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<document id="E6F1B00008BBB5EE03A379CB318A5D83" ID-CLB-Dataset="8391" ID-DOI="10.11646/zootaxa.4920.2.5" ID-GBIF-Dataset="5676f8ca-c69e-461c-894d-2938f1f5cbbc" ID-ISSN="1175-5326" ID-Zenodo-Dep="4475213" ID-ZooBank="0A5D8BEB-2156-4198-B59E-C99565BCB2E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611820307356" checkinUser="plazi" docAuthor="Queiroz-Santos, Luziany, Casagrande, Mirna Martins &amp; Mielke, Olaf Hermann Hendrik" docDate="2021" docId="581E6933BD1EFF8AFF2007FE0FF82412" docLanguage="en" docName="zootaxa.4920.2.5.pdf" docOrigin="Zootaxa 4920 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Iza Walker 1865" docType="treatment" docVersion="7" lastPageNumber="239" masterDocId="A427114BBD1DFF8EFFB701390A7E205F" masterDocTitle="Taxonomic revision of the monotypic genera of Siculodini (Lepidoptera, Thyridoidea, Thyrididae, Siculodinae)" masterLastPageNumber="253" masterPageNumber="235" pageNumber="238" updateTime="1698925725074" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B49DC64D98381675F354BF111DE6B8E5">Taxonomic revision of the monotypic genera of Siculodini (Lepidoptera, Thyridoidea, Thyrididae, Siculodinae)</mods:title>
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<mods:namePart id="CB0AB0F90DC17B3FFA8310F4F54B2482">Queiroz-Santos, Luziany</mods:namePart>
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<mods:affiliation id="B9B71CBC4E4F771FA784DB9118F1D7BB">Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, P. O. Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil. &amp; luzianyqueiroz @ gmail. com; https: // orcid. org / 0000 - 0002 - 3796 - 2823</mods:affiliation>
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<mods:namePart id="49639073A5CC5024AD50A35AE5A418CE">Casagrande, Mirna Martins</mods:namePart>
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<mods:affiliation id="DD2E8AB9CE3E2417CC8887BFF04036AC">Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, P. O. Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil. &amp; mibras @ ufpr. br; https: // orcid. org / 0000 - 0002 - 6076 - 8463</mods:affiliation>
<mods:nameIdentifier id="A988329F8CB09BB78A9C296BD84C66A0" type="email">mibras@ufpr.br</mods:nameIdentifier>
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<mods:namePart id="A1BE800B4F66E0D959FED182A127A874">Mielke, Olaf Hermann Hendrik</mods:namePart>
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<mods:affiliation id="52417D87367734F7E1DF9E987094FC85">Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, P. O. Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil. &amp; omhesp @ ufpr. br; https: // orcid. org / 0000 - 0003 - 3655 - 4606</mods:affiliation>
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<mods:title id="AE5A36CB86A06A6B9D2C7934106602CD">Zootaxa</mods:title>
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<mods:date id="B89CDE9CB2E8A1BF454F756BBA7C6A29">2021</mods:date>
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<treatment id="581E6933BD1EFF8AFF2007FE0FF82412" ID-DOI="http://doi.org/10.5281/zenodo.4519943" ID-GBIF-Taxon="177309994" ID-Zenodo-Dep="4519943" LSID="urn:lsid:plazi:treatment:581E6933BD1EFF8AFF2007FE0FF82412" httpUri="http://treatment.plazi.org/id/581E6933BD1EFF8AFF2007FE0FF82412" lastPageId="4" lastPageNumber="239" pageId="3" pageNumber="238">
<subSubSection id="98AD8BAEBD1EFF8DFF2007FE0B1926BF" box="[151,359,1734,1761]" pageId="3" pageNumber="238" type="nomenclature">
<paragraph id="D008D825BD1EFF8DFF2007FE0B1926BF" blockId="3.[151,359,1734,1798]" box="[151,359,1734,1761]" pageId="3" pageNumber="238">
<heading id="8B406F49BD1EFF8DFF2007FE0B1926BF" bold="true" box="[151,359,1734,1761]" fontSize="11" level="1" pageId="3" pageNumber="238" reason="1">
<taxonomicName id="17B7A3A6BD1EFF8DFF2007FE0B1926BF" ID-CoL="92CLR" authority="Walker, 1865" authorityName="Walker" authorityYear="1865" box="[151,359,1734,1761]" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="E2C30437BD1EFF8DFF2007FE0B1926BF" bold="true" box="[151,359,1734,1761]" pageId="3" pageNumber="238">
<emphasis id="E2C30437BD1EFF8DFF2007FE0AC226BE" bold="true" box="[151,188,1735,1761]" italics="true" pageId="3" pageNumber="238">Iza</emphasis>
<bibRefCitation id="B426A5D4BD1EFF8DFF7407FE0B1926BF" author="Walker, F." box="[195,359,1734,1761]" pageId="3" pageNumber="238" pagination="1 - 706" refId="ref14761" refString="Walker, F. (1865) Catologue of Lepidoptera Heterocera. List of the specimens of Lepidopterous insects in the Collection of the British Museum, 31 (7), 1 - 706." type="journal article" year="1865">Walker, 1865</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="98AD8BAEBD1EFF8DFF2007D20B212759" box="[151,351,1771,1798]" pageId="3" pageNumber="238" type="description">
<paragraph id="D008D825BD1EFF8DFF2007D20B212759" blockId="3.[151,359,1734,1798]" box="[151,351,1771,1798]" pageId="3" pageNumber="238">
(
<figureCitation id="488CC4A0BD1EFF8DFF2807D20A96275A" box="[159,232,1771,1797]" captionStart="FIGURES 1–2" captionStartId="5.[151,264,1743,1767]" captionTargetBox="[151,1436,525,1720]" captionTargetId="figure-168@5.[151,1436,525,1720]" captionTargetPageId="5" captionText="FIGURES 1–2. Hepialodes and Iza species in dorsal (A and C) and ventral (B and D) views. 1. H. follicula, A–B: male from Brazil, Rio de Janeiro, Itatiaia (MZUSP); C–D: female from Brazil, Pará (NHMUK 010921585). 2. I. nubecula, A–B: male from Brazil, Bahia, Camacan (MWM); C–D: female from Brazil, Mato Grosso, Alto Paraguai (MWM 36.272). Scale bar 1cm." figureDoi="http://doi.org/10.5281/zenodo.4475215" httpUri="https://zenodo.org/record/4475215/files/figure.png" pageId="3" pageNumber="238">Figs 2</figureCitation>
,
<figureCitation id="488CC4A0BD1EFF8DFF4307D20B7A275A" box="[244,260,1771,1797]" captionStart="FIGURE 6" captionStartId="10.[151,250,737,761]" captionTargetBox="[151,1435,309,713]" captionTargetId="figure-67@10.[151,1435,309,713]" captionTargetPageId="10" captionText="FIGURE 6. Male genitalia of Iza nubecula from Brazil, Bahia, Camacan (Reserva Serra Bonita) (MVOB 145120). A) ventral view. B) lateral view. C) inner view of valva. D) dorsal view of tegumen and uncus. E) dorsal view of aedeagus. F) lateral view of aedeagus. G) ventral view of aedeagus. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4475221" httpUri="https://zenodo.org/record/4475221/files/figure.png" pageId="3" pageNumber="238">6</figureCitation>
,
<figureCitation id="488CC4A0BD1EFF8DFEA707D20B50275A" box="[272,302,1771,1797]" captionStart="FIGURES 9–12" captionStartId="11.[151,264,1547,1571]" captionTargetBox="[151,1435,181,1524]" captionTargetId="figure-22@11.[151,1435,181,1524]" captionTargetPageId="11" captionText="FIGURES 9–12. Female genitalia of Hepialodes, Iza, Loxiorhiza and Meskea species in lateral view. 9. H. follicula from Brazil, Goiás, Campinaçu (Barra Mansa farm) (DZ 39.216). 10. I. nubecula from Brazil, S„o Paulo, Salesópolis (Boraceia) (DZ 39.288). 11. L. cervinalis from Bolivia, Santa Cruz, Buenavista (NHMUK 010921618). 12. M. dyspteraria from El Salvador, San Salvador (MSFR). Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4475227" httpUri="https://zenodo.org/record/4475227/files/figure.png" pageId="3" pageNumber="238">10</figureCitation>
,
<figureCitation id="488CC4A0BD1EFF8DFE8D07D20B292759" box="[314,343,1771,1798]" captionStart="FIGURE 16" captionStartId="15.[151,250,2003,2027]" captionTargetBox="[177,1410,356,1970]" captionTargetId="figure-98@15.[166,1421,345,1979]" captionTargetPageId="15" captionText="FIGURE 16. Geographic distribution of Iza and Loxiorhiza species in South America." figureDoi="http://doi.org/10.5281/zenodo.4475235" httpUri="https://zenodo.org/record/4475235/files/figure.png" pageId="3" pageNumber="238">16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="98AD8BAEBD1EFF8DFF2006090F0327BE" pageId="3" pageNumber="238" type="reference_group">
<paragraph id="D008D825BD1EFF8DFF2006090F0327BE" blockId="3.[151,1437,1839,2017]" pageId="3" pageNumber="238">
<taxonomicName id="17B7A3A6BD1EFF8DFF2006090B3A2718" ID-CoL="92CLR" authority="Walker, 1865" authorityName="Walker" authorityYear="1865" box="[151,324,1839,1863]" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="E2C30437BD1EFF8DFF2006090AC82718" box="[151,182,1840,1863]" italics="true" pageId="3" pageNumber="238">Iza</emphasis>
<bibRefCitation id="B426A5D4BD1EFF8DFF0C06160B3A2718" author="Walker, F." box="[187,324,1839,1863]" pageId="3" pageNumber="238" pagination="1 - 706" refId="ref14761" refString="Walker, F. (1865) Catologue of Lepidoptera Heterocera. List of the specimens of Lepidopterous insects in the Collection of the British Museum, 31 (7), 1 - 706." type="journal article" year="1865">Walker, 1865</bibRefCitation>
</taxonomicName>
.
<emphasis id="E2C30437BD1EFF8DFEFC061609B52718" bold="true" box="[331,971,1839,1863]" pageId="3" pageNumber="238">Lepidopterous insects in the Collection of the British, 31 (70)</emphasis>
: 521; species included:
<taxonomicName id="17B7A3A6BD1EFF8DFB0C06090B7B2739" ID-CoL="3QHS6" authority="(Guenee, 1858)" baseAuthorityName="Guenee" baseAuthorityYear="1858" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="238" phylum="Arthropoda" rank="species" species="nubecula">
<emphasis id="E2C30437BD1EFF8DFB0C06090F452718" box="[1211,1339,1839,1863]" italics="true" pageId="3" pageNumber="238">Iza nubecula</emphasis>
(
<bibRefCitation id="B426A5D4BD1EFF8DFAF006160A832739" author="Guenee, M. A." pageId="3" pageNumber="238" pagination="1 - 8" refId="ref13536" refString="Guenee, M. A. (1858) s. n. In: Boisduval M. M. &amp; Guenee, M. A., Histoire naturelle des Insectes. Species general des Lepidopteres, Atlas, Uranides, Phalenites, Siculides, Siculides Planche 1. Figs. 1 - 8. Librairie Encyclopedique de Roret, Paris, pp. 58." type="book chapter" year="1858">Guenée, 1858</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="17B7A3A6BD1EFF8DFEA7067608562739" ID-CoL="979Z4" authority="Walker, 1865" authorityName="Walker" authorityYear="1865" box="[272,552,1870,1894]" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="238" phylum="Arthropoda" rank="species" species="botydana">
<emphasis id="E2C30437BD1EFF8DFEA706760BEB2739" box="[272,405,1870,1894]" italics="true" pageId="3" pageNumber="238">Iza botydana</emphasis>
<bibRefCitation id="B426A5D4BD1EFF8DFE2A067708562739" author="Walker, F." box="[413,552,1870,1894]" pageId="3" pageNumber="238" pagination="1 - 706" refId="ref14761" refString="Walker, F. (1865) Catologue of Lepidoptera Heterocera. List of the specimens of Lepidopterous insects in the Collection of the British Museum, 31 (7), 1 - 706." type="journal article" year="1865">Walker, 1865</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="17B7A3A6BD1EFF8DFD830676082D2739" ID-CoL="92CLR" authorityName="Walker" authorityYear="1865" box="[564,595,1871,1894]" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="E2C30437BD1EFF8DFD830676082D2739" box="[564,595,1871,1894]" italics="true" pageId="3" pageNumber="238">Iza</emphasis>
</taxonomicName>
?
<emphasis id="E2C30437BD1EFF8DFDDA067708AA2739" box="[621,724,1870,1894]" italics="true" pageId="3" pageNumber="238">terminalis</emphasis>
<bibRefCitation id="B426A5D4BD1EFF8DFD6B067709182739" author="Walker, F." box="[732,870,1870,1894]" pageId="3" pageNumber="238" pagination="1 - 706" refId="ref14761" refString="Walker, F. (1865) Catologue of Lepidoptera Heterocera. List of the specimens of Lepidopterous insects in the Collection of the British Museum, 31 (7), 1 - 706." type="journal article" year="1865">Walker, 1865</bibRefCitation>
,
<taxonomicName id="17B7A3A6BD1EFF8DFCC506760EF62739" authority="(Guenee, 1858)" baseAuthorityName="Guenee" baseAuthorityYear="1858" box="[882,1160,1870,1894]" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="238" phylum="Arthropoda" rank="species" species="perlula">
<emphasis id="E2C30437BD1EFF8DFCC50676099C2739" box="[882,994,1870,1894]" italics="true" pageId="3" pageNumber="238">Iza perlula</emphasis>
(
<bibRefCitation id="B426A5D4BD1EFF8DFC4606770EFF2739" author="Guenee, M. A." box="[1009,1153,1870,1894]" pageId="3" pageNumber="238" pagination="1 - 8" refId="ref13536" refString="Guenee, M. A. (1858) s. n. In: Boisduval M. M. &amp; Guenee, M. A., Histoire naturelle des Insectes. Species general des Lepidopteres, Atlas, Uranides, Phalenites, Siculides, Siculides Planche 1. Figs. 1 - 8. Librairie Encyclopedique de Roret, Paris, pp. 58." type="book chapter" year="1858">Guenée, 1858</bibRefCitation>
)
</taxonomicName>
.—
<bibRefCitation id="B426A5D4BD1EFF8DFB1106770F442739" author="Guenee, A." box="[1190,1338,1870,1894]" pageId="3" pageNumber="238" pagination="275 - 304" refId="ref13603" refString="Guenee, A. (1877) Monographie de la famille des Siculides. Annales de la Societe Entomologique de France, 7 (5), 275 - 304." type="journal article" year="1877">Guenée, 1877</bibRefCitation>
.
<emphasis id="E2C30437BD1EFF8DFAF2067708EF27DB" bold="true" pageId="3" pageNumber="238">
Annales de la Société Entomologique de
<collectingCountry id="A8A098B5BD1EFF8DFDAC0654081627DA" box="[539,616,1901,1925]" name="France" pageId="3" pageNumber="238">France</collectingCountry>
, (5)
</emphasis>
7: 301.—Druce, 1895,
<emphasis id="E2C30437BD1EFF8DFCC9065409EC27DB" box="[894,914,1901,1924]" italics="true" pageId="3" pageNumber="238">in</emphasis>
Godman &amp; Salvin.
<emphasis id="E2C30437BD1EFF8DFBEC0655083E27FC" bold="true" pageId="3" pageNumber="238">Biologia Centrali-Americana: zoology, botany and archaeology 2</emphasis>
: 186;
<typeStatus id="0F0C6687BD1EFF8DFD3306B508CE27FB" box="[644,688,1932,1956]" pageId="3" pageNumber="238">type</typeStatus>
species designation
<taxonomicName id="17B7A3A6BD1EFF8DFC3206B20E9E27FC" ID-CoL="98CLQ" authority="Guenee, 1858" authorityName="Guenee" authorityYear="1858" box="[901,1248,1931,1955]" class="Insecta" family="Thyrididae" genus="Siculodes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="238" phylum="Arthropoda" rank="species" species="nubecula">
<emphasis id="E2C30437BD1EFF8DFC3206B20E3527FC" box="[901,1099,1931,1955]" italics="true" pageId="3" pageNumber="238">Siculodes nubecula</emphasis>
<bibRefCitation id="B426A5D4BD1EFF8DFBE406B20E9E27FC" author="Guenee, M. A." box="[1107,1248,1931,1955]" pageId="3" pageNumber="238" pagination="1 - 8" refId="ref13536" refString="Guenee, M. A. (1858) s. n. In: Boisduval M. M. &amp; Guenee, M. A., Histoire naturelle des Insectes. Species general des Lepidopteres, Atlas, Uranides, Phalenites, Siculides, Siculides Planche 1. Figs. 1 - 8. Librairie Encyclopedique de Roret, Paris, pp. 58." type="book chapter" year="1858">Guenée, 1858</bibRefCitation>
</taxonomicName>
.—
<bibRefCitation id="B426A5D4BD1EFF8DFB4906B20FE727FC" author="Whalley, P. E. S." box="[1278,1433,1931,1955]" pageId="3" pageNumber="238" pagination="115 - 127" refId="ref14824" refString="Whalley, P. E. S. (1964) Catalogue of the world genera of the Thyrididae (Lepidoptera) with type selection and synonymy. Annals and Magazine of Natural History, Series 13, 7 (74), 115 - 127. https: // doi. org / 10.1080 / 00222936408651447" type="journal article" year="1964">Whalley, 1964</bibRefCitation>
.
<emphasis id="E2C30437BD1EFF8DFF70069308B3279D" bold="true" box="[199,717,1961,1986]" pageId="3" pageNumber="238">Annals and Magazine of Natural History, (13) 7</emphasis>
(74): 121.—
<bibRefCitation id="B426A5D4BD1EFF8DFCF906930E1C279D" author="Whalley, P. E. S. &amp; Heppner, J. B." box="[846,1122,1962,1986]" pageId="3" pageNumber="238" pagination="31 - 34" refId="ref14979" refString="Whalley, P. E. S. &amp; Heppner, J. B. (1995) Thyridoidea. 59. Thyrididae. In: Becker, V. O., Brown, J. W., Brown, R. L., Davis, D. R., Donahue, J. P., Eichlin, T. D., Epstein, M. E., Gielis, C., Heppner, J. B., Miller, S. E., Munroe, E. G., Powell, J. A., Razowski, J., Schaffer, J. C., Shaffer, M., Solis, M. A., Tarmann, G. M. &amp; Whalley, P. E. S. (Eds.), Checklist: Part 2, Hyblaeoidea-Pyraloidea Tortricoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Vol. 3, Gainesville, Washington, Hamburg, Lima, Taipei, Tokyo, pp. 31 - 34." type="book chapter" year="1995">Whalley &amp; Heppner, 1995</bibRefCitation>
,
<emphasis id="E2C30437BD1EFF8DFBDA06930E8D279D" bold="true" box="[1133,1267,1962,1986]" pageId="3" pageNumber="238">Thyridoidea</emphasis>
,
<emphasis id="E2C30437BD1EFF8DFAB706930F6A279E" box="[1280,1300,1962,1985]" italics="true" pageId="3" pageNumber="238">in</emphasis>
Becker
<emphasis id="E2C30437BD1EFF8DFADE06920FE3279D" box="[1385,1437,1962,1986]" italics="true" pageId="3" pageNumber="238">et al.</emphasis>
Checklist: Part 2,
<emphasis id="E2C30437BD1EFF8DFECA06F1096627BF" bold="true" box="[381,792,1992,2016]" pageId="3" pageNumber="238">Hyblaeoidea–Pyraloidea–Tortricoidea</emphasis>
,
<emphasis id="E2C30437BD1EFF8DFC9206F0094727BF" box="[805,825,1993,2016]" italics="true" pageId="3" pageNumber="238">in</emphasis>
Heppner (ed.).
<emphasis id="E2C30437BD1EFF8DFC6F06F10F2E27BF" bold="true" box="[984,1360,1992,2016]" pageId="3" pageNumber="238">Atlas of Neotropical Lepidoptera 3</emphasis>
: 32.
</paragraph>
</subSubSection>
<subSubSection id="98AD8BAEBD19FF8AFF2001AE098F21FE" pageId="4" pageNumber="239" type="type_taxon">
<paragraph id="D008D825BD19FF8AFF2001AE0B9A20B3" blockId="4.[151,1436,150,236]" pageId="4" pageNumber="239">
<emphasis id="E2C30437BD19FF8AFF2001AE0B6320F1" bold="true" box="[151,285,150,175]" pageId="4" pageNumber="239">
<typeStatus id="0F0C6687BD19FF8AFF2001AE0AB220F0" box="[151,204,151,175]" pageId="4" pageNumber="239">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="17B7A3A6BD19FF8AFE9E01AE080020F1" ID-CoL="98CLQ" authority="Guenee, 1858" authorityName="Guenee" authorityYear="1858" box="[297,638,150,175]" class="Insecta" family="Thyrididae" genus="Siculodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="239" phylum="Arthropoda" rank="species" species="nubecula">
<emphasis id="E2C30437BD19FF8AFE9E01AE0B9220F0" box="[297,492,151,175]" italics="true" pageId="4" pageNumber="239">Siculodes nubecula</emphasis>
<bibRefCitation id="B426A5D4BD19FF8AFE4501AE080020F1" author="Guenee, M. A." box="[498,638,150,175]" pageId="4" pageNumber="239" pagination="1 - 8" refId="ref13536" refString="Guenee, M. A. (1858) s. n. In: Boisduval M. M. &amp; Guenee, M. A., Histoire naturelle des Insectes. Species general des Lepidopteres, Atlas, Uranides, Phalenites, Siculides, Siculides Planche 1. Figs. 1 - 8. Librairie Encyclopedique de Roret, Paris, pp. 58." type="book chapter" year="1858">Guenée, 1858</bibRefCitation>
</taxonomicName>
by subsequent designation (
<bibRefCitation id="B426A5D4BD19FF8AFC2E01AE0E5F20F0" author="Guenee, A." box="[921,1057,151,175]" pageId="4" pageNumber="239" pagination="275 - 304" refId="ref13603" refString="Guenee, A. (1877) Monographie de la famille des Siculides. Annales de la Societe Entomologique de France, 7 (5), 275 - 304." type="journal article" year="1877">Guenée 1877</bibRefCitation>
) (“Cette espèce deviendra par la suite le
<typeStatus id="0F0C6687BD19FF8AFF56018F0B732091" box="[225,269,182,206]" pageId="4" pageNumber="239">type</typeStatus>
d’un genre (
<taxonomicName id="17B7A3A6BD19FF8AFE25018F0B882092" ID-CoL="92CLR" authority="Walk." authorityName="Walker" authorityYear="1865" box="[402,502,181,205]" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="E2C30437BD19FF8AFE25018F0BCF2092" box="[402,433,182,205]" italics="true" pageId="4" pageNumber="239">Iza</emphasis>
Walk.
</taxonomicName>
) que je n’ai pas cru devoir adopter dans l’état imparfait de la science”). This fact was later confirmed by Druce (1895).
</paragraph>
<paragraph id="D008D825BD19FF8AFF200023098F21FE" blockId="4.[151,1437,282,1101]" pageId="4" pageNumber="239">
<emphasis id="E2C30437BD19FF8AFF2000230B0B216B" bold="true" box="[151,373,282,308]" pageId="4" pageNumber="239">Taxonomic history</emphasis>
.
<taxonomicName id="17B7A3A6BD19FF8AFE3500220BDA216B" ID-CoL="92CLR" authorityName="Walker" authorityYear="1865" box="[386,420,283,308]" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="E2C30437BD19FF8AFE3500220BDA216B" box="[386,420,283,308]" italics="true" pageId="4" pageNumber="239">Iza</emphasis>
</taxonomicName>
was described to include the following species:
<taxonomicName id="17B7A3A6BD19FF8AFC0B00220E44216B" ID-CoL="98CLQ" authorityName="Guenee" authorityYear="1858" box="[956,1082,282,308]" class="Insecta" family="Thyrididae" genus="Siculodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="239" phylum="Arthropoda" rank="species" species="nubecula">
<emphasis id="E2C30437BD19FF8AFC0B00220E44216B" box="[956,1082,282,308]" italics="true" pageId="4" pageNumber="239">I. nubecula</emphasis>
</taxonomicName>
,
<taxonomicName id="17B7A3A6BD19FF8AFBF000220EBB216B" ID-CoL="979Z4" authorityName="Walker" authorityYear="1865" box="[1095,1221,282,308]" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="239" phylum="Arthropoda" rank="species" species="botydana">
<emphasis id="E2C30437BD19FF8AFBF000220EBB216B" box="[1095,1221,282,308]" italics="true" pageId="4" pageNumber="239">I. botydana</emphasis>
</taxonomicName>
,
<emphasis id="E2C30437BD19FF8AFB6400220E9A216B" box="[1235,1252,283,308]" italics="true" pageId="4" pageNumber="239">I.</emphasis>
?
<emphasis id="E2C30437BD19FF8AFB4B00230F10216B" box="[1276,1390,282,308]" italics="true" pageId="4" pageNumber="239">terminalis</emphasis>
and
<taxonomicName id="17B7A3A6BD19FF8AFF2000060B7E2107" baseAuthorityName="Guenee" baseAuthorityYear="1858" box="[151,256,318,344]" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="239" phylum="Arthropoda" rank="species" species="perlula">
<emphasis id="E2C30437BD19FF8AFF2000060B7E2107" box="[151,256,318,344]" italics="true" pageId="4" pageNumber="239">I. perlula</emphasis>
</taxonomicName>
, but without designation of the
<typeStatus id="0F0C6687BD19FF8AFDDD000608E42106" box="[618,666,319,345]" pageId="4" pageNumber="239">type</typeStatus>
species. Currently, the only species in the genus is the
<typeStatus id="0F0C6687BD19FF8AFABB00060F422106" box="[1292,1340,319,345]" pageId="4" pageNumber="239">type</typeStatus>
species.
<taxonomicName id="17B7A3A6BD19FF8AFF20005A0B572123" ID-CoL="979Z4" authorityName="Walker" authorityYear="1865" box="[151,297,354,380]" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="239" phylum="Arthropoda" rank="species" species="botydana">
<emphasis id="E2C30437BD19FF8AFF20005A0B572123" box="[151,297,354,380]" italics="true" pageId="4" pageNumber="239">Iza botydana</emphasis>
</taxonomicName>
was transferred by
<bibRefCitation id="B426A5D4BD19FF8AFDB0005B09432122" author="Whalley, P. E. S. &amp; Heppner, J. B." box="[519,829,354,381]" pageId="4" pageNumber="239" pagination="31 - 34" refId="ref14979" refString="Whalley, P. E. S. &amp; Heppner, J. B. (1995) Thyridoidea. 59. Thyrididae. In: Becker, V. O., Brown, J. W., Brown, R. L., Davis, D. R., Donahue, J. P., Eichlin, T. D., Epstein, M. E., Gielis, C., Heppner, J. B., Miller, S. E., Munroe, E. G., Powell, J. A., Razowski, J., Schaffer, J. C., Shaffer, M., Solis, M. A., Tarmann, G. M. &amp; Whalley, P. E. S. (Eds.), Checklist: Part 2, Hyblaeoidea-Pyraloidea Tortricoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Vol. 3, Gainesville, Washington, Hamburg, Lima, Taipei, Tokyo, pp. 31 - 34." type="book chapter" year="1995">Whalley &amp; Heppner (1995)</bibRefCitation>
to
<taxonomicName id="17B7A3A6BD19FF8AFCD6005B0ED62123" ID-CoL="92KYY" authority="Kohler, 1924" authorityName="Kohler" authorityYear="1924" box="[865,1192,354,380]" class="Insecta" family="Thyrididae" genus="Pseudendromis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="E2C30437BD19FF8AFCD6005B0E732123" box="[865,1037,354,380]" italics="true" pageId="4" pageNumber="239">Pseudendromis</emphasis>
<bibRefCitation id="B426A5D4BD19FF8AFBA3005B0ED62123" author="Kohler, P." box="[1044,1192,354,380]" pageId="4" pageNumber="239" pagination="1 - 28" refId="ref13919" refString="Kohler, P. (1924) Fauna Argentina. Lepidoptera e collectiohe Alberto Breyer. II. Teil, Heterocera: Systematischer Katalog und Studicn Berichtigungen, Neubcschreibungen. Zeitschrift fur wissenschaftliche Insektenbiologie, Berlin, Supplement 19, 1 - 28." type="journal article" year="1924">Köhler, 1924</bibRefCitation>
</taxonomicName>
as
<emphasis id="E2C30437BD19FF8AFB79005A0F132123" box="[1230,1389,354,381]" italics="true" pageId="4" pageNumber="239">I.? terminalis</emphasis>
and
<taxonomicName id="17B7A3A6BD19FF8AFF2000BE0B7E21FF" baseAuthorityName="Guenee" baseAuthorityYear="1858" box="[151,256,390,416]" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="239" phylum="Arthropoda" rank="species" species="perlula">
<emphasis id="E2C30437BD19FF8AFF2000BE0B7E21FF" box="[151,256,390,416]" italics="true" pageId="4" pageNumber="239">I. perlula</emphasis>
</taxonomicName>
was transferred by
<bibRefCitation id="B426A5D4BD19FF8AFE6B00BE08E721FF" author="Hampson, G. F." box="[476,665,390,417]" pageId="4" pageNumber="239" pagination="603 - 633" refId="ref13634" refString="Hampson, G. F. (1897) On the Classification of the Thyrididae- a Family of the Lepidoptera Phalaenae. Proceedings of the Zoological Society of London, 1897 (1), 603 - 633, fig. 24. https: // doi. org / 10.1111 / j. 1096 - 3642.1897. tb 03111. x" type="journal article" year="1897">Hampson (1897)</bibRefCitation>
to
<taxonomicName id="17B7A3A6BD19FF8AFD0A00BF099321FE" ID-CoL="92LWS" authority="Guenee, 1858" authorityName="Guenee" authorityYear="1858" box="[701,1005,390,417]" class="Insecta" family="Thyrididae" genus="Rhodoneura" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="E2C30437BD19FF8AFD0A00BF093921FF" box="[701,839,390,416]" italics="true" pageId="4" pageNumber="239">Rhodoneura</emphasis>
<bibRefCitation id="B426A5D4BD19FF8AFCF900BF099321FE" author="Guenee, M. A." box="[846,1005,390,417]" pageId="4" pageNumber="239" pagination="1 - 8" refId="ref13536" refString="Guenee, M. A. (1858) s. n. In: Boisduval M. M. &amp; Guenee, M. A., Histoire naturelle des Insectes. Species general des Lepidopteres, Atlas, Uranides, Phalenites, Siculides, Siculides Planche 1. Figs. 1 - 8. Librairie Encyclopedique de Roret, Paris, pp. 58." type="book chapter" year="1858">Guenée, 1858</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="98AD8BAEBD19FF8AFF7000930FF6226F" pageId="4" pageNumber="239" type="diagnosis">
<paragraph id="D008D825BD19FF8AFF7000930FF6226F" blockId="4.[151,1437,282,1101]" pageId="4" pageNumber="239">
<emphasis id="E2C30437BD19FF8AFF7000930B44219B" bold="true" box="[199,314,426,452]" pageId="4" pageNumber="239">Diagnosis</emphasis>
.
<taxonomicName id="17B7A3A6BD19FF8AFEFF00920B14219B" authorityName="Walker" authorityYear="1865" box="[328,362,427,452]" class="Insecta" family="Thyrididae" genus="Iza" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="E2C30437BD19FF8AFEFF00920B14219B" box="[328,362,427,452]" italics="true" pageId="4" pageNumber="239">Iza</emphasis>
</taxonomicName>
is distinguished from the remaining genera of
<taxonomicName id="17B7A3A6BD19FF8AFCC100930997219B" box="[886,1001,426,452]" class="Insecta" family="Thyrididae" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="239" phylum="Arthropoda" rank="tribe" tribe="Siculodini">Siculodini</taxonomicName>
by: (i) wings dark brown ground color with irregular beige patches and bands dorsal and ventrally (
<figureCitation id="488CC4A0BD19FF8AFC8200F7090721B7" box="[821,889,462,488]" captionStart="FIGURES 1–2" captionStartId="5.[151,264,1743,1767]" captionTargetBox="[151,1436,525,1720]" captionTargetId="figure-168@5.[151,1436,525,1720]" captionTargetPageId="5" captionText="FIGURES 1–2. Hepialodes and Iza species in dorsal (A and C) and ventral (B and D) views. 1. H. follicula, A–B: male from Brazil, Rio de Janeiro, Itatiaia (MZUSP); C–D: female from Brazil, Pará (NHMUK 010921585). 2. I. nubecula, A–B: male from Brazil, Bahia, Camacan (MWM); C–D: female from Brazil, Mato Grosso, Alto Paraguai (MWM 36.272). Scale bar 1cm." figureDoi="http://doi.org/10.5281/zenodo.4475215" httpUri="https://zenodo.org/record/4475215/files/figure.png" pageId="4" pageNumber="239">Fig. 2</figureCitation>
); (ii) abdomen dark brown on the dorsal face and beige on the ventral face; (iii) valva triangular; (iv) valva with spiniform process; (v) fultura inferior U-shaped; (vi) aedeagus narrowing distally with spines; (vii) lamella antevaginalis sclerotized and (viii) papillae anales straight.
</paragraph>
</subSubSection>
<subSubSection id="98AD8BAEBD19FF8AFF7003030FF82412" pageId="4" pageNumber="239" type="description">
<paragraph id="D008D825BD19FF8AFF7003030B0D2227" blockId="4.[151,1437,282,1101]" pageId="4" pageNumber="239">
<emphasis id="E2C30437BD19FF8AFF7003030B13220B" bold="true" box="[199,365,570,596]" pageId="4" pageNumber="239">Redescription</emphasis>
. Forewing: length—males 16.6 mm (n = 11), females 25.4 mm (n = 5); slightly concave tornus; retinaculum absent.
</paragraph>
<paragraph id="D008D825BD19FF8AFF7003BB0E7C229F" blockId="4.[151,1437,282,1101]" pageId="4" pageNumber="239">Hindwing: length—males 12.6 mm (n = 11), females 15.8 mm (n = 5); apex right angled and tornus concave; male with sand-colored patch on the costal margin ventrally; frenulum absent.</paragraph>
<paragraph id="D008D825BD19FF8AFF7003F30BD723E2" blockId="4.[151,1437,282,1101]" pageId="4" pageNumber="239">
Male genitalia (
<figureCitation id="488CC4A0BD19FF8AFEC003F30BC222BA" box="[375,444,714,741]" captionStart="FIGURE 6" captionStartId="10.[151,250,737,761]" captionTargetBox="[151,1435,309,713]" captionTargetId="figure-67@10.[151,1435,309,713]" captionTargetPageId="10" captionText="FIGURE 6. Male genitalia of Iza nubecula from Brazil, Bahia, Camacan (Reserva Serra Bonita) (MVOB 145120). A) ventral view. B) lateral view. C) inner view of valva. D) dorsal view of tegumen and uncus. E) dorsal view of aedeagus. F) lateral view of aedeagus. G) ventral view of aedeagus. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4475221" httpUri="https://zenodo.org/record/4475221/files/figure.png" pageId="4" pageNumber="239">Fig. 6</figureCitation>
): tegumen with proximal and distal margins convex in dorsal view; saccus oval in lateral view; uncus triangular in dorsal view, with thin and short setae; gnathos reduced and distally sclerotized, projected disto-ventrally; valva triangular, costa slightly concave, spiniform process in the proximal part, clasper projected dorsally, apex rounded and ventral margin convex, slightly concave at the posterior end of sacculus; fultura inferior U-shaped, broad and thickly sclerotized arms with blunt apex and short setae; aedeagus cylindrical, proximal half thicker, narrowing distally with numerous spines, opening for the vesica extending along the entire posterior half, vesica without cornutus.
</paragraph>
<paragraph id="D008D825BD19FF8AFF7002FF0FF82412" blockId="4.[151,1437,282,1101]" pageId="4" pageNumber="239">
Female genitalia (
<figureCitation id="488CC4A0BD19FF8AFE2202FF0B9223BF" box="[405,492,966,992]" captionStart="FIGURES 9–12" captionStartId="11.[151,264,1547,1571]" captionTargetBox="[151,1435,181,1524]" captionTargetId="figure-22@11.[151,1435,181,1524]" captionTargetPageId="11" captionText="FIGURES 9–12. Female genitalia of Hepialodes, Iza, Loxiorhiza and Meskea species in lateral view. 9. H. follicula from Brazil, Goiás, Campinaçu (Barra Mansa farm) (DZ 39.216). 10. I. nubecula from Brazil, S„o Paulo, Salesópolis (Boraceia) (DZ 39.288). 11. L. cervinalis from Bolivia, Santa Cruz, Buenavista (NHMUK 010921618). 12. M. dyspteraria from El Salvador, San Salvador (MSFR). Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4475227" httpUri="https://zenodo.org/record/4475227/files/figure.png" pageId="4" pageNumber="239">Fig. 10</figureCitation>
): tergum VIII rectangular in lateral view, lamella antevaginalis sclerotized, ostium membranous; posterior apophyses longer than the anterior apophyses; bursa copulatrix with sclerotized wide ductus at anterior 1/4, second 1/4 membranous and narrow, and posterior half membranous and slightly thicker, corpus bursae membranous, without signum; papillae anales straight, taller than long and covered with short and thin setae.
</paragraph>
</subSubSection>
</treatment>
</document>
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