treatments-xml/data/4D/2B/7E/4D2B7E2EFFBAED704ED4FD96FAFF32B1.xml

<document id="C55EC8EDF04AFA6988F30AC07C61E33B" ID-DOI="10.11646/zootaxa.4759.3.2" ID-GBIF-Dataset="91fa4568-747a-4854-92d0-3159264955a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="3741162" ID-ZooBank="435819AF-DFBB-4B32-892F-6A794C57F73B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586156542478" checkinUser="plazi" docAuthor="Vargovitsh, Robert S. &amp; Kahrarian, Morteza" docDate="2020" docId="4D2B7E2EFFBAED704ED4FD96FAFF32B1" docLanguage="en" docName="zootaxa.4759.3.2.pdf" docOrigin="Zootaxa 4759 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arrhopalites persicus Vargovitsh &amp; Kahrarian 2020, sp. nov." docType="treatment" docUuid="A2C5FCE7-32F2-4C72-B53A-8E57D09CC35A" docUuidSource="ZooBank" docVersion="5" lastPageNumber="343" masterDocId="B1120656FFBBED754E43FFF1FFD2324A" masterDocTitle="A new species of Arrhopalites Börner, 1906 (Collembola, Symphypleona, Arrhopalitidae) from Iran with an updated key to A. diversus group of species" masterLastPageNumber="350" masterPageNumber="338" pageNumber="339" updateTime="1698735725467" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="56FE71839E1D0749E89AB56E600F3A62" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="32203B9A4E3FEAC6B39B603551C3B3E5">
<mods:title id="CBE2C765708C3741109D5C063E000358">A new species of Arrhopalites Börner, 1906 (Collembola, Symphypleona, Arrhopalitidae) from Iran with an updated key to A. diversus group of species</mods:title>
</mods:titleInfo>
<mods:name id="5FBC8FC180CEC80413E79FC1D3F01B4A" type="personal">
<mods:role id="AFA566BB5934DD549ACCD6208BEF9EAC">
<mods:roleTerm id="8D20E7E6D05BCCDBC86375F8005B5FA6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D5D546FA469718938EDC0F3E7E026F18">Vargovitsh, Robert S.</mods:namePart>
</mods:name>
<mods:name id="79A68B58C38F12AA0FF6059D42CD9865" type="personal">
<mods:role id="335EFB4CAA4BD107A66D87975A79455A">
<mods:roleTerm id="D8723440D74417731957AF776A83AE3F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="87782F30311C0B87D1C961990796EA5F">Kahrarian, Morteza</mods:namePart>
</mods:name>
<mods:typeOfResource id="88383B0F520CF9DBEA46A9C0601FEDA8">text</mods:typeOfResource>
<mods:relatedItem id="638FC8F9F534302296C80F8A20D9C028" type="host">
<mods:titleInfo id="947EC0544047F42EB3699839796D447A">
<mods:title id="371CF33ED75FEBFF3EF2F253108AB27D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="454E35F6E5D0DF6425E995E887FB83E3">
<mods:date id="44708E19EC6197C709D17A6BEAA5F05F">2020</mods:date>
<mods:detail id="C25DE8D1323AC72F9FBD6175FF853F9E" type="pubDate">
<mods:number id="A378C9622B614B95D8749BE60605273D">2020-04-03</mods:number>
</mods:detail>
<mods:detail id="0EAA6BB504A9D80C43AA9D2AD13A2369" type="volume">
<mods:number id="B772E8BACA630966072DA6A990F938ED">4759</mods:number>
</mods:detail>
<mods:detail id="618704CE4C34305151F6539BD8CBD7C2" type="issue">
<mods:number id="BC957284F5DEA70E6A7FD3C69E1FA571">3</mods:number>
</mods:detail>
<mods:extent id="22227E6B51E65D4A56D431EFF858CEE6" unit="page">
<mods:start id="52AD4AC2D05F60B4A43E18F266932171">338</mods:start>
<mods:end id="0A7AFC2CB9498D53037C28CF402C9319">350</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="29B27AD2E7443CCB45E7132ABE7C1289">journal article</mods:classification>
<mods:identifier id="36C9B591048EBFF7C6FE64F9BF4C6F2D" type="DOI">10.11646/zootaxa.4759.3.2</mods:identifier>
<mods:identifier id="3546A712898AAA9A752B1974F8E5EA50" type="GBIF-Dataset">91fa4568-747a-4854-92d0-3159264955a5</mods:identifier>
<mods:identifier id="2665B6F4A9437D127BC7D072D9DD8A4D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0C347368ADAE043B07ABC8C9B7E99925" type="Zenodo-Dep">3741162</mods:identifier>
<mods:identifier id="F31509DD029DDBFCBED58D65425FD00E" type="ZooBank">435819AF-DFBB-4B32-892F-6A794C57F73B</mods:identifier>
</mods:mods>
<treatment id="4D2B7E2EFFBAED704ED4FD96FAFF32B1" ID-DOI="http://doi.org/10.5281/zenodo.3810489" ID-GBIF-Taxon="163437684" ID-Zenodo-Dep="3810489" LSID="urn:lsid:zoobank.org:act:A2C5FCE7-32F2-4C72-B53A-8E57D09CC35A" httpUri="http://treatment.plazi.org/id/4D2B7E2EFFBAED704ED4FD96FAFF32B1" lastPageId="5" lastPageNumber="343" pageId="1" pageNumber="339">
<subSubSection id="8D989CB3FFBAED744ED4FD96FE2C30C8" box="[151,510,615,642]" pageId="1" pageNumber="339" type="nomenclature">
<paragraph id="C53DCF38FFBAED744ED4FD96FE2C30C8" blockId="1.[151,510,615,678]" box="[151,510,615,642]" pageId="1" pageNumber="339">
<heading id="9E757854FFBAED744ED4FD96FE2C30C8" bold="true" box="[151,510,615,642]" fontSize="11" level="1" pageId="1" pageNumber="339" reason="1">
<emphasis id="F7F6132AFFBAED744ED4FD96FE2C30C8" bold="true" box="[151,510,615,642]" pageId="1" pageNumber="339">
<taxonomicName id="0282B4BBFFBAED744ED4FD96FE4B30C8" authority="Vargovitsh &amp; Kahrarian, 2020" authorityName="Vargovitsh &amp; Kahrarian" authorityYear="2020" box="[151,409,615,642]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="1" pageNumber="339" phylum="Arthropoda" rank="species" species="persicus" status="sp. nov.">
<emphasis id="F7F6132AFFBAED744ED4FD96FE4B30C8" bold="true" box="[151,409,615,642]" italics="true" pageId="1" pageNumber="339">Arrhopalites persicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECC5AE51FFBAED744FE2FD99FE2C30C8" box="[417,510,616,642]" pageId="1" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8D989CB3FFBAED744ED4FD7AFE5C30EC" box="[151,398,651,678]" pageId="1" pageNumber="339" type="description">
<paragraph id="C53DCF38FFBAED744ED4FD7AFE5C30EC" blockId="1.[151,510,615,678]" box="[151,398,651,678]" pageId="1" pageNumber="339">
<figureCitation id="5DB9D3BDFFBAED744ED4FD7AFEDB30EF" box="[151,265,651,677]" captionStart-0="FIGURES 1–12" captionStart-1="FIGURES 13–15" captionStart-2="FIGURES 16–19" captionStart-3="FIGURES 20–25" captionStart-4="FIGURES 26–31" captionStart-5="FIGURES 32–36" captionStartId-0="3.[151,264,1932,1957]" captionStartId-1="4.[151,264,1862,1887]" captionStartId-2="6.[151,264,1958,1983]" captionStartId-3="7.[151,264,1663,1688]" captionStartId-4="8.[151,264,1876,1901]" captionStartId-5="9.[151,264,1946,1971]" captionTargetBox-0="[215,1368,182,1904]" captionTargetBox-1="[151,1436,192,1830]" captionTargetBox-2="[181,1418,200,1922]" captionTargetBox-3="[160,1426,184,1633]" captionTargetBox-4="[162,1425,184,1847]" captionTargetBox-5="[192,1395,929,1917]" captionTargetId-0="figure@3.[211,1376,179,1904]" captionTargetId-1="figure@4.[151,1436,181,1836]" captionTargetId-2="figure@6.[170,1418,181,1933]" captionTargetId-3="figure@7.[157,1430,181,1638]" captionTargetId-4="figure@8.[159,1428,181,1850]" captionTargetId-5="figure@9.[188,1399,927,1921]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." captionText-1="FIGURES 13–15. Arrhopalites persicus sp. nov.: 13, chaetotaxy of foreleg; 14, chaetotaxy of mid leg; 15, chaetotaxy of hind leg." captionText-2="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." captionText-3="FIGURES 20–25. Arrhopalites persicus sp. nov.: 20, type localities on map of Iran: blue mark—Kal Kushk, red mark—Qader Mazd, green mark—Cheshmeh Sorkh; 21, habitus of mounted female; 22, fragment of head dorsum; 23, chaetotaxy of antennal III organ; 24, labial palps; 25, maxillae heads." captionText-4="FIGURES 26–31. Arrhopalites persicus sp. nov.: 26, chaetotaxy of trichobothrial complex; 27, chaetotaxy of central dorsal complex with seta 3 as sensilla; 28–30, foot complex: 28, foreleg; 29, mid leg; 30, hindleg; 31, mucro." captionText-5="FIGURES 32–36. Arrhopalites persicus sp. nov.: 32, dorsal valve of female Abd VI with broadened circumanal setae and 3+3 cuticular spines; 33, appendices anales with enlarged apex; 34–36, setae of great abdomen: 34, sensilla a of mesothorax; 35, dorsal seta m1 of mesothorax; 36, setae of posterior dorsal complex." figureDoi-0="http://doi.org/10.5281/zenodo.3741164" figureDoi-1="http://doi.org/10.5281/zenodo.3741166" figureDoi-2="http://doi.org/10.5281/zenodo.3741168" figureDoi-3="http://doi.org/10.5281/zenodo.3741170" figureDoi-4="http://doi.org/10.5281/zenodo.3741172" figureDoi-5="http://doi.org/10.5281/zenodo.3741174" httpUri-0="https://zenodo.org/record/3741164/files/figure.png" httpUri-1="https://zenodo.org/record/3741166/files/figure.png" httpUri-2="https://zenodo.org/record/3741168/files/figure.png" httpUri-3="https://zenodo.org/record/3741170/files/figure.png" httpUri-4="https://zenodo.org/record/3741172/files/figure.png" httpUri-5="https://zenodo.org/record/3741174/files/figure.png" pageId="1" pageNumber="339">Figs 1–36</figureCitation>
, 
<tableCitation id="8800FA83FFBAED744F57FD7AFE5C30EC" box="[276,398,651,678]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="5.[151,239,298,322]" captionStartId-1="9.[151,239,154,178]" captionStartId-2="10.[151,239,803,827]" captionText-0="TABLE 1. Lengths (in µm) and angles for body parts of type specimens (11 females) of Arrhopalites persicus sp. nov." captionText-1="TABLE 2. Proportions for some body parts of type specimens (11 females) of Arrhopalites persicus sp. nov." captionText-2="TABLE 3. Differences between Arrhopalites persicus sp. nov. and similar congeners from A. diversus group of species" pageId="1" pageNumber="339">Tables 1–3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="8D989CB3FFBAED744ED4FD3CFC943666" pageId="1" pageNumber="339" type="diagnosis">
<paragraph id="C53DCF38FFBAED744ED4FD3CFC943666" blockId="1.[151,1437,717,2041]" pageId="1" pageNumber="339">
<emphasis id="F7F6132AFFBAED744ED4FD3CFEC030A2" bold="true" box="[151,274,717,744]" pageId="1" pageNumber="339">Diagnosis.</emphasis>
Body white or very slightly pigmented, up to 0.8 mm long. Setae of head dorsum slightly spine-like. Antenna 1.2–1.3× of head; Ant IV not subdivided, with 12 whorls of setae. Claw I with tiny inner tooth and short narrow tunica along distal third of outer margin; claw II and III with well-developed inner tooth and broad tunica. Empodium I with tiny corner tooth, apical filament slightly overtopping claws tip; empodium II and III without tooth, shorter (III) or reaching (II) the tip of corresponding claw. Tibiotarsi I–III with 40, 40, 41 setae; seta 
<emphasis id="F7F6132AFFBAED744B09FCACFAAB3132" bold="true" box="[1354,1401,861,888]" pageId="1" pageNumber="339">FSa</emphasis>
on ti-biotarsus III absent. Tenaculum with 1 seta. Manubrium with 5+5 setae. Dens with 3, 2, 1, 1 simple anterior setae; posterior side with 11 setae, 5 of which are strongly spine-like. Mucro with 30–40 teeth on each posterior lamella and with swollen globular tip. Trichobothria 
<emphasis id="F7F6132AFFBAED744CCFFC38FD1531AE" bold="true" box="[652,711,969,996]" pageId="1" pageNumber="339">ABC</emphasis>
form an angle about 90–110 
<superScript id="32F76270FFBAED744A4BFC38FBC2319D" attach="left" box="[1032,1040,969,983]" fontSize="6" pageId="1" pageNumber="339">o</superScript>
and 
<emphasis id="F7F6132AFFBAED744A05FC38FBBF31AE" bold="true" box="[1094,1133,969,996]" pageId="1" pageNumber="339">AB</emphasis>
&gt; 
<emphasis id="F7F6132AFFBAED744A3FFC38FB7131AE" bold="true" box="[1148,1187,969,996]" pageId="1" pageNumber="339">BC</emphasis>
. Seta 
<emphasis id="F7F6132AFFBAED744AABFC3BFB2431AE" bold="true" box="[1256,1270,970,996]" pageId="1" pageNumber="339">3</emphasis>
of central dorsal complex as short bent sensillum. Sixth abdominal segment with 5+5 cuticular spines; circumanal setae long and broadened; appendices anales smooth and acuminate distally.
</paragraph>
</subSubSection>
<subSubSection id="8D989CB3FFBAED744E84FBC7FEEE37F2" pageId="1" pageNumber="339" type="materials_examined">
<paragraph id="C53DCF38FFBAED744E84FBC7FE3E36D2" blockId="1.[151,1437,717,2041]" pageId="1" pageNumber="339">
<emphasis id="F7F6132AFFBAED744E84FBC7FEA5361A" bold="true" box="[199,375,1077,1104]" pageId="1" pageNumber="339">Type material.</emphasis>
<materialsCitation id="75EAC565FFBAED744F3CFBC7FE3E36D2" ID-GBIF-Occurrence="2595809703" collectingDate="2013-12-13" collectorName="M. Kahrarian" country="Iran" county="County" elevation="1516" latitude="33.988667" location="Kal Kushk Village" longLatPrecision="1" longitude="47.189445" municipality="Osmanevand area" pageId="1" pageNumber="339" specimenCount="1" stateProvince="Kermanshah" typeStatus="holotype">
<emphasis id="F7F6132AFFBAED744F3CFBC7FE27361A" bold="true" box="[383,501,1078,1104]" pageId="1" pageNumber="339">
<typeStatus id="1A39719AFFBAED744F3CFBC7FE22361A" box="[383,496,1078,1104]" pageId="1" pageNumber="339" type="holotype">Holotype</typeStatus>
:
</emphasis>
female, soil and leaf litter under oak trees (
<taxonomicName id="0282B4BBFFBAED744DAFFBC6FB04361A" box="[1004,1238,1078,1105]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="339" phylum="Tracheophyta" rank="subSpecies" species="infectoria" subSpecies="veneris">
<emphasis id="F7F6132AFFBAED744DAFFBC6FB04361A" box="[1004,1238,1078,1105]" italics="true" pageId="1" pageNumber="339">Q. infectoria veneris</emphasis>
</taxonomicName>
), 
<location id="C05D99E3FFBAED744AACFBC7FF14363E" LSID="urn:lsid:plazi:treatment:4D2B7E2EFFBAED704ED4FD96FAFF32B1:C05D99E3FFBAED744AACFBC7FF14363E" country="Iran" county="County" latitude="33.988667" longLatPrecision="1" longitude="47.189445" municipality="Osmanevand area" name="Kal Kushk Village" pageId="1" pageNumber="339" stateProvince="Kermanshah">Kal Kushk Village</location>
(
<geoCoordinate id="A0B6A9FFFFBAED744E9BFBABFE51363E" box="[216,387,1113,1140]" degrees="33" direction="north" minutes="59" orientation="latitude" pageId="1" pageNumber="339" precision="1" seconds="19.2" value="33.988667">N 33°59’19.2”</geoCoordinate>
, 
<geoCoordinate id="A0B6A9FFFFBAED744FD1FBAAFDEA363E" box="[402,568,1114,1141]" degrees="47" direction="east" minutes="11" orientation="longitude" pageId="1" pageNumber="339" precision="1" seconds="22.0" value="47.189445">E 47°11’22.0”</geoCoordinate>
, elev. 
<quantity id="027A62DDFFBAED744CC7FBA8FD0D363F" box="[644,735,1113,1141]" metricMagnitude="3" metricUnit="m" metricValue="1.516" pageId="1" pageNumber="339" unit="m" value="1516.0">
<elevation id="4EAF280BFFBAED744CC7FBA8FD0D363F" box="[644,735,1113,1141]" metricMagnitude="3" metricUnit="m" metricValue="1.516" pageId="1" pageNumber="339" unit="m" value="1516.0">1516 m</elevation>
</quantity>
), 
<collectingMunicipality id="25595542FFBAED744CB5FBABFC11363F" box="[758,963,1114,1141]" pageId="1" pageNumber="339">Osmanevand area</collectingMunicipality>
, 
<collectingRegion id="074601DAFFBAED744D92FBABFB8D363E" box="[977,1119,1114,1140]" country="Iran" name="Kermanshah" pageId="1" pageNumber="339">Kermanshah</collectingRegion>
<collectingCounty id="2C5CB7B4FFBAED744A2BFBABFB6E363E" box="[1128,1212,1114,1140]" pageId="1" pageNumber="339">County</collectingCounty>
, 
<collectingRegion id="074601DAFFBAED744A89FBABFA88363E" box="[1226,1370,1114,1140]" country="Iran" name="Kermanshah" pageId="1" pageNumber="339">Kermanshah</collectingRegion>
, 
<collectingCountry id="BD958FA8FFBAED744B2BFBAAFA4A363F" box="[1384,1432,1115,1141]" name="Iran" pageId="1" pageNumber="339">Iran</collectingCountry>
, 
<date id="B13CE9F8FFBAED744ED4FB8FFEC836D2" box="[151,282,1150,1176]" pageId="1" pageNumber="339" value="2013-12-13">
<collectingDate id="A1781010FFBAED744ED4FB8FFEC836D2" box="[151,282,1150,1176]" pageId="1" pageNumber="339" value="2013-12-13">13.xii.2013</collectingDate>
</date>
, 
<collectorName id="6877AAEEFFBAED744F66FB8EFE6E36D2" box="[293,444,1150,1177]" pageId="1" pageNumber="339">M. Kahrarian</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="C53DCF38FFBAED744E84FB53FCF33706" blockId="1.[151,1437,717,2041]" pageId="1" pageNumber="339">
<materialsCitation id="75EAC565FFBAED744E84FB53FD0B36F6" ID-GBIF-Occurrence="2595809704" box="[199,729,1186,1213]" pageId="1" pageNumber="339" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="F7F6132AFFBAED744E84FB53FEED36F6" bold="true" box="[199,319,1186,1212]" pageId="1" pageNumber="339">
<typeStatus id="1A39719AFFBAED744E84FB53FEED36F6" box="[199,319,1186,1212]" pageId="1" pageNumber="339" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="D38404B1FFBAED744F0DFB53FE6136F6" box="[334,435,1186,1212]" pageId="1" pageNumber="339" type="female">1 female</specimenCount>
, same data as in 
<typeStatus id="1A39719AFFBAED744C30FB53FD0B36F6" box="[627,729,1186,1212]" pageId="1" pageNumber="339" type="holotype">holotype</typeStatus>
</materialsCitation>
. 
<materialsCitation id="75EAC565FFBAED744CA7FB53FCC9374E" ID-GBIF-Occurrence="2595809705" collectingDate="2013-12-13" collectorName="M. Kahrarian" country="Iran" county="County" elevation="1682" latitude="34.004333" location="Qader Mazd Village" longLatPrecision="1" longitude="47.20261" municipality="Osmanevand area" pageId="1" pageNumber="339" specimenCount="9" specimenCount-female="8" stateProvince="Kermanshah" typeStatus="paratype">
<specimenCount id="D38404B1FFBAED744CA7FB53FC8636F6" box="[740,852,1186,1212]" pageId="1" pageNumber="339" type="female">8 females</specimenCount>
, 
<specimenCount id="D38404B1FFBAED744D23FB53FC7036F6" box="[864,930,1186,1212]" pageId="1" pageNumber="339" type="generic">1 juv.</specimenCount>
, soil and leaf litter under oak trees (
<taxonomicName id="0282B4BBFFBAED744B79FB52FEFA36AA" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="339" phylum="Tracheophyta" rank="subSpecies" species="infectoria" subSpecies="veneris">
<emphasis id="F7F6132AFFBAED744B79FB52FEFA36AA" italics="true" pageId="1" pageNumber="339">Q. infectoria veneris</emphasis>
</taxonomicName>
), 
<location id="C05D99E3FFBAED744F02FB37FDF536AA" LSID="urn:lsid:plazi:treatment:4D2B7E2EFFBAED704ED4FD96FAFF32B1:C05D99E3FFBAED744F02FB37FDF536AA" box="[321,551,1222,1248]" country="Iran" county="County" latitude="34.004333" longLatPrecision="1" longitude="47.20261" municipality="Osmanevand area" name="Qader Mazd Village" pageId="1" pageNumber="339" stateProvince="Kermanshah">Qader Mazd Village</location>
(
<geoCoordinate id="A0B6A9FFFFBAED744C7BFB37FD0736AA" box="[568,725,1221,1248]" degrees="34" direction="north" minutes="0" orientation="latitude" pageId="1" pageNumber="339" precision="1" seconds="15.6" value="34.004333">N 34°0’15.6”</geoCoordinate>
, 
<geoCoordinate id="A0B6A9FFFFBAED744CA1FB36FCA936AA" box="[738,891,1222,1249]" degrees="47" direction="east" minutes="12" orientation="longitude" pageId="1" pageNumber="339" precision="1" seconds="9.4" value="47.20261">E 47°12’9.4”</geoCoordinate>
, elev. 
<quantity id="027A62DDFFBAED744D86FB37FBCD36AB" box="[965,1055,1222,1249]" metricMagnitude="3" metricUnit="m" metricValue="1.682" pageId="1" pageNumber="339" unit="m" value="1682.0">
<elevation id="4EAF280BFFBAED744D86FB37FBCD36AB" box="[965,1055,1222,1249]" metricMagnitude="3" metricUnit="m" metricValue="1.682" pageId="1" pageNumber="339" unit="m" value="1682.0">1682 m</elevation>
</quantity>
), 
<collectingMunicipality id="25595542FFBAED744A76FB37FAD336AB" box="[1077,1281,1222,1249]" pageId="1" pageNumber="339">Osmanevand area</collectingMunicipality>
, 
<collectingRegion id="074601DAFFBAED744B4CFB37FA4F36AA" box="[1295,1437,1222,1248]" country="Iran" name="Kermanshah" pageId="1" pageNumber="339">Kermanshah</collectingRegion>
<collectingCounty id="2C5CB7B4FFBAED744ED4FB1BFF39374E" box="[151,235,1258,1284]" pageId="1" pageNumber="339">County</collectingCounty>
, 
<collectingRegion id="074601DAFFBAED744EB6FB1BFE57374E" box="[245,389,1258,1284]" country="Iran" name="Kermanshah" pageId="1" pageNumber="339">Kermanshah</collectingRegion>
, 
<collectingCountry id="BD958FA8FFBAED744FCCFB1AFE6D374F" box="[399,447,1259,1285]" name="Iran" pageId="1" pageNumber="339">Iran</collectingCountry>
, 
<date id="B13CE9F8FFBAED744F8AFB1BFD9E374E" box="[457,588,1258,1284]" pageId="1" pageNumber="339" value="2013-12-13">
<collectingDate id="A1781010FFBAED744F8AFB1BFD9E374E" box="[457,588,1258,1284]" pageId="1" pageNumber="339" value="2013-12-13">13.xii.2013</collectingDate>
</date>
, 
<collectorName id="6877AAEEFFBAED744C15FB1AFD3E374E" box="[598,748,1258,1285]" pageId="1" pageNumber="339">M. Kahrarian</collectorName>
leg.
</materialsCitation>
<materialsCitation id="75EAC565FFBAED744D62FB1BFCF33706" ID-GBIF-Occurrence="2595809701" collectingDate="2013-12-20" collectorName="M. Kahrarian" country="Iran" county="County" elevation="1937" latitude="33.98764" location="Cheshmeh Sorkh Village" longLatPrecision="1" longitude="47.27717" municipality="Osmanevand area" pageId="1" pageNumber="339" specimenCount="1" specimenCount-female="1" stateProvince="Kermanshah" typeStatus="paratype">
<specimenCount id="D38404B1FFBAED744D62FB1BFC56374E" box="[801,900,1258,1284]" pageId="1" pageNumber="339" type="female">1 female</specimenCount>
, soil and leaf litter under oak trees (
<taxonomicName id="0282B4BBFFBAED744B4DFB1AFF3A3762" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="339" phylum="Tracheophyta" rank="subSpecies" species="infectoria" subSpecies="veneris">
<emphasis id="F7F6132AFFBAED744B4DFB1AFF3A3762" italics="true" pageId="1" pageNumber="339">Q. infectoria veneris</emphasis>
</taxonomicName>
), 
<location id="C05D99E3FFBAED744F43FAFFFDCB3762" LSID="urn:lsid:plazi:treatment:4D2B7E2EFFBAED704ED4FD96FAFF32B1:C05D99E3FFBAED744F43FAFFFDCB3762" box="[256,537,1294,1320]" country="Iran" county="County" latitude="33.98764" longLatPrecision="1" longitude="47.27717" municipality="Osmanevand area" name="Cheshmeh Sorkh Village" pageId="1" pageNumber="339" stateProvince="Kermanshah">Cheshmeh Sorkh Village</location>
(
<geoCoordinate id="A0B6A9FFFFBAED744C6BFAFFFD003762" box="[552,722,1293,1320]" degrees="33" direction="north" minutes="59" orientation="latitude" pageId="1" pageNumber="339" precision="1" seconds="15.5" value="33.98764">N 33°59’15.5”</geoCoordinate>
, 
<geoCoordinate id="A0B6A9FFFFBAED744C9DFAFEFC513762" box="[734,899,1294,1321]" degrees="47" direction="east" minutes="16" orientation="longitude" pageId="1" pageNumber="339" precision="1" seconds="37.8" value="47.27717">E 47°16’37.8”</geoCoordinate>
, elev. 
<quantity id="027A62DDFFBAED744D89FAFFFBF13763" box="[970,1059,1294,1321]" metricMagnitude="3" metricUnit="m" metricValue="1.9369999999999998" pageId="1" pageNumber="339" unit="m" value="1937.0">
<elevation id="4EAF280BFFBAED744D89FAFFFBF13763" box="[970,1059,1294,1321]" metricMagnitude="3" metricUnit="m" metricValue="1.9369999999999998" pageId="1" pageNumber="339" unit="m" value="1937.0">1937 m</elevation>
</quantity>
), 
<collectingMunicipality id="25595542FFBAED744A7BFAFFFAD03763" box="[1080,1282,1294,1321]" pageId="1" pageNumber="339">Osmanevand area</collectingMunicipality>
, 
<collectingRegion id="074601DAFFBAED744B4CFAFFFA4F3762" box="[1295,1437,1294,1320]" country="Iran" name="Kermanshah" pageId="1" pageNumber="339">Kermanshah</collectingRegion>
<collectingCounty id="2C5CB7B4FFBAED744ED4FAC3FF393706" box="[151,235,1330,1356]" pageId="1" pageNumber="339">County</collectingCounty>
, 
<collectingRegion id="074601DAFFBAED744EB5FAC3FE543706" box="[246,390,1330,1356]" country="Iran" name="Kermanshah" pageId="1" pageNumber="339">Kermanshah</collectingRegion>
, 
<collectingCountry id="BD958FA8FFBAED744FD2FAC2FE133707" box="[401,449,1331,1357]" name="Iran" pageId="1" pageNumber="339">Iran</collectingCountry>
, 
<date id="B13CE9F8FFBAED744F8FFAC3FD9D3706" box="[460,591,1330,1356]" pageId="1" pageNumber="339" value="2013-12-20">
<collectingDate id="A1781010FFBAED744F8FFAC3FD9D3706" box="[460,591,1330,1356]" pageId="1" pageNumber="339" value="2013-12-20">20.xii.2013</collectingDate>
</date>
, 
<collectorName id="6877AAEEFFBAED744C19FAC2FD233706" box="[602,753,1330,1357]" pageId="1" pageNumber="339">M. Kahrarian</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="C53DCF38FFBAED744E84FAA7FEEE37F2" blockId="1.[151,1437,717,2041]" pageId="1" pageNumber="339">
<materialsCitation id="75EAC565FFBAED744E84FAA7FEEE37F2" ID-GBIF-Occurrence="2595809702" country="Ukraine" location="National Academy of Sciences" municipality="Schmalhausen Institute of Zoology" pageId="1" pageNumber="339" specimenCount="11" stateProvince="Kyiv" typeStatus="holotype">
<typeStatus id="1A39719AFFBAED744E84FAA7FEFD373A" box="[199,303,1366,1392]" pageId="1" pageNumber="339" type="holotype">Holotype</typeStatus>
(slide 25-6) and 
<specimenCount id="D38404B1FFBAED744FBAFAA7FD5A373B" box="[505,648,1366,1393]" pageId="1" pageNumber="339" type="generic" typeStatus="paratypes">11 paratypes</specimenCount>
(slides 25-7, 27-1, 27-2, 27-3, 27-4, 27-5, 27-7, 27-9, 27-10, 27-11, 29-5) are deposited in the collection of the 
<collectingMunicipality id="25595542FFBAED744CC6FA8BFBC137DE" box="[645,1043,1402,1428]" pageId="1" pageNumber="339">Schmalhausen Institute of Zoology</collectingMunicipality>
, 
<location id="C05D99E3FFBAED744A63FA8BFAAE37DE" LSID="urn:lsid:plazi:treatment:4D2B7E2EFFBAED704ED4FD96FAFF32B1:C05D99E3FFBAED744A63FA8BFAAE37DE" box="[1056,1404,1402,1428]" country="Ukraine" municipality="Schmalhausen Institute of Zoology" name="National Academy of Sciences" pageId="1" pageNumber="339" stateProvince="Kyiv">National Academy of Sciences</location>
of 
<collectingCountry id="BD958FA8FFBAED744ED4FA6FFF2637F2" box="[151,244,1438,1464]" name="Ukraine" pageId="1" pageNumber="339">Ukraine</collectingCountry>
, 
<collectingRegion id="074601DAFFBAED744EBCFA6FFEE537F2" box="[255,311,1438,1464]" country="Ukraine" name="Kyiv" pageId="1" pageNumber="339">Kiev</collectingRegion>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8D989CB3FFBAED704E84FA30FC71329D" lastPageId="5" lastPageNumber="343" pageId="1" pageNumber="339" type="description">
<paragraph id="C53DCF38FFBAED744E84FA30FE83344A" blockId="1.[151,1437,717,2041]" pageId="1" pageNumber="339">
<emphasis id="F7F6132AFFBAED744E84FA30FE643796" bold="true" box="[199,438,1473,1500]" pageId="1" pageNumber="339">Description. Female</emphasis>
: Body length up to about 0.8 mm, habitus as in 
<figureCitation id="5DB9D3BDFFBAED744D88FA33FBCE3796" box="[971,1052,1474,1500]" captionStart="FIGURES 20–25" captionStartId="7.[151,264,1663,1688]" captionTargetBox="[160,1426,184,1633]" captionTargetId="figure@7.[157,1430,181,1638]" captionTargetPageId="7" captionText="FIGURES 20–25. Arrhopalites persicus sp. nov.: 20, type localities on map of Iran: blue mark—Kal Kushk, red mark—Qader Mazd, green mark—Cheshmeh Sorkh; 21, habitus of mounted female; 22, fragment of head dorsum; 23, chaetotaxy of antennal III organ; 24, labial palps; 25, maxillae heads." figureDoi="http://doi.org/10.5281/zenodo.3741170" httpUri="https://zenodo.org/record/3741170/files/figure.png" pageId="1" pageNumber="339">Fig. 21</figureCitation>
, white or with slight diffuse traces of pigmentation.
</paragraph>
<paragraph id="C53DCF38FFBAED744E84F9FBFC1934FF" blockId="1.[151,1437,717,2041]" pageId="1" pageNumber="339">
<emphasis id="F7F6132AFFBAED744E84F9FBFED5346E" bold="true" box="[199,263,1546,1572]" pageId="1" pageNumber="339">Head</emphasis>
(
<figureCitation id="5DB9D3BDFFBAED744F56F9FBFE89346E" box="[277,347,1546,1572]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="1" pageNumber="339">Figs 1</figureCitation>
, 
<figureCitation id="5DB9D3BDFFBAED744F2BF9FBFE5A346E" box="[360,392,1546,1572]" captionStart="FIGURES 20–25" captionStartId="7.[151,264,1663,1688]" captionTargetBox="[160,1426,184,1633]" captionTargetId="figure@7.[157,1430,181,1638]" captionTargetPageId="7" captionText="FIGURES 20–25. Arrhopalites persicus sp. nov.: 20, type localities on map of Iran: blue mark—Kal Kushk, red mark—Qader Mazd, green mark—Cheshmeh Sorkh; 21, habitus of mounted female; 22, fragment of head dorsum; 23, chaetotaxy of antennal III organ; 24, labial palps; 25, maxillae heads." figureDoi="http://doi.org/10.5281/zenodo.3741170" httpUri="https://zenodo.org/record/3741170/files/figure.png" pageId="1" pageNumber="339">22</figureCitation>
). Eyes 1+1, about 4 μm in diameter, unpigmented, sometimes hardly noticeable. Clypeal area: row 
<emphasis id="F7F6132AFFBAED744E88F9DFFF0B3402" bold="true" box="[203,217,1582,1608]" pageId="1" pageNumber="339">a</emphasis>
: 4 + axial + 4 setae, rows 
<emphasis id="F7F6132AFFBAED744C4BF9DFFDCA3402" bold="true" box="[520,536,1582,1608]" pageId="1" pageNumber="339">b</emphasis>
, 
<emphasis id="F7F6132AFFBAED744C64F9DFFDE13402" bold="true" box="[551,563,1582,1608]" pageId="1" pageNumber="339">c</emphasis>
, 
<emphasis id="F7F6132AFFBAED744C01F9DFFD803402" bold="true" box="[578,594,1582,1608]" pageId="1" pageNumber="339">d</emphasis>
and 
<emphasis id="F7F6132AFFBAED744CC9F9DFFD443402" bold="true" box="[650,662,1582,1608]" pageId="1" pageNumber="339">e</emphasis>
with 5+5 setae each, row 
<emphasis id="F7F6132AFFBAED744D81F9DCFC193402" bold="true" box="[962,971,1581,1608]" pageId="1" pageNumber="339">f</emphasis>
with 6+6 setae, median region 
<emphasis id="F7F6132AFFBAED744B71F9DFFA9E3402" bold="true" box="[1330,1356,1582,1608]" pageId="1" pageNumber="339">M</emphasis>
with 3 setae (marked with oval in 
<figureCitation id="5DB9D3BDFFBAED744F8BF9A3FDDA3426" box="[456,520,1618,1644]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="1" pageNumber="339">Fig. 1</figureCitation>
). Interantennal area: row 
<emphasis id="F7F6132AFFBAED744D68F9A3FCE93426" bold="true" box="[811,827,1618,1644]" pageId="1" pageNumber="339">α</emphasis>
: 2+2, row 
<emphasis id="F7F6132AFFBAED744DF5F9A3FC173426" bold="true" box="[950,965,1618,1644]" pageId="1" pageNumber="339">β</emphasis>
: 1 + axial + 1. Dorsal area (rows 
<emphasis id="F7F6132AFFBAED744B7CF9A0FA813426" bold="true" box="[1343,1363,1617,1644]" pageId="1" pageNumber="339">A</emphasis>
to 
<emphasis id="F7F6132AFFBAED744B34F9A3FA593426" bold="true" box="[1399,1419,1618,1644]" pageId="1" pageNumber="339">D</emphasis>
): axial seta present in rows 
<emphasis id="F7F6132AFFBAED744FFFF984FE0234DA" bold="true" box="[444,464,1653,1680]" pageId="1" pageNumber="339">A</emphasis>
, 
<emphasis id="F7F6132AFFBAED744F9CF987FE2034DA" bold="true" box="[479,498,1654,1680]" pageId="1" pageNumber="339">B</emphasis>
and 
<emphasis id="F7F6132AFFBAED744C69F984FDEC34DA" bold="true" box="[554,574,1653,1680]" pageId="1" pageNumber="339">C</emphasis>
; 6(5)+1+6(5) dorsal setae (13 or 
<quantity id="027A62DDFFBAED744DF4F987FC2234DA" box="[951,1008,1654,1680]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="1" pageNumber="339" unit="in" value="11.0">11 in</quantity>
total) usually slightly thickened (
<figureCitation id="5DB9D3BDFFBAED744B29F987FF1134FE" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="1" pageNumber="339">Figs 1, 5</figureCitation>
, 
<figureCitation id="5DB9D3BDFFBAED744E8CF96BFF3D34FE" box="[207,239,1690,1716]" captionStart="FIGURES 20–25" captionStartId="7.[151,264,1663,1688]" captionTargetBox="[160,1426,184,1633]" captionTargetId="figure@7.[157,1430,181,1638]" captionTargetPageId="7" captionText="FIGURES 20–25. Arrhopalites persicus sp. nov.: 20, type localities on map of Iran: blue mark—Kal Kushk, red mark—Qader Mazd, green mark—Cheshmeh Sorkh; 21, habitus of mounted female; 22, fragment of head dorsum; 23, chaetotaxy of antennal III organ; 24, labial palps; 25, maxillae heads." figureDoi="http://doi.org/10.5281/zenodo.3741170" httpUri="https://zenodo.org/record/3741170/files/figure.png" pageId="1" pageNumber="339">22</figureCitation>
). Chaetotaxy of the head posterior side as in 
<figureCitation id="5DB9D3BDFFBAED744CA5F96BFCFA34FE" box="[742,808,1690,1716]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="1" pageNumber="339">Fig. 1</figureCitation>
(dotted lines).
</paragraph>
<paragraph id="C53DCF38FFBAED744E84F94CFB433492" blockId="1.[151,1437,717,2041]" box="[199,1169,1725,1753]" pageId="1" pageNumber="339">
<emphasis id="F7F6132AFFBAED744E84F94CFDC43492" bold="true" box="[199,534,1725,1752]" pageId="1" pageNumber="339">Chaetotaxy of mouth region</emphasis>
. Labrum (
<figureCitation id="5DB9D3BDFFBAED744CCFF94FFD1C3492" box="[652,718,1726,1752]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="1" pageNumber="339">Fig. 1</figureCitation>
): labral / prelabral chaetotaxy: 4 5 5 / 6.
</paragraph>
<paragraph id="C53DCF38FFBAED744E84F913FD6135C7" blockId="1.[151,1437,717,2041]" pageId="1" pageNumber="339">
Labium: submentum with 4 setae, mentum with 5 setae; labial palp (
<figureCitation id="5DB9D3BDFFBAED744DFBF913FC2C34B6" box="[952,1022,1762,1788]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="1" pageNumber="339">Figs 3</figureCitation>
, 
<figureCitation id="5DB9D3BDFFBAED744A49F913FBF534B6" box="[1034,1063,1762,1788]" captionStart="FIGURES 20–25" captionStartId="7.[151,264,1663,1688]" captionTargetBox="[160,1426,184,1633]" captionTargetId="figure@7.[157,1430,181,1638]" captionTargetPageId="7" captionText="FIGURES 20–25. Arrhopalites persicus sp. nov.: 20, type localities on map of Iran: blue mark—Kal Kushk, red mark—Qader Mazd, green mark—Cheshmeh Sorkh; 21, habitus of mounted female; 22, fragment of head dorsum; 23, chaetotaxy of antennal III organ; 24, labial palps; 25, maxillae heads." figureDoi="http://doi.org/10.5281/zenodo.3741170" httpUri="https://zenodo.org/record/3741170/files/figure.png" pageId="1" pageNumber="339">24</figureCitation>
) with 4 proximal setae, 5 papillae (A, B, C, D, E) with deeply inserted setae and with 12 associated guard setae (a1, b1-4, d1-3, e1-4 of which b1 and b2 are blunt and thickened) and 3 hypostomal setae (H, h1, h2) one of which is distinctly curved. Maxilla as in 
<figureCitation id="5DB9D3BDFFBAED744B33F8DBFF643522" captionStart="FIGURES 20–25" captionStartId="7.[151,264,1663,1688]" captionTargetBox="[160,1426,184,1633]" captionTargetId="figure@7.[157,1430,181,1638]" captionTargetPageId="7" captionText="FIGURES 20–25. Arrhopalites persicus sp. nov.: 20, type localities on map of Iran: blue mark—Kal Kushk, red mark—Qader Mazd, green mark—Cheshmeh Sorkh; 21, habitus of mounted female; 22, fragment of head dorsum; 23, chaetotaxy of antennal III organ; 24, labial palps; 25, maxillae heads." figureDoi="http://doi.org/10.5281/zenodo.3741170" httpUri="https://zenodo.org/record/3741170/files/figure.png" pageId="1" pageNumber="339">Fig. 25</figureCitation>
. Apical seta of the maxillary outer lobe (
<figureCitation id="5DB9D3BDFFBAED744C3FF8BFFD123522" box="[636,704,1870,1896]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="1" pageNumber="339">Fig. 2</figureCitation>
) with short and thin subparallel branch at the base, sublobal plate with three sublobal hairs. Oral fold with 2 setae.
</paragraph>
<paragraph id="C53DCF38FFBAED774E84F864FCD932B1" blockId="1.[151,1437,717,2041]" lastBlockId="2.[151,1437,152,2015]" lastPageId="2" lastPageNumber="340" pageId="1" pageNumber="339">
<emphasis id="F7F6132AFFBAED744E84F864FEFC35FA" bold="true" box="[199,302,1941,1968]" pageId="1" pageNumber="339">Antenna</emphasis>
(
<figureCitation id="5DB9D3BDFFBAED744F7FF867FE5635FA" box="[316,388,1942,1968]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="1" pageNumber="339">Figs 4</figureCitation>
, 
<figureCitation id="5DB9D3BDFFBAED744FCCF867FE7E35FA" box="[399,428,1942,1968]" captionStart="FIGURES 20–25" captionStartId="7.[151,264,1663,1688]" captionTargetBox="[160,1426,184,1633]" captionTargetId="figure@7.[157,1430,181,1638]" captionTargetPageId="7" captionText="FIGURES 20–25. Arrhopalites persicus sp. nov.: 20, type localities on map of Iran: blue mark—Kal Kushk, red mark—Qader Mazd, green mark—Cheshmeh Sorkh; 21, habitus of mounted female; 22, fragment of head dorsum; 23, chaetotaxy of antennal III organ; 24, labial palps; 25, maxillae heads." figureDoi="http://doi.org/10.5281/zenodo.3741170" httpUri="https://zenodo.org/record/3741170/files/figure.png" pageId="1" pageNumber="339">23</figureCitation>
) short, about 1.3× of head. Mean length ratio of antennal segments I: II: III: IV = 1: 2.1: 2.8: 5.3 (1: 2.4: 3.1: 5.6 in 
<typeStatus id="1A39719AFFBAED744F97F84BFDE9359F" box="[468,571,1978,2005]" pageId="1" pageNumber="339" type="holotype">holotype</typeStatus>
). Ant I with 7 setae, 
<emphasis id="F7F6132AFFBAED744D58F84BFCF9359E" bold="true" box="[795,811,1978,2004]" pageId="1" pageNumber="339">p</emphasis>
as microseta (3.6 μm). Ant II with 14 setae one of which is more than 2× longer than others. Ant III subbasally moderately swollen, with following chaetotaxy: 17 simple setae of which 
<emphasis id="F7F6132AFFB9ED774F01FF69FEBC32F9" bold="true" box="[322,366,152,179]" pageId="2" pageNumber="340">Api</emphasis>
and 
<emphasis id="F7F6132AFFB9ED774FE5FF69FE0432F9" bold="true" box="[422,470,152,179]" pageId="2" pageNumber="340">Ape</emphasis>
especially thin, 2 sense rods in shallow pits (4.4 μm in 
<typeStatus id="1A39719AFFB9ED774A0FFF68FB6232FE" box="[1100,1200,153,180]" pageId="2" pageNumber="340" type="holotype">holotype</typeStatus>
) and 
<emphasis id="F7F6132AFFB9ED774AB3FF69FAC832F9" bold="true" box="[1264,1306,152,179]" pageId="2" pageNumber="340">Aai</emphasis>
as small (3 μm in 
<typeStatus id="1A39719AFFB9ED774EA3FF4CFE963292" box="[224,324,189,216]" pageId="2" pageNumber="340" type="holotype">holotype</typeStatus>
) bent and blunt sensillum (
<figureCitation id="5DB9D3BDFFB9ED774C35FF4CFD18329D" box="[630,714,189,215]" captionStart="FIGURES 20–25" captionStartId="7.[151,264,1663,1688]" captionTargetBox="[160,1426,184,1633]" captionTargetId="figure@7.[157,1430,181,1638]" captionTargetPageId="7" captionText="FIGURES 20–25. Arrhopalites persicus sp. nov.: 20, type localities on map of Iran: blue mark—Kal Kushk, red mark—Qader Mazd, green mark—Cheshmeh Sorkh; 21, habitus of mounted female; 22, fragment of head dorsum; 23, chaetotaxy of antennal III organ; 24, labial palps; 25, maxillae heads." figureDoi="http://doi.org/10.5281/zenodo.3741170" httpUri="https://zenodo.org/record/3741170/files/figure.png" pageId="2" pageNumber="340">Fig. 23</figureCitation>
). Ant IV short and undivided, bears 12 densely settled whorls of setae without clear empty spaces between them (
<figureCitation id="5DB9D3BDFFB9ED774CFAFF10FD2C32B1" box="[697,766,225,251]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="2" pageNumber="340">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="C53DCF38FFB9ED774E84FEF4FB8533BD" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="340">
<emphasis id="F7F6132AFFB9ED774E84FEF4FF2D3355" bold="true" box="[199,255,261,287]" pageId="2" pageNumber="340">Legs</emphasis>
. Foreleg (
<figureCitation id="5DB9D3BDFFB9ED774F36FEF4FE1C3355" box="[373,462,261,287]" captionStart="FIGURES 13–15" captionStartId="4.[151,264,1862,1887]" captionTargetBox="[151,1436,192,1830]" captionTargetId="figure@4.[151,1436,181,1836]" captionTargetPageId="4" captionText="FIGURES 13–15. Arrhopalites persicus sp. nov.: 13, chaetotaxy of foreleg; 14, chaetotaxy of mid leg; 15, chaetotaxy of hind leg." figureDoi="http://doi.org/10.5281/zenodo.3741166" httpUri="https://zenodo.org/record/3741166/files/figure.png" pageId="2" pageNumber="340">Figs 13</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774F9FFEF4FE283355" box="[476,506,261,287]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">16</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774C4BFEF4FDFA3355" box="[520,552,261,287]" captionStart="FIGURES 26–31" captionStartId="8.[151,264,1876,1901]" captionTargetBox="[162,1425,184,1847]" captionTargetId="figure@8.[159,1428,181,1850]" captionTargetPageId="8" captionText="FIGURES 26–31. Arrhopalites persicus sp. nov.: 26, chaetotaxy of trichobothrial complex; 27, chaetotaxy of central dorsal complex with seta 3 as sensilla; 28–30, foot complex: 28, foreleg; 29, mid leg; 30, hindleg; 31, mucro." figureDoi="http://doi.org/10.5281/zenodo.3741172" httpUri="https://zenodo.org/record/3741172/files/figure.png" pageId="2" pageNumber="340">28</figureCitation>
): precoxae 1, 2 and coxa with 1, 0, 1 setae respectively (
<figureCitation id="5DB9D3BDFFB9ED774AF4FEF4FADF336A" box="[1207,1293,261,288]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">Fig. 16</figureCitation>
). Trochanter with 3 anterior and 1 posterior setae. Femur with 12 setae, 
<emphasis id="F7F6132AFFB9ED774D07FED8FCB23309" bold="true" box="[836,864,297,323]" pageId="2" pageNumber="340">a4</emphasis>
turned perpendicularly to the longitudinal axis of the segment, 
<emphasis id="F7F6132AFFB9ED774F6DFEBCFE99332D" bold="true" box="[302,331,333,359]" pageId="2" pageNumber="340">p3</emphasis>
as microseta. Tibiotarsus with 40 setae: whorls I–II with 7, whorls III–IV with 8, whorl V with 7 (sometimes 6) setae and region 
<emphasis id="F7F6132AFFB9ED774FB8FE80FDD933C1" bold="true" box="[507,523,369,395]" pageId="2" pageNumber="340">F</emphasis>
with 3 primary setae (
<emphasis id="F7F6132AFFB9ED774D4BFE80FCC633C1" bold="true" box="[776,788,369,395]" pageId="2" pageNumber="340">e</emphasis>
, 
<emphasis id="F7F6132AFFB9ED774D60FE80FCEF33C1" bold="true" box="[803,829,369,395]" pageId="2" pageNumber="340">ae</emphasis>
, 
<emphasis id="F7F6132AFFB9ED774D08FE80FCB533C1" bold="true" box="[843,871,369,395]" pageId="2" pageNumber="340">pe</emphasis>
). Pretarsus with 1 anterior and 1 posterior setulae. Foot complex (
<figureCitation id="5DB9D3BDFFB9ED774F02FE64FE4733E5" box="[321,405,405,431]" captionStart="FIGURES 26–31" captionStartId="8.[151,264,1876,1901]" captionTargetBox="[162,1425,184,1847]" captionTargetId="figure@8.[159,1428,181,1850]" captionTargetPageId="8" captionText="FIGURES 26–31. Arrhopalites persicus sp. nov.: 26, chaetotaxy of trichobothrial complex; 27, chaetotaxy of central dorsal complex with seta 3 as sensilla; 28–30, foot complex: 28, foreleg; 29, mid leg; 30, hindleg; 31, mucro." figureDoi="http://doi.org/10.5281/zenodo.3741172" httpUri="https://zenodo.org/record/3741172/files/figure.png" pageId="2" pageNumber="340">Fig. 28</figureCitation>
). Claw: with short narrow tunica along distal third of outer margin (sometimes unnoticable); with tiny inner tooth, about 5× shorter than tibiotarsus. Empodium: thin, with hardly noticeable tiny corner tooth or without, 1.1–1.2× shorter than claw; tip of empodial filament overtopping tip of claw.
</paragraph>
<paragraph id="C53DCF38FFB9ED774E84FDF0FD0C30B9" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="340">
Mid leg (
<figureCitation id="5DB9D3BDFFB9ED774F6CFDF0FE543051" box="[303,390,513,539]" captionStart="FIGURES 13–15" captionStartId="4.[151,264,1862,1887]" captionTargetBox="[151,1436,192,1830]" captionTargetId="figure@4.[151,1436,181,1836]" captionTargetPageId="4" captionText="FIGURES 13–15. Arrhopalites persicus sp. nov.: 13, chaetotaxy of foreleg; 14, chaetotaxy of mid leg; 15, chaetotaxy of hind leg." figureDoi="http://doi.org/10.5281/zenodo.3741166" httpUri="https://zenodo.org/record/3741166/files/figure.png" pageId="2" pageNumber="340">Figs 14</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774FD1FDF0FE623051" box="[402,432,513,539]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">16</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774FFEFDF0FE0F3051" box="[445,477,513,539]" captionStart="FIGURES 26–31" captionStartId="8.[151,264,1876,1901]" captionTargetBox="[162,1425,184,1847]" captionTargetId="figure@8.[159,1428,181,1850]" captionTargetPageId="8" captionText="FIGURES 26–31. Arrhopalites persicus sp. nov.: 26, chaetotaxy of trichobothrial complex; 27, chaetotaxy of central dorsal complex with seta 3 as sensilla; 28–30, foot complex: 28, foreleg; 29, mid leg; 30, hindleg; 31, mucro." figureDoi="http://doi.org/10.5281/zenodo.3741172" httpUri="https://zenodo.org/record/3741172/files/figure.png" pageId="2" pageNumber="340">29</figureCitation>
): precoxae 1 and 2 with 1, 1 setae respectively, precoxal process present, coxa with 3 setae and microsensillum (
<figureCitation id="5DB9D3BDFFB9ED774F85FDD4FDC9300A" box="[454,539,549,576]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">Fig. 16</figureCitation>
). Trochanter with 4 setae including 2 simple anterior and 1 posterior setae and anterior trochanteral organ. Femur with 13 setae, 1 posterior as microseta. Tibiotarsus with 40 setae: distal whorl I with 7, whorls II–IV with 8, whorl V with 6 setae, basal region 
<emphasis id="F7F6132AFFB9ED774D2FFD9CFCAE30CD" bold="true" box="[876,892,621,647]" pageId="2" pageNumber="340">F</emphasis>
with 3 
<emphasis id="F7F6132AFFB9ED774D97FD9CFC2730CD" bold="true" box="[980,1013,621,647]" pageId="2" pageNumber="340">FP</emphasis>
setae. Foot complex (
<figureCitation id="5DB9D3BDFFB9ED774AB3FD9CFA9730CD" box="[1264,1349,621,647]" captionStart="FIGURES 26–31" captionStartId="8.[151,264,1876,1901]" captionTargetBox="[162,1425,184,1847]" captionTargetId="figure@8.[159,1428,181,1850]" captionTargetPageId="8" captionText="FIGURES 26–31. Arrhopalites persicus sp. nov.: 26, chaetotaxy of trichobothrial complex; 27, chaetotaxy of central dorsal complex with seta 3 as sensilla; 28–30, foot complex: 28, foreleg; 29, mid leg; 30, hindleg; 31, mucro." figureDoi="http://doi.org/10.5281/zenodo.3741172" httpUri="https://zenodo.org/record/3741172/files/figure.png" pageId="2" pageNumber="340">Fig. 29</figureCitation>
). Claw: with well-developed tunica, distinct inner tooth, 4.7–5.5× shorter than tibiotarsus (5.1 in 
<typeStatus id="1A39719AFFB9ED774ADFFD60FAD130E6" box="[1180,1283,657,684]" pageId="2" pageNumber="340" type="holotype">holotype</typeStatus>
). Empodium: broader than in foreleg, without corner tooth, 1.3–1.7× (1.5 in 
<typeStatus id="1A39719AFFB9ED774D0DFD44FC60309A" box="[846,946,693,720]" pageId="2" pageNumber="340" type="holotype">holotype</typeStatus>
) shorter than claw; tip of empodial filament visually not reaching or almost reaching tip of claw.
</paragraph>
<paragraph id="C53DCF38FFB9ED774E84FD0CFAA03181" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="340">
Hind leg (
<figureCitation id="5DB9D3BDFFB9ED774F77FD0CFE58315D" box="[308,394,764,791]" captionStart="FIGURES 13–15" captionStartId="4.[151,264,1862,1887]" captionTargetBox="[151,1436,192,1830]" captionTargetId="figure@4.[151,1436,181,1836]" captionTargetPageId="4" captionText="FIGURES 13–15. Arrhopalites persicus sp. nov.: 13, chaetotaxy of foreleg; 14, chaetotaxy of mid leg; 15, chaetotaxy of hind leg." figureDoi="http://doi.org/10.5281/zenodo.3741166" httpUri="https://zenodo.org/record/3741166/files/figure.png" pageId="2" pageNumber="340">Figs 15</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774FD6FD0CFE61315D" box="[405,435,765,791]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">16</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774FFEFD0CFE0F315D" box="[445,477,765,791]" captionStart="FIGURES 26–31" captionStartId="8.[151,264,1876,1901]" captionTargetBox="[162,1425,184,1847]" captionTargetId="figure@8.[159,1428,181,1850]" captionTargetPageId="8" captionText="FIGURES 26–31. Arrhopalites persicus sp. nov.: 26, chaetotaxy of trichobothrial complex; 27, chaetotaxy of central dorsal complex with seta 3 as sensilla; 28–30, foot complex: 28, foreleg; 29, mid leg; 30, hindleg; 31, mucro." figureDoi="http://doi.org/10.5281/zenodo.3741172" httpUri="https://zenodo.org/record/3741172/files/figure.png" pageId="2" pageNumber="340">30</figureCitation>
): precoxae 1 and 2 with 1, 1 setae respectively, process on precoxa 1 present, coxa with 3 setae and microsensillum (
<figureCitation id="5DB9D3BDFFB9ED774F97FCD0FDF53176" box="[468,551,801,828]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">Fig. 16</figureCitation>
). Trochanter with 4 setae: anterior trochanteral organ, 2 anterior and 1 posterior simple setae. Femur with 12 setae, 2 posterior ones as microsetae. Tibiotarsus with 41 setae: whorl I–IV with 8, whorl V with 6 setae, region F with 3 primary setae 
<emphasis id="F7F6132AFFB9ED774C9CFC98FCD231C9" bold="true" box="[735,768,873,899]" pageId="2" pageNumber="340">FP</emphasis>
present and secondary seta 
<emphasis id="F7F6132AFFB9ED774A7AFC99FBBA31C9" bold="true" box="[1081,1128,872,899]" pageId="2" pageNumber="340">FSa</emphasis>
absent. Foot complex (
<figureCitation id="5DB9D3BDFFB9ED774B2CFC98FF6531ED" captionStart="FIGURES 26–31" captionStartId="8.[151,264,1876,1901]" captionTargetBox="[162,1425,184,1847]" captionTargetId="figure@8.[159,1428,181,1850]" captionTargetPageId="8" captionText="FIGURES 26–31. Arrhopalites persicus sp. nov.: 26, chaetotaxy of trichobothrial complex; 27, chaetotaxy of central dorsal complex with seta 3 as sensilla; 28–30, foot complex: 28, foreleg; 29, mid leg; 30, hindleg; 31, mucro." figureDoi="http://doi.org/10.5281/zenodo.3741172" httpUri="https://zenodo.org/record/3741172/files/figure.png" pageId="2" pageNumber="340">Fig. 30</figureCitation>
). Claw: with broad tunica, with distinct inner tooth, 5.4–6.3× shorter than tibiotarsus (6.3 in 
<typeStatus id="1A39719AFFB9ED774A96FC7CFAEE31E2" box="[1237,1340,909,936]" pageId="2" pageNumber="340" type="holotype">holotype</typeStatus>
). Empodium: broad, corner tooth absent, 1.5–2.1× shorter than claw; tip of empodial filament not reaching tip of claw.
</paragraph>
<paragraph id="C53DCF38FFB9ED774E84FC24FA4E31A5" blockId="2.[151,1437,152,2015]" box="[199,1436,981,1008]" pageId="2" pageNumber="340">
Mean lengths ratio of tibiotarsi I: II: III = 1: 1: 1.3 (same as in 
<typeStatus id="1A39719AFFB9ED774D3AFC24FC0931BA" box="[889,987,981,1008]" pageId="2" pageNumber="340" type="holotype">holotype</typeStatus>
). Tibiotarsus I about 2× shorter than head.
</paragraph>
<paragraph id="C53DCF38FFB9ED774E84FC08FCF83659" blockId="2.[151,1437,152,2015]" box="[199,810,1017,1044]" pageId="2" pageNumber="340">
<emphasis id="F7F6132AFFB9ED774E84FC08FE8E3659" bold="true" box="[199,348,1017,1043]" pageId="2" pageNumber="340">Ventral tube</emphasis>
(
<figureCitation id="5DB9D3BDFFB9ED774F2FFC08FE6F365E" box="[364,445,1017,1044]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">Fig. 16</figureCitation>
) with 1+1 subapical microsetae.
</paragraph>
<paragraph id="C53DCF38FFB9ED774E84FBECFAC9367D" blockId="2.[151,1437,152,2015]" box="[199,1307,1053,1079]" pageId="2" pageNumber="340">
<emphasis id="F7F6132AFFB9ED774E84FBECFE9E367D" bold="true" box="[199,332,1053,1079]" pageId="2" pageNumber="340">Tenaculum</emphasis>
(
<figureCitation id="5DB9D3BDFFB9ED774F18FBECFE7D367D" box="[347,431,1053,1079]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">Fig. 18</figureCitation>
): each ramus with 3 teeth and a basal process; anterior lobe with 1 apical seta.
</paragraph>
<paragraph id="C53DCF38FFB9ED774E84FBB0FD183682" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="340">
<emphasis id="F7F6132AFFB9ED774E84FBB0FEDC3611" bold="true" box="[199,270,1089,1115]" pageId="2" pageNumber="340">Furca</emphasis>
(
<figureCitation id="5DB9D3BDFFB9ED774F5FFBB0FEA03611" box="[284,370,1089,1115]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">Figs 17</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774F3EFBB0FE4E3611" box="[381,412,1089,1115]" captionStart="FIGURES 26–31" captionStartId="8.[151,264,1876,1901]" captionTargetBox="[162,1425,184,1847]" captionTargetId="figure@8.[159,1428,181,1850]" captionTargetPageId="8" captionText="FIGURES 26–31. Arrhopalites persicus sp. nov.: 26, chaetotaxy of trichobothrial complex; 27, chaetotaxy of central dorsal complex with seta 3 as sensilla; 28–30, foot complex: 28, foreleg; 29, mid leg; 30, hindleg; 31, mucro." figureDoi="http://doi.org/10.5281/zenodo.3741172" httpUri="https://zenodo.org/record/3741172/files/figure.png" pageId="2" pageNumber="340">31</figureCitation>
). Manubrium with 5+5 posterior setae. Dens with 18 setae (7 anterior and 11 posterior). 3, 2, 1, 1 thin setae on anterior side, 
<emphasis id="F7F6132AFFB9ED774FB6FB94FDDC3635" bold="true" box="[501,526,1125,1151]" pageId="2" pageNumber="340">Ia</emphasis>
– not modified (
<figureCitation id="5DB9D3BDFFB9ED774C8BFB94FCCE3635" box="[712,796,1125,1151]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="2" pageNumber="340">Fig. 10</figureCitation>
). Posterior side with 5 distinctly spine-like setae: 
<emphasis id="F7F6132AFFB9ED774B0BFB94FA8D3635" bold="true" box="[1352,1375,1125,1151]" pageId="2" pageNumber="340">Ie</emphasis>
(
<figureCitation id="5DB9D3BDFFB9ED774B2CFB94FF6636E9" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="2" pageNumber="340">Fig. 11</figureCitation>
), 
<emphasis id="F7F6132AFFB9ED774E8BFB79FF0936E9" bold="true" box="[200,219,1160,1187]" pageId="2" pageNumber="340">Ii</emphasis>
and 
<emphasis id="F7F6132AFFB9ED774F4CFB78FE5336E9" bold="true" box="[271,385,1161,1187]" pageId="2" pageNumber="340">Ipe–IIIpe</emphasis>
. Mucro densely serrated with 30–40 teeth on each lateral lamella, with globular apex (
<figureCitation id="5DB9D3BDFFB9ED774B7EFB78FA5D36E9" box="[1341,1423,1161,1187]" captionStart="FIGURES 26–31" captionStartId="8.[151,264,1876,1901]" captionTargetBox="[162,1425,184,1847]" captionTargetId="figure@8.[159,1428,181,1850]" captionTargetPageId="8" captionText="FIGURES 26–31. Arrhopalites persicus sp. nov.: 26, chaetotaxy of trichobothrial complex; 27, chaetotaxy of central dorsal complex with seta 3 as sensilla; 28–30, foot complex: 28, foreleg; 29, mid leg; 30, hindleg; 31, mucro." figureDoi="http://doi.org/10.5281/zenodo.3741172" httpUri="https://zenodo.org/record/3741172/files/figure.png" pageId="2" pageNumber="340">Fig. 31</figureCitation>
). Dens 1.3–1.7× as long as mucro (1.5 in 
<typeStatus id="1A39719AFFB9ED774C14FB5CFD6C3682" box="[599,702,1197,1224]" pageId="2" pageNumber="340" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="C53DCF38FFB9ED774E84FB21FF3E3419" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="340">
<emphasis id="F7F6132AFFB9ED774E84FB21FE5636A1" bold="true" box="[199,388,1232,1259]" pageId="2" pageNumber="340">Great abdomen</emphasis>
(
<figureCitation id="5DB9D3BDFFB9ED774FD5FB20FE3D36A1" box="[406,495,1233,1259]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">Figs 16</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774FBEFB20FD9B36A1" box="[509,585,1233,1259]" captionStart="FIGURES 26–31" captionStartId="8.[151,264,1876,1901]" captionTargetBox="[162,1425,184,1847]" captionTargetId="figure@8.[159,1428,181,1850]" captionTargetPageId="8" captionText="FIGURES 26–31. Arrhopalites persicus sp. nov.: 26, chaetotaxy of trichobothrial complex; 27, chaetotaxy of central dorsal complex with seta 3 as sensilla; 28–30, foot complex: 28, foreleg; 29, mid leg; 30, hindleg; 31, mucro." figureDoi="http://doi.org/10.5281/zenodo.3741172" httpUri="https://zenodo.org/record/3741172/files/figure.png" pageId="2" pageNumber="340">26, 27</figureCitation>
). Segments Th II and III bearing single short and bent sensillum in row 
<emphasis id="F7F6132AFFB9ED774BCDFB20FA4E36A1" bold="true" box="[1422,1436,1233,1259]" pageId="2" pageNumber="340">a</emphasis>
(
<figureCitation id="5DB9D3BDFFB9ED774EE3FB04FF233745" box="[160,241,1269,1295]" captionStart="FIGURES 32–36" captionStartId="9.[151,264,1946,1971]" captionTargetBox="[192,1395,929,1917]" captionTargetId="figure@9.[188,1399,927,1921]" captionTargetPageId="9" captionText="FIGURES 32–36. Arrhopalites persicus sp. nov.: 32, dorsal valve of female Abd VI with broadened circumanal setae and 3+3 cuticular spines; 33, appendices anales with enlarged apex; 34–36, setae of great abdomen: 34, sensilla a of mesothorax; 35, dorsal seta m1 of mesothorax; 36, setae of posterior dorsal complex." figureDoi="http://doi.org/10.5281/zenodo.3741174" httpUri="https://zenodo.org/record/3741174/files/figure.png" pageId="2" pageNumber="340">Fig. 34</figureCitation>
) and 3 setae in row 
<emphasis id="F7F6132AFFB9ED774F97FB04FE393745" bold="true" box="[468,491,1269,1295]" pageId="2" pageNumber="340">m</emphasis>
with 
<emphasis id="F7F6132AFFB9ED774C6FFB04FD833745" bold="true" box="[556,593,1269,1295]" pageId="2" pageNumber="340">m1</emphasis>
(
<figureCitation id="5DB9D3BDFFB9ED774C23FB04FD7B3745" box="[608,681,1269,1295]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="2" pageNumber="340">Figs 6</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774CF6FB05FD003745" box="[693,722,1268,1295]" captionStart="FIGURES 32–36" captionStartId="9.[151,264,1946,1971]" captionTargetBox="[192,1395,929,1917]" captionTargetId="figure@9.[188,1399,927,1921]" captionTargetPageId="9" captionText="FIGURES 32–36. Arrhopalites persicus sp. nov.: 32, dorsal valve of female Abd VI with broadened circumanal setae and 3+3 cuticular spines; 33, appendices anales with enlarged apex; 34–36, setae of great abdomen: 34, sensilla a of mesothorax; 35, dorsal seta m1 of mesothorax; 36, setae of posterior dorsal complex." figureDoi="http://doi.org/10.5281/zenodo.3741174" httpUri="https://zenodo.org/record/3741174/files/figure.png" pageId="2" pageNumber="340">35</figureCitation>
) about as long as setae of posterior complex (
<figureCitation id="5DB9D3BDFFB9ED774A97FB04FACF3745" box="[1236,1309,1269,1295]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="2" pageNumber="340">Figs 7</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774B69FB04FA98375A" box="[1322,1354,1269,1296]" captionStart="FIGURES 32–36" captionStartId="9.[151,264,1946,1971]" captionTargetBox="[192,1395,929,1917]" captionTargetId="figure@9.[188,1399,927,1921]" captionTargetPageId="9" captionText="FIGURES 32–36. Arrhopalites persicus sp. nov.: 32, dorsal valve of female Abd VI with broadened circumanal setae and 3+3 cuticular spines; 33, appendices anales with enlarged apex; 34–36, setae of great abdomen: 34, sensilla a of mesothorax; 35, dorsal seta m1 of mesothorax; 36, setae of posterior dorsal complex." figureDoi="http://doi.org/10.5281/zenodo.3741174" httpUri="https://zenodo.org/record/3741174/files/figure.png" pageId="2" pageNumber="340">36</figureCitation>
). Abd I bears single row with 5 setae. Trichobothrial complex (
<figureCitation id="5DB9D3BDFFB9ED774D46FAE8FC8E3779" box="[773,860,1305,1331]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">Figs 16</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774D2AFAE8FC5B377E" box="[873,905,1305,1332]" captionStart="FIGURES 26–31" captionStartId="8.[151,264,1876,1901]" captionTargetBox="[162,1425,184,1847]" captionTargetId="figure@8.[159,1428,181,1850]" captionTargetPageId="8" captionText="FIGURES 26–31. Arrhopalites persicus sp. nov.: 26, chaetotaxy of trichobothrial complex; 27, chaetotaxy of central dorsal complex with seta 3 as sensilla; 28–30, foot complex: 28, foreleg; 29, mid leg; 30, hindleg; 31, mucro." figureDoi="http://doi.org/10.5281/zenodo.3741172" httpUri="https://zenodo.org/record/3741172/files/figure.png" pageId="2" pageNumber="340">26</figureCitation>
): 
<emphasis id="F7F6132AFFB9ED774DDDFAE9FC0B3779" bold="true" box="[926,985,1304,1331]" pageId="2" pageNumber="340">ABC</emphasis>
<taxonomicName id="0282B4BBFFB9ED774DA3FAE8FB8F3779" box="[992,1117,1305,1331]" form="about" pageId="2" pageNumber="340" rank="form">form about</taxonomicName>
right or a little obtuse angle (91°–111°) and 
<emphasis id="F7F6132AFFB9ED774F04FACDFEBC371D" bold="true" box="[327,366,1340,1367]" pageId="2" pageNumber="340">AB</emphasis>
is 1.3–1.9× longer than 
<emphasis id="F7F6132AFFB9ED774C3FFACDFD71371D" bold="true" box="[636,675,1340,1367]" pageId="2" pageNumber="340">BC</emphasis>
(1.5 in 
<typeStatus id="1A39719AFFB9ED774CB4FACCFC8C3712" box="[759,862,1341,1368]" pageId="2" pageNumber="340" type="holotype">holotype</typeStatus>
); seta 
<emphasis id="F7F6132AFFB9ED774DE1FACCFC62371D" bold="true" box="[930,944,1341,1367]" pageId="2" pageNumber="340">a</emphasis>
stronger and longer than others of complex; seta 
<emphasis id="F7F6132AFFB9ED774E89FA90FF083731" bold="true" box="[202,218,1377,1403]" pageId="2" pageNumber="340">p</emphasis>
is located approximately at the level of trichobothrium 
<emphasis id="F7F6132AFFB9ED774D0CFA90FCB03731" bold="true" box="[847,866,1377,1403]" pageId="2" pageNumber="340">B</emphasis>
; seta 
<emphasis id="F7F6132AFFB9ED774DE7FA90FC103731" bold="true" box="[932,962,1377,1403]" pageId="2" pageNumber="340">b1</emphasis>
lies on line 
<emphasis id="F7F6132AFFB9ED774A0EFA91FBA63731" bold="true" box="[1101,1140,1376,1403]" pageId="2" pageNumber="340">BC</emphasis>
, closer to 
<emphasis id="F7F6132AFFB9ED774AA9FA91FB2C3731" bold="true" box="[1258,1278,1376,1403]" pageId="2" pageNumber="340">C</emphasis>
; microseta 
<emphasis id="F7F6132AFFB9ED774BC1FA90FA4E3731" bold="true" box="[1410,1436,1377,1403]" pageId="2" pageNumber="340">c1</emphasis>
(4.5–6.8 μm) lies in front of trichobothrium 
<emphasis id="F7F6132AFFB9ED774CC5FA75FD4837D5" bold="true" box="[646,666,1412,1439]" pageId="2" pageNumber="340">C</emphasis>
and seta 
<emphasis id="F7F6132AFFB9ED774D47FA74FCCC37D5" bold="true" box="[772,798,1413,1439]" pageId="2" pageNumber="340">c2</emphasis>
—below 
<emphasis id="F7F6132AFFB9ED774DC5FA75FC4837D5" bold="true" box="[902,922,1412,1439]" pageId="2" pageNumber="340">C</emphasis>
. Posterior lateral complex with 6 setae in two rows and furca base complex with 8 setae in two rows, neosminthuroid seta absent. Central dorsal complex with usual setae 
<emphasis id="F7F6132AFFB9ED774F5FFA3CFEF837AD" bold="true" box="[284,298,1485,1511]" pageId="2" pageNumber="340">1</emphasis>
and 
<emphasis id="F7F6132AFFB9ED774F26FA3CFEA137AD" bold="true" box="[357,371,1485,1511]" pageId="2" pageNumber="340">2</emphasis>
and curved sensillum 
<emphasis id="F7F6132AFFB9ED774C34FA3CFD5737AD" bold="true" box="[631,645,1485,1511]" pageId="2" pageNumber="340">3</emphasis>
(
<figureCitation id="5DB9D3BDFFB9ED774CD5FA3CFD3E37AD" box="[662,748,1485,1511]" captionStart="FIGURES 26–31" captionStartId="8.[151,264,1876,1901]" captionTargetBox="[162,1425,184,1847]" captionTargetId="figure@8.[159,1428,181,1850]" captionTargetPageId="8" captionText="FIGURES 26–31. Arrhopalites persicus sp. nov.: 26, chaetotaxy of trichobothrial complex; 27, chaetotaxy of central dorsal complex with seta 3 as sensilla; 28–30, foot complex: 28, foreleg; 29, mid leg; 30, hindleg; 31, mucro." figureDoi="http://doi.org/10.5281/zenodo.3741172" httpUri="https://zenodo.org/record/3741172/files/figure.png" pageId="2" pageNumber="340">Fig. 27</figureCitation>
). Posterior dorsal complex with 21–25 relatively short setae (
<figureCitation id="5DB9D3BDFFB9ED774EDCFA00FF353441" box="[159,231,1521,1547]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="2" pageNumber="340">Figs 7</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774EB0FA00FEC23446" box="[243,272,1521,1548]" captionStart="FIGURES 32–36" captionStartId="9.[151,264,1946,1971]" captionTargetBox="[192,1395,929,1917]" captionTargetId="figure@9.[188,1399,927,1921]" captionTargetPageId="9" captionText="FIGURES 32–36. Arrhopalites persicus sp. nov.: 32, dorsal valve of female Abd VI with broadened circumanal setae and 3+3 cuticular spines; 33, appendices anales with enlarged apex; 34–36, setae of great abdomen: 34, sensilla a of mesothorax; 35, dorsal seta m1 of mesothorax; 36, setae of posterior dorsal complex." figureDoi="http://doi.org/10.5281/zenodo.3741174" httpUri="https://zenodo.org/record/3741174/files/figure.png" pageId="2" pageNumber="340">36</figureCitation>
) arranged in 3 longitudinal rows, the longest of which (among the most posterior setae of rows 
<emphasis id="F7F6132AFFB9ED774B79FA00FA413441" bold="true" box="[1338,1427,1521,1547]" pageId="2" pageNumber="340">dI–dIII</emphasis>
) = 20–29 μm (24 μm in 
<typeStatus id="1A39719AFFB9ED774FD9F9E4FE2C347A" box="[410,510,1557,1584]" pageId="2" pageNumber="340" type="holotype">holotype</typeStatus>
) and is somewhat shorter or subequal to hind claw (
<tableCitation id="8800FA83FFB9ED774A02F9E4FB4A3465" box="[1089,1176,1557,1583]" captionStart="TABLE 2" captionStartId="9.[151,239,154,178]" captionText="TABLE 2. Proportions for some body parts of type specimens (11 females) of Arrhopalites persicus sp. nov." pageId="2" pageNumber="340">Table 2</tableCitation>
). Ventral complex with 2 setae.
</paragraph>
<paragraph id="C53DCF38FFB9ED774E84F9ADFB5934D1" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="340">
<emphasis id="F7F6132AFFB9ED774E84F9ADFE52343D" bold="true" box="[199,384,1628,1655]" pageId="2" pageNumber="340">Small abdomen</emphasis>
. Fifth abdominal segment (
<figureCitation id="5DB9D3BDFFB9ED774CF6F9ACFCD4343D" box="[693,774,1629,1655]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">Fig. 19</figureCitation>
) with 2 setae and trichobothrium 
<emphasis id="F7F6132AFFB9ED774A3CF9ACFB41343D" bold="true" box="[1151,1171,1629,1655]" pageId="2" pageNumber="340">D</emphasis>
in row 
<emphasis id="F7F6132AFFB9ED774AABF9ACFB24343D" bold="true" box="[1256,1270,1629,1655]" pageId="2" pageNumber="340">a</emphasis>
, and 2 setae in row 
<emphasis id="F7F6132AFFB9ED774E89F970FF0834D1" bold="true" box="[202,218,1665,1691]" pageId="2" pageNumber="340">p</emphasis>
. Genital field usually with 3+3 microsetae along anterior margin of genital opening.
</paragraph>
<paragraph id="C53DCF38FFB9ED774E84F954FE4435F1" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="340">
Sixth abdominal segment (
<figureCitation id="5DB9D3BDFFB9ED774FACF954FD9634F5" box="[495,580,1701,1727]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[181,1418,200,1922]" captionTargetId="figure@6.[170,1418,181,1933]" captionTargetPageId="6" captionText="FIGURES 16–19. Arrhopalites persicus sp. nov.: 16, chaetotaxy of great abdomen; 17, chaetotaxy of furca; 18, tenaculum; 19, chaetotaxy of small abdomen." figureDoi="http://doi.org/10.5281/zenodo.3741168" httpUri="https://zenodo.org/record/3741168/files/figure.png" pageId="2" pageNumber="340">Figs 19</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774C0CF954FDBE34F5" box="[591,620,1701,1727]" captionStart="FIGURES 32–36" captionStartId="9.[151,264,1946,1971]" captionTargetBox="[192,1395,929,1917]" captionTargetId="figure@9.[188,1399,927,1921]" captionTargetPageId="9" captionText="FIGURES 32–36. Arrhopalites persicus sp. nov.: 32, dorsal valve of female Abd VI with broadened circumanal setae and 3+3 cuticular spines; 33, appendices anales with enlarged apex; 34–36, setae of great abdomen: 34, sensilla a of mesothorax; 35, dorsal seta m1 of mesothorax; 36, setae of posterior dorsal complex." figureDoi="http://doi.org/10.5281/zenodo.3741174" httpUri="https://zenodo.org/record/3741174/files/figure.png" pageId="2" pageNumber="340">32</figureCitation>
) with 5+5 cuticular spines: 3+3 on dorsal valve and 2+2 on each of lateral valves; 
<emphasis id="F7F6132AFFB9ED774EAFF938FED734A9" bold="true" box="[236,261,1737,1763]" pageId="2" pageNumber="340">s2</emphasis>
and 
<emphasis id="F7F6132AFFB9ED774F79F938FE8034A9" bold="true" box="[314,338,1737,1763]" pageId="2" pageNumber="340">s3</emphasis>
on dorsal valve usually larger than others (
<tableCitation id="8800FA83FFB9ED774D6EF938FC5234A9" box="[813,896,1737,1763]" captionStart="TABLE 1" captionStartId="5.[151,239,298,322]" captionText="TABLE 1. Lengths (in µm) and angles for body parts of type specimens (11 females) of Arrhopalites persicus sp. nov." pageId="2" pageNumber="340">Table 1</tableCitation>
). Dorsal valve with 11 + 2 axial + 11 setae. Each of lateral valves bears 18 setae. Circumanal row with 5+1+5 modified setae (
<emphasis id="F7F6132AFFB9ED774DBCF91CFBE2354D" bold="true" box="[1023,1072,1773,1799]" pageId="2" pageNumber="340">ms1</emphasis>
, 
<emphasis id="F7F6132AFFB9ED774A7DF91CFB1E354D" bold="true" box="[1086,1228,1773,1799]" pageId="2" pageNumber="340">mps1–mps3</emphasis>
and 
<emphasis id="F7F6132AFFB9ED774B47F91DFA59354D" bold="true" box="[1284,1419,1772,1799]" pageId="2" pageNumber="340">mpi1–mpi2</emphasis>
): long (the longest = 40–81 μm, that is about 2–3× longer than posterior setae of great abdomen), broadened but without distinct lamellation or basal serration (
<figureCitation id="5DB9D3BDFFB9ED774CE7F8C4FCDB3505" box="[676,777,1845,1871]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="2" pageNumber="340">Figs 8, 9</figureCitation>
). Appendices anales (
<figureCitation id="5DB9D3BDFFB9ED774DBDF8C4FB873505" box="[1022,1109,1845,1871]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1932,1957]" captionTargetBox="[215,1368,182,1904]" captionTargetId="figure@3.[211,1376,179,1904]" captionTargetPageId="3" captionText="FIGURES 1–12. Arrhopalites persicus sp. nov.: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales." figureDoi="http://doi.org/10.5281/zenodo.3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="2" pageNumber="340">Figs 12</figureCitation>
, 
<figureCitation id="5DB9D3BDFFB9ED774A22F8C4FBAC3505" box="[1121,1150,1845,1871]" captionStart="FIGURES 32–36" captionStartId="9.[151,264,1946,1971]" captionTargetBox="[192,1395,929,1917]" captionTargetId="figure@9.[188,1399,927,1921]" captionTargetPageId="9" captionText="FIGURES 32–36. Arrhopalites persicus sp. nov.: 32, dorsal valve of female Abd VI with broadened circumanal setae and 3+3 cuticular spines; 33, appendices anales with enlarged apex; 34–36, setae of great abdomen: 34, sensilla a of mesothorax; 35, dorsal seta m1 of mesothorax; 36, setae of posterior dorsal complex." figureDoi="http://doi.org/10.5281/zenodo.3741174" httpUri="https://zenodo.org/record/3741174/files/figure.png" pageId="2" pageNumber="340">33</figureCitation>
) (18–37 μm) smooth and apically acuminate; sitting on globular basal papilla. However, under highest magnification in dorso-lateral position few hardly noticeable toothlets on apex were observed (
<figureCitation id="5DB9D3BDFFB9ED774D4BF88CFC8935DD" box="[776,859,1917,1943]" captionStart="FIGURES 32–36" captionStartId="9.[151,264,1946,1971]" captionTargetBox="[192,1395,929,1917]" captionTargetId="figure@9.[188,1399,927,1921]" captionTargetPageId="9" captionText="FIGURES 32–36. Arrhopalites persicus sp. nov.: 32, dorsal valve of female Abd VI with broadened circumanal setae and 3+3 cuticular spines; 33, appendices anales with enlarged apex; 34–36, setae of great abdomen: 34, sensilla a of mesothorax; 35, dorsal seta m1 of mesothorax; 36, setae of posterior dorsal complex." figureDoi="http://doi.org/10.5281/zenodo.3741174" httpUri="https://zenodo.org/record/3741174/files/figure.png" pageId="2" pageNumber="340">Fig. 33</figureCitation>
). In total, normally 60 setae and 10 cuticular spines on Abd VI are present.
</paragraph>
<paragraph id="C53DCF38FFB9ED774E84F834FEBD3595" blockId="2.[151,1437,152,2015]" box="[199,367,1989,2015]" pageId="2" pageNumber="340">
<emphasis id="F7F6132AFFB9ED774E84F834FED63595" bold="true" box="[199,260,1989,2015]" pageId="2" pageNumber="340">Male</emphasis>
not seen.
</paragraph>
<caption id="91FD9FB0FFB8ED764ED4F87DFC1435A7" ID-DOI="http://doi.org/10.5281/zenodo.3741164" ID-Zenodo-Dep="3741164" httpUri="https://zenodo.org/record/3741164/files/figure.png" pageId="3" pageNumber="341" startId="3.[151,264,1932,1957]" targetBox="[215,1368,182,1904]" targetPageId="3">
<paragraph id="C53DCF38FFB8ED764ED4F87DFC1435A7" blockId="3.[151,1436,1932,2029]" pageId="3" pageNumber="341">
<emphasis id="F7F6132AFFB8ED764ED4F87DFE9535EF" bold="true" box="[151,327,1932,1957]" pageId="3" pageNumber="341">FIGURES 1–12.</emphasis>
<taxonomicName id="0282B4BBFFB8ED764F0DF87CFDFA35EE" authority="Vargovitsh &amp; Kahrarian, 2020" authorityName="Vargovitsh &amp; Kahrarian" authorityYear="2020" box="[334,552,1933,1957]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="341" phylum="Arthropoda" rank="species" species="persicus" status="sp. nov.">
<emphasis id="F7F6132AFFB8ED764F0DF87CFDFA35EE" box="[334,552,1933,1957]" italics="true" pageId="3" pageNumber="341">Arrhopalites persicus</emphasis>
</taxonomicName>
<emphasis id="F7F6132AFFB8ED764C6CF87CFDAD35EF" bold="true" box="[559,639,1933,1957]" pageId="3" pageNumber="341">
<taxonomicNameLabel id="ECC5AE51FFB8ED764C6CF87CFDAD35EF" box="[559,639,1933,1957]" pageId="3" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 1, chaetotaxy of head; 2, maxillary outer lobe; 3, chaetotaxy of labial palp; 4, chaetotaxy of antenna; 5–11, shape of setae: 5, seta of head vertex; 6, m1 of mesothorax; 7, dIII-1 of great abdomen; 8, ms1 of Abd VI; 9, mps3 of abd VI; 10, Ia of dens; 11, Ie of dens; 12, appendices anales.
</paragraph>
</caption>
<caption id="91FD9FB0FFBFED714ED4F8B7FF6F35C9" ID-DOI="http://doi.org/10.5281/zenodo.3741166" ID-Zenodo-Dep="3741166" httpUri="https://zenodo.org/record/3741166/files/figure.png" pageId="4" pageNumber="342" startId="4.[151,264,1862,1887]" targetBox="[151,1436,192,1830]" targetPageId="4">
<paragraph id="C53DCF38FFBFED714ED4F8B7FF6F35C9" blockId="4.[151,1436,1862,1923]" pageId="4" pageNumber="342">
<emphasis id="F7F6132AFFBFED714ED4F8B7FE863515" bold="true" box="[151,340,1862,1887]" pageId="4" pageNumber="342">FIGURES 13–15.</emphasis>
<taxonomicName id="0282B4BBFFBFED714F18F8B7FDE73514" authority="Vargovitsh &amp; Kahrarian, 2020" authorityName="Vargovitsh &amp; Kahrarian" authorityYear="2020" box="[347,565,1862,1886]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="342" phylum="Arthropoda" rank="species" species="persicus" status="sp. nov.">
<emphasis id="F7F6132AFFBFED714F18F8B7FDE73514" box="[347,565,1862,1886]" italics="true" pageId="4" pageNumber="342">Arrhopalites persicus</emphasis>
</taxonomicName>
<emphasis id="F7F6132AFFBFED714C7FF8B6FD5F3515" bold="true" box="[572,653,1863,1887]" pageId="4" pageNumber="342">
<taxonomicNameLabel id="ECC5AE51FFBFED714C7FF8B6FD5F3515" box="[572,653,1863,1887]" pageId="4" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 13, chaetotaxy of foreleg; 14, chaetotaxy of mid leg; 15, chaetotaxy of hind leg.
</paragraph>
</caption>
<paragraph id="C53DCF38FFBFED714E84F841FAF235A5" blockId="4.[151,1437,1968,2031]" pageId="4" pageNumber="342">
<emphasis id="F7F6132AFFBFED714E84F841FE9F3581" bold="true" box="[199,333,1968,1995]" pageId="4" pageNumber="342">Variability.</emphasis>
Two cases of teratology were observed: in two females (
<typeStatus id="1A39719AFFBFED714D8BF840FBEB3581" box="[968,1081,1969,1995]" pageId="4" pageNumber="342" type="paratype">paratypes</typeStatus>
: slide 27-3 and 27-10) seta 
<emphasis id="F7F6132AFFBFED714B2FF840FA4F3581" bold="true" box="[1388,1437,1969,1995]" pageId="4" pageNumber="342">ms1</emphasis>
distinctly bifurcated and in one female (
<typeStatus id="1A39719AFFBFED714C15F824FD6E35A5" box="[598,700,2005,2031]" pageId="4" pageNumber="342" type="paratype">paratype</typeStatus>
: slide 27-9) claw of left hind leg bears two inner teeth.
</paragraph>
<paragraph id="C53DCF38FFBEED704E84FF69FC71329D" blockId="5.[151,1436,152,251]" pageId="5" pageNumber="343">
<emphasis id="F7F6132AFFBEED704E84FF69FDDB32F9" bold="true" box="[199,521,152,179]" pageId="5" pageNumber="343">Bionomy and distribution.</emphasis>
Specimens of 
<taxonomicName id="0282B4BBFFBEED704CFAFF68FC7F32F9" authority="Vargovitsh &amp; Kahrarian, 2020" authorityName="Vargovitsh &amp; Kahrarian" authorityYear="2020" box="[697,941,153,179]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="343" phylum="Arthropoda" rank="species" species="persicus" status="sp. nov.">
<emphasis id="F7F6132AFFBEED704CFAFF68FC7F32F9" box="[697,941,153,179]" italics="true" pageId="5" pageNumber="343">Arrhopalites persicus</emphasis>
</taxonomicName>
<emphasis id="F7F6132AFFBEED704DFBFF68FBC732F9" bold="true" box="[952,1045,153,179]" pageId="5" pageNumber="343">
<taxonomicNameLabel id="ECC5AE51FFBEED704DFBFF68FBC732F9" box="[952,1045,153,179]" pageId="5" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were sampled from three sites in western 
<collectingCountry id="BD958FA8FFBEED704EB6FF4CFEF0329D" box="[245,290,189,215]" name="Iran" pageId="5" pageNumber="343">Iran</collectingCountry>
(
<figureCitation id="5DB9D3BDFFBEED704F71FF4CFE51329D" box="[306,387,189,215]" captionStart="FIGURES 20–25" captionStartId="7.[151,264,1663,1688]" captionTargetBox="[160,1426,184,1633]" captionTargetId="figure@7.[157,1430,181,1638]" captionTargetPageId="7" captionText="FIGURES 20–25. Arrhopalites persicus sp. nov.: 20, type localities on map of Iran: blue mark—Kal Kushk, red mark—Qader Mazd, green mark—Cheshmeh Sorkh; 21, habitus of mounted female; 22, fragment of head dorsum; 23, chaetotaxy of antennal III organ; 24, labial palps; 25, maxillae heads." figureDoi="http://doi.org/10.5281/zenodo.3741170" httpUri="https://zenodo.org/record/3741170/files/figure.png" pageId="5" pageNumber="343">Fig. 20</figureCitation>
) from upper soil layer and litter under oak trees.
</paragraph>
</subSubSection>
<subSubSection id="8D989CB3FFBEED704E84FF10FAFF32B1" box="[199,1325,225,251]" pageId="5" pageNumber="343" type="etymology">
<paragraph id="C53DCF38FFBEED704E84FF10FAFF32B1" blockId="5.[151,1436,152,251]" box="[199,1325,225,251]" pageId="5" pageNumber="343">
<emphasis id="F7F6132AFFBEED704E84FF10FE9C32B1" bold="true" box="[199,334,225,251]" pageId="5" pageNumber="343">Etymology.</emphasis>
The species name is an adjective derived from 
<collectingCountry id="BD958FA8FFBEED704D21FF10FC7B32B1" box="[866,937,225,251]" name="Iran" pageId="5" pageNumber="343">Persia</collectingCountry>
, the Latin name pertaining to 
<collectingCountry id="BD958FA8FFBEED704ABAFF10FAF832B1" box="[1273,1322,225,251]" name="Iran" pageId="5" pageNumber="343">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>