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<mods:title id="E11C8770BF514F25AEE7692CB44D36D2">Brachiopods from Late Jurassic-Early Cretaceous hydrocarbon seep deposits, central Spitsbergen, Svalbard</mods:title>
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<mods:affiliation id="04ECBB9691163E692F30A2D2CDE3B86B">Natural History Museum, University of Oslo, P. O. Box 1172, NO- 0318 Oslo, Norway. &amp; Current address: Institute of Paleobiology, Polish Academy of Sciences, ul. Twarda 51 / 55, 00 - 818 Warszawa, Poland.</mods:affiliation>
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<mods:namePart id="D0A3E260BB2E21283A3FF41108442A33">Little, Crispin T. S.</mods:namePart>
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<treatment id="466F6C336256FFB19685418FFBC83740" ID-DOI="http://doi.org/10.5281/zenodo.4952552" ID-GBIF-Taxon="184851316" ID-Zenodo-Dep="4952552" LSID="urn:lsid:plazi:treatment:466F6C336256FFB19685418FFBC83740" httpUri="http://treatment.plazi.org/id/466F6C336256FFB19685418FFBC83740" lastPageNumber="515" pageId="14" pageNumber="515">
<subSubSection id="86DC8EAE6256FFB19685418FFE8333F5" box="[151,343,924,950]" pageId="14" pageNumber="515" type="nomenclature">
<paragraph id="CE79DD256256FFB19685418FFE8333F5" blockId="14.[151,343,924,983]" box="[151,343,924,950]" pageId="14" pageNumber="515">
<heading id="95316A496256FFB19685418FFE8333F5" bold="true" box="[151,343,924,950]" fontSize="11" level="1" pageId="14" pageNumber="515" reason="1">
<emphasis id="FCB201376256FFB19685418FFE8333F5" bold="true" box="[151,343,924,950]" pageId="14" pageNumber="515">
<taxonomicName id="09C6A6A66256FFB19685418FFECA33F5" authorityName="Middlemiss" authorityYear="1959" box="[151,286,924,950]" class="Rhynchonellata" family="Sellithyrididae" genus="Cyrtothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="515" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376256FFB19685418FFECA33F5" bold="true" box="[151,286,924,950]" italics="true" pageId="14" pageNumber="515">Cyrtothyris</emphasis>
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? sp.
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</heading>
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<subSubSection id="86DC8EAE6256FFB1968541ADFCFA3478" pageId="14" pageNumber="515" type="description">
<paragraph id="CE79DD256256FFB1968541ADFECA3394" blockId="14.[151,343,924,983]" box="[151,286,958,983]" pageId="14" pageNumber="515">
<figureCitation id="56FDC1A06256FFB1968541ADFECA3394" box="[151,286,958,983]" captionStart="FIGURE 7" captionStartId="15.[151,250,1827,1849]" captionTargetBox="[229,1337,195,1772]" captionTargetId="figure-120@15.[228,1338,193,1778]" captionTargetPageId="15" captionText="FIGURE 7. 14. Cyrtothyris aff. cyrta (Walker). PMO 224.895, seep 9, dorsal, ventral, lateral, and anterior views. 58. Cyrtothyris? sp. PMO 224.870, seep 3, dorsal, ventral, lateral, and anterior views. 920. Praelongithyris? aff. borealis Owen. 912. PMO 224.907, seep 1, dorsal, ventral, lateral, and anterior views. 1316. PMO 224.898, seep 9, dorsal, ventral, lateral, and anterior views. Resembles juvenile Cyrtothyris. 1720. PMO 224.886, seep 1, dorsal, ventral, lateral, and anterior views. Form resembling Moutonithyris moutoniana (dOrbigny). 2124. Rouillieria cf. michalkowii (Fahrenkohl). PMO 224.901, seep 9, dorsal, ventral, lateral, and anterior views.Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951778" httpUri="https://zenodo.org/record/4951778/files/figure.png" pageId="14" pageNumber="515">Fig. 7.57.8</figureCitation>
</paragraph>
<paragraph id="CE79DD256256FFB196854615FB64345F" blockId="14.[151,1200,1030,1083]" box="[151,1200,1030,1052]" pageId="14" pageNumber="515">
1976? v. “
<taxonomicName id="09C6A6A66256FFB1970B4615FE58345F" authorityName="Middlemiss" authorityYear="1959" box="[281,396,1030,1052]" class="Rhynchonellata" family="Sellithyrididae" genus="Cyrtothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="515" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376256FFB1970B4615FE58345F" box="[281,396,1030,1052]" italics="true" pageId="14" pageNumber="515">Cyrtothyris</emphasis>
</taxonomicName>
<emphasis id="FCB201376256FFB1978C4614FE33345F" box="[414,487,1031,1052]" italics="true" pageId="14" pageNumber="515">maynci</emphasis>
new species—Owen, p. 13, pl. 2, fig. 1ac, pl. 3, fig. 4ac, text-fig. 7.
</paragraph>
<paragraph id="CE79DD256256FFB196854636FCFA3478" blockId="14.[151,1200,1030,1083]" box="[151,814,1061,1083]" pageId="14" pageNumber="515">
<date id="BA78FBE56256FFB196854636FF233478" box="[151,247,1061,1083]" pageId="14" pageNumber="515" value="2011-05">
2011
<taxonomicName id="09C6A6A66256FFB196F74635FF233478" box="[229,247,1062,1083]" pageId="14" pageNumber="515" rank="variety" variety="partim">v.</taxonomicName>
</date>
partim
<taxonomicName id="09C6A6A66256FFB1975F4636FE143478" authorityName="Middlemiss" authorityYear="1959" box="[333,448,1061,1083]" class="Rhynchonellata" family="Sellithyrididae" genus="Cyrtothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="515" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376256FFB1975F4636FE143478" box="[333,448,1061,1083]" italics="true" pageId="14" pageNumber="515">Cyrtothyris</emphasis>
</taxonomicName>
sp.—Hammer
<emphasis id="FCB201376256FFB194764635FD4C3478" box="[612,664,1061,1083]" italics="true" pageId="14" pageNumber="515">et al.</emphasis>
, p. 20, table 2.
</paragraph>
</subSubSection>
<subSubSection id="86DC8EAE6256FFB196854671FAB73438" box="[151,1379,1122,1147]" pageId="14" pageNumber="515" type="materials_examined">
<paragraph id="CE79DD256256FFB196854671FAB73438" blockId="14.[151,1437,1122,1795]" box="[151,1379,1122,1147]" pageId="14" pageNumber="515">
<emphasis id="FCB201376256FFB196854671FE113438" bold="true" box="[151,453,1122,1147]" inLineHeading="true" pageId="14" pageNumber="515" reason="1">Material and occurrence.</emphasis>
Seep 3 (PMO 224.870, PMO 224.904?, PMO 224.919?), seep 8 (PMO 227.428?).
</paragraph>
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<subSubSection id="86DC8EAE6256FFB196D54695FEF23480" pageId="14" pageNumber="515" type="description">
<paragraph id="CE79DD256256FFB196D54695FEF23480" blockId="14.[151,1437,1122,1795]" pageId="14" pageNumber="515">
<emphasis id="FCB201376256FFB196D54695FE8C34DC" bold="true" box="[199,344,1158,1183]" inLineHeading="true" pageId="14" pageNumber="515" reason="1">Description.</emphasis>
Rounded, circular, to gently elongate outline, equibiconvex lateral profile, rectimarginate anterior commissure.
</paragraph>
</subSubSection>
<subSubSection id="86DC8EAE6256FFB196D546DDFBC83740" pageId="14" pageNumber="515" type="discussion">
<paragraph id="CE79DD256256FFB196D546DDFA9035FC" blockId="14.[151,1437,1122,1795]" pageId="14" pageNumber="515">
<emphasis id="FCB201376256FFB196D546DDFE9834A4" bold="true" box="[199,332,1230,1255]" inLineHeading="true" pageId="14" pageNumber="515" reason="1">Discussion.</emphasis>
The material from Spitsbergen reaches larger maximum dimensions than “
<taxonomicName id="09C6A6A66256FFB192A746DDFAE134A4" authorityName="Middlemiss" authorityYear="1959" box="[1205,1333,1230,1255]" class="Rhynchonellata" family="Sellithyrididae" genus="Cyrtothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="515" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376256FFB192A746DDFAE134A4" box="[1205,1333,1230,1255]" italics="true" pageId="14" pageNumber="515">Cyrtothyris</emphasis>
</taxonomicName>
<emphasis id="FCB201376256FFB1935946DCFA4834A5" box="[1355,1436,1231,1254]" italics="true" pageId="14" pageNumber="515">maynci</emphasis>
described from
<collectingCountry id="B6D19DB56256FFB1975F46E1FE163548" box="[333,450,1266,1291]" name="Greenland" pageId="14" pageNumber="515">Greenland</collectingCountry>
(
<bibRefCitation id="AA57A0D46256FFB197C646E1FD8B3548" author="Owen, E. F." box="[468,607,1266,1291]" pageId="14" pageNumber="515" pagination="1 - 19" refId="ref17259" refString="Owen, E. F. (1976) Some Lower Cretaceous brachiopods from East Greenland. Meddelelser om GrOnland, 171 (3), 1 - 19." type="journal article" year="1976">Owen 1976</bibRefCitation>
). Broken ventral umbones hamper identification (e.g.
<figureCitation id="56FDC1A06256FFB192C746E1FAB53548" box="[1237,1377,1266,1291]" captionStart="FIGURE 7" captionStartId="15.[151,250,1827,1849]" captionTargetBox="[229,1337,195,1772]" captionTargetId="figure-120@15.[228,1338,193,1778]" captionTargetPageId="15" captionText="FIGURE 7. 14. Cyrtothyris aff. cyrta (Walker). PMO 224.895, seep 9, dorsal, ventral, lateral, and anterior views. 58. Cyrtothyris? sp. PMO 224.870, seep 3, dorsal, ventral, lateral, and anterior views. 920. Praelongithyris? aff. borealis Owen. 912. PMO 224.907, seep 1, dorsal, ventral, lateral, and anterior views. 1316. PMO 224.898, seep 9, dorsal, ventral, lateral, and anterior views. Resembles juvenile Cyrtothyris. 1720. PMO 224.886, seep 1, dorsal, ventral, lateral, and anterior views. Form resembling Moutonithyris moutoniana (dOrbigny). 2124. Rouillieria cf. michalkowii (Fahrenkohl). PMO 224.901, seep 9, dorsal, ventral, lateral, and anterior views.Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951778" httpUri="https://zenodo.org/record/4951778/files/figure.png" pageId="14" pageNumber="515">Fig. 7.57.8</figureCitation>
) and referral to
<taxonomicName id="09C6A6A66256FFB197004705FE45356C" authorityName="Middlemiss" authorityYear="1959" box="[274,401,1302,1327]" class="Rhynchonellata" family="Sellithyrididae" genus="Cyrtothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="515" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376256FFB197004705FE45356C" box="[274,401,1302,1327]" italics="true" pageId="14" pageNumber="515">Cyrtothyris</emphasis>
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must be considered tentative. Owen commented that the
<collectingCountry id="B6D19DB56256FFB192384705FB4B356C" box="[1066,1183,1302,1327]" name="Greenland" pageId="14" pageNumber="515">Greenland</collectingCountry>
form has an incipient uniplicate anterior commissure. The Spitsbergen specimens show a rectimarginate anterior commissure to possibly weakly uniplicate (
<figureCitation id="56FDC1A06256FFB19768474DFE0E3534" box="[378,474,1374,1399]" captionStart="FIGURE 7" captionStartId="15.[151,250,1827,1849]" captionTargetBox="[229,1337,195,1772]" captionTargetId="figure-120@15.[228,1338,193,1778]" captionTargetPageId="15" captionText="FIGURE 7. 14. Cyrtothyris aff. cyrta (Walker). PMO 224.895, seep 9, dorsal, ventral, lateral, and anterior views. 58. Cyrtothyris? sp. PMO 224.870, seep 3, dorsal, ventral, lateral, and anterior views. 920. Praelongithyris? aff. borealis Owen. 912. PMO 224.907, seep 1, dorsal, ventral, lateral, and anterior views. 1316. PMO 224.898, seep 9, dorsal, ventral, lateral, and anterior views. Resembles juvenile Cyrtothyris. 1720. PMO 224.886, seep 1, dorsal, ventral, lateral, and anterior views. Form resembling Moutonithyris moutoniana (dOrbigny). 2124. Rouillieria cf. michalkowii (Fahrenkohl). PMO 224.901, seep 9, dorsal, ventral, lateral, and anterior views.Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951778" httpUri="https://zenodo.org/record/4951778/files/figure.png" pageId="14" pageNumber="515">Fig. 7.8</figureCitation>
, anterior commissure damaged). The
<collectingCountry id="B6D19DB56256FFB195B0474DFBC33534" box="[930,1047,1374,1399]" name="Greenland" pageId="14" pageNumber="515">Greenland</collectingCountry>
material is from the Falskebugt Member (
<bibRefCitation id="AA57A0D46256FFB197174791FE0C35D8" author="Harper, D. A. T. &amp; Alsen, P. &amp; Owen, E. F. &amp; Sandy, M. R." box="[261,472,1410,1435]" pageId="14" pageNumber="515" pagination="213 - 225" refId="ref14982" refString="Harper, D. A. T., Alsen, P., Owen, E. F. &amp; Sandy, M. R. (2005) Early Cretaceous brachiopods from North-East Greenland: biofacies and biogeography. Bulletin of the Geological Society of Denmark, 52, 213 - 225." type="journal article" year="2005">
Harper
<emphasis id="FCB201376256FFB1974F4790FE4335D8" box="[349,407,1410,1435]" italics="true" pageId="14" pageNumber="515">et al.</emphasis>
2005
</bibRefCitation>
) of Valanginian age whereas the Spitsbergen specimens are from the Upper Volgian.
<taxonomicName id="09C6A6A66256FFB1968547B5FEC335FC" authorityName="Middlemiss" authorityYear="1959" box="[151,279,1446,1471]" class="Rhynchonellata" family="Sellithyrididae" genus="Cyrtothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="515" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376256FFB1968547B5FEC335FC" box="[151,279,1446,1471]" italics="true" pageId="14" pageNumber="515">Cyrtothyris</emphasis>
</taxonomicName>
has not been recorded from the Upper Jurassic and these specimens may represent a new form.
</paragraph>
<paragraph id="CE79DD256256FFB196D547D9FE50369C" blockId="14.[151,1437,1122,1795]" pageId="14" pageNumber="515">
Some of the Spitsbergen brachiopods are morphologically close to
<taxonomicName id="09C6A6A66256FFB195E647D9FBA735A0" authorityName="Middlemiss" authorityYear="1959" box="[1012,1139,1482,1507]" class="Rhynchonellata" family="Sellithyrididae" genus="Cyrtothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="515" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376256FFB195E647D9FBA735A0" box="[1012,1139,1482,1507]" italics="true" pageId="14" pageNumber="515">Cyrtothyris</emphasis>
</taxonomicName>
and
<taxonomicName id="09C6A6A66256FFB192AA47D9FAE535A0" authorityName="Makridin. However" authorityYear="1960" box="[1208,1329,1482,1507]" class="Rhynchonellata" family="Terebratulidae" genus="Rouillieria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="515" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376256FFB192AA47D9FAE535A0" box="[1208,1329,1482,1507]" italics="true" pageId="14" pageNumber="515">Rouillieria</emphasis>
</taxonomicName>
. Species referred to these two genera may be morphologically very similar and details of the ventral umbo and profile can be important in distinguishing between them (as discussed by
<bibRefCitation id="AA57A0D46256FFB195244401FC2A3668" author="Middlemiss, F. A." box="[822,1022,1554,1579]" pageId="14" pageNumber="515" pagination="21 - 104" refId="ref16391" refString="Middlemiss, F. A. (1976) Lower Cretaceous Terebratulidina of northern England and Germany and their geological background. Geologisches Jahrbuch, 30, 21 - 104." type="journal article" year="1976">Middlemiss 1976</bibRefCitation>
). Several specimens from seep 9 are now considered to be
<taxonomicName id="09C6A6A66256FFB1979F4425FD39360C" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[397,749,1590,1615]" class="Rhynchonellata" family="Loboidothyrididae" genus="Seductorithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="515" phylum="Brachiopoda" rank="species" species="septemtrionalis">
<emphasis id="FCB201376256FFB1979F4425FD39360C" box="[397,749,1590,1615]" italics="true" pageId="14" pageNumber="515">Seductorithyris septemtrionalis</emphasis>
</taxonomicName>
; these have an erect beak and lack the produced ventral beak of
<taxonomicName id="09C6A6A66256FFB196A54449FEE23630" authorityName="Middlemiss" authorityYear="1959" box="[183,310,1626,1651]" class="Rhynchonellata" family="Sellithyrididae" genus="Cyrtothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="515" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376256FFB196A54449FEE23630" box="[183,310,1626,1651]" italics="true" pageId="14" pageNumber="515">Cyrtothyris</emphasis>
</taxonomicName>
, and are typically less inflated in profile than the latter. However,
<specimenCount id="D8C016AC6256FFB192204448FB193630" box="[1074,1229,1626,1651]" pageId="14" pageNumber="515" type="generic">one specimen</specimenCount>
from seep 9 (
<figureCitation id="56FDC1A06256FFB1937D4449FF3A36D4" captionStart="FIGURE 7" captionStartId="15.[151,250,1827,1849]" captionTargetBox="[229,1337,195,1772]" captionTargetId="figure-120@15.[228,1338,193,1778]" captionTargetPageId="15" captionText="FIGURE 7. 14. Cyrtothyris aff. cyrta (Walker). PMO 224.895, seep 9, dorsal, ventral, lateral, and anterior views. 58. Cyrtothyris? sp. PMO 224.870, seep 3, dorsal, ventral, lateral, and anterior views. 920. Praelongithyris? aff. borealis Owen. 912. PMO 224.907, seep 1, dorsal, ventral, lateral, and anterior views. 1316. PMO 224.898, seep 9, dorsal, ventral, lateral, and anterior views. Resembles juvenile Cyrtothyris. 1720. PMO 224.886, seep 1, dorsal, ventral, lateral, and anterior views. Form resembling Moutonithyris moutoniana (dOrbigny). 2124. Rouillieria cf. michalkowii (Fahrenkohl). PMO 224.901, seep 9, dorsal, ventral, lateral, and anterior views.Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951778" httpUri="https://zenodo.org/record/4951778/files/figure.png" pageId="14" pageNumber="515">Fig. 7.17.4</figureCitation>
; PMO 224.895) with a suberect ventral umbo and incipiently uniplicate anterior commissure, is referred to
<taxonomicName id="09C6A6A66256FFB1968544B1FE2E36F8" authority="(Walker)" baseAuthorityName="Walker" baseAuthorityYear="1868" box="[151,506,1698,1723]" class="Rhynchonellata" family="Sellithyrididae" genus="Cyrtothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="515" phylum="Brachiopoda" rank="species" species="cyrta">
<emphasis id="FCB201376256FFB1968544B1FEC336F8" box="[151,279,1698,1723]" italics="true" pageId="14" pageNumber="515">Cyrtothyris</emphasis>
aff.
<emphasis id="FCB201376256FFB1975D44B0FE5D36F9" box="[335,393,1699,1722]" italics="true" pageId="14" pageNumber="515">cyrta</emphasis>
(Walker)
</taxonomicName>
.
<bibRefCitation id="AA57A0D46256FFB1941444B1FD0B36F8" author="Middlemiss, F. A." box="[518,735,1698,1723]" pageId="14" pageNumber="515" pagination="21 - 104" refId="ref16391" refString="Middlemiss, F. A. (1976) Lower Cretaceous Terebratulidina of northern England and Germany and their geological background. Geologisches Jahrbuch, 30, 21 - 104." type="journal article" year="1976">Middlemiss (1976)</bibRefCitation>
considers that this species has a rectimarginate to uniplicate anterior commissure.
</paragraph>
<paragraph id="CE79DD256256FFB196D544F9FBC83740" blockId="14.[151,1437,1122,1795]" box="[199,1052,1770,1795]" pageId="14" pageNumber="515">
<emphasis id="FCB201376256FFB196D544F9FD153740" bold="true" box="[199,705,1770,1795]" inLineHeading="true" pageId="14" pageNumber="515" reason="1">Stratigraphic and geographic distribution.</emphasis>
Upper Volgian of Spitsbergen.
</paragraph>
</subSubSection>
</treatment>
</document>