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<document id="657A7CA08110A32FB032AEEF0EDBE6E9" ID-DOI="10.5281/zenodo.3674855" ID-GBIF-Dataset="ac53680f-6358-4d6b-8880-ccf3dd73297e" ID-ISSN="1942-1354" ID-Zenodo-Dep="3674855" ID-ZooBank="F9BABD1C-B18B-47FD-9B87-F44C49FD9D30" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1582111424063" checkinUser="felipe" docAuthor="F, Guillermo González, Hanley, Guy A. &amp; Gordon, Robert D." docDate="2019" docId="4E5F8790C756952E8D90D4FD22A6FB08" docLanguage="en" docName="InsectaMundi.0714.1-32.pdf.imf" docOrigin="Insecta Mundi 714 (714)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Cryptognatha nodiceps Marshall 1912" docType="treatment" docVersion="8" lastPageNumber="8" masterDocId="B266FFE8C75E95278D50D20C2369FF89" masterDocTitle="South American Coccinellidae (Coleoptera), Part XIX: Overview of Cryptognathini and systematic revision of South American Cryptognatha Mulsant" masterLastPageNumber="32" masterPageNumber="1" pageNumber="7" updateTime="1698719361767" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="1C150688CEFE816F128B5FF7DE11D52F">South American Coccinellidae (Coleoptera), Part XIX: Overview of Cryptognathini and systematic revision of South American Cryptognatha Mulsant</mods:title>
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<mods:namePart id="2DD032E927FBD2B1D7041FB58E1F4D24">F, Guillermo González</mods:namePart>
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<mods:namePart id="F5620B0AAD4804047BD69863D5233B6D">Hanley, Guy A.</mods:namePart>
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<mods:namePart id="B9ED6B5330EC1750CE21856C612A8C4F">Gordon, Robert D.</mods:namePart>
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<mods:title id="EBB404CE2A5F346D69000A078A782B2F">Insecta Mundi</mods:title>
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<mods:date id="A8BA7EA0A2BC0BF04530987CB6C555BA">2019</mods:date>
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<treatment id="4E5F8790C756952E8D90D4FD22A6FB08" ID-DOI="http://doi.org/10.5281/zenodo.3680676" ID-GBIF-Taxon="162173262" ID-Zenodo-Dep="3680676" LSID="urn:lsid:plazi:treatment:4E5F8790C756952E8D90D4FD22A6FB08" httpUri="http://treatment.plazi.org/id/4E5F8790C756952E8D90D4FD22A6FB08" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<subSubSection id="8EEC650DC756952F8D90D4FD21F5F880" box="[192,668,1777,1801]" pageId="8" pageNumber="7" type="nomenclature">
<paragraph id="C6493686C756952F8D90D4FD21F5F880" blockId="8.[192,668,1777,1801]" box="[192,668,1777,1801]" pageId="8" pageNumber="7">
<heading id="9D0181EAC756952F8D90D4FD21F5F880" bold="true" box="[192,668,1777,1801]" fontSize="10" level="2" pageId="8" pageNumber="7" reason="3">
<emphasis id="F482EA94C756952F8D90D4FD21F5F880" bold="true" box="[192,668,1777,1801]" pageId="8" pageNumber="7">
2.
<taxonomicName id="01F64D05C756952F8D8ED4FD21F5F880" authority="Marshall" authorityName="Marshall" authorityYear="1912" box="[222,668,1777,1801]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="nodiceps">
<emphasis id="F482EA94C756952F8D8ED4FD2173F880" bold="true" box="[222,538,1777,1801]" italics="true" pageId="8" pageNumber="7">Cryptognatha nodiceps</emphasis>
Marshall
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC756952F8D90D52C27CFF8BF" box="[192,1190,1824,1846]" pageId="8" pageNumber="7" type="reference_group">
<paragraph id="C6493686C756952F8D90D52C27CFF8BF" blockId="8.[192,1190,1824,1846]" box="[192,1190,1824,1846]" pageId="8" pageNumber="7">
<treatmentCitationGroup id="E6E611A8C756952F8D90D52C27CFF8BF" box="[192,1190,1824,1846]" pageId="8" pageNumber="7">
<taxonomicName id="01F64D05C756952F8D90D52C21C9F8BF" authority="Marshall 1912: 321" authorityName="Marshall" authorityPageNumber="321" authorityYear="1912" box="[192,672,1824,1846]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="nodiceps">
<emphasis id="F482EA94C756952F8D90D52C22A9F8BC" box="[192,448,1824,1845]" italics="true" pageId="8" pageNumber="7">Cryptognatha nodiceps</emphasis>
<treatmentCitation id="47571097C756952F8C97D52C21C9F8BF" author="Marshall, G. A. K." box="[455,672,1824,1846]" page="321" pageId="8" pageNumber="7" year="1912">
<bibRefCitation id="A2674B77C756952F8C97D52C21C9F8BF" author="Marshall, G. A. K." box="[455,672,1824,1846]" pageId="8" pageNumber="7" pagination="320 - 322" refId="ref19624" refString="Marshall, G. A. K. 1912. Three new species of Neotropical Coccinellidae. Annals and Magazine of Natural History, ser. 8, 10: 320 - 322." type="journal article" year="1912">Marshall 1912: 321</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="47571097C756952F8FFBD52C20F3F8BF" author="Korshefsky" box="[683,922,1824,1846]" page="219" pageId="8" pageNumber="7" year="1931">Korshefsky 1931: 219</treatmentCitation>
;
<treatmentCitation id="47571097C756952F8EF5D52C27CBF8BF" author="Blackwelder, E." box="[933,1186,1824,1846]" page="449" pageId="8" pageNumber="7" year="1945">
<bibRefCitation id="A2674B77C756952F8EF5D52C27CBF8BF" author="Blackwelder, E." box="[933,1186,1824,1846]" pageId="8" pageNumber="7" pagination="343 - 550" refId="ref18961" refString="Blackwelder, E. 1945. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 3. Bulletin of the United States National Museum 185: 343 - 550." type="journal article" year="1945">Blackwelder 1945: 449</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC756952E8D90D54327D3FCD8" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7" type="description">
<paragraph id="C6493686C756952E8D90D543263EFD1B" blockId="8.[192,1440,1871,1929]" lastBlockId="9.[192,1441,217,658]" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<emphasis id="F482EA94C756952F8D90D54322D5F8E1" bold="true" box="[192,444,1871,1896]" pageId="8" pageNumber="7">Description. Male.</emphasis>
Length
<quantity id="010E9B63C756952F8F76D55C21ECF8EE" box="[550,645,1872,1896]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="7" unit="mm" value="2.6">2.6 mm</quantity>
, width
<quantity id="010E9B63C756952F8FB2D55C202AF8EE" box="[738,835,1872,1896]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="7" unit="mm" value="2.6">2.6 mm</quantity>
. Dorsal surface entirely shiny. Color yellow (
<figureCitation id="5ECD2A03C756952F8820D55C23A6F800" captionStart="Figures 120" captionStartId="25.[193,288,1788,1810]" captionTargetBox="[205,1418,300,1672]" captionTargetId="figure@25.[194,1440,283,1714]" captionTargetPageId="25" captionText="Figures 120. Cryptognatha spp. 17) Cryptognatha auriculata. 1) Habitus. 2) Frons. 3) Proleg. 4) Phallobase ventral. 5) Phallobase lateral. 6) Penis. 7) Genital plates. 814) Cryptognatha nodiceps. 8) Habitus. 9) Frons. 10) Proleg. 11) Phallobase ventral. 12) Phallobase lateral. 13) Penis. 14) Spermathecal capsule. 1520) Cryptognatha gemellata. 15) Habitus. 16) Frons. 17) Spermathecal capsule. 18) Phallobase ventral. 19) Phallobase lateral. 20) Penis." figureDoi="http://doi.org/10.5281/zenodo.3674857" httpUri="https://zenodo.org/record/3674857/files/figure.png" pageId="8" pageNumber="7">Fig. 8</figureCitation>
); pronotum with 5 pale brown spots, 2 on disc, 3 along posterior margin; elytron with irregular black spot occupying most of disc, and elongate longitudinal spot close to sutural margin anterior to middle (
<figureCitation id="5ECD2A03C757952E8D99D2F52279FE9B" box="[201,272,249,274]" captionStart="Figures 120" captionStartId="25.[193,288,1788,1810]" captionTargetBox="[205,1418,300,1672]" captionTargetId="figure@25.[194,1440,283,1714]" captionTargetPageId="25" captionText="Figures 120. Cryptognatha spp. 17) Cryptognatha auriculata. 1) Habitus. 2) Frons. 3) Proleg. 4) Phallobase ventral. 5) Phallobase lateral. 6) Penis. 7) Genital plates. 814) Cryptognatha nodiceps. 8) Habitus. 9) Frons. 10) Proleg. 11) Phallobase ventral. 12) Phallobase lateral. 13) Penis. 14) Spermathecal capsule. 1520) Cryptognatha gemellata. 15) Habitus. 16) Frons. 17) Spermathecal capsule. 18) Phallobase ventral. 19) Phallobase lateral. 20) Penis." figureDoi="http://doi.org/10.5281/zenodo.3674857" httpUri="https://zenodo.org/record/3674857/files/figure.png" pageId="9" pageNumber="8">Fig. 8</figureCitation>
). Head without punctures; pronotal punctures small, separated by less than a diameter; elytral punctures slightly larger than on pronotum, separated by less than a diameter to twice a diameter; prosternal punctures small, separated by less than a diameter; meso- and metasternal punctures large, separated by a diameter or less on mesosternum, widely separated on median 1/3 of metasternum, lateral 1/3 without punctures; basal abdominal ventrite with coarse punctures separated by about a diameter medially, lateral 1/3 of ventrite and ventrites 24 lacking punctation, ventrite 5 with dense, fine punctation. Head with frons parallel-sided, medially depressed, twice as wide as eye, apex truncate medially, lateral 1/8 slightly retracted, angle acute (
<figureCitation id="5ECD2A03C757952E8E16D3D520E6FE7B" box="[838,911,473,498]" captionStart="Figures 120" captionStartId="25.[193,288,1788,1810]" captionTargetBox="[205,1418,300,1672]" captionTargetId="figure@25.[194,1440,283,1714]" captionTargetPageId="25" captionText="Figures 120. Cryptognatha spp. 17) Cryptognatha auriculata. 1) Habitus. 2) Frons. 3) Proleg. 4) Phallobase ventral. 5) Phallobase lateral. 6) Penis. 7) Genital plates. 814) Cryptognatha nodiceps. 8) Habitus. 9) Frons. 10) Proleg. 11) Phallobase ventral. 12) Phallobase lateral. 13) Penis. 14) Spermathecal capsule. 1520) Cryptognatha gemellata. 15) Habitus. 16) Frons. 17) Spermathecal capsule. 18) Phallobase ventral. 19) Phallobase lateral. 20) Penis." figureDoi="http://doi.org/10.5281/zenodo.3674857" httpUri="https://zenodo.org/record/3674857/files/figure.png" pageId="9" pageNumber="8">Fig. 9</figureCitation>
); eye canthus long, about 1/2 width of eye. Prosternum with lateral carina on each side extended from apex of intercoxal process nearly to midpoint of prosternum. Epipleuron strongly descending externally. Postcoxal line on basal abdominal ventrite evenly curved, extended 3/4 distance to rear margin of ventrite. Genitalia long, penis guide nearly as long as paramere, parallel-sided with apical 1/8 narrowed to rounded apex; paramere slender, parallelsided, apex rounded (
<figureCitation id="5ECD2A03C757952E8C95D075212EFD1B" box="[453,583,633,658]" captionStart="Figures 120" captionStartId="25.[193,288,1788,1810]" captionTargetBox="[205,1418,300,1672]" captionTargetId="figure@25.[194,1440,283,1714]" captionTargetPageId="25" captionText="Figures 120. Cryptognatha spp. 17) Cryptognatha auriculata. 1) Habitus. 2) Frons. 3) Proleg. 4) Phallobase ventral. 5) Phallobase lateral. 6) Penis. 7) Genital plates. 814) Cryptognatha nodiceps. 8) Habitus. 9) Frons. 10) Proleg. 11) Phallobase ventral. 12) Phallobase lateral. 13) Penis. 14) Spermathecal capsule. 1520) Cryptognatha gemellata. 15) Habitus. 16) Frons. 17) Spermathecal capsule. 18) Phallobase ventral. 19) Phallobase lateral. 20) Penis." figureDoi="http://doi.org/10.5281/zenodo.3674857" httpUri="https://zenodo.org/record/3674857/files/figure.png" pageId="9" pageNumber="8">Fig. 11, 12</figureCitation>
); penis slender, equal in width throughout, apex acute (
<figureCitation id="5ECD2A03C757952E89A3D0752623FD1B" box="[1267,1354,633,658]" captionStart="Figures 120" captionStartId="25.[193,288,1788,1810]" captionTargetBox="[205,1418,300,1672]" captionTargetId="figure@25.[194,1440,283,1714]" captionTargetPageId="25" captionText="Figures 120. Cryptognatha spp. 17) Cryptognatha auriculata. 1) Habitus. 2) Frons. 3) Proleg. 4) Phallobase ventral. 5) Phallobase lateral. 6) Penis. 7) Genital plates. 814) Cryptognatha nodiceps. 8) Habitus. 9) Frons. 10) Proleg. 11) Phallobase ventral. 12) Phallobase lateral. 13) Penis. 14) Spermathecal capsule. 1520) Cryptognatha gemellata. 15) Habitus. 16) Frons. 17) Spermathecal capsule. 18) Phallobase ventral. 19) Phallobase lateral. 20) Penis." figureDoi="http://doi.org/10.5281/zenodo.3674857" httpUri="https://zenodo.org/record/3674857/files/figure.png" pageId="9" pageNumber="8">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="C6493686C757952E8D90D0A5214BFC8B" blockId="9.[192,1440,681,770]" pageId="9" pageNumber="8">
<emphasis id="F482EA94C757952E8D90D0A52245FD48" bold="true" box="[192,300,681,705]" pageId="9" pageNumber="8">Female.</emphasis>
Similar to male except head weakly depressed, dark yellow; pronotum brownish yellow medially, lacking brown spots; genitalia with spermathecal capsule slender, evenly curved from ramus to acute apex of cornu (
<figureCitation id="5ECD2A03C757952E8CEDD0E5217DFC8B" box="[445,532,745,770]" captionStart="Figures 120" captionStartId="25.[193,288,1788,1810]" captionTargetBox="[205,1418,300,1672]" captionTargetId="figure@25.[194,1440,283,1714]" captionTargetPageId="25" captionText="Figures 120. Cryptognatha spp. 17) Cryptognatha auriculata. 1) Habitus. 2) Frons. 3) Proleg. 4) Phallobase ventral. 5) Phallobase lateral. 6) Penis. 7) Genital plates. 814) Cryptognatha nodiceps. 8) Habitus. 9) Frons. 10) Proleg. 11) Phallobase ventral. 12) Phallobase lateral. 13) Penis. 14) Spermathecal capsule. 1520) Cryptognatha gemellata. 15) Habitus. 16) Frons. 17) Spermathecal capsule. 18) Phallobase ventral. 19) Phallobase lateral. 20) Penis." figureDoi="http://doi.org/10.5281/zenodo.3674857" httpUri="https://zenodo.org/record/3674857/files/figure.png" pageId="9" pageNumber="8">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="C6493686C757952E8D90D11527D3FCD8" blockId="9.[192,1440,793,849]" pageId="9" pageNumber="8">
<emphasis id="F482EA94C757952E8D90D1152222FCB8" bold="true" box="[192,331,793,817]" pageId="9" pageNumber="8">Variation.</emphasis>
Length
<quantity id="010E9B63C757952E8CE6D116210CFCB8" box="[438,613,794,818]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="9" pageNumber="8" unit="mm" value="2.65" valueMax="2.9" valueMin="2.4">2.4 to 2.9 mm</quantity>
, width 2.0 to
<quantity id="010E9B63C757952E8E42D116201DFCB8" box="[786,884,794,818]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="9" pageNumber="8" unit="mm" value="2.7">2.7 mm</quantity>
. Elytral color highly variable from typical to having disc entirely black with reddish yellow spot near suture posterior to middle.
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC757952E8D90D1652131FC08" box="[192,600,873,897]" pageId="9" pageNumber="8" type="distribution">
<paragraph id="C6493686C757952E8D90D1652131FC08" blockId="9.[192,600,873,897]" box="[192,600,873,897]" pageId="9" pageNumber="8">
<emphasis id="F482EA94C757952E8D90D1652212FC08" bold="true" box="[192,379,873,897]" pageId="9" pageNumber="8">
<typeStatus id="194D8824C757952E8D90D165226DFC08" box="[192,260,873,897]" pageId="9" pageNumber="8">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="BEE17616C757952E8CD2D1652299FC08" box="[386,496,873,897]" name="Trinidad and Tobago" pageId="9" pageNumber="8">Trinidad</collectingCountry>
, Cedros.
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC757952E8D90D1952167FC38" box="[192,526,921,945]" pageId="9" pageNumber="8" type="materials_examined">
<paragraph id="C6493686C757952E8D90D1952167FC38" blockId="9.[192,526,921,945]" box="[192,526,921,945]" pageId="9" pageNumber="8">
<materialsCitation id="769E3CDBC757952E8D90D1952167FC38" ID-GBIF-Occurrence="2569343964" box="[192,526,921,945]" pageId="9" pageNumber="8">
<emphasis id="F482EA94C757952E8D90D19522CEFC38" bold="true" box="[192,423,921,945]" pageId="9" pageNumber="8">
<typeStatus id="194D8824C757952E8D90D195226DFC38" box="[192,260,921,945]" pageId="9" pageNumber="8">Type</typeStatus>
depository.
</emphasis>
BMNH.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC757952E8D90D1C52070FC68" box="[192,793,969,994]" pageId="9" pageNumber="8" type="distribution">
<paragraph id="C6493686C757952E8D90D1C52070FC68" blockId="9.[192,793,969,994]" box="[192,793,969,994]" pageId="9" pageNumber="8">
<emphasis id="F482EA94C757952E8D90D1C52146FC68" bold="true" box="[192,559,969,993]" pageId="9" pageNumber="8">Geographical distribution.</emphasis>
<collectingCountry id="BEE17616C757952E8F66D1C621F3FC6B" box="[566,666,970,994]" name="Guyana" pageId="9" pageNumber="8">Guyana</collectingCountry>
,
<collectingCountry id="BEE17616C757952E8FF6D1C5207CFC68" box="[678,789,969,993]" name="Trinidad and Tobago" pageId="9" pageNumber="8">Trinidad</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC757952E8D90D1F522DCFBBB" pageId="9" pageNumber="8" type="materials_examined">
<paragraph id="C6493686C757952E8D90D1F522DCFBBB" blockId="9.[192,1440,1017,1074]" pageId="9" pageNumber="8">
<materialsCitation id="769E3CDBC757952E8D90D1F522DCFBBB" ID-GBIF-Occurrence="2569343978" pageId="9" pageNumber="8">
<emphasis id="F482EA94C757952E8D90D1F52282FB98" bold="true" box="[192,491,1017,1041]" pageId="9" pageNumber="8">Specimens examined.</emphasis>
44. Distributed throughout
<collectingCountry id="BEE17616C757952E8E1CD1F520D3FB98" box="[844,954,1017,1041]" name="Trinidad and Tobago" pageId="9" pageNumber="8">Trinidad</collectingCountry>
, also a single specimen examined from Blairmont,
<collectingCountry id="BEE17616C757952E8C1CD61622D9FBBB" box="[332,432,1050,1074]" name="Guyana" pageId="9" pageNumber="8">Guyana</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC757952E8D90D64522A6FB08" pageId="9" pageNumber="8" type="discussion">
<paragraph id="C6493686C757952E8D90D64522A6FB08" blockId="9.[192,1440,1097,1153]" pageId="9" pageNumber="8">
<emphasis id="F482EA94C757952E8D90D645222AFBE8" bold="true" box="[192,323,1097,1121]" pageId="9" pageNumber="8">Remarks.</emphasis>
This species has a distinctive dorsal color pattern, but male genitalia appear identical to those of
<taxonomicName id="01F64D05C757952E8C76D66622A0FB08" authorityName="Mulsant" authorityYear="1850" box="[294,457,1129,1153]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="auriculata">
<emphasis id="F482EA94C757952E8C76D6662250FB08" box="[294,313,1130,1153]" italics="true" pageId="9" pageNumber="8">C</emphasis>
.
<emphasis id="F482EA94C757952E8C18D66522A0FB08" box="[328,457,1129,1153]" italics="true" pageId="9" pageNumber="8">auriculata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>