treatments-xml/data/46/6F/6C/466F6C336251FFB396854766FBEE3486.xml
2024-06-21 12:35:10 +02:00

568 lines
90 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="CB762C9FF335F425E6D661EAC7042465" ID-CLB-Dataset="4889" ID-DOI="10.11646/zootaxa.3884.6.1" ID-GBIF-Dataset="d69a4a4f-fbba-436e-b053-4a771735b9a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="4951742" ID-ZooBank="BD4F285D-358C-4350-88EA-7FA94513D930" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623727472316" checkinUser="felipe" docAuthor="Sandy, Michael R., Hryniewicz, Krzysztof, Hammer, Øyvind, Nakrem, Hans Arne &amp; Little, Crispin T. S." docDate="2014" docId="466F6C336251FFB396854766FBEE3486" docLanguage="en" docName="zt03884p532.pdf" docOrigin="Zootaxa 3884 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptilorhynchia mclachlani Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="513" masterDocId="BA56144B6258FFBF96124213FFD43043" masterDocTitle="Brachiopods from Late Jurassic-Early Cretaceous hydrocarbon seep deposits, central Spitsbergen, Svalbard" masterLastPageNumber="532" masterPageNumber="501" pageNumber="510" updateTime="1699016241816" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="53D0E0D7DEB8E7CEFD2801CDA7ABA7BE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="946598A8E6761416515DA4A239A305E0">
<mods:title id="1C4909E4656C0727CB1B74A7321A94FE">Brachiopods from Late Jurassic-Early Cretaceous hydrocarbon seep deposits, central Spitsbergen, Svalbard</mods:title>
</mods:titleInfo>
<mods:name id="365ABFE9F8F13BA738F165D3BED770B6" type="personal">
<mods:role id="C1701C793231D3D534A6F0EB1F2CD0EB">
<mods:roleTerm id="B924FA811CB38E1603EA4B99D19D713C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C79EA1A34E87346A8CA4B6817A6C91DB">Sandy, Michael R.</mods:namePart>
<mods:affiliation id="004D0F826FFAA8B96F2FF4733498BEA6">Department of Geology, University of Dayton, Dayton, OH 45469 - 2364, USA. E-mail: msandy 1 @ udayton. edu</mods:affiliation>
</mods:name>
<mods:name id="52827A0549E857F07C428007F3F975C6" type="personal">
<mods:role id="5A3F542773044E51076E9F67FC7BFB34">
<mods:roleTerm id="F4C50D01A5AF5F800A4C080737AA22DF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="65A33725A140D522A7D25E865F179F68">Hryniewicz, Krzysztof</mods:namePart>
<mods:affiliation id="E85DC4793AC9DF1F627A810F873E6FA0">Natural History Museum, University of Oslo, P. O. Box 1172, NO- 0318 Oslo, Norway. &amp; Current address: Institute of Paleobiology, Polish Academy of Sciences, ul. Twarda 51 / 55, 00 - 818 Warszawa, Poland.</mods:affiliation>
</mods:name>
<mods:name id="41EA8924EAC890D7431E34D793F7593C" type="personal">
<mods:role id="4496463DB6418EE141B181AB84A1FB6E">
<mods:roleTerm id="40151358CB39B1217C4CC09E538AE85E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="06CD1C148998C54B2C2D2544594560AD">Hammer, Øyvind</mods:namePart>
<mods:affiliation id="88A44D47FAD44E1FD3935316C1EDC072">Natural History Museum, University of Oslo, P. O. Box 1172, NO- 0318 Oslo, Norway.</mods:affiliation>
</mods:name>
<mods:name id="02DEBB75EBFC93FAD2AC62E4A793696A" type="personal">
<mods:role id="6BC942A7F6FE9B856CEA967DD242FA4F">
<mods:roleTerm id="10511DD5C9479F954B84DADCF36AEF79">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="50BEFA6BDF0B7B93B4F623956F965D4E">Nakrem, Hans Arne</mods:namePart>
<mods:affiliation id="09265AAABBDB0BF904B49C4F6EB65124">Natural History Museum, University of Oslo, P. O. Box 1172, NO- 0318 Oslo, Norway.</mods:affiliation>
</mods:name>
<mods:name id="795DEFEB6262EABC6979A77BF86A8992" type="personal">
<mods:role id="6F6DF4006E5115AD93E1A3A15719A357">
<mods:roleTerm id="0EE56F37BCF5FD28CA277AEBD9BE8C57">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CE7FC6C201F231A0C94C12CC51AB0A4F">Little, Crispin T. S.</mods:namePart>
<mods:affiliation id="2CF2E589FF0B66E5F6F7CBE656BF6746">School of Earth and Environment, University of Leeds, Leeds LS 2 9 JT, United Kingdom.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="2CF3CF9D5715CC1FAAC1649571C8E1D5">text</mods:typeOfResource>
<mods:relatedItem id="DFAE76DE1D60C7ECC0E7BAB8CA0153CF" type="host">
<mods:titleInfo id="244A35895442001A3B6C1BA7D7F99DF1">
<mods:title id="4A3369B4869334E272340B67ED4CCB82">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="73484AFB8931DFA735B0AE9315EE8D2C">
<mods:date id="6A2039E53AF3B59B53054C85F5049E6D">2014</mods:date>
<mods:detail id="200576B3CA9926CB1BE9B1B4C488E553" type="pubDate">
<mods:number id="2D6363B1A398B26CEF8794618C20A084">2014-11-19</mods:number>
</mods:detail>
<mods:detail id="7EB33535A3D9718445297FF3F8FFC3B6" type="volume">
<mods:number id="CF3A37E1A5CE8930648635A7FDB1B707">3884</mods:number>
</mods:detail>
<mods:detail id="B214A3DFBB6A4E1FCAE2E376861709F8" type="issue">
<mods:number id="B1634A18AFDDC322376F0C55190A4EEA">6</mods:number>
</mods:detail>
<mods:extent id="7AA746AE8733C4C87C897F2A3667405A" unit="page">
<mods:start id="26834A3DBC1EA4737ECE44229D6D854A">501</mods:start>
<mods:end id="6989F998EDA42D643E29FA3D6CBBE68C">532</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E2C05A1587650E9F603FB9413A578ADC">journal article</mods:classification>
<mods:identifier id="CC083E99B0B06CAC76F02E6DD782DC03" type="CLB-Dataset">4889</mods:identifier>
<mods:identifier id="078F2FF877BDEE9128CD548A71570B69" type="DOI">10.11646/zootaxa.3884.6.1</mods:identifier>
<mods:identifier id="E4EF5958BF92F19274ABDA4A2457C8D5" type="GBIF-Dataset">d69a4a4f-fbba-436e-b053-4a771735b9a0</mods:identifier>
<mods:identifier id="FFB997ACF084AF8684CC0408E683A245" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="487CE978CC690BE0E9359852E098B011" type="Zenodo-Dep">4951742</mods:identifier>
<mods:identifier id="A08D226910C86BAE9C2530E6EC7CA4D0" type="ZooBank">BD4F285D-358C-4350-88EA-7FA94513D930</mods:identifier>
</mods:mods>
<treatment id="466F6C336251FFB396854766FBEE3486" ID-DOI="http://doi.org/10.5281/zenodo.5236100" ID-GBIF-Taxon="184851313" ID-Zenodo-Dep="5236100" LSID="urn:lsid:plazi:treatment:466F6C336251FFB396854766FBEE3486" httpUri="http://treatment.plazi.org/id/466F6C336251FFB396854766FBEE3486" lastPageId="12" lastPageNumber="513" pageId="9" pageNumber="510">
<subSubSection id="86DC8EAE6251FFB696854766FDE435CC" box="[151,560,1397,1423]" pageId="9" pageNumber="510" type="nomenclature">
<paragraph id="CE79DD256251FFB696854766FDE435CC" blockId="9.[151,560,1397,1456]" box="[151,560,1397,1423]" pageId="9" pageNumber="510">
<heading id="95316A496251FFB696854766FDE435CC" bold="true" box="[151,560,1397,1423]" fontSize="11" level="1" pageId="9" pageNumber="510" reason="1">
<emphasis id="FCB201376251FFB696854766FDE435CC" bold="true" box="[151,560,1397,1423]" pageId="9" pageNumber="510">
<taxonomicName id="09C6A6A66251FFB696854766FE1F35CC" authority="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little, 2014" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[151,459,1397,1423]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="species" species="mclachlani" status="sp. nov.">
<emphasis id="FCB201376251FFB696854766FE1F35CC" bold="true" box="[151,459,1397,1423]" italics="true" pageId="9" pageNumber="510">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E781BC4C6251FFB697C14766FDE435CC" box="[467,560,1397,1423]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86DC8EAE6251FFB696854784FC973657" pageId="9" pageNumber="510" type="description">
<paragraph id="CE79DD256251FFB696854784FEB535F3" blockId="9.[151,560,1397,1456]" box="[151,353,1431,1456]" pageId="9" pageNumber="510">
<figureCitation id="56FDC1A06251FFB696854784FE9C35F3" box="[151,328,1431,1456]" captionStart="FIGURE 3" captionStartId="7.[151,250,1756,1778]" captionTargetBox="[261,1312,207,1726]" captionTargetId="figure-152@7.[260,1316,193,1726]" captionTargetPageId="7" captionText="FIGURE 3. 112. Pseudomonticlarella varia Smirnova. 14. PMO 227.416, seep 13, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 4). 58. PMO 227.419, seep 7, dorsal, ventral, lateral, and anterior views. 912. PMO 227.418, seep 13, dorsal, ventral, lateral, and anterior views. 1332. Ptilorhynchia mclachlani sp. nov. 1316. PMO 224.876, seep 9, dorsal, ventral, lateral, and anterior views. 1720, PMO 224.879, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5B). 2124. PMO 227.431, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5A). 2528. PMO 224.871, seep 9, dorsal, ventral, lateral, and anterior views. 2932. Holotype, PMO 224.882, seep 9, dorsal, ventral, lateral, and anterior views. Dotted line delimits photographs at the same magnification. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951754" httpUri="https://zenodo.org/record/4951754/files/figure.png" pageId="9" pageNumber="510">Figs. 3.133.32</figureCitation>
,
<figureCitation id="56FDC1A06251FFB697414784FEB535F3" box="[339,353,1431,1456]" captionStart="FIGURE 5" captionStartId="11.[151,250,1820,1842]" captionTargetBox="[238,1340,193,1782]" captionTargetId="figure-142@11.[238,1340,193,1798]" captionTargetPageId="11" captionText="FIGURE 5. A and B, transverse serial sections from two specimens of Ptilorhynchia mclachlani sp. nov., seep 9. A) Specimen PMO 227.431, sections above third dotted line. Dotted lines delimit groups of sections drawn at the same magnification. Dimensions of specimen: L 15.3+; W 18.3; T 9.8 mm. Serial sections prepared perpendicular to maximum length of specimen; numbering of sections starts at 0.0 mm, the initial section through the specimen. B) Specimen PMO 224.879, sections below third dotted line. Dimensions of sectioned specimen: L 12.9; W 12.8+; T 9.0 mm. Serial sections prepared perpendicular to length of dorsal valve; numbering of sections starts at 0.0 mm, the initial section through the specimen. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951764" httpUri="https://zenodo.org/record/4951764/files/figure.png" pageId="9" pageNumber="510">5</figureCitation>
</paragraph>
<paragraph id="CE79DD256251FFB6968547F3FC5335B6" blockId="9.[151,903,1504,1556]" box="[151,903,1504,1526]" pageId="9" pageNumber="510">
<date id="BA78FBE56251FFB6968547F3FF2135B6" box="[151,245,1504,1526]" pageId="9" pageNumber="510" value="2011-05">
2011
<taxonomicName id="09C6A6A66251FFB696F147F3FF2135B6" box="[227,245,1504,1525]" pageId="9" pageNumber="510" rank="variety" variety="partim">v.</taxonomicName>
</date>
partim
<taxonomicName id="09C6A6A66251FFB6975947F3FE1335B5" authorityName="Wisniewska" authorityYear="1932" box="[331,455,1504,1526]" class="Rhynchonellata" family="Basiliolidae" genus="Lacunosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376251FFB6975947F3FE1335B5" box="[331,455,1504,1526]" italics="true" pageId="9" pageNumber="510">Lacunosella</emphasis>
</taxonomicName>
sp.—Hammer
<emphasis id="FCB201376251FFB6947947F2FD4A35B5" box="[619,670,1504,1526]" italics="true" pageId="9" pageNumber="510">et al.</emphasis>
, p. 21, table 2, fig. 7R.
</paragraph>
<paragraph id="CE79DD256251FFB6968547EDFC973657" blockId="9.[151,903,1504,1556]" box="[151,835,1534,1556]" pageId="9" pageNumber="510">
<date id="BA78FBE56251FFB6968547EDFF213657" box="[151,245,1534,1556]" pageId="9" pageNumber="510" value="2011-05">
2011
<taxonomicName id="09C6A6A66251FFB696F147ECFF213657" box="[227,245,1535,1556]" pageId="9" pageNumber="510" rank="variety" variety="partim">v.</taxonomicName>
</date>
partim
<taxonomicName id="09C6A6A66251FFB6975947EDFE013657" authorityName="Wisniewska" authorityYear="1932" box="[331,469,1534,1556]" class="Rhynchonellata" family="Norellidae" genus="Monticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376251FFB6975947EDFE013657" box="[331,469,1534,1556]" italics="true" pageId="9" pageNumber="510">Monticlarella</emphasis>
</taxonomicName>
sp.—Hammer
<emphasis id="FCB201376251FFB6946B47ECFD793657" box="[633,685,1534,1556]" italics="true" pageId="9" pageNumber="510">et al.</emphasis>
, p. 21, table 2.
</paragraph>
</subSubSection>
<subSubSection id="86DC8EAE6251FFB696854452FBBE3685" pageId="9" pageNumber="510" type="materials_examined">
<paragraph id="CE79DD256251FFB696854452FDA136E1" blockId="9.[151,1437,1601,2022]" pageId="9" pageNumber="510">
<emphasis id="FCB201376251FFB696854452FE103619" bold="true" box="[151,452,1601,1626]" inLineHeading="true" pageId="9" pageNumber="510" reason="1">Material and occurrence.</emphasis>
<typeStatus id="117D63876251FFB697D84452FDE23619" box="[458,566,1601,1626]" pageId="9" pageNumber="510" type="holotype">Holotype</typeStatus>
: PMO 224.882;
<typeStatus id="117D63876251FFB694FF4451FC893619" box="[749,861,1602,1626]" pageId="9" pageNumber="510" type="paratype">paratypes</typeStatus>
: PMO 227.431, PMO 227.432, PMO 224.871873, PMO 224.875, PMO 224.883, PMO 217.199, PMO 224.914 (crushed). Juvenile specimens: PMO 224.876879, PMO 227.433. All specimens from seep 9.
</paragraph>
<paragraph id="CE79DD256251FFB696D544BEFBBE3685" blockId="9.[151,1437,1601,2022]" box="[199,1130,1709,1734]" pageId="9" pageNumber="510">
<materialsCitation id="7EAED7786251FFB696D544BEFBB33685" ID-GBIF-Occurrence="3337015301" box="[199,1127,1709,1734]" latitude="78.313866" location="Knorringfjellet" longLatPrecision="1" longitude="16.183027" municipality="central Spitsbergen" pageId="9" pageNumber="510" specimenCount="1" typeStatus="holotype">
<emphasis id="FCB201376251FFB696D544BEFEB13685" bold="true" box="[199,357,1709,1734]" inLineHeading="true" pageId="9" pageNumber="510" reason="1">
<typeStatus id="117D63876251FFB696D544BEFED63685" box="[199,258,1709,1734]" pageId="9" pageNumber="510">Type</typeStatus>
locality.
</emphasis>
<location id="CB198BFE6251FFB6977E44BEFDC33685" LSID="urn:lsid:plazi:treatment:466F6C336251FFB396854766FBEE3486:CB198BFE6251FFB6977E44BEFDC33685" box="[364,535,1709,1734]" latitude="78.313866" longLatPrecision="1" longitude="16.183027" municipality="central Spitsbergen" name="Knorringfjellet" pageId="9" pageNumber="510">Knorringfjellet</location>
,
<collectingMunicipality id="2E1D475F6251FFB6943044BEFD283685" box="[546,764,1709,1734]" pageId="9" pageNumber="510">central Spitsbergen</collectingMunicipality>
,
<geoCoordinate id="ABF2BBE26251FFB6951544BEFC673685" box="[775,947,1709,1734]" degrees="78" direction="north" minutes="18" orientation="latitude" pageId="9" pageNumber="510" precision="1" seconds="49.9" value="78.313866">N78° 18 49.9”</geoCoordinate>
<geoCoordinate id="ABF2BBE26251FFB695A844BEFBB33685" box="[954,1127,1709,1734]" degrees="16" direction="east" minutes="10" orientation="longitude" pageId="9" pageNumber="510" precision="1" seconds="58.9" value="16.183027">E16° 10 58.9”</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86DC8EAE6251FFB696D544C2FD85374D" pageId="9" pageNumber="510" type="etymology">
<paragraph id="CE79DD256251FFB696D544C2FD85374D" blockId="9.[151,1437,1601,2022]" pageId="9" pageNumber="510">
<emphasis id="FCB201376251FFB696D544C2FE9A36A9" bold="true" box="[199,334,1745,1770]" inLineHeading="true" pageId="9" pageNumber="510" reason="1">Etymology.</emphasis>
For the late Mr. Archie McLachlan, Department of Geology, Queen Mary College, University of London (see
<bibRefCitation id="AA57A0D46251FFB6973B44E6FE3B374D" author="Middlemiss, F. A." box="[297,495,1781,1806]" pageId="9" pageNumber="510" pagination="165 - 170" refId="ref16482" refString="Middlemiss, F. A. (1997) A brief history of Geology at Queen Mary. Proceedings of the Geologists' Association, 108, 165 - 170. http: // dx. doi. org / 10.1016 / S 0016 - 7878 (97) 80023 - 5" type="journal article" year="1997">Middlemiss 1997</bibRefCitation>
, p. 169).
</paragraph>
</subSubSection>
<subSubSection id="86DC8EAE6251FFB696D5450AFAF83772" box="[199,1324,1817,1842]" pageId="9" pageNumber="510" type="description">
<paragraph id="CE79DD256251FFB696D5450AFAF83772" blockId="9.[151,1437,1601,2022]" box="[199,1324,1817,1842]" pageId="9" pageNumber="510">
<emphasis id="FCB201376251FFB696D5450AFDC53771" bold="true" box="[199,529,1817,1842]" inLineHeading="true" pageId="9" pageNumber="510" reason="1">
Dimensions of the
<typeStatus id="117D63876251FFB697B1450AFDD83771" box="[419,524,1817,1842]" pageId="9" pageNumber="510" type="holotype">holotype</typeStatus>
.
</emphasis>
PMO 224.882—Length
<quantity id="093E70C06251FFB6953A450AFC5A3772" box="[808,910,1817,1842]" metricMagnitude="-2" metricUnit="m" metricValue="1.71" pageId="9" pageNumber="510" unit="mm" value="17.1">17.1 mm</quantity>
, width
<quantity id="093E70C06251FFB695F3450AFB933772" box="[993,1095,1817,1842]" metricMagnitude="-2" metricUnit="m" metricValue="1.89" pageId="9" pageNumber="510" unit="mm" value="18.9">18.9 mm</quantity>
, thickness
<quantity id="093E70C06251FFB692D0450AFAFC3772" box="[1218,1320,1817,1842]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="9" pageNumber="510" unit="mm" value="11.6">11.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="86DC8EAE6251FFB596D5452EFED43096" lastPageId="10" lastPageNumber="511" pageId="9" pageNumber="510" type="diagnosis">
<paragraph id="CE79DD256251FFB596D5452EFED43096" blockId="9.[151,1437,1601,2022]" lastBlockId="10.[151,1437,151,2013]" lastPageId="10" lastPageNumber="511" pageId="9" pageNumber="510">
<emphasis id="FCB201376251FFB696D5452EFE953715" bold="true" box="[199,321,1853,1878]" inLineHeading="true" pageId="9" pageNumber="510" reason="1">Diagnosis.</emphasis>
Subtriangular in outline, generally evenly biconvex in profile. Valve surface smooth except for ornament of costae which may be traceable over most of the length of the valves but are most strongly developed at the anterior commissure. Number of costae on valves variable, with typically 34 costae in the sulcus of the dorsal valve and 45 on the corresponding fold of the ventral valve. Dental lamellae present in ventral valve. Welldeveloped septalium in dorsal valve, outer hinge plates horizontal from which inner socket ridges are deflected. Crura develop at top of Y-shape of septalium, where inner and outer hinge plates meet. Anteriorly crura arch ventrally.
</paragraph>
</subSubSection>
<subSubSection id="86DC8EAE6252FFB396D542CCFBEE3486" lastPageId="12" lastPageNumber="513" pageId="10" pageNumber="511" type="discussion">
<paragraph id="CE79DD256252FFB596D542CCFDD73126" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="511">
<emphasis id="FCB201376252FFB596D542CCFE9830BB" bold="true" box="[199,332,223,248]" inLineHeading="true" pageId="10" pageNumber="511" reason="1">Discussion.</emphasis>
The beak of the ventral valve is frequently damaged/incomplete in the specimens. As a result, the dental lamellae are usually clearly visible in the umbo of the ventral valve. Juvenile specimens have costae developed over both valves, some larger specimens have costae developed primarily in the sulcus of the ventral valve and at the anterior margin.
</paragraph>
<paragraph id="CE79DD256252FFB596D5437CFA993193" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="511">
The material from Spitsbergen shows variability in outline and number of costae. The costae are traceable over most of the length of the valves.
<bibRefCitation id="AA57A0D46252FFB5943A4387FD1631EE" author="Owen, E. F." box="[552,706,404,429]" pageId="10" pageNumber="511" pagination="1 - 8" refId="ref17162" refString="Owen, E. F. (1972) A rhynchonelloid brachiopod from the Upper Jurassic rocks of British Columbia. Geological Survey of Canada Paper, 72 - 26, 1 - 8." type="journal article" year="1972">Owen (1972)</bibRefCitation>
described a variable species that he referred to
<taxonomicName id="09C6A6A66252FFB593104387FF273193" authorityName="Owen, USGS Mesozoic" authorityYear="1972" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="jeletzkyi">
<emphasis id="FCB201376252FFB593104387FF273193" italics="true" pageId="10" pageNumber="511">Ptilorhynchia jeletzkyi</emphasis>
</taxonomicName>
from the latest Jurassic of
<collectingRegion id="0C0213C76252FFB5943243A4FD363193" box="[544,738,439,464]" country="Canada" name="British Columbia" pageId="10" pageNumber="511">British Columbia</collectingRegion>
which is very similar to the material from Spitsbergen.
</paragraph>
<paragraph id="CE79DD256252FFB596D543CFFDD9337B" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="511">
<bibRefCitation id="AA57A0D46252FFB596D543CFFE8C31B6" author="Owen, E. F." box="[199,344,476,501]" pageId="10" pageNumber="511" pagination="1 - 8" refId="ref17162" refString="Owen, E. F. (1972) A rhynchonelloid brachiopod from the Upper Jurassic rocks of British Columbia. Geological Survey of Canada Paper, 72 - 26, 1 - 8." type="journal article" year="1972">Owen (1972</bibRefCitation>
, p. 2) comments on
<taxonomicName id="09C6A6A66252FFB5945A43CFFC9331B6" authorityName="Owen, USGS Mesozoic" authorityYear="1972" box="[584,839,476,501]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="jeletzkyi">
<emphasis id="FCB201376252FFB5945A43CFFC9331B6" box="[584,839,476,501]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia jeletzkyi</emphasis>
</taxonomicName>
“The shell surface is smooth, exhibiting no sign of striae, but often developing strong marginal plicae which originate midway between the umbones and anterior commissure. These plicae number from one to four...”. However, these strong plicae are not obvious in figured specimens (
<bibRefCitation id="AA57A0D46252FFB5970E4054FE713223" author="Owen, E. F." box="[284,421,583,608]" pageId="10" pageNumber="511" pagination="1 - 8" refId="ref17162" refString="Owen, E. F. (1972) A rhynchonelloid brachiopod from the Upper Jurassic rocks of British Columbia. Geological Survey of Canada Paper, 72 - 26, 1 - 8." type="journal article" year="1972">Owen 1972</bibRefCitation>
, pl. 1, figs. 14) or a specimen referred herein to
<taxonomicName id="09C6A6A66252FFB595F64054FB363223" authorityName="Owen, USGS Mesozoic" authorityYear="1972" box="[996,1250,583,608]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="jeletzkyi">
<emphasis id="FCB201376252FFB595F64054FB363223" box="[996,1250,583,608]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia jeletzkyi</emphasis>
</taxonomicName>
from the Upper Jurassic (KimmeridgianTithonian?) of
<collectingRegion id="0C0213C76252FFB59472407FFD7A32C6" box="[608,686,620,645]" country="United States of America" name="Alaska" pageId="10" pageNumber="511">Alaska</collectingRegion>
(
<figureCitation id="56FDC1A06252FFB594AD407FFC9B32C6" box="[703,847,620,645]" captionStart="FIGURE 6" captionStartId="13.[151,250,1796,1818]" captionTargetBox="[152,1430,194,1774]" captionTargetId="figure-113@13.[150,1437,193,1774]" captionTargetPageId="13" captionText="FIGURE 6. 14. Ptilorhynchia aff. plumasensis Crickmay, USGS Mesozoic Locality 32032 (=80Ace119), Karluk Quadrangle, Alaska, Shelikof Formation, Callovian, Middle Jurassic, dorsal, ventral, lateral, and anterior views. 58. Ptilorhynchia jeletzkyi Owen, USGS Mesozoic locality 14496, cooccurs with Buchia mosquensis (von Buch), probably Naknek series, KimmeridgianTithonian(?), Upper Jurassic, McCarthy Quadrangle, Alaska. Collector, Fred H. Moffit, 1928. Dorsal, ventral, lateral, and anterior views.917. Ptilorhynchia obscuricostata Dagys. 912. PMO 224.880, seep 9, dorsal, ventral, lateral, and anterior views. 1317. PMO 224.881, seep 9, dorsal, ventral, lateral, anterior and posterior views. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951772" httpUri="https://zenodo.org/record/4951772/files/figure.png" pageId="10" pageNumber="511">Fig. 6.56.8</figureCitation>
). This specimen is slightly damaged at the anterior commissure and the lateral margin, but does show the generally smooth shell ornament and the development of marginal costae (3 costae on the dorsal fold,
<quantity id="093E70C06252FFB5948C40A7FD1F328E" box="[670,715,692,717]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="511" unit="in" value="2.0">2 in</quantity>
the ventral sulcus). The number of costae in the sulcus (at the anterior commissure) in
<taxonomicName id="09C6A6A66252FFB597BE40C4FD7D32B3" authorityName="Owen, USGS Mesozoic" authorityYear="1972" box="[428,681,727,752]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="jeletzkyi">
<emphasis id="FCB201376252FFB597BE40C4FD7D32B3" box="[428,681,727,752]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia jeletzkyi</emphasis>
</taxonomicName>
ranges from one to 4 or 5 (
<bibRefCitation id="AA57A0D46252FFB595F240C4FBBC32B3" author="Owen, E. F." box="[992,1128,727,752]" pageId="10" pageNumber="511" pagination="1 - 8" refId="ref17162" refString="Owen, E. F. (1972) A rhynchonelloid brachiopod from the Upper Jurassic rocks of British Columbia. Geological Survey of Canada Paper, 72 - 26, 1 - 8." type="journal article" year="1972">Owen 1972</bibRefCitation>
, fig. 4, pl. 1, figs. 1c, 24).
<taxonomicName id="09C6A6A66252FFB5968540EFFE413356" authorityName="Owen, USGS Mesozoic" authorityYear="1972" box="[151,405,764,789]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="jeletzkyi">
<emphasis id="FCB201376252FFB5968540EFFE413356" box="[151,405,764,789]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia jeletzkyi</emphasis>
</taxonomicName>
is reported from the
<taxonomicName id="09C6A6A66252FFB5949B40EFFC873356" box="[649,851,764,789]" class="Magnoliopsida" family="Rubiaceae" genus="Buchia" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="511" phylum="Tracheophyta" rank="species" species="fisheriana">
<emphasis id="FCB201376252FFB5949B40EFFC873356" box="[649,851,764,789]" italics="true" pageId="10" pageNumber="511">Buchia fisheriana</emphasis>
</taxonomicName>
and
<taxonomicName id="09C6A6A66252FFB5959E40EEFC2F3356" box="[908,1019,764,789]" class="Magnoliopsida" family="Rubiaceae" genus="Buchia" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="511" phylum="Tracheophyta" rank="species" species="piochii">
<emphasis id="FCB201376252FFB5959E40EEFC493357" box="[908,925,765,788]" italics="true" pageId="10" pageNumber="511">B</emphasis>
.
<emphasis id="FCB201376252FFB595BF40EFFC2F3356" box="[941,1019,764,789]" italics="true" pageId="10" pageNumber="511">piochii</emphasis>
</taxonomicName>
zones (i.e., Upper Tithonian, in part equivalent to the Upper Volgian).
</paragraph>
<paragraph id="CE79DD256252FFB596D54157FBC434DC" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="511">
In
<taxonomicName id="09C6A6A66252FFB596F54157FDD1331E" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[231,517,836,861]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376252FFB596F54157FDD1331E" box="[231,517,836,861]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
, the specimens may show well-marked costae which may extend across most of the length of the specimen (e.g.
<figureCitation id="56FDC1A06252FFB597ED4174FDBC33C3" box="[511,616,871,896]" captionStart="FIGURE 3" captionStartId="7.[151,250,1756,1778]" captionTargetBox="[261,1312,207,1726]" captionTargetId="figure-152@7.[260,1316,193,1726]" captionTargetPageId="7" captionText="FIGURE 3. 112. Pseudomonticlarella varia Smirnova. 14. PMO 227.416, seep 13, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 4). 58. PMO 227.419, seep 7, dorsal, ventral, lateral, and anterior views. 912. PMO 227.418, seep 13, dorsal, ventral, lateral, and anterior views. 1332. Ptilorhynchia mclachlani sp. nov. 1316. PMO 224.876, seep 9, dorsal, ventral, lateral, and anterior views. 1720, PMO 224.879, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5B). 2124. PMO 227.431, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5A). 2528. PMO 224.871, seep 9, dorsal, ventral, lateral, and anterior views. 2932. Holotype, PMO 224.882, seep 9, dorsal, ventral, lateral, and anterior views. Dotted line delimits photographs at the same magnification. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951754" httpUri="https://zenodo.org/record/4951754/files/figure.png" pageId="10" pageNumber="511">Fig. 3.29</figureCitation>
). This costate shell ornament is not typical of
<taxonomicName id="09C6A6A66252FFB5927F4174FAD233C3" authorityName="Crickmay" authorityYear="1933" box="[1133,1286,871,896]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376252FFB5927F4174FAD233C3" box="[1133,1286,871,896]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia</emphasis>
</taxonomicName>
, which tends to show costation developed at the shell margin (e.g. species figured by
<bibRefCitation id="AA57A0D46252FFB595EE419FFB5F33E6" author="Dagys, A. S." box="[1020,1163,908,933]" pageId="10" pageNumber="511" pagination="1 - 167" refId="ref14289" refString="Dagys, A. S. (1968) Jurskiye i rannemelovye brakhiopody Severa Sibiri [Jurassic and Early Cretaceous brachiopods from northern Siberia]. Akademia Nauk SSSR Sibiroskoe Otdelenie Institut Geologii I Geofiziki (IGIG) Trudy [Institute of Geology and Geophysics, Academy of Sciences of the USSR, Siberian Branch, Transactions], 41, 1 - 167. [In Russian]" type="journal article" year="1968">Dagys 1968</bibRefCitation>
;
<bibRefCitation id="AA57A0D46252FFB5928E419FFAF333E6" author="Owen, E. F." box="[1180,1319,908,933]" pageId="10" pageNumber="511" pagination="1 - 8" refId="ref17162" refString="Owen, E. F. (1972) A rhynchonelloid brachiopod from the Upper Jurassic rocks of British Columbia. Geological Survey of Canada Paper, 72 - 26, 1 - 8." type="journal article" year="1972">Owen 1972</bibRefCitation>
; Owen &amp; Manceñido 2002; and
<taxonomicName id="09C6A6A66252FFB5978B41A2FDC7338B" authorityName="Owen, USGS Mesozoic" authorityYear="1972" box="[409,531,943,968]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="jeletzkyi">
<emphasis id="FCB201376252FFB5978B41A2FE73338B" box="[409,423,945,968]" italics="true" pageId="10" pageNumber="511">P</emphasis>
.
<emphasis id="FCB201376252FFB597A541BCFDC7338B" box="[439,531,943,968]" italics="true" pageId="10" pageNumber="511">jeletzkyi</emphasis>
</taxonomicName>
herein,
<figureCitation id="56FDC1A06252FFB5946041BCFD2A338B" box="[626,766,943,968]" captionStart="FIGURE 6" captionStartId="13.[151,250,1796,1818]" captionTargetBox="[152,1430,194,1774]" captionTargetId="figure-113@13.[150,1437,193,1774]" captionTargetPageId="13" captionText="FIGURE 6. 14. Ptilorhynchia aff. plumasensis Crickmay, USGS Mesozoic Locality 32032 (=80Ace119), Karluk Quadrangle, Alaska, Shelikof Formation, Callovian, Middle Jurassic, dorsal, ventral, lateral, and anterior views. 58. Ptilorhynchia jeletzkyi Owen, USGS Mesozoic locality 14496, cooccurs with Buchia mosquensis (von Buch), probably Naknek series, KimmeridgianTithonian(?), Upper Jurassic, McCarthy Quadrangle, Alaska. Collector, Fred H. Moffit, 1928. Dorsal, ventral, lateral, and anterior views.917. Ptilorhynchia obscuricostata Dagys. 912. PMO 224.880, seep 9, dorsal, ventral, lateral, and anterior views. 1317. PMO 224.881, seep 9, dorsal, ventral, lateral, anterior and posterior views. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951772" httpUri="https://zenodo.org/record/4951772/files/figure.png" pageId="10" pageNumber="511">Fig. 6.56.8</figureCitation>
, and a specimen identified as
<taxonomicName id="09C6A6A66252FFB5924941A2FAE5338B" authorityName="Crickmay, USGS Mesozoic Locality" authorityYear="1933" box="[1115,1329,943,968]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="plumasensis">
<emphasis id="FCB201376252FFB5924941A2FBBD338B" box="[1115,1129,945,968]" italics="true" pageId="10" pageNumber="511">P</emphasis>
. aff.
<emphasis id="FCB201376252FFB592BA41BCFAE5338B" box="[1192,1329,943,968]" italics="true" pageId="10" pageNumber="511">plumasensis</emphasis>
</taxonomicName>
from the Middle Jurassic of
<collectingRegion id="0C0213C76252FFB5976141C7FE1033AE" box="[371,452,980,1005]" country="United States of America" name="Alaska" pageId="10" pageNumber="511">Alaska</collectingRegion>
,
<figureCitation id="56FDC1A06252FFB597C341C7FD8E33AE" box="[465,602,980,1005]" captionStart="FIGURE 6" captionStartId="13.[151,250,1796,1818]" captionTargetBox="[152,1430,194,1774]" captionTargetId="figure-113@13.[150,1437,193,1774]" captionTargetPageId="13" captionText="FIGURE 6. 14. Ptilorhynchia aff. plumasensis Crickmay, USGS Mesozoic Locality 32032 (=80Ace119), Karluk Quadrangle, Alaska, Shelikof Formation, Callovian, Middle Jurassic, dorsal, ventral, lateral, and anterior views. 58. Ptilorhynchia jeletzkyi Owen, USGS Mesozoic locality 14496, cooccurs with Buchia mosquensis (von Buch), probably Naknek series, KimmeridgianTithonian(?), Upper Jurassic, McCarthy Quadrangle, Alaska. Collector, Fred H. Moffit, 1928. Dorsal, ventral, lateral, and anterior views.917. Ptilorhynchia obscuricostata Dagys. 912. PMO 224.880, seep 9, dorsal, ventral, lateral, and anterior views. 1317. PMO 224.881, seep 9, dorsal, ventral, lateral, anterior and posterior views. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951772" httpUri="https://zenodo.org/record/4951772/files/figure.png" pageId="10" pageNumber="511">Fig. 6.16.4</figureCitation>
close to the
<typeStatus id="117D63876252FFB594E241C7FCF433AF" box="[752,800,980,1004]" pageId="10" pageNumber="511">type</typeStatus>
species from the Middle Jurassic Hinchman Formation (Callovian) of
<collectingRegion id="0C0213C76252FFB5972941E4FE793453" box="[315,429,1015,1040]" country="United States of America" name="California" pageId="10" pageNumber="511">California</collectingRegion>
(
<bibRefCitation id="AA57A0D46252FFB597A841E4FDB83453" author="Crickmay, C. H." box="[442,620,1015,1040]" pageId="10" pageNumber="511" pagination="871 - 893" refId="ref14234" refString="Crickmay, C. H. (1933) Attempt to zone the North American Jurassic on the basis of its brachiopods. Bulletin of the Geological Society of America, 44 (5), 871 - 893. http: // dx. doi. org / 10.1130 / GSAB- 44 - 871" type="journal article" year="1933">Crickmay 1933</bibRefCitation>
; see also
<bibRefCitation id="AA57A0D46252FFB594C741E4FC8E3453" author="Owen, E. F." box="[725,858,1015,1040]" pageId="10" pageNumber="511" pagination="1 - 8" refId="ref17162" refString="Owen, E. F. (1972) A rhynchonelloid brachiopod from the Upper Jurassic rocks of British Columbia. Geological Survey of Canada Paper, 72 - 26, 1 - 8." type="journal article" year="1972">Owen 1972</bibRefCitation>
).
<bibRefCitation id="AA57A0D46252FFB5957F41E4FBA13453" author="MacFarlan, D. A. B. &amp; Hasibuan, F. &amp; Grant-Mackie, J. A." box="[877,1141,1015,1040]" pageId="10" pageNumber="511" pagination="149 - 177" refId="ref15700" refString="MacFarlan, D. A. B., Hasibuan, F. &amp; Grant-Mackie, J. A. (2011) Mesozoic brachiopods of Misool archipelago, eastern Indonesia. Memoirs of the Association of Australasian Palaeontologists, 41, 149 - 177." type="journal article" year="2011">
MacFarlan
<emphasis id="FCB201376252FFB595FF41EAFBC93453" box="[1005,1053,1015,1040]" italics="true" pageId="10" pageNumber="511">et al</emphasis>
. (2011)
</bibRefCitation>
claim the
<typeStatus id="117D63876252FFB592FB41EBFACD3453" box="[1257,1305,1016,1040]" pageId="10" pageNumber="511">type</typeStatus>
horizon for
<taxonomicName id="09C6A6A66252FFB59685460EFEE93476" authorityName="Crickmay, USGS Mesozoic Locality" authorityYear="1933" box="[151,317,1052,1077]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="plumasensis">
<emphasis id="FCB201376252FFB59685460EFF713477" box="[151,165,1053,1076]" italics="true" pageId="10" pageNumber="511">P</emphasis>
.
<emphasis id="FCB201376252FFB596A6460FFEE93476" box="[180,317,1052,1077]" italics="true" pageId="10" pageNumber="511">plumasensis</emphasis>
</taxonomicName>
to be the Upper Jurassic Knoxville Formation, but their reason for this revised assignation was not given). Most of the specimens from Spitsbergen display four costae on the dorsal fold and three costae in the ventral sulcus. Due to the distinctive nature of the costation it is considered desirable to refer the Spitsbergen material to a new species rather than to the morphologically similar
<taxonomicName id="09C6A6A66252FFB59580469AFBDD34E3" box="[914,1033,1159,1184]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="jeletzyki">
<emphasis id="FCB201376252FFB59580469AFC7434E3" box="[914,928,1161,1184]" italics="true" pageId="10" pageNumber="511">P</emphasis>
.
<emphasis id="FCB201376252FFB595BF4694FBDD34E3" box="[941,1033,1159,1184]" italics="true" pageId="10" pageNumber="511">jeletzyki</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="CE79DD256252FFB596D546BFFD5636A3" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="511">
The weakly costate
<taxonomicName id="09C6A6A66252FFB597A646BFFD023486" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[436,726,1196,1221]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376252FFB597A646BFFD023486" box="[436,726,1196,1221]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
resembles some species referred to
<taxonomicName id="09C6A6A66252FFB5929546BFFAF53486" authorityName="Crickmay" authorityYear="1933" box="[1159,1313,1196,1221]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376252FFB5929546BFFAF53486" box="[1159,1313,1196,1221]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia</emphasis>
</taxonomicName>
by
<bibRefCitation id="AA57A0D46252FFB5934646BFFF3134AB" author="Dagys, A. S." pageId="10" pageNumber="511" pagination="1 - 167" refId="ref14289" refString="Dagys, A. S. (1968) Jurskiye i rannemelovye brakhiopody Severa Sibiri [Jurassic and Early Cretaceous brachiopods from northern Siberia]. Akademia Nauk SSSR Sibiroskoe Otdelenie Institut Geologii I Geofiziki (IGIG) Trudy [Institute of Geology and Geophysics, Academy of Sciences of the USSR, Siberian Branch, Transactions], 41, 1 - 167. [In Russian]" type="journal article" year="1968">Dagys (1968)</bibRefCitation>
. Of these, the Spitsbergen specimens are closest to
<taxonomicName id="09C6A6A66252FFB5953A46C2FC7834AB" authorityName="Dagys" authorityYear="1968" box="[808,940,1231,1256]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="lenaensis">
<emphasis id="FCB201376252FFB5953A46C2FCE234AB" box="[808,822,1233,1256]" italics="true" pageId="10" pageNumber="511">P</emphasis>
.
<emphasis id="FCB201376252FFB5955646DCFC7834AB" box="[836,940,1231,1256]" italics="true" pageId="10" pageNumber="511">lenaensis</emphasis>
</taxonomicName>
in the nature of the costation of the anterior commissure but the overall size of the latter is smaller, and the number of costae on the fold and sulcus fewer than in the Spitsbergen specimens. Matching closely in outline and costation is
<taxonomicName id="09C6A6A66252FFB595F2470AFAC93573" authority="Dagys (1968" authorityName="Dagys" authorityYear="1968" box="[992,1309,1303,1328]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="anadyrensis">
<emphasis id="FCB201376252FFB595F2470AFC3A3573" box="[992,1006,1305,1328]" italics="true" pageId="10" pageNumber="511">P</emphasis>
.
<emphasis id="FCB201376252FFB595EF4704FB573573" box="[1021,1155,1303,1328]" italics="true" pageId="10" pageNumber="511">anadyrensis</emphasis>
<bibRefCitation id="AA57A0D46252FFB59299470BFAC93573" author="Dagys, A. S." box="[1163,1309,1303,1328]" pageId="10" pageNumber="511" pagination="1 - 167" refId="ref14289" refString="Dagys, A. S. (1968) Jurskiye i rannemelovye brakhiopody Severa Sibiri [Jurassic and Early Cretaceous brachiopods from northern Siberia]. Akademia Nauk SSSR Sibiroskoe Otdelenie Institut Geologii I Geofiziki (IGIG) Trudy [Institute of Geology and Geophysics, Academy of Sciences of the USSR, Siberian Branch, Transactions], 41, 1 - 167. [In Russian]" type="journal article" year="1968">Dagys (1968</bibRefCitation>
</taxonomicName>
, pl. 4, figs. 3, 4) although the nature of the costation appears more angular at the anterior commissure when compared to
<taxonomicName id="09C6A6A66252FFB59685474CFE60353B" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[151,436,1375,1400]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376252FFB59685474CFE60353B" box="[151,436,1375,1400]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
. The specimens of these species figured by Dagys appear to all be decorticated. The age assignment of
<taxonomicName id="09C6A6A66252FFB597564796FE1935DE" authorityName="Dagys" authorityYear="1968" box="[324,461,1412,1437]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="lenaensis">
<emphasis id="FCB201376252FFB597564796FE8635DF" box="[324,338,1413,1436]" italics="true" pageId="10" pageNumber="511">P</emphasis>
.
<emphasis id="FCB201376252FFB597774797FE1935DE" box="[357,461,1412,1437]" italics="true" pageId="10" pageNumber="511">lenaensis</emphasis>
</taxonomicName>
is Valanginian (
<bibRefCitation id="AA57A0D46252FFB594834797FCCA35DE" author="Dagys, A. S." box="[657,798,1412,1437]" pageId="10" pageNumber="511" pagination="1 - 167" refId="ref14289" refString="Dagys, A. S. (1968) Jurskiye i rannemelovye brakhiopody Severa Sibiri [Jurassic and Early Cretaceous brachiopods from northern Siberia]. Akademia Nauk SSSR Sibiroskoe Otdelenie Institut Geologii I Geofiziki (IGIG) Trudy [Institute of Geology and Geophysics, Academy of Sciences of the USSR, Siberian Branch, Transactions], 41, 1 - 167. [In Russian]" type="journal article" year="1968">Dagys 1968</bibRefCitation>
) so it is slightly younger in age than the Spitsbergen specimens while
<taxonomicName id="09C6A6A66252FFB5974947BAFE2B3583" authorityName="Dagys" authorityYear="1968" box="[347,511,1447,1472]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="anadyrensis">
<emphasis id="FCB201376252FFB5974947BAFEBD3583" box="[347,361,1449,1472]" italics="true" pageId="10" pageNumber="511">P</emphasis>
.
<emphasis id="FCB201376252FFB5976B47B4FE2B3583" box="[377,511,1447,1472]" italics="true" pageId="10" pageNumber="511">anadyrensis</emphasis>
</taxonomicName>
was given as of uncertain age, referred questionably to the BathonianCallovian (
<bibRefCitation id="AA57A0D46252FFB5968C47DFFEFE35A6" author="Dagys, A. S." box="[158,298,1484,1509]" pageId="10" pageNumber="511" pagination="1 - 167" refId="ref14289" refString="Dagys, A. S. (1968) Jurskiye i rannemelovye brakhiopody Severa Sibiri [Jurassic and Early Cretaceous brachiopods from northern Siberia]. Akademia Nauk SSSR Sibiroskoe Otdelenie Institut Geologii I Geofiziki (IGIG) Trudy [Institute of Geology and Geophysics, Academy of Sciences of the USSR, Siberian Branch, Transactions], 41, 1 - 167. [In Russian]" type="journal article" year="1968">Dagys 1968</bibRefCitation>
, table 1, p. 145). A single rhynchonellide specimen (internal cast of a ventral valve, approximately
<quantity id="093E70C06252FFB5939C47DFFF173644" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="511" unit="mm" value="8.0">8 mm</quantity>
x
<quantity id="093E70C06252FFB596F647FCFEFD364B" box="[228,297,1519,1544]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="511" unit="mm" value="8.0">8 mm</quantity>
) from undifferentiated BerriasianValanginian of the Barents Sea (
<bibRefCitation id="AA57A0D46252FFB5923D47FCFB29364B" author="Arhus, N. &amp; Kelly, S. R. A. &amp; Collins, J. S. H. &amp; Sandy, M. R." box="[1071,1277,1519,1544]" pageId="10" pageNumber="511" pagination="165 - 194" refId="ref13801" refString="Arhus, N., Kelly, S. R. A., Collins, J. S. H. &amp; Sandy, M. R. (1990) Systematic palaeontology and biostratigraphy of two Early Cretaceous condensed sections from the Barents Sea. Polar Research, 8, 165 - 194. http: // dx. doi. org / 10.3402 / polar. v 8 i 2.6811" type="journal article" year="1990">
Århus
<emphasis id="FCB201376252FFB5926D47E2FB6E364B" box="[1151,1210,1519,1544]" italics="true" pageId="10" pageNumber="511">et al.</emphasis>
1990
</bibRefCitation>
) identified as
<taxonomicName id="09C6A6A66252FFB596854407FEB63668" box="[151,354,1556,1581]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="undetermined">
<emphasis id="FCB201376252FFB596854407FEE5366E" box="[151,305,1556,1581]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia</emphasis>
sp.
</taxonomicName>
, was tentatively referred to this genus.
<bibRefCitation id="AA57A0D46252FFB595474407FBAC366E" author="MacFarlan, D. A. B. &amp; Hasibuan, F. &amp; Grant-Mackie, J. A." box="[853,1144,1556,1581]" pageId="10" pageNumber="511" pagination="149 - 177" refId="ref15700" refString="MacFarlan, D. A. B., Hasibuan, F. &amp; Grant-Mackie, J. A. (2011) Mesozoic brachiopods of Misool archipelago, eastern Indonesia. Memoirs of the Association of Australasian Palaeontologists, 41, 149 - 177." type="journal article" year="2011">
MacFarlan
<emphasis id="FCB201376252FFB595CC4406FBC3366E" box="[990,1047,1556,1581]" italics="true" pageId="10" pageNumber="511">et al</emphasis>
. (2011)
</bibRefCitation>
described
<taxonomicName id="09C6A6A66252FFB593104407FEED3613" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="pugnaciformis">
<emphasis id="FCB201376252FFB593104407FEED3613" italics="true" pageId="10" pageNumber="511">Ptilorhynchia pugnaciformis</emphasis>
</taxonomicName>
from the Lower Cretaceous of the Misool Archipelago, eastern
<collectingCountry id="B6D19DB56252FFB5923B4424FB4E3613" box="[1065,1178,1591,1616]" name="Indonesia" pageId="10" pageNumber="511">Indonesia</collectingCountry>
. Like other species of
<taxonomicName id="09C6A6A66252FFB59685444FFEE53636" authorityName="Crickmay" authorityYear="1933" box="[151,305,1628,1653]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376252FFB59685444FFEE53636" box="[151,305,1628,1653]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia</emphasis>
</taxonomicName>
already discussed, the number of costae for this species is considered to be variable (between 36 per valve; MacFarlan
<emphasis id="FCB201376252FFB597704492FE4536DB" box="[354,401,1663,1688]" italics="true" pageId="10" pageNumber="511">et al</emphasis>
., table 1). The posterior profile of the dorsal valve in
<taxonomicName id="09C6A6A66252FFB595F54492FB7036DB" box="[999,1188,1663,1688]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="pugnaciformis">
<emphasis id="FCB201376252FFB595F54492FC2136DB" box="[999,1013,1665,1688]" italics="true" pageId="10" pageNumber="511">P</emphasis>
.
<emphasis id="FCB201376252FFB59210446CFB7036DB" box="[1026,1188,1663,1688]" italics="true" pageId="10" pageNumber="511">pugnaciformis</emphasis>
</taxonomicName>
is not as rounded as it is in
<taxonomicName id="09C6A6A66252FFB596DC44B6FEB336FE" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[206,359,1700,1725]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376252FFB596DC44B6FF0836FF" box="[206,220,1701,1724]" italics="true" pageId="10" pageNumber="511">P</emphasis>
.
<emphasis id="FCB201376252FFB596F944B7FEB336FE" box="[235,359,1700,1725]" italics="true" pageId="10" pageNumber="511">mclachlani</emphasis>
</taxonomicName>
or
<taxonomicName id="09C6A6A66252FFB5979C44B6FDD236FE" authorityName="Owen, USGS Mesozoic" authorityYear="1972" box="[398,518,1700,1725]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="jeletzkyi">
<emphasis id="FCB201376252FFB5979C44B6FDD236FE" box="[398,518,1700,1725]" italics="true" pageId="10" pageNumber="511">P. jeletzkyi</emphasis>
</taxonomicName>
; in
<taxonomicName id="09C6A6A66252FFB5942144B7FD0136FE" box="[563,725,1700,1725]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="pugnaciformis">
<emphasis id="FCB201376252FFB5942144B7FD0136FE" box="[563,725,1700,1725]" italics="true" pageId="10" pageNumber="511">pugnaciformis</emphasis>
</taxonomicName>
the profile may be cynocephalous and therefore resembling, at least in profile,
<taxonomicName id="09C6A6A66252FFB5975444D4FD5636A3" authority="Buckman." box="[326,642,1735,1760]" class="Rhynchonellata" family="Rhynchonellidae" genus="Homoeorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376252FFB5975444D4FDDC36A3" box="[326,520,1735,1760]" italics="true" pageId="10" pageNumber="511">Homoeorhynchia</emphasis>
Buckman.
</taxonomicName>
</paragraph>
<paragraph id="CE79DD256252FFB396D544FFFE6F318C" blockId="10.[151,1437,151,2013]" lastBlockId="12.[151,1437,151,1221]" lastPageId="12" lastPageNumber="513" pageId="10" pageNumber="511">
Small multicostate specimens are considered juveniles of
<taxonomicName id="09C6A6A66252FFB5955844FFFBB23746" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[842,1126,1772,1797]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376252FFB5955844FFFBB23746" box="[842,1126,1772,1797]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
. These specimens had been tentatively referred to
<taxonomicName id="09C6A6A66252FFB5979D451CFDFC376B" authorityName="Wisniewska" authorityYear="1932" box="[399,552,1807,1832]" class="Rhynchonellata" family="Norellidae" genus="Monticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376252FFB5979D451CFDFC376B" box="[399,552,1807,1832]" italics="true" pageId="10" pageNumber="511">Monticlarella</emphasis>
</taxonomicName>
by Sandy in
<bibRefCitation id="AA57A0D46252FFB594A84503FC79376B" author="Hammer, O. &amp; Nakrem, H. A. &amp; Little, C. T. S. &amp; Hryniewicz, K. &amp; Sandy, M. R. &amp; Hurum, J. H. &amp; Druckenmiller, P. &amp; Knutsen, E. M. &amp; Hoyberget, M." box="[698,941,1807,1832]" pageId="10" pageNumber="511" pagination="15 - 26" refId="ref14853" refString="Hammer, O., Nakrem, H. A., Little, C. T. S., Hryniewicz, K., Sandy, M. R., Hurum, J. H., Druckenmiller, P., Knutsen, E. M. &amp; Hoyberget, M. (2011) Hydrocarbon seeps from close to the Jurassic - Cretaceous boundary, Svalbard. Palaeogeography, Palaeoecology, Palaeoclimatology, 306, 15 - 26. http: // dx. doi. org / 10.1016 / j. palaeo. 2011.03.019" type="journal article" year="2011">
Hammer
<emphasis id="FCB201376252FFB595314502FC8F376B" box="[803,859,1807,1832]" italics="true" pageId="10" pageNumber="511">et al.</emphasis>
(2011)
</bibRefCitation>
; they have a similar outline to
<taxonomicName id="09C6A6A66252FFB59311451CFE4A370E" authority="(Quenstedt)" baseAuthorityName="Quenstedt" baseAuthorityYear="1858" class="Rhynchonellata" family="Norellidae" genus="Monticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="triloboides">
<emphasis id="FCB201376252FFB59311451CFEC4370E" italics="true" pageId="10" pageNumber="511">Monticlarella triloboides</emphasis>
(Quenstedt)
</taxonomicName>
in
<bibRefCitation id="AA57A0D46252FFB597D44527FD89370E" author="Childs, A." box="[454,605,1844,1869]" pageId="10" pageNumber="511" pagination="1 - 119" refId="ref14079" refString="Childs, A. (1969) Upper Jurassic rhynchonellid brachiopods from Northwestern Europe. British Museum (Natural History) Bulletin (Geology), 6, 1 - 119. [Supplement]" type="journal article" year="1969">Childs (1969</bibRefCitation>
, pl. I, figs. 57). However, the nature of the costation is consistent with
<taxonomicName id="09C6A6A66252FFB596854544FE6D3733" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[151,441,1879,1904]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376252FFB596854544FE6D3733" box="[151,441,1879,1904]" italics="true" pageId="10" pageNumber="511">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
and the broad uniplicate fold develops more strongly as the brachiopod approaches maturity, with a concomitant deflection of the lateral commissure (
<figureCitation id="56FDC1A06252FFB59597456FFB7D37D6" box="[901,1193,1916,1941]" captionStart="FIGURE 3" captionStartId="7.[151,250,1756,1778]" captionTargetBox="[261,1312,207,1726]" captionTargetId="figure-152@7.[260,1316,193,1726]" captionTargetPageId="7" captionText="FIGURE 3. 112. Pseudomonticlarella varia Smirnova. 14. PMO 227.416, seep 13, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 4). 58. PMO 227.419, seep 7, dorsal, ventral, lateral, and anterior views. 912. PMO 227.418, seep 13, dorsal, ventral, lateral, and anterior views. 1332. Ptilorhynchia mclachlani sp. nov. 1316. PMO 224.876, seep 9, dorsal, ventral, lateral, and anterior views. 1720, PMO 224.879, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5B). 2124. PMO 227.431, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5A). 2528. PMO 224.871, seep 9, dorsal, ventral, lateral, and anterior views. 2932. Holotype, PMO 224.882, seep 9, dorsal, ventral, lateral, and anterior views. Dotted line delimits photographs at the same magnification. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951754" httpUri="https://zenodo.org/record/4951754/files/figure.png" pageId="10" pageNumber="511">Fig. 3.213.24, 3.293.32</figureCitation>
cf. for juveniles,
<figureCitation id="56FDC1A06252FFB59362456FFE4737FA" captionStart="FIGURE 3" captionStartId="7.[151,250,1756,1778]" captionTargetBox="[261,1312,207,1726]" captionTargetId="figure-152@7.[260,1316,193,1726]" captionTargetPageId="7" captionText="FIGURE 3. 112. Pseudomonticlarella varia Smirnova. 14. PMO 227.416, seep 13, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 4). 58. PMO 227.419, seep 7, dorsal, ventral, lateral, and anterior views. 912. PMO 227.418, seep 13, dorsal, ventral, lateral, and anterior views. 1332. Ptilorhynchia mclachlani sp. nov. 1316. PMO 224.876, seep 9, dorsal, ventral, lateral, and anterior views. 1720, PMO 224.879, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5B). 2124. PMO 227.431, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5A). 2528. PMO 224.871, seep 9, dorsal, ventral, lateral, and anterior views. 2932. Holotype, PMO 224.882, seep 9, dorsal, ventral, lateral, and anterior views. Dotted line delimits photographs at the same magnification. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951754" httpUri="https://zenodo.org/record/4951754/files/figure.png" pageId="10" pageNumber="511">Fig. 3.133.20, 3.253.28</figureCitation>
). Dental lamellae and a median septum can be readily discerned in the majority of the smaller specimens; characters that if present in
<taxonomicName id="09C6A6A66252FFB594AE45D7FC8D379E" authorityName="Wisniewska" authorityYear="1932" box="[700,857,1988,2013]" class="Rhynchonellata" family="Norellidae" genus="Monticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="10" pageNumber="511" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376252FFB594AE45D7FC8D379E" box="[700,857,1988,2013]" italics="true" pageId="10" pageNumber="511">Monticlarella</emphasis>
</taxonomicName>
are typically poorly developed (e.g.
<bibRefCitation id="AA57A0D46252FFB5931145D7FA58379E" author="Owen, E. F." box="[1283,1420,1988,2013]" pageId="10" pageNumber="511" pagination="17 - 32" refId="ref17093" refString="Owen, E. F. (1968) A further study of some Cretaceous rhynchonelloid brachiopods. Bulletin of the Indian Geologists' Association, 1, 17 - 32." type="journal article" year="1968">Owen 1968</bibRefCitation>
; Manceñido
<emphasis id="FCB201376254FFB3970F428AFE8C30F3" box="[285,344,151,176]" italics="true" pageId="12" pageNumber="513">et al.</emphasis>
2002, p. 1316). (From the palaeobiographic point of view,
<taxonomicName id="09C6A6A66254FFB392124284FB4D30F3" authorityName="Wisniewska" authorityYear="1932" box="[1024,1177,151,176]" class="Rhynchonellata" family="Norellidae" genus="Monticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376254FFB392124284FB4D30F3" box="[1024,1177,151,176]" italics="true" pageId="12" pageNumber="513">Monticlarella</emphasis>
</taxonomicName>
has not been recorded from Boreal faunas of the Russian Platform or Siberia, whereas several species have been referred to
<taxonomicName id="09C6A6A66254FFB3931042AFFA4F3096" authorityName="Crickmay" authorityYear="1933" box="[1282,1435,188,213]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376254FFB3931042AFFA4F3096" box="[1282,1435,188,213]" italics="true" pageId="12" pageNumber="513">Ptilorhynchia</emphasis>
</taxonomicName>
from these regions.) Serial sections were taken of one of the smaller-sized specimens of
<taxonomicName id="09C6A6A66254FFB3926C42CCFA4830BB" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[1150,1436,223,248]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376254FFB3926C42CCFA4830BB" box="[1150,1436,223,248]" italics="true" pageId="12" pageNumber="513">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
to clarify the relationships among the material (
<figureCitation id="56FDC1A06254FFB394BD4317FCDC315E" box="[687,776,260,285]" captionStart="FIGURE 5" captionStartId="11.[151,250,1820,1842]" captionTargetBox="[238,1340,193,1782]" captionTargetId="figure-142@11.[238,1340,193,1798]" captionTargetPageId="11" captionText="FIGURE 5. A and B, transverse serial sections from two specimens of Ptilorhynchia mclachlani sp. nov., seep 9. A) Specimen PMO 227.431, sections above third dotted line. Dotted lines delimit groups of sections drawn at the same magnification. Dimensions of specimen: L 15.3+; W 18.3; T 9.8 mm. Serial sections prepared perpendicular to maximum length of specimen; numbering of sections starts at 0.0 mm, the initial section through the specimen. B) Specimen PMO 224.879, sections below third dotted line. Dimensions of sectioned specimen: L 12.9; W 12.8+; T 9.0 mm. Serial sections prepared perpendicular to length of dorsal valve; numbering of sections starts at 0.0 mm, the initial section through the specimen. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951764" httpUri="https://zenodo.org/record/4951764/files/figure.png" pageId="12" pageNumber="513">Fig. 5B</figureCitation>
). The serial sections show remnants of a dental lamella on the right side of the specimen (
<figureCitation id="56FDC1A06254FFB394154334FDB53103" box="[519,609,295,320]" captionStart="FIGURE 5" captionStartId="11.[151,250,1820,1842]" captionTargetBox="[238,1340,193,1782]" captionTargetId="figure-142@11.[238,1340,193,1798]" captionTargetPageId="11" captionText="FIGURE 5. A and B, transverse serial sections from two specimens of Ptilorhynchia mclachlani sp. nov., seep 9. A) Specimen PMO 227.431, sections above third dotted line. Dotted lines delimit groups of sections drawn at the same magnification. Dimensions of specimen: L 15.3+; W 18.3; T 9.8 mm. Serial sections prepared perpendicular to maximum length of specimen; numbering of sections starts at 0.0 mm, the initial section through the specimen. B) Specimen PMO 224.879, sections below third dotted line. Dimensions of sectioned specimen: L 12.9; W 12.8+; T 9.0 mm. Serial sections prepared perpendicular to length of dorsal valve; numbering of sections starts at 0.0 mm, the initial section through the specimen. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951764" httpUri="https://zenodo.org/record/4951764/files/figure.png" pageId="12" pageNumber="513">Fig. 5B</figureCitation>
, section 0.1), a well-developed septalium (0.30.7) and crura that arch ventrally towards the anterior of the brachiopod (1.01.6). The crura develop as discrete structures at the junction between the outer hinge plates and the inner hinge plates of the septalial structure (0.51.0). These serial sections are very similar in overall aspect to those taken from the larger specimen (
<figureCitation id="56FDC1A06254FFB392014387FBBB31EE" box="[1043,1135,404,429]" captionStart="FIGURE 5" captionStartId="11.[151,250,1820,1842]" captionTargetBox="[238,1340,193,1782]" captionTargetId="figure-142@11.[238,1340,193,1798]" captionTargetPageId="11" captionText="FIGURE 5. A and B, transverse serial sections from two specimens of Ptilorhynchia mclachlani sp. nov., seep 9. A) Specimen PMO 227.431, sections above third dotted line. Dotted lines delimit groups of sections drawn at the same magnification. Dimensions of specimen: L 15.3+; W 18.3; T 9.8 mm. Serial sections prepared perpendicular to maximum length of specimen; numbering of sections starts at 0.0 mm, the initial section through the specimen. B) Specimen PMO 224.879, sections below third dotted line. Dimensions of sectioned specimen: L 12.9; W 12.8+; T 9.0 mm. Serial sections prepared perpendicular to length of dorsal valve; numbering of sections starts at 0.0 mm, the initial section through the specimen. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951764" httpUri="https://zenodo.org/record/4951764/files/figure.png" pageId="12" pageNumber="513">Fig. 5A</figureCitation>
) and both are considered
<taxonomicName id="09C6A6A66254FFB3968543A4FE673193" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[151,435,439,464]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376254FFB3968543A4FE673193" box="[151,435,439,464]" italics="true" pageId="12" pageNumber="513">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="9AB98DAD6253FFB49685450FFAA13788" ID-DOI="http://doi.org/10.5281/zenodo.4951764" ID-Zenodo-Dep="4951764" httpUri="https://zenodo.org/record/4951764/files/figure.png" pageId="11" pageNumber="512" startId="11.[151,250,1820,1842]" targetBox="[238,1340,193,1782]" targetPageId="11">
<paragraph id="CE79DD256253FFB49685450FFAA13788" blockId="11.[151,1437,1820,1995]" pageId="11" pageNumber="512">
<emphasis id="FCB201376253FFB49685450FFEC63771" bold="true" box="[151,274,1820,1842]" pageId="11" pageNumber="512">FIGURE 5.</emphasis>
A and B, transverse serial sections from two specimens of
<taxonomicName id="09C6A6A66253FFB49575450FFBBC3771" authority="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little, 2014" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[871,1128,1820,1842]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="11" pageNumber="512" phylum="Brachiopoda" rank="species" species="mclachlani" status="sp. nov.">
<emphasis id="FCB201376253FFB49575450FFBBC3771" box="[871,1128,1820,1842]" italics="true" pageId="11" pageNumber="512">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
<emphasis id="FCB201376253FFB4927C450FFB6B3771" bold="true" box="[1134,1215,1820,1842]" pageId="11" pageNumber="512">
<taxonomicNameLabel id="E781BC4C6253FFB4927C450FFB6B3771" box="[1134,1215,1820,1842]" pageId="11" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, seep 9. A) Specimen PMO 227.431, sections above third dotted line. Dotted lines delimit groups of sections drawn at the same magnification. Dimensions of specimen: L 15.3+; W 18.3; T 9.8 mm. Serial sections prepared perpendicular to maximum length of specimen; numbering of sections starts at 0.0 mm, the initial section through the specimen. B) Specimen PMO 224.879, sections below third dotted line. Dimensions of sectioned specimen: L 12.9; W 12.8+; T 9.0 mm. Serial sections prepared perpendicular to length of dorsal valve; numbering of sections starts at 0.0 mm, the initial section through the specimen. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="CE79DD256254FFB396D543CFFF3D341B" blockId="12.[151,1437,151,1221]" pageId="12" pageNumber="513">
In comparing the serial sections of
<taxonomicName id="09C6A6A66254FFB3944243CFFCB831B6" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[592,876,476,501]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376254FFB3944243CFFCB831B6" box="[592,876,476,501]" italics="true" pageId="12" pageNumber="513">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
(
<figureCitation id="56FDC1A06254FFB3956E43CFFC6A31B6" box="[892,958,476,501]" captionStart="FIGURE 5" captionStartId="11.[151,250,1820,1842]" captionTargetBox="[238,1340,193,1782]" captionTargetId="figure-142@11.[238,1340,193,1798]" captionTargetPageId="11" captionText="FIGURE 5. A and B, transverse serial sections from two specimens of Ptilorhynchia mclachlani sp. nov., seep 9. A) Specimen PMO 227.431, sections above third dotted line. Dotted lines delimit groups of sections drawn at the same magnification. Dimensions of specimen: L 15.3+; W 18.3; T 9.8 mm. Serial sections prepared perpendicular to maximum length of specimen; numbering of sections starts at 0.0 mm, the initial section through the specimen. B) Specimen PMO 224.879, sections below third dotted line. Dimensions of sectioned specimen: L 12.9; W 12.8+; T 9.0 mm. Serial sections prepared perpendicular to length of dorsal valve; numbering of sections starts at 0.0 mm, the initial section through the specimen. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951764" httpUri="https://zenodo.org/record/4951764/files/figure.png" pageId="12" pageNumber="513">Fig. 5</figureCitation>
) with
<taxonomicName id="09C6A6A66254FFB3921543CFFAD731B6" authorityName="Owen, USGS Mesozoic" authorityYear="1972" box="[1031,1283,476,501]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="species" species="jeletzkyi">
<emphasis id="FCB201376254FFB3921543CFFAD731B6" box="[1031,1283,476,501]" italics="true" pageId="12" pageNumber="513">Ptilorhynchia jeletzkyi</emphasis>
</taxonomicName>
(
<bibRefCitation id="AA57A0D46254FFB3930343CFFA4C31B6" author="Owen, E. F." box="[1297,1432,476,501]" pageId="12" pageNumber="513" pagination="1 - 8" refId="ref17162" refString="Owen, E. F. (1972) A rhynchonelloid brachiopod from the Upper Jurassic rocks of British Columbia. Geological Survey of Canada Paper, 72 - 26, 1 - 8." type="journal article" year="1972">Owen 1972</bibRefCitation>
, fig. 1) there are similarities in the development of the septalium and the crura on the inner margins of the hinge plates, and the crura then arch ventrally anteriorly. The larger Spitsbergen specimen (
<figureCitation id="56FDC1A06254FFB3924C4037FB62327E" box="[1118,1206,548,573]" captionStart="FIGURE 5" captionStartId="11.[151,250,1820,1842]" captionTargetBox="[238,1340,193,1782]" captionTargetId="figure-142@11.[238,1340,193,1798]" captionTargetPageId="11" captionText="FIGURE 5. A and B, transverse serial sections from two specimens of Ptilorhynchia mclachlani sp. nov., seep 9. A) Specimen PMO 227.431, sections above third dotted line. Dotted lines delimit groups of sections drawn at the same magnification. Dimensions of specimen: L 15.3+; W 18.3; T 9.8 mm. Serial sections prepared perpendicular to maximum length of specimen; numbering of sections starts at 0.0 mm, the initial section through the specimen. B) Specimen PMO 224.879, sections below third dotted line. Dimensions of sectioned specimen: L 12.9; W 12.8+; T 9.0 mm. Serial sections prepared perpendicular to length of dorsal valve; numbering of sections starts at 0.0 mm, the initial section through the specimen. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951764" httpUri="https://zenodo.org/record/4951764/files/figure.png" pageId="12" pageNumber="513">Fig. 5A</figureCitation>
) did not capture the dental lamellae of the ventral valve, and this is also true when comparing the serial sections of species referred to
<taxonomicName id="09C6A6A66254FFB39685407FFEE532C6" authorityName="Crickmay" authorityYear="1933" box="[151,305,620,645]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376254FFB39685407FFEE532C6" box="[151,305,620,645]" italics="true" pageId="12" pageNumber="513">Ptilorhynchia</emphasis>
</taxonomicName>
by
<bibRefCitation id="AA57A0D46254FFB39777407FFDDC32C6" author="Dagys, A. S." box="[357,520,620,645]" pageId="12" pageNumber="513" pagination="1 - 167" refId="ref14289" refString="Dagys, A. S. (1968) Jurskiye i rannemelovye brakhiopody Severa Sibiri [Jurassic and Early Cretaceous brachiopods from northern Siberia]. Akademia Nauk SSSR Sibiroskoe Otdelenie Institut Geologii I Geofiziki (IGIG) Trudy [Institute of Geology and Geophysics, Academy of Sciences of the USSR, Siberian Branch, Transactions], 41, 1 - 167. [In Russian]" type="journal article" year="1968">Dagys (1968)</bibRefCitation>
. The presence of well-developed dental lamellae in the ventral valves of the Canadian (
<bibRefCitation id="AA57A0D46254FFB39700409CFE4932EB" author="Owen, E. F." box="[274,413,655,680]" pageId="12" pageNumber="513" pagination="1 - 8" refId="ref17162" refString="Owen, E. F. (1972) A rhynchonelloid brachiopod from the Upper Jurassic rocks of British Columbia. Geological Survey of Canada Paper, 72 - 26, 1 - 8." type="journal article" year="1972">Owen 1972</bibRefCitation>
), Siberian (
<bibRefCitation id="AA57A0D46254FFB39430409CFD7A32EB" author="Dagys, A. S." box="[546,686,655,680]" pageId="12" pageNumber="513" pagination="1 - 167" refId="ref14289" refString="Dagys, A. S. (1968) Jurskiye i rannemelovye brakhiopody Severa Sibiri [Jurassic and Early Cretaceous brachiopods from northern Siberia]. Akademia Nauk SSSR Sibiroskoe Otdelenie Institut Geologii I Geofiziki (IGIG) Trudy [Institute of Geology and Geophysics, Academy of Sciences of the USSR, Siberian Branch, Transactions], 41, 1 - 167. [In Russian]" type="journal article" year="1968">Dagys 1968</bibRefCitation>
) and Indonesian (
<bibRefCitation id="AA57A0D46254FFB3956D409CFB5532EB" author="MacFarlan, D. A. B. &amp; Hasibuan, F. &amp; Grant-Mackie, J. A." box="[895,1153,655,680]" pageId="12" pageNumber="513" pagination="149 - 177" refId="ref15700" refString="MacFarlan, D. A. B., Hasibuan, F. &amp; Grant-Mackie, J. A. (2011) Mesozoic brachiopods of Misool archipelago, eastern Indonesia. Memoirs of the Association of Australasian Palaeontologists, 41, 149 - 177." type="journal article" year="2011">
MacFarlan
<emphasis id="FCB201376254FFB392164082FBEC32EB" box="[1028,1080,655,680]" italics="true" pageId="12" pageNumber="513">et al</emphasis>
. 2011
</bibRefCitation>
) material is a difference from the Spitsbergen material. The ventral umbo of the larger specimen selected for serial sectioning (
<figureCitation id="56FDC1A06254FFB3937D40A7FEDC32B3" captionStart="FIGURE 3" captionStartId="7.[151,250,1756,1778]" captionTargetBox="[261,1312,207,1726]" captionTargetId="figure-152@7.[260,1316,193,1726]" captionTargetPageId="7" captionText="FIGURE 3. 112. Pseudomonticlarella varia Smirnova. 14. PMO 227.416, seep 13, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 4). 58. PMO 227.419, seep 7, dorsal, ventral, lateral, and anterior views. 912. PMO 227.418, seep 13, dorsal, ventral, lateral, and anterior views. 1332. Ptilorhynchia mclachlani sp. nov. 1316. PMO 224.876, seep 9, dorsal, ventral, lateral, and anterior views. 1720, PMO 224.879, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5B). 2124. PMO 227.431, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5A). 2528. PMO 224.871, seep 9, dorsal, ventral, lateral, and anterior views. 2932. Holotype, PMO 224.882, seep 9, dorsal, ventral, lateral, and anterior views. Dotted line delimits photographs at the same magnification. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951754" httpUri="https://zenodo.org/record/4951754/files/figure.png" pageId="12" pageNumber="513">Fig. 3.213.24</figureCitation>
) was damaged, with the outline of dental lamellae visible, and their traces can also be determined in the duplicate cast taken of this specimen. In addition, dental lamellae are clearly visible in the damaged ventral umbones of other specimens. Traces of dental lamellae were not seen in the serial sections and acetate peels (for specimen in
<figureCitation id="56FDC1A06254FFB3973A4157FE57331E" box="[296,387,836,861]" captionStart="FIGURE 5" captionStartId="11.[151,250,1820,1842]" captionTargetBox="[238,1340,193,1782]" captionTargetId="figure-142@11.[238,1340,193,1798]" captionTargetPageId="11" captionText="FIGURE 5. A and B, transverse serial sections from two specimens of Ptilorhynchia mclachlani sp. nov., seep 9. A) Specimen PMO 227.431, sections above third dotted line. Dotted lines delimit groups of sections drawn at the same magnification. Dimensions of specimen: L 15.3+; W 18.3; T 9.8 mm. Serial sections prepared perpendicular to maximum length of specimen; numbering of sections starts at 0.0 mm, the initial section through the specimen. B) Specimen PMO 224.879, sections below third dotted line. Dimensions of sectioned specimen: L 12.9; W 12.8+; T 9.0 mm. Serial sections prepared perpendicular to length of dorsal valve; numbering of sections starts at 0.0 mm, the initial section through the specimen. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951764" httpUri="https://zenodo.org/record/4951764/files/figure.png" pageId="12" pageNumber="513">Fig. 5A</figureCitation>
). This specimen was decorticated and the calcite shell of the dental lamellae apparently lost through decortication or weathering in the umbonal region of the ventral valve. Most of the external shell of the specimen was absent too. Of the serial sections of species described by Dagys,
<taxonomicName id="09C6A6A66254FFB39209419EFAEF33E6" authority="Dagys (1968" authorityName="Dagys" authorityYear="1968" box="[1051,1339,908,933]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="species" species="lenaensis">
<emphasis id="FCB201376254FFB39209419EFBFD33E7" box="[1051,1065,909,932]" italics="true" pageId="12" pageNumber="513">P</emphasis>
.
<emphasis id="FCB201376254FFB3922A419FFB7433E6" box="[1080,1184,908,933]" italics="true" pageId="12" pageNumber="513">lenaensis</emphasis>
<bibRefCitation id="AA57A0D46254FFB392BA419FFAEF33E6" author="Dagys, A. S." box="[1192,1339,908,933]" pageId="12" pageNumber="513" pagination="1 - 167" refId="ref14289" refString="Dagys, A. S. (1968) Jurskiye i rannemelovye brakhiopody Severa Sibiri [Jurassic and Early Cretaceous brachiopods from northern Siberia]. Akademia Nauk SSSR Sibiroskoe Otdelenie Institut Geologii I Geofiziki (IGIG) Trudy [Institute of Geology and Geophysics, Academy of Sciences of the USSR, Siberian Branch, Transactions], 41, 1 - 167. [In Russian]" type="journal article" year="1968">Dagys (1968</bibRefCitation>
</taxonomicName>
, fig. 43) and
<taxonomicName id="09C6A6A66254FFB396D541A2FDD7338B" authority="Dagys (1968" authorityName="Dagys" authorityYear="1968" box="[199,515,943,968]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="species" species="anadyrensis">
<emphasis id="FCB201376254FFB396D541A2FF01338B" box="[199,213,945,968]" italics="true" pageId="12" pageNumber="513">P</emphasis>
.
<emphasis id="FCB201376254FFB396F141BCFEBD338B" box="[227,361,943,968]" italics="true" pageId="12" pageNumber="513">anadyrensis</emphasis>
<bibRefCitation id="AA57A0D46254FFB3976341A3FDD7338B" author="Dagys, A. S." box="[369,515,943,968]" pageId="12" pageNumber="513" pagination="1 - 167" refId="ref14289" refString="Dagys, A. S. (1968) Jurskiye i rannemelovye brakhiopody Severa Sibiri [Jurassic and Early Cretaceous brachiopods from northern Siberia]. Akademia Nauk SSSR Sibiroskoe Otdelenie Institut Geologii I Geofiziki (IGIG) Trudy [Institute of Geology and Geophysics, Academy of Sciences of the USSR, Siberian Branch, Transactions], 41, 1 - 167. [In Russian]" type="journal article" year="1968">Dagys (1968</bibRefCitation>
</taxonomicName>
, fig. 39) appear closest to those of
<taxonomicName id="09C6A6A66254FFB3959C41BCFB7F338B" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[910,1195,943,968]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376254FFB3959C41BCFB7F338B" box="[910,1195,943,968]" italics="true" pageId="12" pageNumber="513">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
(
<figureCitation id="56FDC1A06254FFB392AE41BCFB2A338B" box="[1212,1278,943,968]" captionStart="FIGURE 5" captionStartId="11.[151,250,1820,1842]" captionTargetBox="[238,1340,193,1782]" captionTargetId="figure-142@11.[238,1340,193,1798]" captionTargetPageId="11" captionText="FIGURE 5. A and B, transverse serial sections from two specimens of Ptilorhynchia mclachlani sp. nov., seep 9. A) Specimen PMO 227.431, sections above third dotted line. Dotted lines delimit groups of sections drawn at the same magnification. Dimensions of specimen: L 15.3+; W 18.3; T 9.8 mm. Serial sections prepared perpendicular to maximum length of specimen; numbering of sections starts at 0.0 mm, the initial section through the specimen. B) Specimen PMO 224.879, sections below third dotted line. Dimensions of sectioned specimen: L 12.9; W 12.8+; T 9.0 mm. Serial sections prepared perpendicular to length of dorsal valve; numbering of sections starts at 0.0 mm, the initial section through the specimen. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951764" httpUri="https://zenodo.org/record/4951764/files/figure.png" pageId="12" pageNumber="513">Fig. 5</figureCitation>
) based on the development of the septalium, hinge plates, and derivation of the crura from the inner margins of the hinge plates. It should be noted that plates 2 and
<quantity id="093E70C06254FFB3943E41E4FD8C3453" box="[556,600,1015,1040]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="12" pageNumber="513" unit="in" value="5.0">5 in</quantity>
<bibRefCitation id="AA57A0D46254FFB3944D41EBFD333453" author="Dagys, A. S." box="[607,743,1015,1040]" pageId="12" pageNumber="513" pagination="1 - 167" refId="ref14289" refString="Dagys, A. S. (1968) Jurskiye i rannemelovye brakhiopody Severa Sibiri [Jurassic and Early Cretaceous brachiopods from northern Siberia]. Akademia Nauk SSSR Sibiroskoe Otdelenie Institut Geologii I Geofiziki (IGIG) Trudy [Institute of Geology and Geophysics, Academy of Sciences of the USSR, Siberian Branch, Transactions], 41, 1 - 167. [In Russian]" type="journal article" year="1968">Dagys 1968</bibRefCitation>
were printed in the wrong order. The taxonomic descriptions and caption for plate 5 refers to
<taxonomicName id="09C6A6A66254FFB397E8460EFDB53476" authorityName="Dagys" authorityYear="1968" box="[506,609,1052,1077]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="species" species="glabra">
<emphasis id="FCB201376254FFB397E8460EFDDC3477" box="[506,520,1053,1076]" italics="true" pageId="12" pageNumber="513">P</emphasis>
.
<emphasis id="FCB201376254FFB39404460FFDB53476" box="[534,609,1052,1077]" italics="true" pageId="12" pageNumber="513">glabra</emphasis>
</taxonomicName>
,
<taxonomicName id="09C6A6A66254FFB3947C460EFD253476" authorityName="Dagys" authorityYear="1968" box="[622,753,1052,1077]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="species" species="lenaensis">
<emphasis id="FCB201376254FFB3947C460EFDA83477" box="[622,636,1053,1076]" italics="true" pageId="12" pageNumber="513">P</emphasis>
.
<emphasis id="FCB201376254FFB3949B460FFD253476" box="[649,753,1052,1077]" italics="true" pageId="12" pageNumber="513">lenaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="09C6A6A66254FFB39535460EFC3C3476" box="[807,1000,1052,1077]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="species" species="obscuricostata">
<emphasis id="FCB201376254FFB39535460EFCE13477" box="[807,821,1053,1076]" italics="true" pageId="12" pageNumber="513">P</emphasis>
.
<emphasis id="FCB201376254FFB39551460FFC3C3476" box="[835,1000,1052,1077]" italics="true" pageId="12" pageNumber="513">obscuricostata</emphasis>
</taxonomicName>
. However, these species are figured on plate 2.
</paragraph>
<paragraph id="CE79DD256254FFB396D54677FCF934E3" blockId="12.[151,1437,151,1221]" pageId="12" pageNumber="513">
Serial sections of the Spitsbergen material (
<figureCitation id="56FDC1A06254FFB394C54677FCF5343E" box="[727,801,1124,1149]" captionStart="FIGURE 5" captionStartId="11.[151,250,1820,1842]" captionTargetBox="[238,1340,193,1782]" captionTargetId="figure-142@11.[238,1340,193,1798]" captionTargetPageId="11" captionText="FIGURE 5. A and B, transverse serial sections from two specimens of Ptilorhynchia mclachlani sp. nov., seep 9. A) Specimen PMO 227.431, sections above third dotted line. Dotted lines delimit groups of sections drawn at the same magnification. Dimensions of specimen: L 15.3+; W 18.3; T 9.8 mm. Serial sections prepared perpendicular to maximum length of specimen; numbering of sections starts at 0.0 mm, the initial section through the specimen. B) Specimen PMO 224.879, sections below third dotted line. Dimensions of sectioned specimen: L 12.9; W 12.8+; T 9.0 mm. Serial sections prepared perpendicular to length of dorsal valve; numbering of sections starts at 0.0 mm, the initial section through the specimen. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951764" httpUri="https://zenodo.org/record/4951764/files/figure.png" pageId="12" pageNumber="513">Fig. 5</figureCitation>
) lack the falciform crura characteristic of the genus
<taxonomicName id="09C6A6A66254FFB396854694FEF434E3" authorityName="Wisniewska" authorityYear="1932" box="[151,288,1159,1184]" class="Rhynchonellata" family="Basiliolidae" genus="Lacunosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="12" pageNumber="513" phylum="Brachiopoda" rank="genus">
<emphasis id="FCB201376254FFB396854694FEF434E3" box="[151,288,1159,1184]" italics="true" pageId="12" pageNumber="513">Lacunosella</emphasis>
</taxonomicName>
recorded from
<collectingCountry id="B6D19DB56254FFB397DF4694FD9634E3" box="[461,578,1159,1184]" name="Greenland" pageId="12" pageNumber="513">Greenland</collectingCountry>
(
<bibRefCitation id="AA57A0D46254FFB394424694FD0334E3" author="Owen, E. F." box="[592,727,1159,1184]" pageId="12" pageNumber="513" pagination="1 - 19" refId="ref17259" refString="Owen, E. F. (1976) Some Lower Cretaceous brachiopods from East Greenland. Meddelelser om GrOnland, 171 (3), 1 - 19." type="journal article" year="1976">Owen 1976</bibRefCitation>
, fig. 3).
</paragraph>
<paragraph id="CE79DD256254FFB396D546BFFBEE3486" blockId="12.[151,1437,151,1221]" box="[199,1082,1196,1221]" pageId="12" pageNumber="513">
<emphasis id="FCB201376254FFB396D546BFFD153480" box="[199,705,1196,1221]" inLineHeading="true" pageId="12" pageNumber="513" reason="1">
<emphasis id="FCB201376254FFB396D546BFFD6D3486" bold="true" box="[199,697,1196,1221]" pageId="12" pageNumber="513">Stratigraphic and geographic distribution</emphasis>
.
</emphasis>
Upper Ryazanian of Spitsbergen.
</paragraph>
</subSubSection>
</treatment>
</document>