171 lines
18 KiB
XML
171 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.870.36970" ID-GBIF-Dataset="0c6fd07e-55da-42a2-85d1-4f1068abc419" ID-PMC="PMC6694067" ID-Pensoft-Pub="1313-2970-870-51" ID-Pensoft-UUID="B9AE5951AA745D6BAF3EEB40C2061AF4" ID-PubMed="31423078" ID-ZooBank="BEC530D3DD9A466AAAC705305B20E761" ModsDocID="1313-2970-870-51" checkinTime="1565240931249" checkinUser="pensoft" docAuthor="Sutcharit, Chirasak, Backeljau, Thierry & Panha, Somsak" docDate="2019" docId="43794688E83058DE807AEAACF0B8162B" docLanguage="en" docName="ZooKeys 870: 51-76" docOrigin="ZooKeys 870" docSource="http://dx.doi.org/10.3897/zookeys.870.36970" docTitle="Ganesella capitium Benson 1848" docType="treatment" docVersion="4" id="B9AE5951AA745D6BAF3EEB40C2061AF4" lastPageNumber="51" masterDocId="B9AE5951AA745D6BAF3EEB40C2061AF4" masterDocTitle="Re-description of the type species of the genera Ganesella Blanford, 1863 and Globotrochus Haas, 1935; with description of a new Ganesella species from Thailand (Eupulmonata, Camaenidae)" masterLastPageNumber="76" masterPageNumber="51" pageNumber="51" updateTime="1668167689048" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Re-description of the type species of the genera Ganesella Blanford, 1863 and Globotrochus Haas, 1935; with description of a new Ganesella species from Thailand (Eupulmonata, Camaenidae)</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Backeljau, Thierry</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>870</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>51</mods:start>
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<mods:end>76</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.870.36970</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.870.36970</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-870-51</mods:identifier>
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<mods:identifier type="ZooBank">BEC530D3DD9A466AAAC705305B20E761</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159043269" LSID="urn:lsid:plazi:treatment:43794688E83058DE807AEAACF0B8162B" httpUri="http://treatment.plazi.org/id/43794688E83058DE807AEAACF0B8162B" lastPageNumber="51" pageId="0" pageNumber="51">
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<subSubSection pageId="0" pageNumber="51" type="nomenclature">
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName LSID="43794688-e830-58de-807a-eaacf0b8162b" authority="Benson, 1848" authorityName="Benson" authorityYear="1848" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella capitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="capitium">Ganesella capitium (Benson, 1848)</taxonomicName>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A, B Ganesella capitium A lectotype UMZC I. 102385 / 1 and B paralectotype UMZC I. 102385 / 2 - 3 C-E Ganesella hariola C lectotype UMZC I. 104370 / 1 D paralectotype UMZC I. 104370 / 2 - 4 and E shell from Popa Mountain, Myanmar CUMZ 5134 F-H Ganesella carinella F lectotype SMF 27534 a G paralectotype SMF 27534 b and H shell from Takhli, Nakhonsawan CUMZ 5133." figureDoi="10.3897/zookeys.870.36970.figure2" httpUri="https://binary.pensoft.net/fig/323907" pageId="0" pageNumber="51">Figs 2A, B</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic distribution of Ganesella spp. and Globotrochus spp." figureDoi="10.3897/zookeys.870.36970.figure8" httpUri="https://binary.pensoft.net/fig/323913" pageId="0" pageNumber="51">8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="reference_group">
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix capitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="capitium">Helix capitium</taxonomicName>
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Benson, 1848: 160.
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.14456" author="Hanley, SCT" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="51" refId="B24" refString="Hanley, SCT, Theobald, W, 1870. . https://doi.org/10.5962/bhl.title.14456" url="https://doi.org/10.5962/bhl.title.14456" year="1870">Hanley and Theobald 1870</bibRefCitation>
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: 7, pl. 14, fig. 5.
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<bibRefCitation author="Tryon, GW" journalOrPublisher="The Academy of Natural Sciences of Philadelphia, PA" pageId="0" pageNumber="51" refId="B66" refString="Tryon, GW, 1887. Manual of Conchology, Structure and Systematic, with Illustrations of the Species, Series 2 volume 3. . The Academy of Natural Sciences of Philadelphia, PA" title="Manual of Conchology, Structure and Systematic, with Illustrations of the Species, Series 2 volume 3." year="1887">Tryon 1887</bibRefCitation>
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: 74, pl. 14, fig. 99.
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</paragraph>
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Planispira) capitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="capitium" subGenus="Planispira">Helix (Planispira) capitium</taxonomicName>
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:
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="51" refId="B38" refString="Nevill, G, 1878. . https://doi.org/10.5962/bhl.title.11957" url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
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: 78.
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</paragraph>
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName class="Gastropoda" family="Camaenidae" genus="Trochomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trochomorpha capitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="capitium">Trochomorpha capitium</taxonomicName>
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:
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<bibRefCitation author="Morlet, L" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="51" pagination="121 - 199" refId="B36" refString="Morlet, L, 1889. Catalogue des coquilles recueillies, par M. Pavie dans le Cambodge et le Royaume de Siam, et description ďespeces nouvelles (1). . Journal de Conchyliologie 37: 121 - 199" title="Catalogue des coquilles recueillies, par M. Pavie dans le Cambodge et le Royaume de Siam, et description ďespeces nouvelles (1)." volume="37" year="1889">Morlet 1889</bibRefCitation>
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: 124, 125.
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</paragraph>
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella capitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="capitium">Ganesella capitium</taxonomicName>
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:
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<bibRefCitation author="Pilsbry, HA" journalOrPublisher="The Academy of Natural Sciences of Philadelphia, PA" pageId="0" pageNumber="51" refId="B48" refString="Pilsbry, HA, 1895. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Series 2 volume 9. . The Academy of Natural Sciences of Philadelphia, PA" title="Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Series 2 volume 9." year="1895">Pilsbry 1895</bibRefCitation>
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: 170, pl. 55, fig. 18.
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<bibRefCitation DOI="https://doi.org/10.1093/oxfordjournals.mollus.a065982" author="Blanford, WT" journalOrPublisher="Proceeding of the Malacological Society of London" pageId="0" pageNumber="51" pagination="274 - 284" refId="B11" refString="Blanford, WT, 1903. Note on Mr. W.M. Daly's collections of land and freshwater Mollusca from Siam. . Proceeding of the Malacological Society of London 5: 274 - 284" title="Note on Mr. W. M. Daly's collections of land and freshwater Mollusca from Siam." url="https://doi.org/10.1093/oxfordjournals.mollus.a065982" volume="5" year="1903">Blanford 1903</bibRefCitation>
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: 278.
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12891" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="51" refId="B22" refString="Gude, GK, 1914. Mollusca II (Trochomorphidae-Janellidae). In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, Including Ceylon and Burma. . Taylor and Francis, London" title="Mollusca II (Trochomorphidae-Janellidae). In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, Including Ceylon and Burma." url="https://doi.org/10.5962/bhl.title.12891" year="1914">Gude 1914</bibRefCitation>
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: 196, 197.
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<bibRefCitation pageId="0" pageNumber="51" refId="B71">Zilch 1960</bibRefCitation>
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: 610, fig. 2140.
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<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="51" pagination="197 - 223" refId="B72" refString="Zilch, A, 1966. Die Typen und Typoide des Natur-Museums Senckenberg 34: Mollusca, Camaenidae (4). . Archiv fuer Molluskenkunde 95: 197 - 223" title="Die Typen und Typoide des Natur-Museums Senckenberg 34: Mollusca, Camaenidae (4)." volume="95" year="1966">Zilch 1966</bibRefCitation>
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: 202.
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<bibRefCitation author="Richardson, L" journalOrPublisher="Tryonia" pageId="0" pageNumber="51" pagination="1 - 479" refId="B52" refString="Richardson, L, 1985. Camaenidae: Catalogue of Species. . Tryonia 12: 1 - 479" title="Camaenidae: Catalogue of Species." volume="12" year="1985">Richardson 1985</bibRefCitation>
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: 132, 133.
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</paragraph>
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName class="Gastropoda" family="Camaenidae" genus="Darwininitium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Darwininitium shiwalikianum" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="shiwalikianum">Darwininitium shiwalikianum</taxonomicName>
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Budha & Mordan in
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.175.2755" author="Budha, PB" journalOrPublisher="ZooKeys" pageId="0" pageNumber="51" pagination="19 - 26" refId="B12" refString="Budha, PB, Mordan, PB, Naggs, F, Backeljau, T, 2012. Darwininitium - a new fully pseudosigmurethrous orthurethran genus from Nepal (Gastropoda: Pulmonata: Cerastidae). . ZooKeys 175: 19 - 26" title="Darwininitium - a new fully pseudosigmurethrous orthurethran genus from Nepal (Gastropoda: Pulmonata: Cerastidae)." url="https://doi.org/10.3897/zookeys.175.2755" volume="175" year="2012">Budha et al. 2012</bibRefCitation>
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: 21-23, figs 2-4. Type locality: Kasara near Tamor Lake, Chitwan National Park, Central Nepal. New synonym.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="type locality">
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<paragraph pageId="0" pageNumber="51">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="51">Sicrigali province Bahar Indiae Orientalis [Bihar State, India].</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="materials_examined">
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<paragraph pageId="0" pageNumber="51">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="51">
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<emphasis bold="true" pageId="0" pageNumber="51">Type specimens.</emphasis>
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Three syntypes of
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix capitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="capitium">
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<emphasis italics="true" pageId="0" pageNumber="51">Helix capitium</emphasis>
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</taxonomicName>
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are in
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<normalizedToken originalValue="Benson’s">Benson's</normalizedToken>
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collection. The specimen that closely matched with the measurement in the original description is designated here as the lectotype UMZC I.102385/1 (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A, B Ganesella capitium A lectotype UMZC I. 102385 / 1 and B paralectotype UMZC I. 102385 / 2 - 3 C-E Ganesella hariola C lectotype UMZC I. 104370 / 1 D paralectotype UMZC I. 104370 / 2 - 4 and E shell from Popa Mountain, Myanmar CUMZ 5134 F-H Ganesella carinella F lectotype SMF 27534 a G paralectotype SMF 27534 b and H shell from Takhli, Nakhonsawan CUMZ 5133." figureDoi="10.3897/zookeys.870.36970.figure2" httpUri="https://binary.pensoft.net/fig/323907" pageId="0" pageNumber="51">Fig. 2A</figureCitation>
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, height 13.3 mm, width 13.5 mm), and the other two as paralectotypes UMZC I.102385/2-3 (2 shells;
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A, B Ganesella capitium A lectotype UMZC I. 102385 / 1 and B paralectotype UMZC I. 102385 / 2 - 3 C-E Ganesella hariola C lectotype UMZC I. 104370 / 1 D paralectotype UMZC I. 104370 / 2 - 4 and E shell from Popa Mountain, Myanmar CUMZ 5134 F-H Ganesella carinella F lectotype SMF 27534 a G paralectotype SMF 27534 b and H shell from Takhli, Nakhonsawan CUMZ 5133." figureDoi="10.3897/zookeys.870.36970.figure2" httpUri="https://binary.pensoft.net/fig/323907" pageId="0" pageNumber="51">Fig. 2B</figureCitation>
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, height 14.4 mm, width 13.8 mm).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="materials_examined">
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<paragraph pageId="0" pageNumber="51">Additional material.</paragraph>
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<paragraph pageId="0" pageNumber="51">Bahar Province, India: ZMB ex. Albers coll. 1 lot (1 shell). South India: NHMUK ex. Godwin-Austen coll. no. 501 (2 shells).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="remarks">
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<paragraph pageId="0" pageNumber="51">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="51">
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.175.2755" author="Budha, PB" journalOrPublisher="ZooKeys" pageId="0" pageNumber="51" pagination="19 - 26" refId="B12" refString="Budha, PB, Mordan, PB, Naggs, F, Backeljau, T, 2012. Darwininitium - a new fully pseudosigmurethrous orthurethran genus from Nepal (Gastropoda: Pulmonata: Cerastidae). . ZooKeys 175: 19 - 26" title="Darwininitium - a new fully pseudosigmurethrous orthurethran genus from Nepal (Gastropoda: Pulmonata: Cerastidae)." url="https://doi.org/10.3897/zookeys.175.2755" volume="175" year="2012">Budha et al. (2012)</bibRefCitation>
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described
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<taxonomicName class="Gastropoda" family="Camaenidae" genus="Darwininitium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Darwininitium shiwalikianum" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="shiwalikianum">
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<emphasis italics="true" pageId="0" pageNumber="51">Darwininitium shiwalikianum</emphasis>
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</taxonomicName>
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from Chitwan National Park, Nepal. The holotype has a trochoid, brownish shell with irregular opaque white spots. As such, it appears to be identical to the type specimens of
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<taxonomicName class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella capitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="capitium">
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<emphasis italics="true" pageId="0" pageNumber="51">Ganesella capitium</emphasis>
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</taxonomicName>
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. In addition, the type localities of
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<taxonomicName lsidName="D. shiwalikianum" pageId="0" pageNumber="51" rank="species" species="shiwalikianum">
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<emphasis italics="true" pageId="0" pageNumber="51">D. shiwalikianum</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="G. capitium" pageId="0" pageNumber="51" rank="species" species="capitium">
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<emphasis italics="true" pageId="0" pageNumber="51">G. capitium</emphasis>
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</taxonomicName>
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are geographically quite close to each other. Hence, in line with our earlier conclusions about the genus
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<taxonomicName class="Gastropoda" family="Camaenidae" genus="Darwininitium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Darwininitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="51">Darwininitium</emphasis>
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</taxonomicName>
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, we tentatively regard
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<taxonomicName lsidName="D. shiwalikianum" pageId="0" pageNumber="51" rank="species" species="shiwalikianum">
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<emphasis italics="true" pageId="0" pageNumber="51">D. shiwalikianum</emphasis>
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</taxonomicName>
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as a junior subjective synonym of
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<taxonomicName lsidName="G. capitium" pageId="0" pageNumber="51" rank="species" species="capitium">
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<emphasis italics="true" pageId="0" pageNumber="51">G. capitium</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="51">
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Shell and genitalia have been described in detail by
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.175.2755" author="Budha, PB" journalOrPublisher="ZooKeys" pageId="0" pageNumber="51" pagination="19 - 26" refId="B12" refString="Budha, PB, Mordan, PB, Naggs, F, Backeljau, T, 2012. Darwininitium - a new fully pseudosigmurethrous orthurethran genus from Nepal (Gastropoda: Pulmonata: Cerastidae). . ZooKeys 175: 19 - 26" title="Darwininitium - a new fully pseudosigmurethrous orthurethran genus from Nepal (Gastropoda: Pulmonata: Cerastidae)." url="https://doi.org/10.3897/zookeys.175.2755" volume="175" year="2012">Budha et al. (2012)</bibRefCitation>
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. The unique and distinctive characters of
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<taxonomicName lsidName="G. capitium" pageId="0" pageNumber="51" rank="species" species="capitium">
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<emphasis italics="true" pageId="0" pageNumber="51">G. capitium</emphasis>
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</taxonomicName>
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are its small, relatively elevated, trochoid shell. Aperture open sublaterally. Whorls slightly convex with wide and shallow suture. Last whorl angular with weak peripheral keel. Shell colour brownish with whitish-opaque, irregular, spots or streaks. Genitalia with short atrium; cylindrical, short penis, about half as long as the vagina, and proximally with blackish, spongy tissue. Epiphallus and flagellum together short, about as long as the penis, but flagellum longer than epiphallus. Internal wall of penis with numerous longitudinal pilasters.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |