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<document id="13645E95D9C4DE67D5EB678FF8BF3E6E" ID-DOI="10.5852/ejt.2024.956.2657" ID-ISSN="2118-9773" ID-Zenodo-Dep="13742850" ID-ZooBank="9DCDD521-909D-4436-8495-086A11C50DA0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1725988271530" checkinUser="plazi" docAuthor="Karimbumkara, Seena Narayanan &amp; Priyadarsanan, Dharma Rajan" docDate="2024" docId="557187D2FFFE890C9D72FEB8FABFFCFC" docLanguage="en" docName="EJT.2024.956.1-61.pdf" docOrigin="European Journal of Taxonomy 956" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onitis bhomorensis Karimbumkara &amp; Priyadarsanan, 2024, sp. nov." docType="treatment" docUuid="7D70D754-D82E-4BCA-A3C9-D42645B4F708" docUuidSource="ZooBank" docVersion="3" lastPageNumber="11" masterDocId="A948FFAAFFF689069F01FFB1FFF9FF8A" masterDocTitle="Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India" masterLastPageNumber="61" masterPageNumber="1" pageNumber="9" updateTime="1726236628153" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="3F244A9760481625BFB9FC3E111B6F0A">
<mods:title id="9A7A09E0D5E144E864AE82A2255E74A8">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
</mods:titleInfo>
<mods:name id="F739D3C3618FA8C4F9380E4140360AA8" type="personal">
<mods:role id="5B07D1EA938F74E56929D0C5E45D82CD">
<mods:roleTerm id="977582EDFC64DCECA4CFA82A437748B6">Author</mods:roleTerm>
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<mods:namePart id="57A2DA8A8452268B407ABF99A12D6A8F">Karimbumkara, Seena Narayanan</mods:namePart>
<mods:nameIdentifier id="AC015DA4F5449CCD0D3B0B338FBFD1E7" type="LSID">A6B50103-8D6D-4977-9111-EF8BA4AA0477</mods:nameIdentifier>
<mods:affiliation id="95A0294A4910C191A7B9F12C8C9B7179">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
<mods:nameIdentifier id="2386AC76E13A4C3ADF708EF10AC8C2E3" type="email">seena.narayanan@atree.org</mods:nameIdentifier>
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<mods:name id="AE087C38889CBEA125BF6D4728228D8A" type="personal">
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<mods:namePart id="6F53CEF63D475912CC08D8B4F2108639">Priyadarsanan, Dharma Rajan</mods:namePart>
<mods:nameIdentifier id="04A98171E06FFFA51942F834ABF5E707" type="LSID">03907E93-F783-4720-9F7B-CB662E7FC745</mods:nameIdentifier>
<mods:affiliation id="446B42B0C5667FBA4997267B795C3865">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
<mods:nameIdentifier id="8A0E870F7535F70C86A5EFB974C14D59" type="email">priyan@atree.org</mods:nameIdentifier>
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<mods:typeOfResource id="6A3B10FA711407DF239EBF8D01B4C274">text</mods:typeOfResource>
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<mods:title id="FC6BCF589975DAECD63B707364811CDF">European Journal of Taxonomy</mods:title>
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<mods:part id="F8DCADAD659C739F914CD07FD28045A2">
<mods:date id="DD1D50FBD5A4262EE1529DA438F6DDF0">2024</mods:date>
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<mods:number id="89853AA8F34646DD0C03B1DB6B1F6AC9">2024-09-10</mods:number>
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<mods:detail id="7BED3BA85FB70DFD8CBA8D69EDB75C6C" type="volume">
<mods:number id="E9C35BE8CCA3458867D45F89D952EA78">956</mods:number>
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<mods:start id="1E789E13DF451220B261CFC6BD7B5DF6">1</mods:start>
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<mods:identifier id="2AD02EF6A2CF56890C154D8E35E8B0EC" type="DOI">10.5852/ejt.2024.956.2657</mods:identifier>
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<treatment id="557187D2FFFE890C9D72FEB8FABFFCFC" LSID="urn:lsid:zoobank.org:act:7D70D754-D82E-4BCA-A3C9-D42645B4F708" httpUri="http://treatment.plazi.org/id/557187D2FFFE890C9D72FEB8FABFFCFC" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="95C2654FFFFE890E9D72FEB8FC38FEA9" box="[627,961,264,291]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="DD6736C4FFFE890E9D72FEB8FC38FEA9" blockId="8.[361,1226,264,362]" box="[627,961,264,291]" pageId="8" pageNumber="9">
<heading id="862F81A8FFFE890E9D72FEB8FC38FEA9" box="[627,961,264,291]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
<taxonomicName id="1AD84D47FFFE890E9D72FEB8FCA6FEA9" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[627,863,264,291]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bhomorensis" status="sp. nov.">
<emphasis id="EFACEAD6FFFE890E9D72FEB8FCA6FEA9" bold="true" box="[627,863,264,291]" italics="true" pageId="8" pageNumber="9">Onitis bhomorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFFE890E9C67FEB8FC38FEA9" box="[870,961,265,291]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFE890E9E68FE9DFC0BFEE3" pageId="8" pageNumber="9" type="description">
<paragraph id="DD6736C4FFFE890E9E68FE9DFB33FECD" blockId="8.[361,1226,264,362]" box="[361,1226,300,327]" pageId="8" pageNumber="9">
<uri id="A9493AC6FFFE890E9E68FE9DFB33FECD" box="[361,1226,300,327]" pageId="8" pageNumber="9">
urn:lsid:zoobank.org:act:
<uuid id="A97E0C11FFFE890E9D86FE9DFB33FECD" box="[647,1226,300,327]" pageId="8" pageNumber="9">7D70D754-D82E-4BCA-A3C9-D42645B4F708</uuid>
</uri>
</paragraph>
<paragraph id="DD6736C4FFFE890E9D40FEFEFC0BFEE3" blockId="8.[361,1226,264,362]" box="[577,1010,335,362]" pageId="8" pageNumber="9">
<figureCitation id="45E32A41FFFE890E9D40FEFEFD72FEE3" box="[577,651,335,361]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[245,1340,356,1841]" captionTargetId="figure-1@9.[189,1398,265,1856]" captionTargetPageId="9" captionText="Fig. 2. Onitisbhomorensis sp. nov., holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head and prontum. D. Protibia, arrow showing rudimentary spur. E. Metasternum. F. Pygidium, arrow showing longitudinal groove. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742855" httpUri="https://zenodo.org/record/13742855/files/figure.png" pageId="8" pageNumber="9">Figs 2</figureCitation>
,
<figureCitation id="45E32A41FFFE890E9D99FEFEFD2AFEE3" box="[664,723,335,361]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. E–H. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. I–L. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. M–P. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742867" httpUri="https://zenodo.org/record/13742867/files/figure.png" pageId="8" pageNumber="9">6I–L</figureCitation>
,
<figureCitation id="45E32A41FFFE890E9DDEFEFEFCFFFEE3" box="[735,774,335,361]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="8" pageNumber="9">9A</figureCitation>
,
<figureCitation id="45E32A41FFFE890E9C13FEFEFCBEFEE3" box="[786,839,335,361]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="8" pageNumber="9">14A</figureCitation>
,
<figureCitation id="45E32A41FFFE890E9C52FEFEFC71FEE0" box="[851,904,335,362]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742895" httpUri="https://zenodo.org/record/13742895/files/figure.png" pageId="8" pageNumber="9">15A</figureCitation>
,
<figureCitation id="45E32A41FFFE890E9C95FEFEFC33FEE3" box="[916,970,335,361]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="8" pageNumber="9">16A</figureCitation>
,
<figureCitation id="45E32A41FFFE890E9CD4FEFEFC0BFEE3" box="[981,1010,335,361]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742901" httpUri="https://zenodo.org/record/13742901/files/figure.png" pageId="8" pageNumber="9">17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFE890E9FBCFE21FAC4FC0A" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="DD6736C4FFFE890E9FBCFE21FE3FFE20" blockId="8.[189,454,400,426]" box="[189,454,400,426]" pageId="8" pageNumber="9">
<heading id="862F81A8FFFE890E9FBCFE21FE3FFE20" bold="true" box="[189,454,400,426]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EFACEAD6FFFE890E9FBCFE21FE3FFE20" bold="true" box="[189,454,400,426]" pageId="8" pageNumber="9">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFE890E9FBCFE0EFAC4FC0A" blockId="8.[189,1399,447,896]" pageId="8" pageNumber="9">
<emphasis id="EFACEAD6FFFE890E9FBCFE0EFE58FE53" box="[189,417,447,473]" italics="true" pageId="8" pageNumber="9">Onitisbhomorensis</emphasis>
<taxonomicNameLabel id="F49F57ADFFFE890E9EA6FE71FDFBFE50" box="[423,514,448,474]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
is assigned to Group I as its body is completely black and the frontal carina is interrupted. It is similar to
<taxonomicName id="1AD84D47FFFE890E9D14FE52FD01FE76" authorityName="Janssens" authorityYear="1937" box="[533,760,483,509]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFFE890E9D14FE52FD01FE76" box="[533,760,483,509]" italics="true" pageId="8" pageNumber="9">O. punctatostriatus</emphasis>
</taxonomicName>
in having broad elytral striae. It differs from the latter in the nature of pronotal punctures which are stronger in
<taxonomicName id="1AD84D47FFFE890E9C6AFDB7FBDCFDAA" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[875,1061,518,544]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bhomorensis">
<emphasis id="EFACEAD6FFFE890E9C6AFDB7FBDCFDAA" box="[875,1061,518,544]" italics="true" pageId="8" pageNumber="9">O. bhomorensis</emphasis>
</taxonomicName>
than in
<taxonomicName id="1AD84D47FFFE890E9B8BFDB7FA96FDAA" authorityName="Janssens" authorityYear="1937" box="[1162,1391,518,544]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFFE890E9B8BFDB7FA96FDAA" box="[1162,1391,518,544]" italics="true" pageId="8" pageNumber="9">O. punctatostriatus</emphasis>
</taxonomicName>
; basal foveae are more spaced in the former than the latter; metasternal shield slightly convex, granular anteriorly with reddish hairs in
<taxonomicName id="1AD84D47FFFE890E9D2CFDFCFD1CFDEC" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[557,741,588,615]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bhomorensis">
<emphasis id="EFACEAD6FFFE890E9D2CFDFCFD1CFDEC" box="[557,741,588,615]" italics="true" pageId="8" pageNumber="9">O. bhomorensis</emphasis>
</taxonomicName>
while in
<taxonomicName id="1AD84D47FFFE890E9C53FDFCFBCCFDEC" authorityName="Janssens" authorityYear="1937" box="[850,1077,589,615]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFFE890E9C53FDFCFBCCFDEC" box="[850,1077,589,615]" italics="true" pageId="8" pageNumber="9">O. punctatostriatus</emphasis>
</taxonomicName>
it is flat in the middle, with a weak longitudinal groove and with smooth scattered punctures. Clypeal margin truncated, straight in the middle in the former, while it is slightly reflexed in the latter. The presence of a median groove on the pygidium and the rudimentary spur on the protibia make it different from all other Oriental
<taxonomicName id="1AD84D47FFFE890E9A2EFD07FEE0FD79" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EFACEAD6FFFE890E9A2EFD07FA8EFD5A" box="[1327,1399,694,720]" italics="true" pageId="8" pageNumber="9">Onitis</emphasis>
species.
</taxonomicName>
<emphasis id="EFACEAD6FFFE890E9E1EFD68FDFBFD79" box="[287,514,729,755]" italics="true" pageId="8" pageNumber="9">Onitisbhomorensis</emphasis>
, is compared with
<taxonomicName id="1AD84D47FFFE890E9DDCFD68FC7DFD79" authorityName="Biswas" authorityYear="1980" box="[733,900,729,755]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFFE890E9DDCFD68FC7DFD79" box="[733,900,729,755]" italics="true" pageId="8" pageNumber="9">O. assamensis</emphasis>
</taxonomicName>
, a species described from a nearby locality as that of the former by
<bibRefCitation id="B9494B35FFFE890E9EDCFD4DFD71FC9C" author="Biswas S." box="[477,648,764,790]" pageId="8" pageNumber="9" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas (1980)</bibRefCitation>
based on a single specimen.
<taxonomicName id="1AD84D47FFFE890E9CE2FD4CFB64FC9C" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[995,1181,764,791]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bhomorensis">
<emphasis id="EFACEAD6FFFE890E9CE2FD4CFB64FC9C" box="[995,1181,764,791]" italics="true" pageId="8" pageNumber="9">O. bhomorensis</emphasis>
</taxonomicName>
is smaller and the body is reddish black, while
<taxonomicName id="1AD84D47FFFE890E9D0DFC91FD4AFCB3" authorityName="Biswas" authorityYear="1980" box="[524,691,800,826]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFFE890E9D0DFC91FD4AFCB3" box="[524,691,800,826]" italics="true" pageId="8" pageNumber="9">O. assamensis</emphasis>
</taxonomicName>
is greenish. The frontal carina is entire with a tubercle in the former while the carina is interrupted in the latter. The protibia of
<taxonomicName id="1AD84D47FFFE890E9CBFFCF2FB8EFCD6" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[958,1143,834,861]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bhomorensis">
<emphasis id="EFACEAD6FFFE890E9CBFFCF2FB8EFCD6" box="[958,1143,834,861]" italics="true" pageId="8" pageNumber="9">O. bhomorensis</emphasis>
</taxonomicName>
is with a rudimentary spur and the pygidium is with a vertical groove, both these characters are absent in
<taxonomicName id="1AD84D47FFFE890E9B8FFCD7FACFFC0A" authorityName="Biswas" authorityYear="1980" box="[1166,1334,870,896]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFFE890E9B8FFCD7FACFFC0A" box="[1166,1334,870,896]" italics="true" pageId="8" pageNumber="9">O. assamensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFE890E9FBCFC16FDEDFB99" pageId="8" pageNumber="9" type="etymology">
<paragraph id="DD6736C4FFFE890E9FBCFC16FEBDFC4B" blockId="8.[189,324,935,961]" box="[189,324,935,961]" pageId="8" pageNumber="9">
<heading id="862F81A8FFFE890E9FBCFC16FEBDFC4B" bold="true" box="[189,324,935,961]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EFACEAD6FFFE890E9FBCFC16FEBDFC4B" bold="true" box="[189,324,935,961]" pageId="8" pageNumber="9">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFE890E9FBCFC67FDEDFB99" blockId="8.[189,1399,982,1043]" pageId="8" pageNumber="9">
Named after the bridge “Kolia Bhomora” across Brahmaputra – the widest river in
<collectingCountry id="A5CF7654FFFE890E9B83FC67FB38FC7A" box="[1154,1217,982,1008]" name="India" pageId="8" pageNumber="9">India</collectingCountry>
, near which the
<typeStatus id="02638866FFFE890E9FBCFC48FF09FB99" box="[189,240,1017,1043]" pageId="8" pageNumber="9">type</typeStatus>
specimen was collected.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFE890E9FBCFB8BFDBCFB67" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="DD6736C4FFFE890E9FBCFB8BFE53FBDE" blockId="8.[189,426,1082,1108]" box="[189,426,1082,1108]" pageId="8" pageNumber="9">
<heading id="862F81A8FFFE890E9FBCFB8BFE53FBDE" bold="true" box="[189,426,1082,1108]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EFACEAD6FFFE890E9FBCFB8BFE53FBDE" bold="true" box="[189,426,1082,1108]" pageId="8" pageNumber="9">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="6DB03C99FFFE890E9FEBFBD8FDBCFB67" collectingDate="2012-10-10" collectingMethod="hand-picked / cattle dung" collectionCode="NBAIR" collectorName="Karimbumkara Seena Narayanan" country="India" elevation="74" latitude="26.582407" location="Kolia" longLatPrecision="1" longitude="92.864624" municipality="Tezpur" pageId="8" pageNumber="9" specimenCode="NIM/ NBAIR/COL/ONIT/H-271023A" specimenCount="1" specimenCount-female="1" stateProvince="Assam" typeStatus="holotype">
<paragraph id="DD6736C4FFFE890E9FEBFBD8FEA5FB09" blockId="8.[189,1399,1129,1261]" box="[234,348,1129,1155]" pageId="8" pageNumber="9">
<emphasis id="EFACEAD6FFFE890E9FEBFBD8FEA5FB09" bold="true" box="[234,348,1129,1155]" pageId="8" pageNumber="9">
<typeStatus id="02638866FFFE890E9FEBFBD8FEA5FB09" box="[234,348,1129,1155]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFFE890E9FBCFB3DFDBCFB67" blockId="8.[189,1399,1129,1261]" pageId="8" pageNumber="9">
<collectingCountry id="A5CF7654FFFE890E9FBCFB3DFEF7FB2C" box="[189,270,1164,1190]" name="India" pageId="8" pageNumber="9">INDIA</collectingCountry>
•
<specimenCount id="CBDEFD4DFFFE890E9E33FB3CFEB0FB2D" box="[306,329,1165,1191]" count="1" pageId="8" pageNumber="9" type="female">♀</specimenCount>
;
<collectingRegion id="1F1CF826FFFE890E9E5AFB3DFE57FB2C" box="[347,430,1164,1190]" country="India" name="Assam" pageId="8" pageNumber="9">Assam</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFFE890E9EBFFB3CFDE8FB2D" box="[446,529,1165,1191]" pageId="8" pageNumber="9">Tezpur</collectingMunicipality>
,
<location id="D807601FFFFE890E9D23FB3DFD9AFB2C" LSID="urn:lsid:plazi:treatment:557187D2FFFE890C9D72FEB8FABFFCFC:D807601FFFFE890E9D23FB3DFD9AFB2C" box="[546,611,1164,1190]" country="India" latitude="26.582407" longLatPrecision="1" longitude="92.864624" municipality="Tezpur" name="Kolia" pageId="8" pageNumber="9" stateProvince="Assam">Kolia</location>
bhomora;
<date id="A9661004FFFE890E9DECFB3DFC6AFB2C" box="[749,915,1164,1190]" pageId="8" pageNumber="9" value="2012-10-10">
<collectingDate id="B922E9ECFFFE890E9DECFB3DFC6AFB2C" box="[749,915,1164,1190]" pageId="8" pageNumber="9" value="2012-10-10">10 Oct. 2012</collectingDate>
</date>
;
<geoCoordinate id="B8EC5003FFFE890E9CA5FB3AFB9BFB2D" box="[932,1122,1163,1191]" degrees="26" direction="north" minutes="34" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="56.66" value="26.582407">26°34ʹ56.66ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFFE890E9B72FB3AFAD7FB2D" box="[1139,1326,1163,1191]" degrees="92" direction="east" minutes="51" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="52.65" value="92.864624">92°51ʹ52.65ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFFE890E9FBCFB1EFEC0FB43" box="[189,313,1199,1226]" metricMagnitude="1" metricUnit="m" metricValue="7.4" pageId="8" pageNumber="9" unit="m" value="74.0">
<quantity id="1A209B21FFFE890E9FBCFB1EFF00FB40" box="[189,249,1199,1226]" metricMagnitude="1" metricUnit="m" metricValue="7.4" pageId="8" pageNumber="9" unit="m" value="74.0">74 m</quantity>
a.s.l.
</elevation>
;
<collectorName id="702D5312FFFE890E9E46FB1EFD34FB40" box="[327,717,1199,1226]" pageId="8" pageNumber="9">Karimbumkara Seena Narayanan</collectorName>
leg.;
<collectingMethod id="04994ED3FFFE890E9C11FB1EFC5DFB43" box="[784,932,1199,1225]" pageId="8" pageNumber="9">hand-picked</collectingMethod>
,
<collectingMethod id="04994ED3FFFE890E9CB1FB1EFBCFFB43" box="[944,1078,1199,1225]" pageId="8" pageNumber="9">cattle dung</collectingMethod>
;
<collectionCode id="BBC9AE01FFFE890E9B43FB01FB59FB40" box="[1090,1184,1200,1226]" pageId="8" pageNumber="9">NBAIR</collectionCode>
, Bangalore;
<specimenCode id="8D7E9EBFFFFE890E9A34FB1EFDBBFB67" pageId="8" pageNumber="9">NIM/ NBAIR/COL/ONIT/H-271023A</specimenCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="95C2654FFFFE890C9FBCFAA5FEC0FD69" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="DD6736C4FFFE890E9FBCFAA5FEB7FAA4" blockId="8.[189,334,1300,1326]" box="[189,334,1300,1326]" pageId="8" pageNumber="9">
<heading id="862F81A8FFFE890E9FBCFAA5FEB7FAA4" bold="true" box="[189,334,1300,1326]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EFACEAD6FFFE890E9FBCFAA5FEB7FAA4" bold="true" box="[189,334,1300,1326]" pageId="8" pageNumber="9">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFE890E9FBCFAF2FB3BFAD7" blockId="8.[189,1218,1346,1373]" box="[189,1218,1346,1373]" pageId="8" pageNumber="9">MEASUREMENTS (in mm). TL=18; BW= 10; PL=7; PW=11; EL=10; HL=5; HW=5.1.</paragraph>
<paragraph id="DD6736C4FFFE890E9FEBFA35FE52FA14" blockId="8.[189,1399,1412,2002]" box="[234,427,1412,1438]" pageId="8" pageNumber="9">
<emphasis id="EFACEAD6FFFE890E9FEBFA35FED3FA14" bold="true" box="[234,298,1412,1438]" pageId="8" pageNumber="9">Male</emphasis>
(
<typeStatus id="02638866FFFE890E9E38FA35FE5AFA14" box="[313,419,1412,1438]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="DD6736C4FFFE890C9FBCFA16FE89FE7C" blockId="8.[189,1399,1412,2002]" lastBlockId="10.[189,1399,265,503]" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
Body narrow, elongate oval, reddish black, not very shining, slightly convex. Antennae red with the club reddish yellow, legs reddish black; body devoid of setae on the dorsal side, ventral side covered with reddish hairs except on the middle of metasternum and the abdominal sclerites. Head shining with the clypeus truncate (
<figureCitation id="45E32A41FFFE890E9EF4F9A0FDAAF9A1" box="[501,595,1553,1579]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="8" pageNumber="9">Fig. 9A</figureCitation>
), almost straight anteriorly and at the sides, strongly rugose; clypeal carina present, slightly elevated and curved. The frontal carina is broadly interrupted in the middle with a blunt conical tubercle behind. The vertex punctuation granular, closer in the middle and scattered at the sides appears rugose behind the tubercle. Genae smooth, curved, extended slightly outwards and upwards anteriorly; inner margin slightly elevated and continues behind as a posterior carina. Pronotum moderately convex, sides smoothly margined, except the base which is slightly angulate in the middle; with strong close punctures, closer between the basal foveae; uneven in the middle; deeper and closer anteriorly. Pronotal front angles not very sharp, almost right-angled, the sides straight anteriorly, rounded in the middle, bisinuate behind; hind angles obsolete. A pair of deep basal foveae present at the base on either side of the middle separated by twice its length. Lateral foveae smooth with small punctures in the periphery. Elytra finely and broadly striate, striae devoid of punctures; intervals flat, opaque; sutural interval convex, shining with visible fine scattered punctures. The lateral carinae straight, with minute visible punctures. Pygidium (
<figureCitation id="45E32A41FFFE890E9D13F806FD90F85B" box="[530,617,1975,2001]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[245,1340,356,1841]" captionTargetId="figure-1@9.[189,1398,265,1856]" captionTargetPageId="9" captionText="Fig. 2. Onitisbhomorensis sp. nov., holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head and prontum. D. Protibia, arrow showing rudimentary spur. E. Metasternum. F. Pygidium, arrow showing longitudinal groove. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742855" httpUri="https://zenodo.org/record/13742855/files/figure.png" pageId="8" pageNumber="9">Fig. 2F</figureCitation>
) without hair or setae, slightly shining with imperceptible scattered punctures. A smooth median longitudinal groove extending from base to almost the middle is present. Metasternal shield shining, (
<figureCitation id="45E32A41FFFC890C9D05FE9DFDA2FECC" box="[516,603,300,326]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[245,1340,356,1841]" captionTargetId="figure-1@9.[189,1398,265,1856]" captionTargetPageId="9" captionText="Fig. 2. Onitisbhomorensis sp. nov., holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head and prontum. D. Protibia, arrow showing rudimentary spur. E. Metasternum. F. Pygidium, arrow showing longitudinal groove. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742855" httpUri="https://zenodo.org/record/13742855/files/figure.png" pageId="10" pageNumber="11">Fig. 2E</figureCitation>
) with minute punctures in the middle; anterior part of the metasternal shield slightly convex, with uneven granules and reddish hairs. Sides of the metasternum closely and evenly granular, covered with smooth short reddish hairs. Protibia slender, bears four external teeth, the tip not very long or curved, with a rudimentary spur near the tip (
<figureCitation id="45E32A41FFFC890C9CBBFE27FBE5FE3A" box="[954,1052,406,432]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[245,1340,356,1841]" captionTargetId="figure-1@9.[189,1398,265,1856]" captionTargetPageId="9" captionText="Fig. 2. Onitisbhomorensis sp. nov., holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head and prontum. D. Protibia, arrow showing rudimentary spur. E. Metasternum. F. Pygidium, arrow showing longitudinal groove. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742855" httpUri="https://zenodo.org/record/13742855/files/figure.png" pageId="10" pageNumber="11">Figs 2D</figureCitation>
,
<figureCitation id="45E32A41FFFC890C9B29FE27FBA6FE3A" box="[1064,1119,406,432]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="10" pageNumber="11">16A</figureCitation>
); teeth short, triangular, placed almost equidistantly. The inner margins of femur and tibia fringed. Femur without protrusion on any of the legs.
</paragraph>
<caption id="89A7664CFFFF890F9FBCF839FB19F862" ID-DOI="http://doi.org/10.5281/zenodo.13742855" ID-Zenodo-Dep="13742855" httpUri="https://zenodo.org/record/13742855/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1928,1954]" targetBox="[245,1340,356,1841]" targetPageId="9">
<paragraph id="DD6736C4FFFF890F9FBCF839FB19F862" blockId="9.[189,1398,1928,2025]" pageId="9" pageNumber="10">
<emphasis id="EFACEAD6FFFF890F9FBCF839FEF7F828" bold="true" box="[189,270,1928,1954]" pageId="9" pageNumber="10">Fig. 2.</emphasis>
<emphasis id="EFACEAD6FFFF890F9E19F839FE06F828" box="[280,511,1928,1954]" italics="true" pageId="9" pageNumber="10">Onitisbhomorensis</emphasis>
<taxonomicNameLabel id="F49F57ADFFFF890F9D0BF839FD9EF828" box="[522,615,1928,1954]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A.
<emphasis id="EFACEAD6FFFF890F9BECF839FAFBF828" bold="true" box="[1261,1282,1928,1954]" pageId="9" pageNumber="10">A</emphasis>
. Habitus, dorsal view.
<emphasis id="EFACEAD6FFFF890F9E54F81AFE90F84F" bold="true" box="[341,361,1963,1989]" pageId="9" pageNumber="10">B</emphasis>
. Habitus, ventral view.
<emphasis id="EFACEAD6FFFF890F9D86F81AFD65F84F" bold="true" box="[647,668,1963,1989]" pageId="9" pageNumber="10">C</emphasis>
. Head and prontum.
<emphasis id="EFACEAD6FFFF890F9C9BF81AFC56F84F" bold="true" box="[922,943,1963,1989]" pageId="9" pageNumber="10">D</emphasis>
. Protibia, arrow showing rudimentary spur.
<emphasis id="EFACEAD6FFFF890F9FFCF87FFEE8F862" bold="true" box="[253,273,1998,2024]" pageId="9" pageNumber="10">E</emphasis>
. Metasternum.
<emphasis id="EFACEAD6FFFF890F9EC9F87FFE23F862" bold="true" box="[456,474,1998,2024]" pageId="9" pageNumber="10">F</emphasis>
. Pygidium, arrow showing longitudinal groove. (Images: AIMB).
</paragraph>
</caption>
<paragraph id="DD6736C4FFFC890C9FBCFDAFFD7EFDF4" blockId="10.[189,1399,541,639]" pageId="10" pageNumber="11">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFFC890C9EAFFDAFFDDAFDBD" box="[430,547,541,568]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. E–H. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. I–L. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. M–P. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742867" httpUri="https://zenodo.org/record/13742867/files/figure.png" pageId="10" pageNumber="11">Fig. 6I–L</figureCitation>
). LP=3, Lp=2.5; BP=1.4, BpB =0.7, BpT=0.4. Parameres almost same length as phallobase, broader at the base and further tapers become slender and slightly curved at the tip. Phallobase slightly broader at the base.
</paragraph>
<paragraph id="DD6736C4FFFC890C9FEBFD17FEBCFD4A" blockId="10.[189,325,678,739]" box="[234,325,678,704]" pageId="10" pageNumber="11">
<emphasis id="EFACEAD6FFFC890C9FEBFD17FEBCFD4A" bold="true" box="[234,325,678,704]" pageId="10" pageNumber="11">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFFC890C9FBCFD78FEC0FD69" blockId="10.[189,325,678,739]" box="[189,313,713,739]" pageId="10" pageNumber="11">Unknown.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFC890C9FBCFCBBFABFFCFC" pageId="10" pageNumber="11" type="biology_ecology">
<paragraph id="DD6736C4FFFC890C9FBCFCBBFEE4FCAE" blockId="10.[189,285,778,804]" box="[189,285,778,804]" pageId="10" pageNumber="11">
<heading id="862F81A8FFFC890C9FBCFCBBFEE4FCAE" bold="true" box="[189,285,778,804]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="EFACEAD6FFFC890C9FBCFCBBFEE4FCAE" bold="true" box="[189,285,778,804]" pageId="10" pageNumber="11">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFC890C9FBCFC88FABFFCFC" blockId="10.[189,1398,825,886]" pageId="10" pageNumber="11">
The specimen was collected from cattle dung on the river bed of the Brahmaputra, where the vegetation is dry seasonal grasslands, typical of Brahmaputra flood plains dominated by
<emphasis id="EFACEAD6FFFC890C9B4BFCEDFAF9FCFC" box="[1098,1280,860,886]" italics="true" pageId="10" pageNumber="11">Ziziphusjujuba</emphasis>
trees.
</paragraph>
</subSubSection>
</treatment>
</document>
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