302 lines
30 KiB
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302 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="4ADD25D4A060DA959506865141451AC7" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole podargea Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="9A9D09F6-79B6-40AA-BB1E-BE65E6DC3FB3" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>905</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>235</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161777009" LSID="urn:lsid:zoobank.org:act:9A9D09F6-79B6-40AA-BB1E-BE65E6DC3FB3" httpUri="http://treatment.plazi.org/id/4ADD25D4A060DA959506865141451AC7" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/9A9D09F6-79B6-40AA-BB1E-BE65E6DC3FB3" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole podargea" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="podargea" status="sp. nov.">Pheidole podargea</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">Figs 33A-F</figureCitation>
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<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85R</figureCitation>
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<figureCitation captionStart="Figure 88" captionStartId="F88" captionText="Figure 88. Distribution. Pheidole petax Forel (A). P. podargea sp. nov. (B). P. praegrandis sp. nov. (C). P. ranohirensis sp. nov. (D). P. rugocephala sp. nov. (E). P. rugofitarata sp. nov. (F). P. scabrata Forel (G). P. typhlos sp. nov. (H). P. vatovavensis sp. nov. (I). P. voasara sp. nov. (J). P. vohemarensis sp. nov. (K). P. zavamanira sp. nov. (L)." figureDoi="10.3897/zookeys.905.39592.figure88" httpUri="https://binary.pensoft.net/fig/373135" pageId="0" pageNumber="1">, 88B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Antsiranana;
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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<normalizedToken originalValue="d’">d'</normalizedToken>
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Andavakoera, 21.4 km 75°ENE Ambilobe; 4.6 km 356°N Betsiaka; -13.11833, 49.23; alt. 425 m; 16 Dec 2003; B.L. Fisher leg.; BLF10317, CASENT0487570, middle specimen on the pin (CASC).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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Madagascar. •9w., 8s.; same data as for holotype; CASENT0487566, CASENT0872085, CASENT0487569, CASENT0487568, CASENT0487567 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: •9w., 4s.;
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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de Binara, 7.5 km 230°SW Daraina;
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<geoCoordinate degrees="13.255" direction="south" orientation="latitude" precision="55" value="-13.255">-13.255</geoCoordinate>
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,
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<geoCoordinate degrees="49.61667" direction="east" orientation="longitude" precision="1" value="49.61667">49.61667</geoCoordinate>
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; alt. 375 m; 1 Dec 2003; B.L. Fisher leg.; CASENT0041888, CASENT0043218, CASENT0043224, CASENT0043226, CASENT0043235, CASENT0043241, CASENT0043253, CASENT0043256, CASENT0043259, CASENT0043317, CASENT0043319, CASENT0043331, CASENT0043402 (CASC). •5w., 1s., 2q.;
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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<normalizedToken originalValue="d’Ambre">d'Ambre</normalizedToken>
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, 3.5 km 235°SW Sakaramy;
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<geoCoordinate degrees="12.46889" direction="south" orientation="latitude" precision="1" value="-12.46889">-12.46889</geoCoordinate>
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,
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<geoCoordinate degrees="49.24217" direction="east" orientation="longitude" precision="1" value="49.24217">49.24217</geoCoordinate>
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; alt. 325 m; 26 Jan 2001; Fisher et al. leg.; CASENT0406663, CASENT0406673, CASENT0423867, CASENT0427613, CASENT0427696, CASENT0427699 (CASC). •32w., 9s., 3q.;
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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de
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<normalizedToken originalValue="l’Ankarana">l'Ankarana</normalizedToken>
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, 13.6 km 192°SSW Anivorano Nord;
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<geoCoordinate degrees="12.86361" direction="south" orientation="latitude" precision="1" value="-12.86361">-12.86361</geoCoordinate>
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,
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<geoCoordinate degrees="49.22583" direction="east" orientation="longitude" precision="1" value="49.22583">49.22583</geoCoordinate>
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; alt. 210 m; 16 Feb 2001; Fisher et al. leg.; CASENT0440586, CASENT0440686, CASENT0440729, CASENT0440730, CASENT0440731, CASENT0440735, CASENT0440736, CASENT0440739, CASENT0440740, CASENT0440743, CASENT0440745, CASENT0440746, CASENT0440748, CASENT0440750, CASENT0440752, CASENT0440754, CASENT0440755, CASENT0440756, CASENT0440758, CASENT0440762-CASENT0440765, CASENT0440769, CASENT0440771, CASENT0440773, CASENT0440774, CASENT0440776, CASENT0440780, CASENT0440782, CASENT0440786-CASENT0440788, CASENT0440790, CASENT0440792, CASENT0440803, CASENT0440808, CASENT0440813, CASENT0441204-CASENT0441206, CASENT0441208, CASENT0441211, CASENT0441212, CASENT0441214, CASENT0441216 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Body size moderate: HL: 1.85-2.0 (1.9), HW: 1.55-1.65 (1.59), WL: 1.19-1.35 (1.26); propodeal spines very long (PSL: 0.39-0.46 (0.42)); head in full-face view rectangular, with lateral sides relatively straight, only their posteriormost part slightly convex; sides of the head with moderately dense, moderately long, erect pilosity; occipital lobes shiny, with sparse and thick rugoreticulation; inner hypostomal teeth distinct, closely spaced, triangular, with rounded apex; outer hypostomal teeth distinct, as high as inner hypostomal teeth, lobe-like, with base wide and tops directed slightly inward; inner and outer teeth closely spaced and connected by concavity.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Body size moderate: HL: 0.61-0.7 (0.66), HW: 0.62-0.69 (0.65), WL: 0.81-0.9 (0.86); propodeal spines very long (PSL: 0.25-0.31 (0.28)); scape, when laid back, surpassing posterior head margin by two-fifths of its length; whole head with fine and dense rugo-punctuation, only basal area of frons and genae with sculpture reduced or sometimes absent; mesosoma rugo-punctate, sometimes sculpture weakening on dorsal surface and pronotum.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 10): HL: 1.85-2.0 (1.9); HW: 1.55-1.65 (1.59); SL: 0.67-0.76 (0.72); EL: 0.16-0.2 (0.17); WL: 1.19-1.35 (1.26); PSL: 0.39-0.46 (0.42); MTL: 0.7-0.76 (0.73); PNW: 0.71-0.81 (0.76); PTW: 0.15-0.22 (0.19); PPW: 0.59-0.71 (0.66); CI: 80.1-87.6 (83.4); SI: 42.6-47.7 (45.6); PSLI: 20.9-24.4 (21.8); PPI: 25.4-32.7 (29.1); PNI: 45.5-51.0 (48.2); MTI: 42.6-48.8 (46.4).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view rectangular, with lateral sides relatively straight, only their posteriormost part slightly convex (Fig.
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33B</figureCitation>
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). In lateral view oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with moderately dense, moderately long, erect pilosity. Antennal scrobes indistinct and not delimited by carinulae. Occipital lobes shiny, with sparse and thick rugoreticulation; genae shiny, with moderately dense to dense, fine rugoreticulation; frons with longitudinal, sparse to moderately sparse rugae, on posterior part of frons rugae directed outward and more irregular, interspaces with sparse rugulae, sometimes indistinct; malar area and lateral sides of head shiny, with longitudinal rugoreticulation, surface between rugae with sparse and indistinct rugulae. Centre of clypeus smooth and shiny or with fine longitudinal rugulae, lateral sides with longitudinal rugae; median notch present, wide, and shallow; median longitudinal carina present but sometimes indistinct; lateral longitudinal carinae absent. Scape, when laid back, reaching midlength of head; pilosity suberect to erect (Fig.
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33B, D</figureCitation>
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). Inner hypostomal teeth distinct, closely spaced, triangular, with rounded apex; outer hypostomal teeth distinct, as high as inner hypostomal teeth, lobe-like, with base wide and tops directed slightly inward; inner and outer teeth closely spaced and connected by concavity (Fig.
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<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85R</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum short, angular, and high, with slightly convex posterior declivity; promesonotal groove absent; metanotal groove absent; propodeal spines very long, massive basally, with acute apex; humeral area laterally weakly produced (Fig.
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33D</figureCitation>
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). Surface shiny, with fine, dense rugoreticulation, sometimes sculpture weakening on dorsum. Pilosity sparse, long, and erect (Fig.
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shagreened; peduncle relatively long, with small tooth-like horizontal lobes on its basal part; node triangular, with rounded apex, in rear view node convex; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Shagreened; in dorsal view sides with acute, horn-like, moderately long projections; pilosity long, moderately sparse, and erect (Fig.
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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First gastral tergite shagreened, at least on its basal part; pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
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.
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</emphasis>
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Unicolourous, ocherous to reddish-brown (Fig.
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33D, F</figureCitation>
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).
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</paragraph>
|
||
<caption doi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1" start="Figure 33" startId="F33">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 33.</emphasis>
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole podargea" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="podargea">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole podargea</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., full-face view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
), profile (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
), and dorsal view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
) of paratype minor worker (CASENT0487567) and full-face view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
), profile (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
), and dorsal view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
) of holotype major worker (CASENT0487570).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 0.61-0.7 (0.66); HW: 0.62-0.69 (0.65); SL: 0.63-0.7 (0.67); EL: 0.11-0.12 (0.11); WL: 0.81-0.9 (0.86); PSL: 0.25-0.31 (0.28); MTL: 0.49-0.58 (0.53); PNW: 0.39-0.45 (0.43); PTW: 0.09-0.11 (0.1); PPW: 0.17-0.21 (0.19); CI: 94.3-100.7 (97.2); SI: 97.5-107.9 (103.1); PSLI: 38.8-44.3 (41.7); PPI: 46.9-56.9 (51.6); PNI: 63.4-69.7 (66.2); MTI: 78.4-85.0 (82.3).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
In full-face view square, posterior of eyes slightly convex, anterior of eyes relatively straight, occipital margin straight or indistinctly convex; occipital carina indistinct, weakly developed (Fig.
|
||
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33A</figureCitation>
|
||
). Pilosity moderately dense, long, and erect. Whole head with fine and dense rugo-punctuation, only basal area of frons and genae with sculpture reduced or sometimes absent; antennal sockets with dense carinae curved outward. Clypeus with median longitudinal carina present; two lateral longitudinal carinae absent. Scape, when laid back, surpassing posterior head margin by two-fifths of its length; pilosity suberect to erect (Fig.
|
||
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33A, C</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
In lateral view, promesonotum high, short, and convex; promesonotal groove present, indistinct; metanotal groove shallow and indistinct; propodeal spines very long, massive basally, with acute apex (Fig.
|
||
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33C</figureCitation>
|
||
). Whole surface rugo-punctate, sometimes sculpture weakening on dorsal surface and pronotum. Pilosity sparse, long, and erect (Fig.
|
||
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Peduncle relatively long and thin; node low, triangular; with few long, erect setae (Fig.
|
||
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Short, low, and slightly convex; with few long, erect setae at the anterior edge (Fig.
|
||
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
With few long, erect setae (Fig.
|
||
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Unicolourous, bright brown to brown (Fig.
|
||
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole podargea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0487567) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0487570)." figureDoi="10.3897/zookeys.905.39592.figure33" httpUri="https://binary.pensoft.net/fig/373080" pageId="0" pageNumber="1">33C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Named after Podarge, a harpy from Greek mythology, in reference to long and sharp propodeal spines of minor workers reminiscent of claws.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">The species was collected between 210-800 m in elevation, in rainforest and tropical dry forest. Nests were located in rotten logs.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="comments">
|
||
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
This species is most similar to
|
||
<taxonomicName lsidName="P. aelloea" pageId="0" pageNumber="1" rank="species" species="aelloea">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. aelloea</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName lsidName="P. ocypodea" pageId="0" pageNumber="1" rank="species" species="ocypodea">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. ocypodea</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole podargea" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="podargea">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole podargea</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName lsidName="P. aelloea" pageId="0" pageNumber="1" rank="species" species="aelloea">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. aelloea</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName lsidName="P. ocypodea" pageId="0" pageNumber="1" rank="species" species="ocypodea">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. ocypodea</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by outer hypostomal teeth approximately the same size as the inner ones and lack of smooth sculpture on genae and propodeum.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole podargea" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="podargea">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole podargea</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName lsidName="P. aelloea" pageId="0" pageNumber="1" rank="species" species="aelloea">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. aelloea</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName lsidName="P. ocypodea" pageId="0" pageNumber="1" rank="species" species="ocypodea">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. ocypodea</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by smooth area limited to basal area of frons and genae or the whole head finely rugo-punctate, presence of promesonotal and metanotal groove, and absence of smooth surfaces on mesosoma.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |