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<document id="550658DC872798D672EE36AADE25C299" ID-DOI="10.5852/ejt.2024.956.2657" ID-ISSN="2118-9773" ID-Zenodo-Dep="13742850" ID-ZooBank="9DCDD521-909D-4436-8495-086A11C50DA0" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1725988271530" checkinUser="plazi" docAuthor="Karimbumkara, Seena Narayanan &amp; Priyadarsanan, Dharma Rajan" docDate="2024" docId="557187D2FFEB89189D54F9EBFB5AF9A0" docLanguage="en" docName="EJT.2024.956.1-61.pdf" docOrigin="European Journal of Taxonomy 956" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onitis naviauxi Cambefort 1988" docType="treatment" docVersion="5" lastPageNumber="31" masterDocId="A948FFAAFFF689069F01FFB1FFF9FF8A" masterDocTitle="Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India" masterLastPageNumber="61" masterPageNumber="1" pageNumber="30" updateTime="1726490331196" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0594CD419972E160860A603FFD4AAF68">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<mods:namePart id="49F6C131756F417E1AB011B0E7F83433">Karimbumkara, Seena Narayanan</mods:namePart>
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<mods:affiliation id="D2602FC4000B4791A1686F754A6C774C">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
<mods:nameIdentifier id="BDAE97DAD1C928CD718BF06BC0BD070B" type="email">seena.narayanan@atree.org</mods:nameIdentifier>
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<mods:namePart id="7711F61709A222C94B115BBC30473D12">Priyadarsanan, Dharma Rajan</mods:namePart>
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<mods:affiliation id="5B0495A01A9148A6B52CB5F4C9A23948">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
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<mods:title id="3CC2E83D363C6A7449A762A00CBDA60B">European Journal of Taxonomy</mods:title>
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<mods:date id="B378066280975A2EFF5A572ECE67FB07">2024</mods:date>
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<mods:number id="9808D4D292B29B3F31CE208C5B2CC8AF">956</mods:number>
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<treatment id="557187D2FFEB89189D54F9EBFB5AF9A0" LSID="urn:lsid:plazi:treatment:557187D2FFEB89189D54F9EBFB5AF9A0" httpUri="http://treatment.plazi.org/id/557187D2FFEB89189D54F9EBFB5AF9A0" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="95C2654FFFEB891B9D54F9EBFC26F9FE" box="[597,991,1626,1652]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="DD6736C4FFEB891B9D54F9EBFC26F9FE" blockId="29.[597,991,1626,1687]" box="[597,991,1626,1652]" pageId="29" pageNumber="30">
<heading id="862F81A8FFEB891B9D54F9EBFC26F9FE" box="[597,991,1626,1652]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="30" reason="2">
<taxonomicName id="1AD84D47FFEB891B9D54F9EBFC26F9FE" ID-CoL="49KJD" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[597,991,1626,1652]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFEB891B9D54F9EBFCE9F9FE" bold="true" box="[597,784,1626,1652]" italics="true" pageId="29" pageNumber="30">Onitis naviauxi</emphasis>
<bibRefCitation id="B9494B35FFEB891B9C16F9EBFC26F9FE" author="Cambefort Y." box="[791,991,1626,1652]" pageId="29" pageNumber="30" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEB891B9DA9F9CCFC75F91D" box="[680,908,1661,1687]" pageId="29" pageNumber="30" type="description">
<paragraph id="DD6736C4FFEB891B9DA9F9CCFC75F91D" blockId="29.[597,991,1626,1687]" box="[680,908,1661,1687]" pageId="29" pageNumber="30">
<figureCitation id="45E32A41FFEB891B9DA9F9CCFCD8F91D" box="[680,801,1661,1687]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. A–D. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). E–F. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). G–J. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). K–N. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742861" httpUri="https://zenodo.org/record/13742861/files/figure.png" pageId="29" pageNumber="30">Figs 4E–F</figureCitation>
,
<figureCitation id="45E32A41FFEB891B9C2CF9CCFC9BF91D" box="[813,866,1661,1687]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." figureDoi="http://doi.org/10.5281/zenodo.13742879" httpUri="https://zenodo.org/record/13742879/files/figure.png" pageId="29" pageNumber="30">10D</figureCitation>
,
<figureCitation id="45E32A41FFEB891B9C6EF9CCFC75F91D" box="[879,908,1661,1687]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742907" httpUri="https://zenodo.org/record/13742907/files/figure.png" pageId="29" pageNumber="30">19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEB891B9FBCF90EFD14F8B7" pageId="29" pageNumber="30" type="reference_group">
<paragraph id="DD6736C4FFEB891B9FBCF90EFC7BF952" blockId="29.[189,898,1726,1753]" box="[189,898,1726,1753]" pageId="29" pageNumber="30">
<emphasis id="EFACEAD6FFEB891B9FBCF90EFE88F953" box="[189,369,1727,1753]" italics="true" pageId="29" pageNumber="30">Onitisnaviauxi</emphasis>
<bibRefCitation id="B9494B35FFEB891B9E78F90FFD82F953" author="Cambefort Y." box="[377,635,1726,1753]" pageId="29" pageNumber="30" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988: 189</bibRefCitation>
(original description).
</paragraph>
<paragraph id="DD6736C4FFEB891B9FBCF8B1FD14F8B7" blockId="29.[189,1399,1792,1854]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="FDC811EAFFEB891B9FBCF8B1FD14F8B7" pageId="29" pageNumber="30">
<taxonomicName id="1AD84D47FFEB891B9FBCF8B1FE81F890" ID-CoL="49KJD" authorityName="Cambefort" authorityYear="1988" box="[189,376,1792,1818]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFEB891B9FBCF8B1FE81F890" box="[189,376,1792,1818]" italics="true" pageId="29" pageNumber="30">Onitis naviauxi</emphasis>
</taxonomicName>
–
<treatmentCitation id="5C7910D5FFEB891B9EA5F8B1FD01F891" author="Bezdek A. &amp; Krell F. - T." box="[420,760,1792,1819]" page="159" pageId="29" pageNumber="30" year="2006">
<bibRefCitation id="B9494B35FFEB891B9EA5F8B1FD01F891" author="Bezdek A. &amp; Krell F. - T." box="[420,760,1792,1819]" pageId="29" pageNumber="30" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFEB891B9CE3F8B1FB3FF890" author="Bezdek A." box="[994,1222,1792,1818]" page="179" pageId="29" pageNumber="30" year="2016">
<bibRefCitation id="B9494B35FFEB891B9CE3F8B1FB3FF890" author="Bezdek A." box="[994,1222,1792,1818]" pageId="29" pageNumber="30" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFEB891B9E0EF892FDF3F8B7" author="Schoolmeesters P." box="[271,522,1827,1853]" pageId="29" pageNumber="30" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEB89189FBCF8D5FD4DFECC" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="DD6736C4FFEB891B9FBCF8D5FE3EF8F4" blockId="29.[189,455,1892,1918]" box="[189,455,1892,1918]" pageId="29" pageNumber="30">
<heading id="862F81A8FFEB891B9FBCF8D5FE3EF8F4" bold="true" box="[189,455,1892,1918]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="EFACEAD6FFEB891B9FBCF8D5FE3EF8F4" bold="true" box="[189,455,1892,1918]" pageId="29" pageNumber="30">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEB89189FBCF822FD4DFECC" blockId="29.[189,1399,1939,2001]" lastBlockId="30.[189,1398,265,327]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<bibRefCitation id="B9494B35FFEB891B9FBCF822FE77F827" author="Cambefort Y." box="[189,398,1939,1965]" pageId="29" pageNumber="30" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort (1988)</bibRefCitation>
classified
<emphasis id="EFACEAD6FFEB891B9D07F822FD41F827" box="[518,696,1939,1965]" italics="true" pageId="29" pageNumber="30">Onitisnaviauxi</emphasis>
in Group III of
<bibRefCitation id="B9494B35FFEB891B9C68F825FBDCF827" author="Janssens A." box="[873,1061,1939,1966]" pageId="29" pageNumber="30" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens (1937)</bibRefCitation>
. It is similar to
<taxonomicName id="1AD84D47FFEB891B9BD0F822FA8EF827" authorityName="Biswas" authorityYear="1980" box="[1233,1399,1939,1965]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFEB891B9BD0F822FA8EF827" box="[1233,1399,1939,1965]" italics="true" pageId="29" pageNumber="30">O. assamensis</emphasis>
</taxonomicName>
in having vertex with punctures and clypeal carina being placed away from vertex. It is distinct from the latter in having shorter clypeal carina and the male protibia having 3–4 teeth on the inferomedial ridge, while in
<taxonomicName id="1AD84D47FFE889189E25FE9CFE32FECC" authorityName="Biswas" authorityYear="1980" box="[292,459,301,327]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFE889189E25FE9CFE32FECC" box="[292,459,301,327]" italics="true" pageId="30" pageNumber="31">O. assamensis</emphasis>
</taxonomicName>
there are 6–7 teeth.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE889189FBCFEDEFE7CFD06" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="DD6736C4FFE889189FBCFEDEFD31FE03" blockId="30.[189,712,367,394]" box="[189,712,367,394]" pageId="30" pageNumber="31">
<emphasis id="EFACEAD6FFE889189FBCFEDEFE15FE03" bold="true" box="[189,492,367,393]" pageId="30" pageNumber="31">Type material examined</emphasis>
(based on images)
</paragraph>
<materialsCitation id="6DB03C99FFE889189FEBFE2EFE7FFD8B" collectionCode="MNHN" country="Nepal" determinerName="Y. Cambefort" location="NEPAL" pageId="30" pageNumber="31" specimenCode="EC4807" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="DD6736C4FFE889189FEBFE2EFEA5FE33" blockId="30.[189,1399,415,513]" box="[234,348,415,441]" pageId="30" pageNumber="31">
<emphasis id="EFACEAD6FFE889189FEBFE2EFEA5FE33" bold="true" box="[234,348,415,441]" pageId="30" pageNumber="31">
<typeStatus id="02638866FFE889189FEBFE2EFEA5FE33" box="[234,348,415,441]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE889189FBCFE75FE7FFD8B" blockId="30.[189,1399,415,513]" pageId="30" pageNumber="31">
<collectingCountry id="A5CF7654FFE889189FBCFE75FEE1FE54" box="[189,280,452,478]" name="Nepal" pageId="30" pageNumber="31">NEPAL</collectingCountry>
•
<specimenCount id="CBDEFD4DFFE889189E34FE75FEB7FE54" box="[309,334,452,478]" count="1" pageId="30" pageNumber="31" type="male">♂</specimenCount>
; “
<emphasis id="EFACEAD6FFE889189E6BFE75FDD9FE54" box="[362,544,452,478]" italics="true" pageId="30" pageNumber="31">Onitisnaviauxi</emphasis>
<taxonomicNameLabel id="F49F57ADFFE889189D2BFE75FD92FE54" box="[554,619,452,478]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
<determinerName id="28BBDA00FFE889189D75FE75FCEAFE54" box="[628,787,452,478]" pageId="30" pageNumber="31">Y. Cambefort</determinerName>
det. 1987, Nepal: Sauraha. Alt. 150, 19.4.82 RN”;
<collectionCode id="BBC9AE01FFE889189FBCFE56FEEEFD8B" box="[189,279,487,513]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
<specimenCode id="8D7E9EBFFFE889189E1FFE56FE7BFD8B" box="[286,386,487,513]" collectionCode="EC" pageId="30" pageNumber="31">EC4807</specimenCode>
.
</paragraph>
</materialsCitation>
<paragraph id="DD6736C4FFE889189FEBFD9BFEA5FDCE" blockId="30.[189,1399,554,652]" box="[234,348,554,580]" pageId="30" pageNumber="31">
<materialsCitation id="6DB03C99FFE889189FEBFD9BFEA5FDCE" box="[234,348,554,580]" collectionCode="MNHN" country="Nepal" determinerName="Y. Cambefort" location="NEPAL" pageId="30" pageNumber="31" specimenCode="EC4808" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="EFACEAD6FFE889189FEBFD9BFEA5FDCE" bold="true" box="[234,348,554,580]" pageId="30" pageNumber="31">
<typeStatus id="02638866FFE889189FEBFD9BFEA5FDCE" box="[234,348,554,580]" pageId="30" pageNumber="31" type="paratype">Paratype</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="DD6736C4FFE889189FBCFDFEFE7CFD06" blockId="30.[189,1399,554,652]" pageId="30" pageNumber="31">
<collectingCountry id="A5CF7654FFE889189FBCFDFEFEE1FDE3" box="[189,280,591,617]" name="Nepal" pageId="30" pageNumber="31">NEPAL</collectingCountry>
•
<specimenCount id="CBDEFD4DFFE889189E37FDFEFEB4FDE3" box="[310,333,591,617]" count="1" pageId="30" pageNumber="31" type="female">♀</specimenCount>
; “
<emphasis id="EFACEAD6FFE889189E6DFDFEFDDAFDE3" box="[364,547,591,617]" italics="true" pageId="30" pageNumber="31">Onitisnaviauxi</emphasis>
<taxonomicNameLabel id="F49F57ADFFE889189D2CFDFEFD96FDE3" box="[557,623,591,617]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
<determinerName id="28BBDA00FFE889189D6EFDFEFCF6FDE2" box="[623,783,590,617]" pageId="30" pageNumber="31">Y. Cambefort</determinerName>
det. 1987,
<collectingCountry id="A5CF7654FFE889189C9AFDFFFC1FFDE2" box="[923,998,590,616]" name="Nepal" pageId="30" pageNumber="31">Nepal</collectingCountry>
: Sauraha. Alt. 150, 19.4.82 RN”;
<collectionCode id="BBC9AE01FFE889189FBCFDC3FEEEFD06" box="[189,279,626,652]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
<specimenCode id="8D7E9EBFFFE889189E1FFDC3FE87FD06" box="[286,382,626,652]" collectionCode="EC" pageId="30" pageNumber="31">EC4808</specimenCode>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE889189FBCFD05FBB5FA3D" pageId="30" pageNumber="31" type="description">
<paragraph id="DD6736C4FFE889189FBCFD05FEB7FD44" blockId="30.[189,334,692,718]" box="[189,334,692,718]" pageId="30" pageNumber="31">
<heading id="862F81A8FFE889189FBCFD05FEB7FD44" bold="true" box="[189,334,692,718]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="EFACEAD6FFE889189FBCFD05FEB7FD44" bold="true" box="[189,334,692,718]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE889189FBCFD54FCEAFD75" blockId="30.[189,787,740,767]" box="[189,787,740,767]" pageId="30" pageNumber="31">MEASUREMENTS (in mm). TL=20–25. BW= 10–12.</paragraph>
<paragraph id="DD6736C4FFE889189FEBFC99FED3FCC8" blockId="30.[189,1399,808,1187]" box="[234,298,808,834]" pageId="30" pageNumber="31">
<emphasis id="EFACEAD6FFE889189FEBFC99FED3FCC8" bold="true" box="[234,298,808,834]" pageId="30" pageNumber="31">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE889189FBCFCFCFAEEFB29" blockId="30.[189,1399,808,1187]" pageId="30" pageNumber="31">Bronzy green, shining; head rounded, slightly flattened anteriorly; clypeus with fine, sparse punctuation, slightly rugose behind; clypeal carina present, short; frontal carina narrowly interrupted in the middle, with a fairly strong sharp tubercle behind; vertex with smooth punctures; pronotum with simple, strong and dense punctures on the disc, finer, more sparse anteriorly, almost imperceptible at the sides; base not bordered, basal foveae round and placed apart; scutellum very small; elytra smooth, striae broad, not very deep, intervals minutely and sparsely punctured; pygidium convex, very minutely punctured; metasternal shield with shallow longitudinal groove, sparse fine long hairs, the sides granulate anteriorly. Clypeus reflexed, straight in the middle. Profemur with a strong tooth on the antero-median ridge, protibia short, tridentate on the infero-internal ridge, median femur unarmed, tibia toothed on the dorsal side, posterior femur with a strong tooth on the ridge below, directed towards the back, tibia bilobed at the apex.</paragraph>
<paragraph id="DD6736C4FFE889189FBCFB7DFBB4FA83" blockId="30.[189,1398,1228,1289]" pageId="30" pageNumber="31">
AEDEAGUS (
<figureCitation id="45E32A41FFE889189E4BFB7DFE3EFB6C" box="[330,455,1228,1254]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. A–D. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). E–F. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). G–J. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). K–N. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742861" httpUri="https://zenodo.org/record/13742861/files/figure.png" pageId="30" pageNumber="31">Fig. 4E–F</figureCitation>
). Parameres shorter than the phallobase, phallobase cylindrical, slightly curved; parameres straight above and curved below, largely notched on the underside.
</paragraph>
<paragraph id="DD6736C4FFE889189FEBFA83FEBCFAC6" blockId="30.[189,1398,1330,1463]" box="[234,325,1330,1356]" pageId="30" pageNumber="31">
<emphasis id="EFACEAD6FFE889189FEBFA83FEBCFAC6" bold="true" box="[234,325,1330,1356]" pageId="30" pageNumber="31">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE889189FBCFAE7FBB5FA3D" blockId="30.[189,1398,1330,1463]" pageId="30" pageNumber="31">Similar to male but more elongated head, clypeus parabolic, not much reflexed, rugose; with confluent transverse rugulosity behind the clypeal carina; vertex as in the male, with simple, very sparse punctuation; metasternum with well-marked narrow groove; legs not toothed.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE889189FBCFA6EFB5AF9A0" box="[189,1187,1503,1578]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="DD6736C4FFE889189FBCFA6EFEAFFA73" blockId="30.[189,342,1503,1529]" box="[189,342,1503,1529]" pageId="30" pageNumber="31">
<heading id="862F81A8FFE889189FBCFA6EFEAFFA73" bold="true" box="[189,342,1503,1529]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="EFACEAD6FFE889189FBCFA6EFEAFFA73" bold="true" box="[189,342,1503,1529]" pageId="30" pageNumber="31">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE889189FBCF9A1FB5AF9A0" blockId="30.[189,1187,1552,1578]" box="[189,1187,1552,1578]" pageId="30" pageNumber="31">
<collectingCountry id="A5CF7654FFE889189FBCF9A1FEFAF9A0" box="[189,259,1552,1578]" name="Nepal" pageId="30" pageNumber="31">Nepal</collectingCountry>
(
<bibRefCitation id="B9494B35FFE889189E13F9A1FE2EF9A0" author="Cambefort Y." box="[274,471,1552,1578]" pageId="30" pageNumber="31" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort 1988</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE889189EE2F9A1FD1BF9A0" author="Bezdek A. &amp; Krell F. - T." box="[483,738,1552,1578]" pageId="30" pageNumber="31" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE889189DEFF9A1FC75F9A0" author="Bezdek A." box="[750,908,1552,1578]" pageId="30" pageNumber="31" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE889189C99F9A1FB6EF9A0" author="Schoolmeesters P." box="[920,1175,1552,1578]" pageId="30" pageNumber="31" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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