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<mods:title id="48374D083F226F19DCE8B4562798D9FA">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<mods:namePart id="B062BCC62085B45FE097E5CC9941DC2D">Karimbumkara, Seena Narayanan</mods:namePart>
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<paragraph id="DD6736C4FFE189119D58F8B1FC23F890" blockId="23.[601,986,1791,1853]" box="[601,986,1791,1818]" pageId="23" pageNumber="24">
<heading id="862F81A8FFE189119D58F8B1FC23F890" box="[601,986,1791,1818]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<taxonomicName id="1AD84D47FFE189119D58F8B1FC23F890" ID-CoL="49KGS" authority="(Pallas, 1771)" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[601,986,1791,1818]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="humerosus">
<emphasis id="EFACEAD6FFE189119D58F8B1FCC9F890" bold="true" box="[601,816,1791,1818]" italics="true" pageId="23" pageNumber="24">Onitis humerosus</emphasis>
(
<bibRefCitation id="B9494B35FFE189119C3EF8B1FC2BF890" author="Pallas P. S." box="[831,978,1792,1818]" pageId="23" pageNumber="24" refId="ref30468" refString="Pallas P. S. 1771. Reise durch verschiedene Provinzen des Russischen Reiches in den Jahren 1768 - 1774 I: 1 - 504 (453 - 504). Petersburg Akademische Buchhandlung, St Petersburg." type="book" year="1771">Pallas, 1771</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE189119DCFF892FC9CF8B7" box="[718,869,1827,1853]" pageId="23" pageNumber="24" type="description">
<paragraph id="DD6736C4FFE189119DCFF892FC9CF8B7" blockId="23.[601,986,1791,1853]" box="[718,869,1827,1853]" pageId="23" pageNumber="24">
<figureCitation id="45E32A41FFE189119DCFF892FCC2F8B7" box="[718,827,1827,1853]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." figureDoi="http://doi.org/10.5281/zenodo.13742879" httpUri="https://zenodo.org/record/13742879/files/figure.png" pageId="23" pageNumber="24">Figs 10B</figureCitation>
,
<figureCitation id="45E32A41FFE189119C49F892FC9CF8B7" box="[840,869,1827,1853]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742903" httpUri="https://zenodo.org/record/13742903/files/figure.png" pageId="23" pageNumber="24">18</figureCitation>
</paragraph>
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<paragraph id="DD6736C4FFE189119FBCF8D5FC5CF8F4" blockId="23.[189,933,1892,2025]" box="[189,933,1892,1919]" pageId="23" pageNumber="24">
<emphasis id="EFACEAD6FFE189119FBCF8D5FE32F8F4" box="[189,459,1892,1918]" italics="true" pageId="23" pageNumber="24">Scarabaeushumerosus</emphasis>
<bibRefCitation id="B9494B35FFE189119ED2F8D5FD64F8F4" author="Pallas P. S." box="[467,669,1892,1919]" pageId="23" pageNumber="24" refId="ref30468" refString="Pallas P. S. 1771. Reise durch verschiedene Provinzen des Russischen Reiches in den Jahren 1768 - 1774 I: 1 - 504 (453 - 504). Petersburg Akademische Buchhandlung, St Petersburg." type="book" year="1771">Pallas, 1771: 462</bibRefCitation>
(original description).
</paragraph>
<paragraph id="DD6736C4FFE189119FBCF839FD81F828" blockId="23.[189,933,1892,2025]" box="[189,632,1928,1954]" pageId="23" pageNumber="24">
<emphasis id="EFACEAD6FFE189119FBCF839FE45F828" box="[189,444,1928,1954]" italics="true" pageId="23" pageNumber="24">Scarabaeusmenalcas</emphasis>
<bibRefCitation id="B9494B35FFE189119EC2F839FD88F828" author="Pallas P. S." box="[451,625,1928,1954]" pageId="23" pageNumber="24" refId="ref30506" refString="Pallas P. S. 1781. Icones Insectorum praesertim Rossiae Siberiaeque pecularium quae collegit et Descriptionibus illustravit: 1 - 96. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1781">Pallas, 1781: 4</bibRefCitation>
.
</paragraph>
<paragraph id="DD6736C4FFE189119FBCF81AFD92F84F" blockId="23.[189,933,1892,2025]" box="[189,619,1963,1989]" pageId="23" pageNumber="24">
<emphasis id="EFACEAD6FFE189119FBCF81AFE7BF84F" box="[189,386,1963,1989]" italics="true" pageId="23" pageNumber="24">Onitischevrolati</emphasis>
Lucas 1846: pl. 23.
</paragraph>
<paragraph id="DD6736C4FFE189119FBCF87FFD03F863" blockId="23.[189,933,1892,2025]" box="[189,762,1998,2025]" pageId="23" pageNumber="24">
<emphasis id="EFACEAD6FFE189119FBCF87FFDF9F862" box="[189,512,1998,2024]" italics="true" pageId="23" pageNumber="24">Onitishumerosusviolaceus</emphasis>
<bibRefCitation id="B9494B35FFE189119D09F87FFD0AF862" author="Sahlberg J." box="[520,755,1998,2024]" pageId="23" pageNumber="24" pagination="113 - 128" refId="ref30971" refString="Sahlberg J. 1913. Coleoptera mediterranea orientalia, quae Aegypto, Palaestina, Syria, Caramania atque in Anatolia occidentali anno 1904 (Scarabaeidae). Ofversight af Finska Vetenkaps-Societatens Forhandlingar. Helsingfors 55 (19): 113 - 128." type="journal article" year="1913">Sahlberg, 1913: 119</bibRefCitation>
.
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFEB8FCC4FEAE" blockId="24.[189,858,265,366]" box="[189,829,265,294]" pageId="24" pageNumber="25">
<taxonomicName id="1AD84D47FFEE891E9FBCFEB8FDEBFEA9" authorityName="Mikšić" authorityYear="1950" box="[189,530,265,291]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="humerosus" subSpecies="viridipenni">
<emphasis id="EFACEAD6FFEE891E9FBCFEB8FDEBFEA9" box="[189,530,265,291]" italics="true" pageId="24" pageNumber="25">Onitis humerosus viridipenni</emphasis>
</taxonomicName>
sMikŠić, 1950: 165167.
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFE9FFCA3FEC3" blockId="24.[189,858,265,366]" box="[189,858,302,331]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FBCFE9FFDC2FEC2" box="[189,571,302,328]" italics="true" pageId="24" pageNumber="25">Onitishumerosusviolaceipennis</emphasis>
<bibRefCitation id="B9494B35FFEE891E9D43FE9FFCAFFEC3" author="Miksic, R." box="[578,854,302,331]" pageId="24" pageNumber="25" pagination="165 - 167" refId="ref30176" refString="Miksic, R. (1950). Biljeske o Scarabaeidama palearkticke faune. Glasnik Bioloski Sekcije Hrvatkog Privredno Drustvo Zagreb (2 - B) 2 - 3: 165 - 167." type="book chapter" year="1950">MikŠić, 1950: 165167</bibRefCitation>
.
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFEE5FCF7FEE4" blockId="24.[189,858,265,366]" box="[189,782,340,366]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FBCFEE5FDE5FEE4" box="[189,540,340,366]" italics="true" pageId="24" pageNumber="25">Onitishumerosuscupripennis</emphasis>
<bibRefCitation id="B9494B35FFEE891E9D22FEE5FCF2FEE4" author="Balthasar V." box="[547,779,340,366]" pageId="24" pageNumber="25" refId="ref27200" refString="Balthasar V. 1963. Monographie der Scarabaeidae undAphodiidae der palaeoarktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 2: Coprinae (Onitini, Oniticellini, Onthophagini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague." type="book" year="1963">Balthasar, 1963: 48</bibRefCitation>
.
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFE29FC0EFD32" blockId="24.[189,1399,407,697]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="FDC811EAFFEE891E9FBCFE29FC0EFD32" pageId="24" pageNumber="25">
<taxonomicName id="1AD84D47FFEE891E9FBCFE29FE31FE38" ID-CoL="4V269" authorityName="Pallas" authorityYear="1771" box="[189,456,408,434]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="humerosus">
<emphasis id="EFACEAD6FFEE891E9FBCFE29FE31FE38" box="[189,456,408,434]" italics="true" pageId="24" pageNumber="25">Scarabaeus humerosus</emphasis>
</taxonomicName>
Castelnau 1840: 89 (natural history). —
<treatmentCitation id="5C7910D5FFEE891E9CD6FE29FB36FE38" author="Lansberge G." box="[983,1231,407,434]" page="116" pageId="24" pageNumber="25" year="1875">
<bibRefCitation id="B9494B35FFEE891E9CD6FE29FB36FE38" author="Lansberge G." box="[983,1231,407,434]" pageId="24" pageNumber="25" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge 1875: 116</bibRefCitation>
</treatmentCitation>
(monograph). —
<treatmentCitation id="5C7910D5FFEE891E9E10FE0CFE2FFE5D" author="Bedel L." box="[273,470,445,471]" page="268" pageId="24" pageNumber="25" year="1892">
<bibRefCitation id="B9494B35FFEE891E9E10FE0CFE2FFE5D" author="Bedel L." box="[273,470,445,471]" pageId="24" pageNumber="25" pagination="282 - 288" refId="ref27264" refString="Bedel L. 1892. Revision des Scarabaeus palearctiques. L'Abeille, Journal d'Entomologie 27: 282 - 288." type="journal article" year="1892">Bedel 1892: 268</bibRefCitation>
</treatmentCitation>
, 271 (revision). —
<treatmentCitation id="5C7910D5FFEE891E9DBFFE0CFC84FE5D" author="Reitter E." box="[702,893,445,471]" page="97" pageId="24" pageNumber="25" year="1892">
<bibRefCitation id="B9494B35FFEE891E9DBFFE0CFC84FE5D" author="Reitter E." box="[702,893,445,471]" pageId="24" pageNumber="25" refId="ref30798" refString="Reitter E. 1892. Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen Faunengebietes. Verhandlungen des Naturforschenden Vereines in Brunn. https: // www. zobodat. at / pdf / Verh-naturf-Ver-Bruenn _ 31 _ 0003 - 0109. pdf" type="book" year="1892">Reitter 1892: 97</bibRefCitation>
</treatmentCitation>
(keys). —
<treatmentCitation id="5C7910D5FFEE891E9B05FE0CFB37FE5D" author="Arrow G. J." box="[1028,1230,445,471]" page="390" pageId="24" pageNumber="25" year="1931">
<bibRefCitation id="B9494B35FFEE891E9B05FE0CFB37FE5D" author="Arrow G. J." box="[1028,1230,445,471]" pageId="24" pageNumber="25" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 390</bibRefCitation>
</treatmentCitation>
(identification key, description). —
<treatmentCitation id="5C7910D5FFEE891E9EE4FE52FD5AFE77" author="Porta A." box="[485,675,483,509]" page="414" pageId="24" pageNumber="25" year="1932">
<bibRefCitation id="B9494B35FFEE891E9EE4FE52FD5AFE77" author="Porta A." box="[485,675,483,509]" pageId="24" pageNumber="25" refId="ref30657" refString="Porta A. 1932. Fauna Coleopterorum Italica, Vol. II: Scarabaeidae: 373 - 454. Stabilimento Tipografico piancentino, Piacenza." type="book" year="1932">Porta 1932: 414</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFEE891E9C7CFE52FBA5FE77" author="Balthasar V." box="[893,1116,482,509]" page="91" pageId="24" pageNumber="25" year="1935">
<bibRefCitation id="B9494B35FFEE891E9C7CFE52FBA5FE77" author="Balthasar V." box="[893,1116,482,509]" pageId="24" pageNumber="25" refId="ref27158" refString="Balthasar V. 1935. Scarabaeidae des palaarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Edm. Reitter's Nachf. Emmerich Reitter, Troppau." type="book" year="1935">Balthasar 1935: 91</bibRefCitation>
</treatmentCitation>
(monograph); 1963: 47 (monograph).—
<treatmentCitation id="5C7910D5FFEE891E9EADFDB8FD8CFDA8" author="Janssens A." box="[428,629,520,547]" page="91" pageId="24" pageNumber="25" year="1937">
<bibRefCitation id="B9494B35FFEE891E9EADFDB8FD8CFDA8" author="Janssens A." box="[428,629,520,547]" pageId="24" pageNumber="25" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 91</bibRefCitation>
</treatmentCitation>
(revision). —
<bibRefCitation id="B9494B35FFEE891E9C15FDB9FBC9FDA8" author="Hashmi A. &amp; Tashfeen A." box="[788,1072,520,546]" pageId="24" pageNumber="25" pagination="133 - 170" refId="ref29508" refString="Hashmi A. &amp; Tashfeen A. 1992. Coleoptera of Pakistan. Proceedings of Pakistan Congress of Zoology 12: 133 - 170." type="journal article" year="1992">Hashmi &amp;Tashfeen 1992</bibRefCitation>
(distribution).—
<treatmentCitation id="5C7910D5FFEE891E9BF4FDB9FEECFDC2" author="Awal M. M." page="170" pageId="24" pageNumber="25" year="2006">
<bibRefCitation id="B9494B35FFEE891E9BF4FDB9FEECFDC2" author="Awal M. M." pageId="24" pageNumber="25" pagination="163 - 172" refId="ref27017" refString="Awal M. M. 2006. Preliminary studies on Scarabaeoidea (Coleoptera) fauna of Razavi Khorasan province of Iran. Turkiye Entomoloji Dergisi 30 (3): 163 - 172. Available from: https: // dergipark. org. tr / tr / download / article-file / 64989 [accessed 27 Nov. 2019]" type="journal article" year="2006">Awal 2006: 170</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFEE891E9EE9FD9FFD74FDC2" author="Bai M. &amp; Yang X. &amp; Youwei Z." box="[488,653,557,584]" page="391" pageId="24" pageNumber="25" year="2006">
<bibRefCitation id="B9494B35FFEE891E9EE9FD9FFD74FDC2" author="Bai M. &amp; Yang X. &amp; Youwei Z." box="[488,653,557,584]" pageId="24" pageNumber="25" pagination="389 - 395" refId="ref27077" refString="Bai M., Yang X. &amp; Youwei Z. 2006. A key to species of the genus Onitis Fabricius (Coleoptera: Scarabaeidae: Scarabaeinae) from China, with the description of a new species and a new record for China. Proceedings of the Entomological Society of Washington 108: 389 - 395. Available from: https: // www. biodiversitylibrary. org / part / 69133 [accessed 27 nov. 2019]" type="journal article" year="2006">Bai 2006: 391</bibRefCitation>
</treatmentCitation>
(identification key). —
<treatmentCitation id="5C7910D5FFEE891E9CA7FD9FFB22FDC2" author="Bezdek A. &amp; Krell F. - T." box="[934,1243,557,584]" page="159" pageId="24" pageNumber="25" year="2006">
<bibRefCitation id="B9494B35FFEE891E9CA7FD9FFB22FDC2" author="Bezdek A. &amp; Krell F. - T." box="[934,1243,557,584]" pageId="24" pageNumber="25" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution, synonyms). —
<treatmentCitation id="5C7910D5FFEE891E9E9AFDE2FD44FDE4" author="Bunalski M. &amp; Samin N. &amp; Ghahari H. &amp; Hawkeswood T. J." box="[411,701,595,622]" page="156" pageId="24" pageNumber="25" year="2014">
<bibRefCitation id="B9494B35FFEE891E9E9AFDE2FD44FDE4" author="Bunalski M. &amp; Samin N. &amp; Ghahari H. &amp; Hawkeswood T. J." box="[411,701,595,622]" pageId="24" pageNumber="25" pagination="141 - 170" refId="ref28015" refString="Bunalski M., Samin N., Ghahari H. &amp; Hawkeswood T. J. 2014. Contributions to the knowledge the scarab beetles of Golestan province, Northern Iran with checklist of Iranian Scarabaeoidea (Coleoptera). Polish Journal of Entomology 83: 141 - 170. https: // doi. org / 10.2478 / pjen- 2014 - 0011" type="journal article" year="2014">
Bunalski
<emphasis id="EFACEAD6FFEE891E9D09FDE5FDBBFDE7" box="[520,578,595,621]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2014: 156
</bibRefCitation>
</treatmentCitation>
(list of species). — Siddhiqui
<emphasis id="EFACEAD6FFEE891E9B1CFDE5FBAEFDE7" box="[1053,1111,595,621]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2014: 300 (distribution). —
<treatmentCitation id="5C7910D5FFEE891E9E0FFDC9FE25FD19" author="Bezdek A." box="[270,476,632,659]" page="179" pageId="24" pageNumber="25" year="2016">
<bibRefCitation id="B9494B35FFEE891E9E0FFDC9FE25FD19" author="Bezdek A." box="[270,476,632,659]" pageId="24" pageNumber="25" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution, synonyms). —
<treatmentCitation id="5C7910D5FFEE891E9C2BFDC9FBC1FD18" author="Bunalski M. &amp; Samin N. &amp; Ghahari H." box="[810,1080,632,658]" page="36" pageId="24" pageNumber="25" year="2016">
<bibRefCitation id="B9494B35FFEE891E9C2BFDC9FBC1FD18" author="Bunalski M. &amp; Samin N. &amp; Ghahari H." box="[810,1080,632,658]" pageId="24" pageNumber="25" pagination="31 - 40" refId="ref28079" refString="Bunalski M., Samin N. &amp; Ghahari H. 2016. A contribution to the faunistic study of Scarabaeoidea (Coleoptera) from Mazandaran Province, northern Iran. Wiadomosci Entomologiczne 35 (1): 31 - 40. Available from: https: // sparrow. up. poznan. pl / pte / we / 2016 / 35 _ 3 _ Bunalski _ i _ in. pdf [Accessed 15 Nov. 2018]" type="journal article" year="2016">
Bunalski
<emphasis id="EFACEAD6FFEE891E9C94FDC8FC37FD18" box="[917,974,632,658]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2016: 36
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFEE891E9A04FDC9FEA4FD33" author="Montreuil O." page="265" pageId="24" pageNumber="25" year="2017">
<bibRefCitation id="B9494B35FFEE891E9A04FDC9FEA4FD33" author="Montreuil O." pageId="24" pageNumber="25" pagination="259 - 277" refId="ref30308" refString="Montreuil O. 2017. New records of Geotrupidae and Scarabeidae dung beetles (Insecta, Coleoptera) from Iran. Journal of Entomological Society of Iran 36 (4): 259 - 277." type="journal article" year="2017">Montreuil 2017: 265</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFEE891E9D2BFD2FFCE2FD32" author="Schoolmeesters P." box="[554,795,670,696]" pageId="24" pageNumber="25" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEE891E9FBCFD50FE3AFCCD" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="DD6736C4FFEE891E9FBCFD50FE3FFD71" blockId="24.[189,1398,737,839]" box="[189,454,737,763]" pageId="24" pageNumber="25">
<heading id="862F81A8FFEE891E9FBCFD50FE3FFD71" bold="true" box="[189,454,737,763]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="EFACEAD6FFEE891E9FBCFD50FE3FFD71" bold="true" box="[189,454,737,763]" pageId="24" pageNumber="25">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFCB6FE3AFCCD" blockId="24.[189,1398,737,839]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FBCFCB6FE74FCAB" box="[189,397,775,801]" italics="true" pageId="24" pageNumber="25">Onitishumerosus</emphasis>
is a very distinct species having bright yellow elytra with longitudinal green lines upon elevated sutures.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEE891E9FBCFCC0FED2FB6A" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="DD6736C4FFEE891E9FBCFCC0FE15FC00" blockId="24.[189,492,880,907]" box="[189,492,880,907]" pageId="24" pageNumber="25">
<heading id="862F81A8FFEE891E9FBCFCC0FE15FC00" bold="true" box="[189,492,880,907]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="EFACEAD6FFEE891E9FBCFCC0FE15FC00" bold="true" box="[189,492,880,907]" pageId="24" pageNumber="25">Type material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="6DB03C99FFEE891E9FEBFC10FB16FBA6" collectionCode="MfN" country="Russia" determinerName="MfN Janssens" location="RUSSIA" pageId="24" pageNumber="25" specimenCode="MNHUB 2007" specimenCount="10" specimenCount-female="2" specimenCount-male="3" typeStatus="syntype">
<paragraph id="DD6736C4FFEE891E9FEBFC10FDFFFC31" blockId="24.[189,1398,929,1068]" box="[234,518,929,956]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FEBFC10FEA3FC31" bold="true" box="[234,346,929,955]" pageId="24" pageNumber="25">
<typeStatus id="02638866FFEE891E9FEBFC10FEA3FC31" box="[234,346,929,955]" pageId="24" pageNumber="25" type="syntype">Syntypes</typeStatus>
</emphasis>
(
<specimenCount id="CBDEFD4DFFEE891E9E6AFC10FE07FC31" box="[363,510,929,956]" count="5" pageId="24" pageNumber="25" type="generic">5 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFC76FB16FBA6" blockId="24.[189,1398,929,1068]" pageId="24" pageNumber="25">
<collectingCountry id="A5CF7654FFEE891E9FBCFC76FEDDFC6B" box="[189,292,967,993]" name="Russia" pageId="24" pageNumber="25">RUSSIA</collectingCountry>
<specimenCount id="CBDEFD4DFFEE891E9E3CFC76FE7BFC6B" box="[317,386,967,993]" count="3" pageId="24" pageNumber="25" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFEE891E9E8EFC76FE2BFC6B" box="[399,466,967,993]" count="2" pageId="24" pageNumber="25" type="female">2 ♀♀</specimenCount>
; “
<taxonomicName id="1AD84D47FFEE891E9EEFFC76FD17FC6B" authorityName="Pallas" authorityYear="1781" box="[494,750,967,993]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="menalcas">
<emphasis id="EFACEAD6FFEE891E9EEFFC76FD17FC6B" box="[494,750,967,993]" italics="true" pageId="24" pageNumber="25">Scarabaeus menalcas</emphasis>
</taxonomicName>
;
<determinerName id="28BBDA00FFEE891E9DFCFC76FC5AFC6B" box="[765,931,967,993]" pageId="24" pageNumber="25">MfN Janssens</determinerName>
det. as
<taxonomicName id="1AD84D47FFEE891E9CFFFC76FB32FC6B" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[1022,1227,967,993]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="humerosus">
<emphasis id="EFACEAD6FFEE891E9CFFFC76FB32FC6B" box="[1022,1227,967,993]" italics="true" pageId="24" pageNumber="25">Onitis humerosus</emphasis>
</taxonomicName>
in 1938; Hist.- Coll. (
<taxonomicName id="1AD84D47FFEE891E9E04FC5DFE71FB8C" authorityName="Linnaeus" authorityYear="1758" box="[261,392,1004,1030]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
), Nr. 26753,
<emphasis id="EFACEAD6FFEE891E9D20FC5DFD27FB8C" box="[545,734,1004,1030]" italics="true" pageId="24" pageNumber="25">Onitismenalcas</emphasis>
<bibRefCitation id="B9494B35FFEE891E9DE4FC5DFC8DFB8C" author="Pallas P. S." box="[741,884,1004,1030]" pageId="24" pageNumber="25" refId="ref30506" refString="Pallas P. S. 1781. Icones Insectorum praesertim Rossiae Siberiaeque pecularium quae collegit et Descriptionibus illustravit: 1 - 96. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1781">Pallas, 1781</bibRefCitation>
, Russia austr., Bob., Zool. Mus. Berlin; Red Label: SYNTYPUS,
<taxonomicName id="1AD84D47FFEE891E9EB4FBA3FCB2FBA6" authority="Pallas, 1781" authorityName="Pallas" authorityYear="1781" box="[437,843,1042,1068]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="menalcas">
<emphasis id="EFACEAD6FFEE891E9EB4FBA3FD4CFBA6" box="[437,693,1042,1068]" italics="true" pageId="24" pageNumber="25">Scarabaeus menalcas</emphasis>
<bibRefCitation id="B9494B35FFEE891E9DBDFBA3FCB2FBA6" author="Pallas P. S." box="[700,843,1042,1068]" pageId="24" pageNumber="25" refId="ref30506" refString="Pallas P. S. 1781. Icones Insectorum praesertim Rossiae Siberiaeque pecularium quae collegit et Descriptionibus illustravit: 1 - 96. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1781">Pallas, 1781</bibRefCitation>
</taxonomicName>
, labelled by
<specimenCode id="8D7E9EBFFFEE891E9CE2FBA3FB6EFBA6" box="[995,1175,1042,1068]" pageId="24" pageNumber="25">MNHUB 2007</specimenCode>
”;
<collectionCode id="BBC9AE01FFEE891E9BAFFBA3FB12FBA6" box="[1198,1259,1042,1068]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="24" pageNumber="25" type="Museum">MfN</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="DD6736C4FFEE891E9FEBFBE4FCF5FBE5" blockId="24.[189,1399,1109,1248]" box="[234,780,1109,1136]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FEBFBE4FD98FBE5" bold="true" box="[234,609,1109,1135]" pageId="24" pageNumber="25">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFEE891E9D70FBE7FCFDFBE5" box="[625,772,1109,1136]" count="7" pageId="24" pageNumber="25" type="generic">7 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFBCAFED2FB6A" blockId="24.[189,1399,1109,1248]" pageId="24" pageNumber="25">
<materialsCitation id="6DB03C99FFEE891E9FBCFBCAFE15FB31" collectionCode="BMNH" collectorName="G. R. Crotch." country="Israel" location="Jerusalem" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Jerusalem">
<collectingCountry id="A5CF7654FFEE891E9FBCFBCAFEDAFB1F" box="[189,291,1147,1173]" name="Israel" pageId="24" pageNumber="25">ISRAEL</collectingCountry>
<specimenCount id="CBDEFD4DFFEE891E9E42FBCAFEA5FB1F" box="[323,348,1147,1173]" count="1" pageId="24" pageNumber="25" type="male"></specimenCount>
; “
<taxonomicName id="1AD84D47FFEE891E9E78FBCAFD1FFB1F" authority="Pallas, Mid." authorityName="Pallas, Mid." box="[377,742,1147,1173]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="humerosus">
<emphasis id="EFACEAD6FFEE891E9E78FBCAFDB3FB1F" box="[377,586,1147,1173]" italics="true" pageId="24" pageNumber="25">Onitis humerosus</emphasis>
Pallas, Mid.
</taxonomicName>
East;
<collectingRegion id="1F1CF826FFEE891E9C37FBCAFC57FB1F" box="[822,942,1147,1173]" country="Israel" name="Jerusalem" pageId="24" pageNumber="25">Jerusalem</collectingRegion>
,
<collectorName id="702D5312FFEE891E9CBFFBCAFBA2FB1F" box="[958,1115,1147,1173]" pageId="24" pageNumber="25">G.R. Crotch.</collectorName>
, Nevinson Coll., 1918- 14”;
<collectionCode id="BBC9AE01FFEE891E9FF6FB10FEB6FB31" box="[247,335,1185,1211]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="24" pageNumber="25">BMNH</collectionCode>
(
<collectionCode id="BBC9AE01FFEE891E9E61FB10FE8DFB31" box="[352,372,1185,1211]" pageId="24" pageNumber="25">E</collectionCode>
) 1237143
</materialsCitation>
<materialsCitation id="6DB03C99FFEE891E9D07FB10FEDEFB6A" collectionCode="BMNH" collectorName="Pascoe" country="Israel" location="Onitishumerosus Pallas" municipality="Onitishumerosus Pallas" pageId="24" pageNumber="25" specimenCode="(E) 1237144" specimenCount="1" specimenCount-female="1" stateProvince="Mid. East">
<specimenCount id="CBDEFD4DFFEE891E9D07FB10FDE4FB31" box="[518,541,1185,1211]" count="1" pageId="24" pageNumber="25" type="female"></specimenCount>
; “
<collectingMunicipality id="3D03ACBEFFEE891E9D38FB11FCA1FB30" box="[569,856,1184,1210]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9D38FB11FCFEFB30" box="[569,775,1184,1210]" italics="true" pageId="24" pageNumber="25">Onitishumerosus</emphasis>
Pallas
</collectingMunicipality>
,
<collectingRegion id="1F1CF826FFEE891E9C64FB11FC22FB31" box="[869,987,1184,1211]" pageId="24" pageNumber="25">Mid. East</collectingRegion>
;
<collectorName id="702D5312FFEE891E9CE6FB10FBC2FB31" box="[999,1083,1185,1211]" pageId="24" pageNumber="25">Pascoe</collectorName>
, Coll. 9360;”
<collectionCode id="BBC9AE01FFEE891E9BF0FB10FAB0FB31" box="[1265,1353,1185,1211]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="24" pageNumber="25">BMNH</collectionCode>
<specimenCode id="8D7E9EBFFFEE891E9A50FB10FEDEFB6A" pageId="24" pageNumber="25">(E) 1237144</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEE891E9FBCFAB8FB70F863" pageId="24" pageNumber="25" type="description">
<paragraph id="DD6736C4FFEE891E9FBCFAB8FEB7FAA9" blockId="24.[189,334,1289,1315]" box="[189,334,1289,1315]" pageId="24" pageNumber="25">
<heading id="862F81A8FFEE891E9FBCFAB8FEB7FAA9" bold="true" box="[189,334,1289,1315]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="EFACEAD6FFEE891E9FBCFAB8FEB7FAA9" bold="true" box="[189,334,1289,1315]" pageId="24" pageNumber="25">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFA8AFCCBFADF" blockId="24.[189,818,1338,1365]" box="[189,818,1338,1365]" pageId="24" pageNumber="25">MEASUREMENTS (in mm). TL=12.518.5; BW=710.</paragraph>
<paragraph id="DD6736C4FFEE891E9FEBFACFFED3FA12" blockId="24.[189,1398,1406,1882]" box="[234,298,1406,1432]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FEBFACFFED3FA12" bold="true" box="[234,298,1406,1432]" pageId="24" pageNumber="25">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFA15FE6AF8D0" blockId="24.[189,1398,1406,1882]" pageId="24" pageNumber="25">
Bright metallic green, coppery, indigo-blue or violet, the elytra bright yellow with four narrow longitudinal green lines on the elevated 1
<superScript id="2AAD9B8CFFEE891E9DC2FA78FD36FA5D" attach="none" box="[707,719,1481,1495]" fontSize="6" pageId="24" pageNumber="25">st</superScript>
, 3
<superScript id="2AAD9B8CFFEE891E9DF1FA79FD06FA5C" attach="left" box="[752,767,1480,1494]" fontSize="6" pageId="24" pageNumber="25">rd</superScript>
and 5
<superScript id="2AAD9B8CFFEE891E9C4FFA79FCA5FA5C" attach="left" box="[846,860,1480,1494]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
intervals, pale intervals with light metallic lustre; body oval, convex, not very shining; head closely covered with fine tubercles; clypeus parabolic, rugose with yellow setae, clypeal carina curved, elevated, near to clypeal margin; frontal carina entire with a tubercle behind; pronotum metallic green, unevenly punctured posteriorly, front angles granular, basal foveae deep, close with granules anteriorly; elytra finely striate with sutural, 3
<superScript id="2AAD9B8CFFEE891E9E2EF935FEC7F918" attach="left" box="[303,318,1668,1682]" fontSize="6" pageId="24" pageNumber="25">rd</superScript>
and 5
<superScript id="2AAD9B8CFFEE891E9E89F935FE6FF918" attach="left" box="[392,406,1668,1682]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
intervals elevated, shining, finely punctured, the other intervals opaque, scarcely punctured; pygidium shining, finely, sparsely granular; metasternum with a longitudinal groove, metasternal shield broad, granular and thinly covered with hairs anteriorly, sparingly punctured behind, sides of metasternum closely granular and hairy; protibia elongated, slender with a strong tooth at the middle of the lower surface; hind femur with a very strong hooked tooth near the middle of its upper edge.
</paragraph>
<paragraph id="DD6736C4FFEE891E9FEBF835FEBCF814" blockId="24.[189,1398,1924,2025]" box="[234,325,1924,1950]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FEBF835FEBCF814" bold="true" box="[234,325,1924,1950]" pageId="24" pageNumber="25">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCF818FB70F863" blockId="24.[189,1398,1924,2025]" pageId="24" pageNumber="25">Clypeal carina long, curved, space between the carina and front margin covered with long yellow, closelying hairs; legs short, stout, protibia with four strong teeth and an articulated spur.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEF891F9FBCFEB9FD76FE31" pageId="25" pageNumber="26" type="distribution">
<paragraph id="DD6736C4FFEF891F9FBCFEB9FEAFFEA8" blockId="25.[189,342,264,290]" box="[189,342,264,290]" pageId="25" pageNumber="26">
<heading id="862F81A8FFEF891F9FBCFEB9FEAFFEA8" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="EFACEAD6FFEF891F9FBCFEB9FEAFFEA8" bold="true" box="[189,342,264,290]" pageId="25" pageNumber="26">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEF891F9FBCFE86FD76FE31" blockId="25.[189,1398,311,443]" pageId="25" pageNumber="26">
<collectingCountry id="A5CF7654FFEF891F9FBCFE86FEA9FEDB" box="[189,336,311,337]" name="Afghanistan" pageId="25" pageNumber="26">Afghanistan</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9E5BFE86FE42FEDB" box="[346,443,311,337]" name="Albania" pageId="25" pageNumber="26">Albania</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9EC4FE86FDC9FEDB" box="[453,560,311,337]" name="Armenia" pageId="25" pageNumber="26">Armenia</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9D3AFE86FD7CFEDB" box="[571,645,311,337]" name="China" pageId="25" pageNumber="26">China</collectingCountry>
;
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(
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</paragraph>
</subSubSection>
</treatment>
</document>