treatments-xml/data/4E/5F/87/4E5F8790C74D95338D90D79122C9FCC8.xml
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<document id="A7C0C78370675D6811B50AEC203D478D" ID-DOI="10.5281/zenodo.3674855" ID-GBIF-Dataset="ac53680f-6358-4d6b-8880-ccf3dd73297e" ID-ISSN="1942-1354" ID-Zenodo-Dep="3674855" ID-ZooBank="F9BABD1C-B18B-47FD-9B87-F44C49FD9D30" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1582111424063" checkinUser="felipe" docAuthor="F, Guillermo González, Hanley, Guy A. &amp; Gordon, Robert D." docDate="2019" docId="4E5F8790C74D95338D90D79122C9FCC8" docLanguage="en" docName="InsectaMundi.0714.1-32.pdf.imf" docOrigin="Insecta Mundi 714 (714)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Cryptognatha batesi Crotch" docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="B266FFE8C75E95278D50D20C2369FF89" masterDocTitle="South American Coccinellidae (Coleoptera), Part XIX: Overview of Cryptognathini and systematic revision of South American Cryptognatha Mulsant" masterLastPageNumber="32" masterPageNumber="1" pageNumber="18" updateTime="1698719361767" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="5C5F29D7FCDD267A8BD11F79C0E04DE5">South American Coccinellidae (Coleoptera), Part XIX: Overview of Cryptognathini and systematic revision of South American Cryptognatha Mulsant</mods:title>
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<mods:namePart id="6392C38B38006F7AC7C34A89682C354F">F, Guillermo González</mods:namePart>
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<mods:namePart id="4E54E783562CDB3CE5DDA0725B55B2D9">Hanley, Guy A.</mods:namePart>
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<mods:namePart id="BEF879E89AF5DFC1D89512901C49CE48">Gordon, Robert D.</mods:namePart>
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<mods:title id="1D54620CE25A4FEAF155DD460C8214E5">Insecta Mundi</mods:title>
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<treatment id="4E5F8790C74D95338D90D79122C9FCC8" ID-DOI="http://doi.org/10.5281/zenodo.3680650" ID-GBIF-Taxon="162173276" ID-Zenodo-Dep="3680650" LSID="urn:lsid:plazi:treatment:4E5F8790C74D95338D90D79122C9FCC8" httpUri="http://treatment.plazi.org/id/4E5F8790C74D95338D90D79122C9FCC8" lastPageId="20" lastPageNumber="19" pageId="19" pageNumber="18">
<subSubSection id="8EEC650DC74D95348D90D7912102FA3C" box="[192,619,1436,1461]" pageId="19" pageNumber="18" type="nomenclature">
<paragraph id="C6493686C74D95348D90D7912102FA3C" blockId="19.[192,619,1436,1461]" box="[192,619,1436,1461]" pageId="19" pageNumber="18">
<heading id="9D0181EAC74D95348D90D7912102FA3C" bold="true" box="[192,619,1436,1461]" fontSize="10" level="2" pageId="19" pageNumber="18" reason="3">
<emphasis id="F482EA94C74D95348D90D7912102FA3C" bold="true" box="[192,619,1436,1461]" pageId="19" pageNumber="18">
18.
<taxonomicName id="01F64D05C74D95348DBDD7902102FA3C" authority="Crotch" authorityName="Crotch" box="[237,619,1436,1461]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="F482EA94C74D95348DBDD790216DFA3D" bold="true" box="[237,516,1436,1460]" italics="true" pageId="19" pageNumber="18">Cryptognatha batesi</emphasis>
Crotch
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74D95348D90D7C72653FA68" box="[192,1338,1483,1505]" pageId="19" pageNumber="18" type="reference_group">
<paragraph id="C6493686C74D95348D90D7C72653FA68" blockId="19.[192,1338,1483,1505]" box="[192,1338,1483,1505]" pageId="19" pageNumber="18">
<treatmentCitationGroup id="E6E611A8C74D95348D90D7C72653FA68" box="[192,1338,1483,1505]" pageId="19" pageNumber="18">
<taxonomicName id="01F64D05C74D95348D90D7C7210EFA68" authority="Crotch 1874: 206" authorityName="Crotch" authorityPageNumber="206" authorityYear="1874" box="[192,615,1483,1505]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="F482EA94C74D95348D90D7C722C8FA69" box="[192,417,1483,1504]" italics="true" pageId="19" pageNumber="18">Cryptognatha batesi</emphasis>
<treatmentCitation id="47571097C74D95348CF8D7C7210EFA68" author="Crotch, G. R." box="[424,615,1483,1505]" page="206" pageId="19" pageNumber="18" year="1874">
<bibRefCitation id="A2674B77C74D95348CF8D7C7210EFA68" author="Crotch, G. R." box="[424,615,1483,1505]" pageId="19" pageNumber="18" refId="ref19186" refString="Crotch, G. R. 1874. A revision of the coleopterous family Coccinellidae. E. W. Janson; London. 311 p." type="book" year="1874">Crotch 1874: 206</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="47571097C74D95348F21D7C72005FA68" author="Korschefsky, R." box="[625,876,1483,1505]" page="219" pageId="19" pageNumber="18" year="1931">
<bibRefCitation id="A2674B77C74D95348F21D7C72005FA68" author="Korschefsky, R." box="[625,876,1483,1505]" pageId="19" pageNumber="18" refId="ref19600" refString="Korschefsky, R. 1931. Coccinellidae I. Coleopterorum Catalogus. Part 118. W. Junk; Berlin. 224 p." type="book" year="1931">Korschefsky 1931: 219</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="47571097C74D95348E26D7C7271BFA68" author="Blackwelder, E." box="[886,1138,1483,1505]" page="449" pageId="19" pageNumber="18" year="1945">
<bibRefCitation id="A2674B77C74D95348E26D7C7271BFA68" author="Blackwelder, E." box="[886,1138,1483,1505]" pageId="19" pageNumber="18" pagination="343 - 550" refId="ref18961" refString="Blackwelder, E. 1945. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 3. Bulletin of the United States National Museum 185: 343 - 550." type="journal article" year="1945">Blackwelder 1945: 449</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="47571097C74D9534892ED7C7265FFA68" author="Gordon, R. D." box="[1150,1334,1483,1505]" page="26" pageId="19" pageNumber="18" year="1987">
<bibRefCitation id="A2674B77C74D9534892ED7C7265FFA68" author="Gordon, R. D." box="[1150,1334,1483,1505]" pageId="19" pageNumber="18" pagination="1 - 46" refId="ref19285" refString="Gordon, R. D. 1987. A catalogue of the Crotch collection of Coccinellidae (Coleoptera). Occasional Papers on Systematic Entomology 3: 1 - 46." type="journal article" year="1987">Gordon 1987: 26</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74D95338D90D7F522BBFD98" lastPageId="20" lastPageNumber="19" pageId="19" pageNumber="18" type="description">
<paragraph id="C6493686C74D95338D90D7F521B3FE38" blockId="19.[192,1441,1529,1906]" lastBlockId="20.[192,1440,217,433]" lastPageId="20" lastPageNumber="19" pageId="19" pageNumber="18">
<emphasis id="F482EA94C74D95348D90D7F52153F99B" bold="true" box="[192,570,1529,1554]" pageId="19" pageNumber="18">
Description. Male
<typeStatus id="194D8824C74D95348CEAD7F6215DF99B" box="[442,564,1530,1554]" pageId="19" pageNumber="18" type="holotype">holotype</typeStatus>
.
</emphasis>
Length
<quantity id="010E9B63C74D95348FCFD7F62195F998" box="[671,764,1530,1554]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="19" pageNumber="18" unit="mm" value="3.8">3.8 mm</quantity>
, width
<quantity id="010E9B63C74D95348E05D7F620DAF998" box="[853,947,1530,1554]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="19" pageNumber="18" unit="mm" value="3.4">3.4 mm</quantity>
. Dorsal surface shiny. Color black; head, pronotum pale yellow, pronotum with faint, brown basomedian macula (
<figureCitation id="5ECD2A03C74D9534895FD4162708F9BB" box="[1039,1121,1562,1586]" captionStart="Figures 86102" captionStartId="30.[192,287,1820,1842]" captionTargetBox="[209,1423,326,1699]" captionTargetId="figure@30.[192,1438,309,1740]" captionTargetPageId="30" captionText="Figures 86102. Cryptognatha spp. 8690) Cryptognatha yolandi. 86) Habitus. 87) Frons. 88) Phallobase ventral. 89) Phallobase lateral. 90) Penis. 9198) Cryptognatha pudibunda. 91) Habitus. 92) Frons. 93) Maxillary palpus. 94) Antenna. 95) Phallobase ventral. 96) Phallobase lateral. 97) Penis. 98) Spermathecal capsule. 99102) Cryptognatha batesi. 99) Habitus. 100) Frons. 101) Phallobase lateral. 102) Phallobase ventral." figureDoi="http://doi.org/10.5281/zenodo.3674867" httpUri="https://zenodo.org/record/3674867/files/figure.png" pageId="19" pageNumber="18">Fig. 99</figureCitation>
); venter dark brown except mouthparts, legs yellow; abdomen brown except mediobasal portion of basal ventrite dark brown. Head finely punctured, punctures separated by less than a diameter; pronotal punctures small, separated by a diameter or less; elytral punctures larger than on pronotum, separated by less than a diameter to twice a diameter, becoming gradually larger toward lateral margin; prosternal punctures large, indistinct; mesosternal punctures larger than on prosternum, contiguous, metasternal punctures large, separated by a diameter or less medially, absent in lateral 1/3; basal abdominal ventrite with coarse punctures separated by about a diameter medially, lateral 1/3 of ventrite lacking punctation, ventrites 24 finely punctured, punctures separated by about a diameter, ventrite 5 with dense, fine punctation. Head with frons slightly narrowed from base to clypeus, medially depressed, three times as wide as eye, apex emarginate medially, strong lateral carina present on frons near apex of eye (
<figureCitation id="5ECD2A03C74D953489DDD556279CF8FB" box="[1165,1269,1882,1906]" captionStart="Figures 86102" captionStartId="30.[192,287,1820,1842]" captionTargetBox="[209,1423,326,1699]" captionTargetId="figure@30.[192,1438,309,1740]" captionTargetPageId="30" captionText="Figures 86102. Cryptognatha spp. 8690) Cryptognatha yolandi. 86) Habitus. 87) Frons. 88) Phallobase ventral. 89) Phallobase lateral. 90) Penis. 9198) Cryptognatha pudibunda. 91) Habitus. 92) Frons. 93) Maxillary palpus. 94) Antenna. 95) Phallobase ventral. 96) Phallobase lateral. 97) Penis. 98) Spermathecal capsule. 99102) Cryptognatha batesi. 99) Habitus. 100) Frons. 101) Phallobase lateral. 102) Phallobase ventral." figureDoi="http://doi.org/10.5281/zenodo.3674867" httpUri="https://zenodo.org/record/3674867/files/figure.png" pageId="19" pageNumber="18">Fig. 100</figureCitation>
); eye canthus short, about 1/2 width of eye. Prosternum with lateral carina on each side long, curved, extended from apex of intercoxal process about 2/3 distance to apex of prosternum. Epipleuron moderately descending externally. Postcoxal line on basal abdominal ventrite evenly curved, extended about 3/4 distance to rear margin of ventrite. Genitalia long, penis guide 3/4 as long as paramere, slender, evenly narrowed from base to abruptly rounded apex; paramere long, slender, curved to rounded apex (
<figureCitation id="5ECD2A03C74A953389B3D3552621FEFB" box="[1251,1352,345,370]" captionStart="Figures 86102" captionStartId="30.[192,287,1820,1842]" captionTargetBox="[209,1423,326,1699]" captionTargetId="figure@30.[192,1438,309,1740]" captionTargetPageId="30" captionText="Figures 86102. Cryptognatha spp. 8690) Cryptognatha yolandi. 86) Habitus. 87) Frons. 88) Phallobase ventral. 89) Phallobase lateral. 90) Penis. 9198) Cryptognatha pudibunda. 91) Habitus. 92) Frons. 93) Maxillary palpus. 94) Antenna. 95) Phallobase ventral. 96) Phallobase lateral. 97) Penis. 98) Spermathecal capsule. 99102) Cryptognatha batesi. 99) Habitus. 100) Frons. 101) Phallobase lateral. 102) Phallobase ventral." figureDoi="http://doi.org/10.5281/zenodo.3674867" httpUri="https://zenodo.org/record/3674867/files/figure.png" pageId="20" pageNumber="19">Fig. 101</figureCitation>
); penis long, apex acute; penis capsule slender, elongate, inner arm slightly curved, apex rounded, outer arm slender, apex truncate. Penis not examined.
</paragraph>
<paragraph id="C6493686C74A95338D90D3C522DAFE68" blockId="20.[192,435,457,481]" box="[192,435,457,481]" pageId="20" pageNumber="19">
<emphasis id="F482EA94C74A95338D90D3C52245FE68" bold="true" box="[192,300,457,481]" pageId="20" pageNumber="19">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="C6493686C74A95338D90D3F522BBFD98" blockId="20.[192,466,505,529]" box="[192,466,505,529]" pageId="20" pageNumber="19">
<emphasis id="F482EA94C74A95338D90D3F52222FD98" bold="true" box="[192,331,505,529]" pageId="20" pageNumber="19">Variation.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74A95338D90D0252174FDC8" box="[192,541,553,578]" pageId="20" pageNumber="19" type="distribution">
<paragraph id="C6493686C74A95338D90D0252174FDC8" blockId="20.[192,541,553,578]" box="[192,541,553,578]" pageId="20" pageNumber="19">
<emphasis id="F482EA94C74A95338D90D0252212FDC8" bold="true" box="[192,379,553,577]" pageId="20" pageNumber="19">
<typeStatus id="194D8824C74A95338D90D025226DFDC8" box="[192,260,553,577]" pageId="20" pageNumber="19">Type</typeStatus>
locality.
</emphasis>
Ega (
<collectingCountry id="BEE17616C74A95338C92D0252178FDC8" box="[450,529,553,577]" name="Brazil" pageId="20" pageNumber="19">Brazil</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74A95338D90D055204AFDFB" box="[192,803,601,626]" pageId="20" pageNumber="19" type="materials_examined">
<paragraph id="C6493686C74A95338D90D055204AFDFB" blockId="20.[192,803,601,626]" box="[192,803,601,626]" pageId="20" pageNumber="19">
<materialsCitation id="769E3CDBC74A95338D90D055204AFDFB" ID-GBIF-Occurrence="2569343959" box="[192,803,601,626]" pageId="20" pageNumber="19" typeStatus="holotype">
<emphasis id="F482EA94C74A95338D90D05522E0FDF8" bold="true" box="[192,393,601,625]" pageId="20" pageNumber="19">Type material.</emphasis>
UMZC.
<typeStatus id="194D8824C74A95338CA1D055210BFDF8" box="[497,610,601,625]" pageId="20" pageNumber="19" type="holotype">Holotype</typeStatus>
,
<bibRefCitation id="A2674B77C74A95338F3DD0552049FDFB" author="Gordon, R. D." box="[621,800,601,626]" pageId="20" pageNumber="19" pagination="1 - 46" refId="ref19285" refString="Gordon, R. D. 1987. A catalogue of the Crotch collection of Coccinellidae (Coleoptera). Occasional Papers on Systematic Entomology 3: 1 - 46." type="journal article" year="1987">Gordon (1987)</bibRefCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74A95338D90D08521E0FD28" box="[192,649,649,673]" pageId="20" pageNumber="19" type="distribution">
<paragraph id="C6493686C74A95338D90D08521E0FD28" blockId="20.[192,649,649,673]" box="[192,649,649,673]" pageId="20" pageNumber="19">
<emphasis id="F482EA94C74A95338D90D0852146FD28" bold="true" box="[192,559,649,673]" pageId="20" pageNumber="19">Geographical distribution.</emphasis>
<collectingCountry id="BEE17616C74A95338F66D08521ECFD28" box="[566,645,649,673]" name="Brazil" pageId="20" pageNumber="19">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74A95338D90D0B52114FD58" box="[192,637,697,722]" pageId="20" pageNumber="19" type="materials_examined">
<paragraph id="C6493686C74A95338D90D0B52114FD58" blockId="20.[192,637,697,722]" box="[192,637,697,722]" pageId="20" pageNumber="19">
<materialsCitation id="769E3CDBC74A95338D90D0B52114FD58" ID-GBIF-Occurrence="2569343931" box="[192,637,697,722]" pageId="20" pageNumber="19" typeStatus="holotype">
<emphasis id="F482EA94C74A95338D90D0B52280FD58" bold="true" box="[192,489,697,721]" pageId="20" pageNumber="19">Specimens examined.</emphasis>
1,
<typeStatus id="194D8824C74A95338F5DD0B52110FD58" box="[525,633,697,721]" pageId="20" pageNumber="19" type="holotype">holotype</typeStatus>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74A95338D90D0E522C9FCC8" pageId="20" pageNumber="19" type="discussion">
<paragraph id="C6493686C74A95338D90D0E522C9FCC8" blockId="20.[192,1440,745,833]" pageId="20" pageNumber="19">
<emphasis id="F482EA94C74A95338D90D0E5222AFC88" bold="true" box="[192,323,745,769]" pageId="20" pageNumber="19">Remarks.</emphasis>
This is an unusual species because of large size, striking contrast between black elytra and pale-yellow head, pronotum, emarginate clypeal apex, and the pronounced frontal carinae. Known only from the
<typeStatus id="194D8824C74A95338C60D12522F5FCC8" box="[304,412,809,833]" pageId="20" pageNumber="19" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>