="species" species="togashii">
<emphasis id="DF9D725DC002FF81FAFA2D7CFAB5FCAB" box="[1287,1388,797,818]" italics="true" pageId="5" pageNumber="228">F. togashii</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C002FF81FA852D7CFA5EFCAB" box="[1400,1415,797,818]" pageId="5" pageNumber="228" type="female"></specimenCount>
<specimenCount id="FBEF65C6C002FF81FA7A2D7CFA45FCAB" box="[1415,1436,797,818]" pageId="5" pageNumber="228" type="male"></specimenCount>
</keyLead>
</paragraph>
<paragraph id="ED56AE4FC002FF81FF6A2D5AFA45FC12" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2D5AFA45FC12" pageId="5" pageNumber="228">
- Mesepisternum very narrowly glabrous and slightly expanded beside postspiracular sclerite (
<figureCitation id="75D2B2CAC002FF81FB932D5BFB6DFCC9" box="[1134,1204,826,848]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="5" pageNumber="228">Fig. 8J</figureCitation>
). In female, postocellar area yellow (
<figureCitation id="75D2B2CAC002FF81FEA32D36FE75FCF4" box="[350,428,855,877]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="5" pageNumber="228">Fig. 6H</figureCitation>
), mesepimeron entirely yellow to brown yellow (
<figureCitation id="75D2B2CAC002FF81FC782D36FC09FCF7" box="[901,976,855,878]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (AE, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (FH, N, female; OQ, late instar, final instar and mature final instar larva; holotype) and F. quercivora (IK, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. LQ, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. AD, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="5" pageNumber="228">Fig. 4C</figureCitation>
), and abdomen with terga 1, 2 mostly black and laterotergites 18 yellow (
<figureCitation id="75D2B2CAC002FF81FE202D14FD9EFC12" box="[477,583,885,907]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (AE, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (FH, N, female; OQ, late instar, final instar and mature final instar larva; holotype) and F. quercivora (IK, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. LQ, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. AD, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="5" pageNumber="228">Figs 4B, C</figureCitation>
)...............................................
<taxonomicName id="2AE9D5CCC002FF81FB8B2D14FB2EFC13" authorityName="Hara" authorityYear="2022" box="[1142,1271,885,906]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="228" phylum="Arthropoda" rank="species" species="fulvistriata">
<emphasis id="DF9D725DC002FF81FB8B2D14FB2EFC13" box="[1142,1271,885,906]" italics="true" pageId="5" pageNumber="228">F. fulvistriata</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C002FF81FAFE2D14FACCFC13" box="[1283,1301,885,906]" pageId="5" pageNumber="228" type="female"></specimenCount>
(
<specimenCount id="FBEF65C6C002FF81FADF2D14FAEDFC13" box="[1314,1332,885,906]" pageId="5" pageNumber="228" type="male"></specimenCount>
unknown)
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</paragraph>
</keyStep>
<keyStep id="561DB6EAC002FF81FF6A2DF3FA45FB84" pageId="5" pageNumber="228">
<paragraph id="ED56AE4FC002FF81FF6A2DF3FA45FC7A" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2DF3FA45FC7A" pageId="5" pageNumber="228">
11(9) Pale areas of body yellowish in color (
<figureCitation id="75D2B2CAC002FF81FDA62DF3FD1EFC31" box="[603,711,914,936]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (AE, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (FH, N, female; OQ, late instar, final instar and mature final instar larva; holotype) and F. quercivora (IK, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. LQ, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. AD, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="5" pageNumber="228">Figs 4FH</figureCitation>
). Mesoscutum mostly black. Postocellar area darkened (
<figureCitation id="75D2B2CAC002FF81FB162DF3FAE9FC31" box="[1259,1328,914,936]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="5" pageNumber="228">Fig. 6I</figureCitation>
). Hind trochanter pale yellow (
<figureCitation id="75D2B2CAC002FF81FE4D2DCEFDD9FC5F" box="[432,512,943,966]" captionStart="FIGURE 4" captionStartId="15.[151,250,1696,1720]" captionTargetBox="[151,1435,188,1672]" captionTargetId="figure-19@15.[151,1435,181,1672]" captionTargetPageId="15" captionText="FIGURE 4. Fagineura fulvistriata (AE, female; L, M, late instar and mature final instar larva; holotype), F. flavomaculata (FH, N, female; OQ, late instar, final instar and mature final instar larva; holotype) and F. quercivora (IK, female, holotype). A, F, I, Head, frontal view; B, E, G, J, dorsal view; C, H, K, lateral or ventrolateral view; D, ventral view. LQ, Nakagawa; L, 13 May 2019; M, 20 May 2019; N, 19 April 2019; O, 2 May 2018; P, 8 May 2018; Q, 9 May 2018; photographed by S. Ibuki. AD, Photographed just after killing." figureDoi="http://doi.org/10.5281/zenodo.6367269" httpUri="https://zenodo.org/record/6367269/files/figure.png" pageId="5" pageNumber="228">Fig. 4H</figureCitation>
). Mesoscutum except for posterolateral hollow mostly covered with setiferous punctures (
<figureCitation id="75D2B2CAC002FF81FA882DCEFF27FC7A" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="5" pageNumber="228">Fig. 8B</figureCitation>
)......................................................................
<taxonomicName id="2AE9D5CCC002FF81FBAB2DACFB2EFC7B" authorityName="Hara" authorityYear="2022" box="[1110,1271,973,994]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="228" phylum="Arthropoda" rank="species" species="flavomaculata">
<emphasis id="DF9D725DC002FF81FBAB2DACFB2EFC7B" box="[1110,1271,973,994]" italics="true" pageId="5" pageNumber="228">F. flavomaculata</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C002FF81FAFE2DACFACCFC7B" box="[1283,1301,973,994]" pageId="5" pageNumber="228" type="female"></specimenCount>
(
<specimenCount id="FBEF65C6C002FF81FADF2DACFAEDFC7B" box="[1314,1332,973,994]" pageId="5" pageNumber="228" type="male"></specimenCount>
unknown)
</keyLead>
</paragraph>
<paragraph id="ED56AE4FC002FF81FF6A2D8AFA45FB84" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2D8AFA45FB84" pageId="5" pageNumber="228">
- Pale areas of body mostly reddish in color (
<figureCitation id="75D2B2CAC002FF81FD802D8BFD3DFB99" box="[637,740,1002,1024]" captionStart="FIGURE 5" captionStartId="18.[151,250,1940,1964]" captionTargetBox="[151,1435,394,1917]" captionTargetId="figure-147@18.[151,1435,381,1917]" captionTargetPageId="18" captionText="FIGURE 5. Fagineura quercivora (AC, female; I, early instar larvae; J, final instar larvae) and F. glabella (DH, female, holotype). A, D, H, Dorsal view; B, E, lateral view; C, F, ventral view; G, head, frontal view. I, Nakagawa, 5 May 2016; J, Nakagwa, 14 May 2012. AF, Photographed by F. Ito, just after killing; I, J, photographed by S. Ibuki." figureDoi="http://doi.org/10.5281/zenodo.6367273" httpUri="https://zenodo.org/record/6367273/files/figure.png" pageId="5" pageNumber="228">Figs 5AJ</figureCitation>
). Mesoscutum predominantly pale. Combination of other character states not as above........................................................................................ 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="561DB6EAC002FF81FF6A2A44FA45FB54" pageId="5" pageNumber="228">
<paragraph id="ED56AE4FC002FF81FF6A2A44FA45FBEC" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2A44FA45FBEC" pageId="5" pageNumber="228">
12(11) Postocellar area predominantly or entirely black (
<figureCitation id="75D2B2CAC002FF81FD352A44FCCEFBA2" box="[712,791,1061,1083]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="5" pageNumber="228">Fig. 6K</figureCitation>
). Hind trochanter mostly black (
<figureCitation id="75D2B2CAC002FF81FBAA2A44FB7DFBA2" box="[1111,1188,1061,1083]" captionStart="FIGURE 5" captionStartId="18.[151,250,1940,1964]" captionTargetBox="[151,1435,394,1917]" captionTargetId="figure-147@18.[151,1435,381,1917]" captionTargetPageId="18" captionText="FIGURE 5. Fagineura quercivora (AC, female; I, early instar larvae; J, final instar larvae) and F. glabella (DH, female, holotype). A, D, H, Dorsal view; B, E, lateral view; C, F, ventral view; G, head, frontal view. I, Nakagawa, 5 May 2016; J, Nakagwa, 14 May 2012. AF, Photographed by F. Ito, just after killing; I, J, photographed by S. Ibuki." figureDoi="http://doi.org/10.5281/zenodo.6367273" httpUri="https://zenodo.org/record/6367273/files/figure.png" pageId="5" pageNumber="228">Fig. 5C</figureCitation>
). Mesoscutum except for posterolateral hollow mostly covered with setiferous punctures (
<figureCitation id="75D2B2CAC002FF81FCAC2A23FC47FBC1" box="[849,926,1090,1112]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="5" pageNumber="228">Fig. 8D</figureCitation>
), at most very sparsely punctured on anterior part of lateral lobe (
<figureCitation id="75D2B2CAC002FF81FEA72A3EFE7FFBEF" box="[346,422,1119,1142]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="5" pageNumber="228">Fig. 8C</figureCitation>
). Valvula 3 not concave apically in dorsal view (
<figureCitation id="75D2B2CAC002FF81FC852A3EFC1DFBEF" box="[888,964,1119,1142]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). AH, Dorsal view; IP, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="5" pageNumber="228">Fig. 9G</figureCitation>
)..............
<taxonomicName id="2AE9D5CCC002FF81FB8A2A01FB2FFBEC" baseAuthorityName="Togashi" baseAuthorityYear="1997" box="[1143,1270,1120,1141]" class="Insecta" family="Tenthredinidae" genus="Dineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="228" phylum="Arthropoda" rank="species" species="quercivora">
<emphasis id="DF9D725DC002FF81FB8A2A01FB2FFBEC" box="[1143,1270,1120,1141]" italics="true" pageId="5" pageNumber="228">F. quercivora</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C002FF81FAFE2A01FACCFBEC" box="[1283,1301,1120,1141]" pageId="5" pageNumber="228" type="female"></specimenCount>
(
<specimenCount id="FBEF65C6C002FF81FADF2A01FAEDFBEC" box="[1314,1332,1120,1141]" pageId="5" pageNumber="228" type="male"></specimenCount>
unknown)
</keyLead>
</paragraph>
<paragraph id="ED56AE4FC002FF81FF6A2A1CFA45FB54" blockId="5.[151,1437,151,1230]" pageId="5" pageNumber="228">
<keyLead id="56180D7AC002FF81FF6A2A1CFA45FB54" pageId="5" pageNumber="228">
- Postocellar area red yellow (
<figureCitation id="75D2B2CAC002FF81FE0B2A1CFD98FB0A" box="[502,577,1149,1171]" captionStart="FIGURE 6" captionStartId="19.[151,250,1889,1913]" captionTargetBox="[151,1436,381,1866]" captionTargetId="figure-91@19.[151,1436,381,1866]" captionTargetPageId="19" captionText="FIGURE 6. Head: Fagineura crenativora (A, B, M, O, P, female; A, M, holotype; C, male), F. parva (D, Q, female, holotype; E, male), F. togashii (F, N, R, female; G, male), F. fulvistriata (H, female, holotype), F. flavomaculata (I, female, holotype), F. quercivora (J, K, S, female; J, S, holotype) and F. glabella (L, female, holotype). AL, Dorsal view; M, N, lateral view; O, anteromedial part, anterolateral view; PS, anterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.6367275" httpUri="https://zenodo.org/record/6367275/files/figure.png" pageId="5" pageNumber="228">Fig. 6L</figureCitation>
), at most narrow median line and lateral margin black. Hind trochanter pale yellow (
<figureCitation id="75D2B2CAC002FF81FA882A1CFF27FB29" captionStart="FIGURE 5" captionStartId="18.[151,250,1940,1964]" captionTargetBox="[151,1435,394,1917]" captionTargetId="figure-147@18.[151,1435,381,1917]" captionTargetPageId="18" captionText="FIGURE 5. Fagineura quercivora (AC, female; I, early instar larvae; J, final instar larvae) and F. glabella (DH, female, holotype). A, D, H, Dorsal view; B, E, lateral view; C, F, ventral view; G, head, frontal view. I, Nakagawa, 5 May 2016; J, Nakagwa, 14 May 2012. AF, Photographed by F. Ito, just after killing; I, J, photographed by S. Ibuki." figureDoi="http://doi.org/10.5281/zenodo.6367273" httpUri="https://zenodo.org/record/6367273/files/figure.png" pageId="5" pageNumber="228">Fig. 5F</figureCitation>
). Mesoscutum except for posterolateral hollow widely smooth and glabrous (
<figureCitation id="75D2B2CAC002FF81FC202AFBFBFEFB29" box="[989,1063,1178,1200]" captionStart="FIGURE 8" captionStartId="21.[151,250,1580,1604]" captionTargetBox="[151,1436,166,1564]" captionTargetId="figure-19@21.[151,1436,166,1564]" captionTargetPageId="21" captionText="FIGURE 8. Thorax, claw and anteromiddle part of fore wing: Fagineura crenativora (A, G, O, W, X, female; O, W, holotype), F. parva (H, male; P, female, holotype), F. togashii (F, I, Q, female), F. fulvistriata (J, R, Y, female; J, R, holotype), F. flavomaculata (B, K, S, female, holotype), F. quercivora (C, D, L, M, T, U, female; C, L, T, holotype) and F. glabella (E, N, V, female, holotype).AE, Mesoscutum, dorsolateral view; F, anterodorsal part of mesepisternum, anteroventral view; GN, dorsal part of mesepisternum, anterolateral view; OV, claw, anterior or posterior view; WY, anteromiddle part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6367281" httpUri="https://zenodo.org/record/6367281/files/figure.png" pageId="5" pageNumber="228">Fig. 8E</figureCitation>
). Valvula 3 slightly concave apically in dorsal view (
<figureCitation id="75D2B2CAC002FF81FEA32AD6FE73FB57" box="[350,426,1207,1230]" captionStart="FIGURE 9" captionStartId="22.[151,250,1554,1578]" captionTargetBox="[151,1437,561,1531]" captionTargetId="figure-204@22.[151,1437,561,1531]" captionTargetPageId="22" captionText="FIGURE 9. Apex of female abdomen: Fagineura crenativora (A, I), F. parva (B, J, holotype), F. togashii (C, K), F. fulvistriata (D, L; D, holotype), F. flavomaculata (E, M; E, holotype), F. quercivora (F, G, N, O; F, N, holotype) and F. glabella (H, P, holotype). AH, Dorsal view; IP, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6367285" httpUri="https://zenodo.org/record/6367285/files/figure.png" pageId="5" pageNumber="228">Fig. 9H</figureCitation>
).............................................................
<taxonomicName id="2AE9D5CCC002FF81FB6C2AD9FB2EFB55" authorityName="Hara" authorityYear="2022" box="[1169,1271,1207,1229]" class="Insecta" family="Tenthredinidae" genus="Fagineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="228" phylum="Arthropoda" rank="species" species="glabella">
<emphasis id="DF9D725DC002FF81FB6C2AD9FB2EFB55" box="[1169,1271,1207,1229]" italics="true" pageId="5" pageNumber="228">F. glabella</emphasis>
</taxonomicName>
,
<specimenCount id="FBEF65C6C002FF81FAFE2AD9FACCFB54" box="[1283,1301,1208,1229]" pageId="5" pageNumber="228" type="female"></specimenCount>
(
<specimenCount id="FBEF65C6C002FF81FADF2AD9FAEDFB54" box="[1314,1332,1208,1229]" pageId="5" pageNumber="228" type="male"></specimenCount>
unknown)
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</subSection>
</treatment>
</document>