treatments-xml/data/47/39/C1/4739C1B3B00F5DF4980532C80AD75CEC.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1072.67935" ID-GBIF-Dataset="0a9353f0-1f24-45f0-8ddc-bd941470a567" ID-PMC="PMC8626413" ID-Pensoft-Pub="1313-2970-1072-1" ID-Pensoft-UUID="DDB04884FF7C52BD9D1814C0EA84E496" ID-PubMed="34899006" ID-ZooBank="9F578799E05F44118E2D378E3C17F3E3" ModsDocID="1313-2970-1072-1" checkinTime="1637381493323" checkinUser="pensoft" docAuthor="Li, Ya, Li, Shuqiang &amp; Lin, Yucheng" docDate="2021" docId="4739C1B3B00F5DF4980532C80AD75CEC" docLanguage="en" docName="ZooKeys 1072: 1-47" docOrigin="ZooKeys 1072" docPubDate="2021-11-19" docSource="http://dx.doi.org/10.3897/zookeys.1072.67935" docTitle="Patu Marples 1951" docType="treatment" docVersion="4" id="DDB04884FF7C52BD9D1814C0EA84E496" lastPageNumber="1" masterDocId="DDB04884FF7C52BD9D1814C0EA84E496" masterDocTitle="Taxonomic study on fourteen symphytognathid species from Asia (Araneae, Symphytognathidae)" masterLastPageNumber="47" masterPageNumber="1" pageNumber="1" updateTime="1668151161727" updateUser="ExternalLinkService">
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<mods:title>Taxonomic study on fourteen symphytognathid species from Asia (Araneae, Symphytognathidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Li, Ya</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1097-6192</mods:nameIdentifier>
<mods:affiliation>Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
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<mods:namePart>Li, Shuqiang</mods:namePart>
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<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart>Lin, Yucheng</mods:namePart>
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<mods:affiliation>Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China &amp; The Sichuan Key Laboratory for Conservation Biology of Endangered Wildlife, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
<mods:nameIdentifier type="email">linyucheng@scu.edu.cn</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-11-19</mods:number>
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<mods:number>1072</mods:number>
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<mods:end>47</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1072.67935</mods:url>
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<treatment ID-GBIF-Taxon="190343152" LSID="urn:lsid:plazi:treatment:4739C1B3B00F5DF4980532C80AD75CEC" httpUri="http://treatment.plazi.org/id/4739C1B3B00F5DF4980532C80AD75CEC" lastPageNumber="1" pageId="0" pageNumber="1">
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Genus 
<taxonomicName LSID="4739C1B3-B00F-5DF4-9805-32C80AD75CEC" authority="Marples, 1951" authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Patu Marples, 1951</taxonomicName>
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<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Patu</taxonomicName>
Marples, 1951: 47.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Patu</taxonomicName>
Forster, 1959: 318.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Patu</taxonomicName>
Forster &amp; Platnick, 1977: 15.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu vitiensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vitiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Patu vitiensis</emphasis>
</taxonomicName>
Marples, 1951 by original designation, from Fiji.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Patu</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName authorityName="Gertsch" authorityYear="1941" class="Arachnida" family="Symphytognathidae" genus="Anapistula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anapistula" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Anapistula</emphasis>
</taxonomicName>
Gertsch, 1941 by having 6 eyes vs. four or lacking and from 
<taxonomicName authorityName="Simon" authorityYear="1905" class="Arachnida" family="Symphytognathidae" genus="Anapogonia" kingdom="Animalia" lsidName="Anapogonia" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Anapogonia</emphasis>
</taxonomicName>
Simon, 1905, tentatively placed in 
<taxonomicName authorityName="Hickman" authorityYear="1931" class="Arachnida" family="Symphytognathidae" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Symphytognathidae</taxonomicName>
(Platnick and Forster 1989: 76), by the chelicerae fused at the mid-line vs. unfused. 
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Patu</emphasis>
</taxonomicName>
differs from 
<taxonomicName authorityName="Balogh &amp; Loksa" authorityYear="1968" class="Arachnida" family="Symphytognathidae" genus="Globignatha" kingdom="Animalia" lsidName="Globignatha" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Globignatha</emphasis>
</taxonomicName>
Balogh &amp; Loksa, 1968 and 
<taxonomicName authorityName="Hickman" authorityYear="1931" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphytognatha" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Symphytognatha</emphasis>
</taxonomicName>
Hickman, 1931 by the chelicerae fused only at mid-line vs. almost fully fused, see 
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4638.2.8" author="Lin, Y" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="291 - 295" refId="B10" refString="Lin, Y, 2019. First report of the spider genus Symphytognatha from Asia (Araneae, Symphytognathidae). Zootaxa 4638 (2): 291 - 295, DOI: https://doi.org/10.11646/zootaxa.4638.2.8" title="First report of the spider genus Symphytognatha from Asia (Araneae, Symphytognathidae)." url="https://doi.org/10.11646/zootaxa.4638.2.8" volume="4638" year="2019">Lin 2019</bibRefCitation>
: fig. 1H. It differs from 
<taxonomicName class="Arachnida" family="Symphytognathidae" genus="Curimagua" kingdom="Animalia" lsidName="Curimagua" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Curimagua</emphasis>
</taxonomicName>
Forster &amp; Platnick, 1977 by having 6 eyes in diads and the female lacking palps (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Patu nagarat sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure8" httpUri="https://binary.pensoft.net/fig/612450" pageId="0" pageNumber="1">8A</figureCitation>
and 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Patu nagarat sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure8" httpUri="https://binary.pensoft.net/fig/612450" pageId="0" pageNumber="1">C</figureCitation>
) vs. 6 eyes in triads and female palps reduced to vestiges (
<bibRefCitation author="Forster, RR" journalOrPublisher="American Museum Novitates" pageId="0" pageNumber="1" pagination="1 - 29" refId="B2" refString="Forster, RR, Platnick, NI, 1977. A review of the spider family Symphytognathidae (Arachnida, Araneae). American Museum Novitates 2619: 1 - 29" title="A review of the spider family Symphytognathidae (Arachnida, Araneae)." volume="2619" year="1977">Forster and Platnick 1977</bibRefCitation>
: figs 40 and 63). 
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Patu</emphasis>
</taxonomicName>
differs from 
<taxonomicName authorityName="Simon" authorityYear="1899" class="Arachnida" family="Symphytognathidae" genus="Iardinis" kingdom="Animalia" lsidName="Iardinis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Iardinis</emphasis>
</taxonomicName>
Simon, 1899 (
<taxonomicName lsidName="I. martensi" pageId="0" pageNumber="1" rank="species" species="martensi">
<emphasis italics="true" pageId="0" pageNumber="1">I. martensi</emphasis>
</taxonomicName>
Brignoli, 1978 from Nepal and 
<taxonomicName lsidName="I. mussardi" pageId="0" pageNumber="1" rank="species" species="mussardi">
<emphasis italics="true" pageId="0" pageNumber="1">I. mussardi</emphasis>
</taxonomicName>
Brignoli, 1980 from India) by having clasping spines on tibia II on the male, but lacking in the latter and from 
<taxonomicName class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Crassignatha</emphasis>
</taxonomicName>
Wunderlich, 1995 and 
<taxonomicName authorityName="S. Li &amp; Lin" authorityYear="2021" family="Symphytognathidae" genus="Swilda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Swilda" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Swilda</emphasis>
</taxonomicName>
gen. nov. by lacking a latero-posterior abdominal scutum in the male and the rod-shaped or oval spermathecae (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Patu catba sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D male palp, prolateral E male palp, retrolateral. Abbreviations: Co = conductor; Cy = cymbium; E = embolus; Fe = femur; MA = median apophysis; Pa = patella; T = tegulum; Ti = tibia; TS = male clasping spines on tibia II. Scale bars: 0.50 (A-C); 0.10 (D, E)." figureDoi="10.3897/zookeys.1072.67935.figure1" httpUri="https://binary.pensoft.net/fig/612443" pageId="0" pageNumber="1">1C</figureCitation>
, 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Patu nagarat sp. nov. A male palp, prolateral B male palp, retrolateral C epigyne, ventral D epigyne, lateral E vulva, ventral F vulva, dorsal. Abbreviations: CD = copulatory ducts; CO = copulatory opening; Cy = cymbium; E = embolus; FD = fertilisation ducts; Fe = femur; MA = median apophysis; Pa = patella; Pl = parmula; S = spermathecae; T = tegulum; Ti = tibia; TP = tegular process. Scale bars: 0.10 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure9" httpUri="https://binary.pensoft.net/fig/612451" pageId="0" pageNumber="1">9F</figureCitation>
and 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Patu nigeri A female habitus, dorsal B female habitus, ventral C female habitus, lateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Abbreviations: CD = copulatory ducts; CO = copulatory opening; FD = fertilisation ducts; Pl = parmula; S = spermathecae. Scale bars: 0.50 (A-C); 0.10 (D-G)." figureDoi="10.3897/zookeys.1072.67935.figure10" httpUri="https://binary.pensoft.net/fig/612452" pageId="0" pageNumber="1">10G</figureCitation>
) vs. having an abdominal scutum and spherical spermathecae (Figs 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Swilda longtou A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure19" httpUri="https://binary.pensoft.net/fig/612461" pageId="0" pageNumber="1">19A</figureCitation>
, 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Swilda spinathoraxi A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure21" httpUri="https://binary.pensoft.net/fig/612463" pageId="0" pageNumber="1">21A</figureCitation>
and 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Swilda spinathoraxi A male palp, prolateral B male palp, retrolateral C epigyne, ventral D vulva, ventral E vulva, dorsal. Abbreviations: CA = cymbial apophysis; CD = copulatory ducts; Co = conductor; CO = copulatory opening; Cy = cymbium; CP 1 = proximal cymbial process; CP 2 = distal cymbial process; E = embolus; FD = fertilisation ducts; Fe = femur; MA = median apophysis; Pa = patella; S = spermathecae; T = tegulum; Ti = tibia. Scale bars: 0.10 (A-E)." figureDoi="10.3897/zookeys.1072.67935.figure22" httpUri="https://binary.pensoft.net/fig/612464" pageId="0" pageNumber="1">22E</figureCitation>
; Li, Lin and 
<bibRefCitation DOI="https://doi.org/10.11865/zs.202011" author="Li, S" journalOrPublisher="Zoological Systematics" pageId="0" pageNumber="1" pagination="73 - 77" refId="B8" refString="Li, S, 2020. Spider taxonomy for an advanced China. Zoological Systematics 45 (2): 73 - 77, DOI: https://doi.org/10.11865/zs.202011" title="Spider taxonomy for an advanced China." url="https://doi.org/10.11865/zs.202011" volume="45" year="2020">Li 2020</bibRefCitation>
: figs 16C and 22D). 
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Patu</emphasis>
</taxonomicName>
is similar to 
<taxonomicName authorityName="S. Li &amp; Lin" authorityYear="2021" family="Symphytognathidae" genus="Kirinua" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kirinua" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Kirinua</emphasis>
</taxonomicName>
gen. nov. by the absence of a latero-posterior abdominal scutum in the male and the carapace surface lacking granular or spinous ornaments in both sexes, but it can be distinguished by the male having sulci and pores on the clypeus, rather than a pair of pocket-shaped pits in the latter and the male palpal cymbium lacks accessorial structures (e.g. primary conductor, cymbial process or apophysis); females can be distinguished by having rod-shaped or oval spermathecae rather than spherical or subspherical spermathecae in the latter.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Tiny, total length 0.40-0.80. Carapace round in male, pear-shaped in female dorsally, nearly triangular laterally (Figs 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Patu jidanweishi A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure6" httpUri="https://binary.pensoft.net/fig/612448" pageId="0" pageNumber="1">6A</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Patu dakou sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure2" httpUri="https://binary.pensoft.net/fig/612444" pageId="0" pageNumber="1">2D</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Patu nagarat sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure8" httpUri="https://binary.pensoft.net/fig/612450" pageId="0" pageNumber="1">8A</figureCitation>
and 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Patu nagarat sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure8" httpUri="https://binary.pensoft.net/fig/612450" pageId="0" pageNumber="1">8D</figureCitation>
). Six eyes in 3 diads, ocular base black, AME absent, lateral eyes adjacent, cephalic part raised (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Patu dakou sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure2" httpUri="https://binary.pensoft.net/fig/612444" pageId="0" pageNumber="1">2C</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Patu dakou sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure2" httpUri="https://binary.pensoft.net/fig/612444" pageId="0" pageNumber="1">2F</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Patu nagarat sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure8" httpUri="https://binary.pensoft.net/fig/612450" pageId="0" pageNumber="1">8C</figureCitation>
and 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Patu nagarat sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure8" httpUri="https://binary.pensoft.net/fig/612450" pageId="0" pageNumber="1">8F</figureCitation>
). Clypeus concave, with modified sulci and pores (fig. 69A-D in 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="9 - 195" refId="B16" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" volume="11" year="2009">Miller et al. 2009</bibRefCitation>
). Female lacking palps. Chelicerae fused at middle, with a single tooth (fig. 69E and F in 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="9 - 195" refId="B16" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" volume="11" year="2009">Miller et al. 2009</bibRefCitation>
). Labium wider than long, fused to sternum (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Patu dakou sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure2" httpUri="https://binary.pensoft.net/fig/612444" pageId="0" pageNumber="1">2B</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Patu dakou sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure2" httpUri="https://binary.pensoft.net/fig/612444" pageId="0" pageNumber="1">2E</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Patu nagarat sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure8" httpUri="https://binary.pensoft.net/fig/612450" pageId="0" pageNumber="1">8B</figureCitation>
and 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Patu nagarat sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure8" httpUri="https://binary.pensoft.net/fig/612450" pageId="0" pageNumber="1">8E</figureCitation>
). Sternum heart shaped, truncated posteriorly. Male tibia II with 1-2 clasping spines subdistally (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Patu catba sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D male palp, prolateral E male palp, retrolateral. Abbreviations: Co = conductor; Cy = cymbium; E = embolus; Fe = femur; MA = median apophysis; Pa = patella; T = tegulum; Ti = tibia; TS = male clasping spines on tibia II. Scale bars: 0.50 (A-C); 0.10 (D, E)." figureDoi="10.3897/zookeys.1072.67935.figure1" httpUri="https://binary.pensoft.net/fig/612443" pageId="0" pageNumber="1">1C</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Patu dakou sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure2" httpUri="https://binary.pensoft.net/fig/612444" pageId="0" pageNumber="1">2C</figureCitation>
, 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Patu jidanweishi A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure6" httpUri="https://binary.pensoft.net/fig/612448" pageId="0" pageNumber="1">6C</figureCitation>
and 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Patu nagarat sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure8" httpUri="https://binary.pensoft.net/fig/612450" pageId="0" pageNumber="1">8C</figureCitation>
). Abdomen globular dorsally, subovoid laterally, without latero-posterior abdominal scutum (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Patu catba sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D male palp, prolateral E male palp, retrolateral. Abbreviations: Co = conductor; Cy = cymbium; E = embolus; Fe = femur; MA = median apophysis; Pa = patella; T = tegulum; Ti = tibia; TS = male clasping spines on tibia II. Scale bars: 0.50 (A-C); 0.10 (D, E)." figureDoi="10.3897/zookeys.1072.67935.figure1" httpUri="https://binary.pensoft.net/fig/612443" pageId="0" pageNumber="1">1C</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Patu damtao sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D male palp, prolateral E male left palp, retrolateral. Abbreviations: Co = conductor; Cy = cymbium; E = embolus; Pa = patella; T = tegulum; Ti = tibia; TS = male clasping spines on tibia II. Scale bars: 0.50 (A-C); 0.10 (D, E)." figureDoi="10.3897/zookeys.1072.67935.figure4" httpUri="https://binary.pensoft.net/fig/612446" pageId="0" pageNumber="1">4C</figureCitation>
and 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Patu nagarat sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure8" httpUri="https://binary.pensoft.net/fig/612450" pageId="0" pageNumber="1">8C</figureCitation>
). Spinnerets without annular plates. Colulus absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male palp</emphasis>
(Figs 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Patu damtao sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D male palp, prolateral E male left palp, retrolateral. Abbreviations: Co = conductor; Cy = cymbium; E = embolus; Pa = patella; T = tegulum; Ti = tibia; TS = male clasping spines on tibia II. Scale bars: 0.50 (A-C); 0.10 (D, E)." figureDoi="10.3897/zookeys.1072.67935.figure4" httpUri="https://binary.pensoft.net/fig/612446" pageId="0" pageNumber="1">4D, E</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Patu nagarat sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure8" httpUri="https://binary.pensoft.net/fig/612450" pageId="0" pageNumber="1">8A</figureCitation>
and 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Patu nagarat sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Abbreviation: TS = male clasping spines on tibia II. Scale bars: 0.50 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure8" httpUri="https://binary.pensoft.net/fig/612450" pageId="0" pageNumber="1">B</figureCitation>
): bulb nearly ovate, large, not less than ~ 
<normalizedToken originalValue="¼">1/4</normalizedToken>
size of carapace. Cymbium membranous, translucent, wrapping around bulb prolaterally, without modified teeth, processes or apophyses. Conductor usually absent (if present, long, finger-like, starting at dorsal side of bulb, close to embolic base, see Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Patu catba sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D male palp, prolateral E male palp, retrolateral. Abbreviations: Co = conductor; Cy = cymbium; E = embolus; Fe = femur; MA = median apophysis; Pa = patella; T = tegulum; Ti = tibia; TS = male clasping spines on tibia II. Scale bars: 0.50 (A-C); 0.10 (D, E)." figureDoi="10.3897/zookeys.1072.67935.figure1" httpUri="https://binary.pensoft.net/fig/612443" pageId="0" pageNumber="1">1D</figureCitation>
and 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Patu damtao sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D male palp, prolateral E male left palp, retrolateral. Abbreviations: Co = conductor; Cy = cymbium; E = embolus; Pa = patella; T = tegulum; Ti = tibia; TS = male clasping spines on tibia II. Scale bars: 0.50 (A-C); 0.10 (D, E)." figureDoi="10.3897/zookeys.1072.67935.figure4" httpUri="https://binary.pensoft.net/fig/612446" pageId="0" pageNumber="1">4D</figureCitation>
). Tegulum cup shaped, with 1 or 2 protrusions (median apophysis and tegular process) (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Patu catba sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D male palp, prolateral E male palp, retrolateral. Abbreviations: Co = conductor; Cy = cymbium; E = embolus; Fe = femur; MA = median apophysis; Pa = patella; T = tegulum; Ti = tibia; TS = male clasping spines on tibia II. Scale bars: 0.50 (A-C); 0.10 (D, E)." figureDoi="10.3897/zookeys.1072.67935.figure1" httpUri="https://binary.pensoft.net/fig/612443" pageId="0" pageNumber="1">1D</figureCitation>
, 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Patu jidanweishi A male palp, prolateral B male palp, retrolateral C epigyne, ventral D epigyne, lateral E vulva, ventral F vulva, dorsal. Abbreviations: CD = copulatory ducts; Cy = cymbium; E = embolus; FD = fertilisation ducts; Fe = femur; Pa = patella; S = spermathecae; Sp = scape; T = tegulum; Ti = tibia; TP = tegular process. Scale bars: 0.10 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure7" httpUri="https://binary.pensoft.net/fig/612449" pageId="0" pageNumber="1">7A</figureCitation>
and 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Patu nagarat sp. nov. A male palp, prolateral B male palp, retrolateral C epigyne, ventral D epigyne, lateral E vulva, ventral F vulva, dorsal. Abbreviations: CD = copulatory ducts; CO = copulatory opening; Cy = cymbium; E = embolus; FD = fertilisation ducts; Fe = femur; MA = median apophysis; Pa = patella; Pl = parmula; S = spermathecae; T = tegulum; Ti = tibia; TP = tegular process. Scale bars: 0.10 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure9" httpUri="https://binary.pensoft.net/fig/612451" pageId="0" pageNumber="1">9A</figureCitation>
). Embolus long, slender, tubular, coiling into at least 2 loops within tegulum, distal part of embolus embedded inside bulb (Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Patu dakou sp. nov. A male palp, prolateral B male palp, retrolateral C epigyne, ventral D epigyne, lateral E vulva, ventral F vulva, dorsal. Abbreviations: CD = copulatory ducts; CO = copulatory opening; Cy = cymbium; E = embolus; FD = fertilisation ducts; Fe = femur; MA = median apophysis; Pa = patella; Pl = parmula; S = spermathecae; T = tegulum; Ti = tibia. Scale bars: 0.10 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure3" httpUri="https://binary.pensoft.net/fig/612445" pageId="0" pageNumber="1">3A, B</figureCitation>
, 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu xiaoxiao A male palp, prolateral B male palp, retrolateral C epigyne, ventral D vulva, ventral E vulva, dorsal. Abbreviations: CD = copulatory ducts; Co = conductor; CO = copulatory opening; E = embolus; FD = fertilisation ducts; Fe = femur; MA = median apophysis; Pa = patella; Pl = parmula; S = spermathecae; T = tegulum; Ti = tibia; TP = tegular process. Scale bars: 0.10 (A-E)." figureDoi="10.3897/zookeys.1072.67935.figure14" httpUri="https://binary.pensoft.net/fig/612456" pageId="0" pageNumber="1">14A and B</figureCitation>
) or extends and twists at tip of bulb (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Patu catba sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D male palp, prolateral E male palp, retrolateral. Abbreviations: Co = conductor; Cy = cymbium; E = embolus; Fe = femur; MA = median apophysis; Pa = patella; T = tegulum; Ti = tibia; TS = male clasping spines on tibia II. Scale bars: 0.50 (A-C); 0.10 (D, E)." figureDoi="10.3897/zookeys.1072.67935.figure1" httpUri="https://binary.pensoft.net/fig/612443" pageId="0" pageNumber="1">1D, E</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Patu damtao sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D male palp, prolateral E male left palp, retrolateral. Abbreviations: Co = conductor; Cy = cymbium; E = embolus; Pa = patella; T = tegulum; Ti = tibia; TS = male clasping spines on tibia II. Scale bars: 0.50 (A-C); 0.10 (D, E)." figureDoi="10.3897/zookeys.1072.67935.figure4" httpUri="https://binary.pensoft.net/fig/612446" pageId="0" pageNumber="1">4D, E</figureCitation>
, 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Patu jidanweishi A male palp, prolateral B male palp, retrolateral C epigyne, ventral D epigyne, lateral E vulva, ventral F vulva, dorsal. Abbreviations: CD = copulatory ducts; Cy = cymbium; E = embolus; FD = fertilisation ducts; Fe = femur; Pa = patella; S = spermathecae; Sp = scape; T = tegulum; Ti = tibia; TP = tegular process. Scale bars: 0.10 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure7" httpUri="https://binary.pensoft.net/fig/612449" pageId="0" pageNumber="1">7A, B</figureCitation>
, 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Patu nagarat sp. nov. A male palp, prolateral B male palp, retrolateral C epigyne, ventral D epigyne, lateral E vulva, ventral F vulva, dorsal. Abbreviations: CD = copulatory ducts; CO = copulatory opening; Cy = cymbium; E = embolus; FD = fertilisation ducts; Fe = femur; MA = median apophysis; Pa = patella; Pl = parmula; S = spermathecae; T = tegulum; Ti = tibia; TP = tegular process. Scale bars: 0.10 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure9" httpUri="https://binary.pensoft.net/fig/612451" pageId="0" pageNumber="1">9A</figureCitation>
and 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Patu nagarat sp. nov. A male palp, prolateral B male palp, retrolateral C epigyne, ventral D epigyne, lateral E vulva, ventral F vulva, dorsal. Abbreviations: CD = copulatory ducts; CO = copulatory opening; Cy = cymbium; E = embolus; FD = fertilisation ducts; Fe = femur; MA = median apophysis; Pa = patella; Pl = parmula; S = spermathecae; T = tegulum; Ti = tibia; TP = tegular process. Scale bars: 0.10 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure9" httpUri="https://binary.pensoft.net/fig/612451" pageId="0" pageNumber="1">B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Epigyne</emphasis>
(Figs 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Patu jidanweishi A male palp, prolateral B male palp, retrolateral C epigyne, ventral D epigyne, lateral E vulva, ventral F vulva, dorsal. Abbreviations: CD = copulatory ducts; Cy = cymbium; E = embolus; FD = fertilisation ducts; Fe = femur; Pa = patella; S = spermathecae; Sp = scape; T = tegulum; Ti = tibia; TP = tegular process. Scale bars: 0.10 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure7" httpUri="https://binary.pensoft.net/fig/612449" pageId="0" pageNumber="1">7E, F</figureCitation>
, 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Patu nagarat sp. nov. A male palp, prolateral B male palp, retrolateral C epigyne, ventral D epigyne, lateral E vulva, ventral F vulva, dorsal. Abbreviations: CD = copulatory ducts; CO = copulatory opening; Cy = cymbium; E = embolus; FD = fertilisation ducts; Fe = femur; MA = median apophysis; Pa = patella; Pl = parmula; S = spermathecae; T = tegulum; Ti = tibia; TP = tegular process. Scale bars: 0.10 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure9" httpUri="https://binary.pensoft.net/fig/612451" pageId="0" pageNumber="1">9E, F</figureCitation>
, 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu xiaoxiao A male palp, prolateral B male palp, retrolateral C epigyne, ventral D vulva, ventral E vulva, dorsal. Abbreviations: CD = copulatory ducts; Co = conductor; CO = copulatory opening; E = embolus; FD = fertilisation ducts; Fe = femur; MA = median apophysis; Pa = patella; Pl = parmula; S = spermathecae; T = tegulum; Ti = tibia; TP = tegular process. Scale bars: 0.10 (A-E)." figureDoi="10.3897/zookeys.1072.67935.figure14" httpUri="https://binary.pensoft.net/fig/612456" pageId="0" pageNumber="1">14D</figureCitation>
and 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Patu nagarat sp. nov. A male palp, prolateral B male palp, retrolateral C epigyne, ventral D epigyne, lateral E vulva, ventral F vulva, dorsal. Abbreviations: CD = copulatory ducts; CO = copulatory opening; Cy = cymbium; E = embolus; FD = fertilisation ducts; Fe = femur; MA = median apophysis; Pa = patella; Pl = parmula; S = spermathecae; T = tegulum; Ti = tibia; TP = tegular process. Scale bars: 0.10 (A-F)." figureDoi="10.3897/zookeys.1072.67935.figure9" httpUri="https://binary.pensoft.net/fig/612451" pageId="0" pageNumber="1">E</figureCitation>
): weakly sclerotised. Scape or parmula short, tongue-shaped or long, finger-like. Spermathecae long, ovate or kidney-shaped, separated by less than 2 lengths. Copulatory openings separated. Copulatory ducts membranous or faintly sclerotised, usually partially or completely surround spermathecae (exceptions are 
<taxonomicName lsidName="P. nigeri" pageId="0" pageNumber="1" rank="species" species="nigeri">
<emphasis italics="true" pageId="0" pageNumber="1">P. nigeri</emphasis>
</taxonomicName>
Lin &amp; S. Li, 2009 and 
<taxonomicName lsidName="P. qiqi" pageId="0" pageNumber="1" rank="species" species="qiqi">
<emphasis italics="true" pageId="0" pageNumber="1">P. qiqi</emphasis>
</taxonomicName>
). Fertilisation ducts short, thin, typically originate from the lateral or anterior side of spermathecae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="composition">
<paragraph pageId="0" pageNumber="1">Composition.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="S. Li &amp; Lin" authorityYear="2021" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu catba" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="catba">
<emphasis italics="true" pageId="0" pageNumber="1">Patu catba</emphasis>
</taxonomicName>
sp. nov. (♂), 
<taxonomicName lsidName="P. dakou" pageId="0" pageNumber="1" rank="species" species="dakou">
<emphasis italics="true" pageId="0" pageNumber="1">P. dakou</emphasis>
</taxonomicName>
sp. nov. (♂♀), 
<taxonomicName lsidName="P. damtao" pageId="0" pageNumber="1" rank="species" species="damtao">
<emphasis italics="true" pageId="0" pageNumber="1">P. damtao</emphasis>
</taxonomicName>
sp. nov. (♂), 
<taxonomicName lsidName="P. digua" pageId="0" pageNumber="1" rank="species" species="digua">
<emphasis italics="true" pageId="0" pageNumber="1">P. digua</emphasis>
</taxonomicName>
Forster &amp; Platnick, 1977 (♂♀), 
<taxonomicName lsidName="P. eberhardi" pageId="0" pageNumber="1" rank="species" species="eberhardi">
<emphasis italics="true" pageId="0" pageNumber="1">P. eberhardi</emphasis>
</taxonomicName>
Forster &amp; Platnick, 1977 (♂♀), 
<taxonomicName lsidName="P. jiangzhou" pageId="0" pageNumber="1" rank="species" species="jiangzhou">
<emphasis italics="true" pageId="0" pageNumber="1">P. jiangzhou</emphasis>
</taxonomicName>
sp. nov. (♀), 
<taxonomicName lsidName="P. jidanweishi" pageId="0" pageNumber="1" rank="species" species="jidanweishi">
<emphasis italics="true" pageId="0" pageNumber="1">P. jidanweishi</emphasis>
</taxonomicName>
Miller, Griswold &amp; Yin, 2009 (♂♀), 
<taxonomicName lsidName="P. marplesi" pageId="0" pageNumber="1" rank="species" species="marplesi">
<emphasis italics="true" pageId="0" pageNumber="1">P. marplesi</emphasis>
</taxonomicName>
Forster, 1959 (♂), 
<taxonomicName lsidName="P. nagarat" pageId="0" pageNumber="1" rank="species" species="nagarat">
<emphasis italics="true" pageId="0" pageNumber="1">P. nagarat</emphasis>
</taxonomicName>
sp. nov. (♂♀), 
<taxonomicName lsidName="P. nigeri" pageId="0" pageNumber="1" rank="species" species="nigeri">
<emphasis italics="true" pageId="0" pageNumber="1">P. nigeri</emphasis>
</taxonomicName>
(♂♀), 
<taxonomicName lsidName="P. putao" pageId="0" pageNumber="1" rank="species" species="putao">
<emphasis italics="true" pageId="0" pageNumber="1">P. putao</emphasis>
</taxonomicName>
sp. nov. (♀), 
<taxonomicName lsidName="P. qiqi" pageId="0" pageNumber="1" rank="species" species="qiqi">
<emphasis italics="true" pageId="0" pageNumber="1">P. qiqi</emphasis>
</taxonomicName>
Miller, Griswold &amp; Yin, 2009 (♂♀), 
<taxonomicName lsidName="P. saladito" pageId="0" pageNumber="1" rank="species" species="saladito">
<emphasis italics="true" pageId="0" pageNumber="1">P. saladito</emphasis>
</taxonomicName>
Forster &amp; Platnick, 1977 (♀), 
<taxonomicName lsidName="P. samoensis" pageId="0" pageNumber="1" rank="species" species="samoensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. samoensis</emphasis>
</taxonomicName>
Marples, 1951 (♂♀), 
<taxonomicName lsidName="P. silho" pageId="0" pageNumber="1" rank="species" species="silho">
<emphasis italics="true" pageId="0" pageNumber="1">P. silho</emphasis>
</taxonomicName>
Saaristo, 1996 (♂♀), 
<taxonomicName lsidName="P. vitiensis" pageId="0" pageNumber="1" rank="species" species="vitiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. vitiensis</emphasis>
</taxonomicName>
Marples, 1951 (♂♀), 
<taxonomicName lsidName="P. woodwardi" pageId="0" pageNumber="1" rank="species" species="woodwardi">
<emphasis italics="true" pageId="0" pageNumber="1">P. woodwardi</emphasis>
</taxonomicName>
Forster, 1959 (♂♀) and 
<taxonomicName lsidName="P. xiaoxiao" pageId="0" pageNumber="1" rank="species" species="xiaoxiao">
<emphasis italics="true" pageId="0" pageNumber="1">P. xiaoxiao</emphasis>
</taxonomicName>
Miller, Griswold &amp; Yin, 2009 (♂♀). Li, Lin and 
<bibRefCitation DOI="https://doi.org/10.11865/zs.202011" author="Li, S" journalOrPublisher="Zoological Systematics" pageId="0" pageNumber="1" pagination="73 - 77" refId="B8" refString="Li, S, 2020. Spider taxonomy for an advanced China. Zoological Systematics 45 (2): 73 - 77, DOI: https://doi.org/10.11865/zs.202011" title="Spider taxonomy for an advanced China." url="https://doi.org/10.11865/zs.202011" volume="45" year="2020">Li (2020)</bibRefCitation>
have previously suggested that the other two Asian species, 
<taxonomicName lsidName="P. bispina" pageId="0" pageNumber="1" rank="species" species="bispina">
<emphasis italics="true" pageId="0" pageNumber="1">P. bispina</emphasis>
</taxonomicName>
Lin, Pham &amp; S. Li, 2009 from Vietnam and 
<taxonomicName lsidName="P. kishidai" pageId="0" pageNumber="1" rank="species" species="kishidai">
<emphasis italics="true" pageId="0" pageNumber="1">P. kishidai</emphasis>
</taxonomicName>
Shinkai, 2009 from Japan, do not belong to this genus and should be transferred to 
<taxonomicName class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Crassignatha</emphasis>
</taxonomicName>
. Based on the above diagnosis of 
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Patu</emphasis>
</taxonomicName>
, we formally propose two new combinations: 
<taxonomicName class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha bispina" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bispina">
<emphasis italics="true" pageId="0" pageNumber="1">Crassignatha bispina</emphasis>
</taxonomicName>
comb. nov. and 
<taxonomicName lsidName="C. kishidai" pageId="0" pageNumber="1" rank="species" species="kishidai">
<emphasis italics="true" pageId="0" pageNumber="1">C. kishidai</emphasis>
</taxonomicName>
comb. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">China (Guangxi, Hainan, and Yunnan), Colombia, Fiji, Myanmar, New Guinea, Samoa, Seychelles, Thailand, Vietnam.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Of the male 
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Patu</emphasis>
</taxonomicName>
species described here, the embolus is either embedded within the tegulum or not, the conductor is present or absent and the tegular process is present or absent. The similarities of the palps are the nearly ovate bulb and the cymbium lacking any teeth, processes or apophyses. In the females, the epigyne and vulva distinctly differ in the type, shape and size of posterior process of the epigyne (scape or parmula) and in the texture, length and course of the copulatory ducts. The similarities of the vulvae are the ovate or short, club-shaped spermathecae.
</paragraph>
</subSubSection>
</treatment>
</document>