406 lines
58 KiB
XML
406 lines
58 KiB
XML
<document id="ED692FBE477FA3130DC553C41A4C1017" ID-DOI="10.1080/00222930903094621" ID-ISSN="1464-5262" ID-Zenodo-Dep="5217087" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629293244201" checkinUser="felipe" docAuthor="Yanes, Yurena, Martín, Javier, Moro, Leopoldo, Alonso, María R. & Ibáñez, Miguel" docDate="2009" docId="4014878B4565793240D08030FE39FB98" docLanguage="en" docName="JNatHist.43.35-36.2179-2207.pdf" docOrigin="Journal of Natural History 43 (35 - 36)" docSource="http://dx.doi.org/10.1080/00222930903094621" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Napaeus tafadaensis Yanes & Martín & Moro & Alonso & Ibáñez 2009, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2200" masterDocId="BC2DFFF345777925413C8541FFAFFF94" masterDocTitle="On the relationships of the genus Napaeus (Gastropoda: Pulmonata: Enidae) with the descriptions of four new species from the Canary Islands" masterLastPageNumber="2207" masterPageNumber="2179" pageNumber="2195" updateTime="1699067505050" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="18B2650A76A3742841BCA2AD930B9118" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5C7D9F9A4F3D2A7F0F43102ED7D0C563">
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<mods:title id="D8A9DD5749BAA7EBC42D4DF6382A7718">On the relationships of the genus Napaeus (Gastropoda: Pulmonata: Enidae) with the descriptions of four new species from the Canary Islands</mods:title>
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<mods:name id="21B520CFDE6BF99D8512E4A982233E6E" type="personal">
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<mods:namePart id="7C7F7416BD658CA6749EFC7D8966BA4E">Yanes, Yurena</mods:namePart>
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</mods:name>
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<mods:roleTerm id="A9AD05F8939560C7EF57638DB09B04B3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C823A197A90C9F638DC596AB3EFF1FA0">Martín, Javier</mods:namePart>
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</mods:name>
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<mods:roleTerm id="A44DCAB07541FE3F6FED5305F40D3066">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D7E8373F81D35DB5A50BB089513BA9FB">Moro, Leopoldo</mods:namePart>
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</mods:name>
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<mods:roleTerm id="0740F4EDD9CAFCDCD5BD70C1ACDAD0A2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6383606FE5959402D891101378C0B3FF">Alonso, María R.</mods:namePart>
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</mods:name>
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<mods:name id="AB096C46726B92B0A3B958C042683510" type="personal">
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<mods:roleTerm id="75CB6D0427465F364010BFC9215D0C0A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AD5C0107A44734D7291B31D3338EAA6F">Ibáñez, Miguel</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8D831FA77BEAED299A9F1985A5F556CE">text</mods:typeOfResource>
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<mods:title id="DECF7CEB7E4B414010F7C689CEFCE7A4">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="68A23428687B164D75D32623236E7F5F">
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<mods:date id="F994EBA991E637E5AB4F95E92570963E">2009</mods:date>
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<mods:detail id="222B64CDD7F4CD8FDF7DF289B78F163F" type="pubDate">
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<mods:number id="517C687F17CD4210D7EAA9CBF6AE3000">2009-08-26</mods:number>
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<mods:detail id="B7432462A2EA618804AAABCA80676BA7" type="volume">
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<mods:number id="ED4D8D5F18DC9B6401C1C9427E4EA27C">43</mods:number>
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<mods:detail id="0917B9A17D23BF082A4B4F1CA0B69340" type="issue">
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<mods:number id="4591A230A11DD23A2418D1A348924F61">35 - 36</mods:number>
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<mods:start id="053D9C5857F0FBE1CB5BAA9E324B3A33">2179</mods:start>
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<mods:end id="D8DD3618627184564D56F27826FA5B38">2207</mods:end>
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<mods:url id="35BB4A2DDEAEE742F7680F47F4B70498">http://dx.doi.org/10.1080/00222930903094621</mods:url>
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<mods:classification id="169FE6896840EB0777F8C4BCFA2866DD">journal article</mods:classification>
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<mods:identifier id="388D5589DD60EFC63807BDBAC2D5EFE8" type="DOI">10.1080/00222930903094621</mods:identifier>
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<mods:identifier id="741D63B6E6351BECCD11B86F38EAFFAC" type="ISSN">1464-5262</mods:identifier>
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<treatment id="4014878B4565793240D08030FE39FB98" LSID="urn:lsid:plazi:treatment:4014878B4565793240D08030FE39FB98" httpUri="http://treatment.plazi.org/id/4014878B4565793240D08030FE39FB98" lastPageId="23" lastPageNumber="2200" pageId="18" pageNumber="2195">
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<subSubSection id="80A765164565793740D08030FCD4FA1D" box="[492,891,1393,1417]" pageId="18" pageNumber="2195" type="nomenclature">
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<paragraph id="C802369D4565793740D08030FCD4FA1D" blockId="18.[492,891,1393,1417]" box="[492,891,1393,1417]" pageId="18" pageNumber="2195">
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<heading id="934A81F14565793740D08030FCD4FA1D" box="[492,891,1393,1417]" centered="true" fontSize="36" level="2" pageId="18" pageNumber="2195" reason="7">
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<taxonomicName id="0FBD4D1E4565793740D08030FCB3FA1D" ID-CoL="45NR3" authority="Yanes" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[492,796,1393,1417]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="2195" phylum="Mollusca" rank="species" species="tafadaensis" status="sp. nov.">
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<emphasis id="FAC9EA8F4565793740D08030FD60FA1D" bold="true" box="[492,719,1393,1417]" italics="true" pageId="18" pageNumber="2195">Napaeus tafadaensis</emphasis>
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Yanes
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</taxonomicName>
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<taxonomicNameLabel id="E1FA57F445657937421E8033FCD4FA1D" box="[802,891,1394,1417]" pageId="18" pageNumber="2195" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="80A765164565793241E880F8FE39FB98" lastPageId="23" lastPageNumber="2200" pageId="18" pageNumber="2195" type="materials_examined">
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<paragraph id="C802369D4565793741E880F8FEC0FA45" blockId="18.[212,367,1465,1489]" box="[212,367,1465,1489]" pageId="18" pageNumber="2195">
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<emphasis id="FAC9EA8F4565793741E880F8FEC0FA45" box="[212,367,1465,1489]" italics="true" pageId="18" pageNumber="2195">Type material</emphasis>
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</paragraph>
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||
<paragraph id="C802369D4565793741E880A0FE45F9AD" blockId="18.[212,1172,1505,1689]" pageId="18" pageNumber="2195">
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<materialsCitation id="78D53CC04565793741E880A0FE45F9AD" collectingDate="1991-04-06" collectionCode="UTM" collectorName="Leg. M. Ibanez" elevation="600" location="Anaga" municipality="Tafada" pageId="18" pageNumber="2195" specimenCode="MT 0398, RCS8761" specimenCount="1" stateProvince="Montana" typeStatus="holotype">
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<typeStatus id="1706883F4565793741E880A0FE90FA6D" box="[212,319,1505,1529]" pageId="18" pageNumber="2195" type="holotype">Holotype</typeStatus>
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||
(
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<figureCitation id="50862A1845657937406F80A0FE74FA6D" box="[339,475,1505,1529]" captionStart="Figure 3" captionStartId="13.[144,211,1424,1446]" captionTargetBox="[150,1096,172,1387]" captionTargetId="figure-92@13.[144,1104,167,1394]" captionTargetPageId="13" captionText="Figure 3. Napaeus shell drawings. (A) N. consecoanus; (B) N. bertheloti; (C) N. subsimplex; (D) N. savinosa (syntype, NHM 1895.2.2.223); (E) N. inflatiusculus; (F) N. severus; (G) N. baeticatus; (H) N. baeticatus morphotype subbeticatus; (J) N. baeticatus morphotype halmyris; (K) N. interpunctatus; (L) N. servus; (M) N. tabidus; (N) N. tafadaensis sp. nov. (holotype, TFMC MT 0398); (O) N. tafadaensis sp. nov. morphotype “irregular” (N. tafadaensis paratype, AIT); (P) N. badiosus." figureDoi="http://doi.org/10.5281/zenodo.5217095" httpUri="https://zenodo.org/record/5217095/files/figure.png" pageId="18" pageNumber="2195">Figures 3N</figureCitation>
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,
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<figureCitation id="50862A184565793740D080A0FDA0FA6D" box="[492,527,1505,1529]" captionStart="Figure 7" captionStartId="17.[144,211,1168,1190]" captionTargetBox="[144,1104,168,1139]" captionTargetId="figure-197@17.[144,1104,168,1139]" captionTargetPageId="17" captionText="Figure 7. Holotype shells (A–D) and a specimen (E). (A) N. bajamarensis sp. nov.; (B) N. tafadaensis sp. nov.; (C) N. ornamentatus sp. nov.; (D) N. teobaldoi sp. nov.; (E) N. ornamentatus, paratype from Mirador de Taguluche (L. Moro photograph)." figureDoi="http://doi.org/10.5281/zenodo.5217105" httpUri="https://zenodo.org/record/5217105/files/figure.png" pageId="18" pageNumber="2195">7B</figureCitation>
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): TFMC (
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<specimenCode id="981B9EE64565793743B180A0FCAFFA6D" box="[653,768,1505,1529]" collectionCode="MT" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="18" pageNumber="2195">MT 0398</specimenCode>
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||
). Type locality:
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<collectingRegion id="0A79F87F4565793742F980A3FB80FA6D" box="[965,1071,1506,1529]" country="United States of America" name="Montana" pageId="18" pageNumber="2195">Montaña</collectingRegion>
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<collectingMunicipality id="2866ACE745657937450780A0FB20FA6D" box="[1083,1167,1505,1529]" pageId="18" pageNumber="2195">Tafada</collectingMunicipality>
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,
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<location id="CD6260464565793741E88340FEB0F98D" LSID="urn:lsid:plazi:treatment:4014878B4565793240D08030FE39FB98:CD6260464565793741E88340FEB0F98D" box="[212,287,1537,1561]" municipality="Tafada" name="Anaga" pageId="18" pageNumber="2195" stateProvince="Montana">Anaga</location>
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massif (Tenerife;
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<collectionCode id="AEACAE5845657937433B8343FDE6F98D" box="[519,585,1538,1561]" pageId="18" pageNumber="2195">UTM</collectionCode>
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||
coordinates: 28
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||
<specimenCode id="981B9EE645657937422E8340FCD0F98D" box="[786,895,1537,1561]" collectionCode="RCS" country="United Kingdom" httpUri="http://grbio.org/cool/bha5-99ng" name="Royal College of Surgeons" pageId="18" pageNumber="2195">RCS8761</specimenCode>
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||
;
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||
<quantity id="0F459B784565793742A88340FC76F98D" box="[916,985,1537,1561]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="2195" unit="m" value="600.0">
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||
<elevation id="4390D1AE4565793742A88340FC76F98D" box="[916,985,1537,1561]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="2195" unit="m" value="600.0">600 m</elevation>
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</quantity>
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altitude).
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<collectorName id="6548534B45657937455D8343FEE4F9AD" pageId="18" pageNumber="2195">Leg. M. Ibáñez</collectorName>
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,
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<date id="BC03105D45657937406A8360FE49F9AD" box="[342,486,1569,1593]" pageId="18" pageNumber="2195" value="1991-04-06">
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||
<collectingDate id="AC47E9B545657937406A8360FE49F9AD" box="[342,486,1569,1593]" pageId="18" pageNumber="2195" value="1991-04-06">6 April 1991</collectingDate>
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</date>
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||
.
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="C802369D4565793741C28303FC2DF90D" blockId="18.[212,1172,1505,1689]" pageId="18" pageNumber="2195">
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<materialsCitation id="78D53CC04565793741C28303FCD1F90D" collectionCode="UTM, CML" location="Tenerife" pageId="18" pageNumber="2195" specimenCode="RCS7859, 28, RCS8356, RCS8761, MT 0134" specimenCount="8" typeStatus="paratype">
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<typeStatus id="1706883F4565793741C28303FEDDF9CE" box="[254,370,1602,1626]" pageId="18" pageNumber="2195" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="DEBBFD144565793740418300FDABF9CD" box="[381,516,1601,1625]" pageId="18" pageNumber="2195" type="generic">8 specimens</specimenCount>
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and 11 shells, collected between 1991 and 2008, from the northeast of
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<location id="CD62604645657937405F8320FE6EF9ED" LSID="urn:lsid:plazi:treatment:4014878B4565793240D08030FE39FB98:CD62604645657937405F8320FE6EF9ED" box="[355,449,1633,1657]" name="Tenerife" pageId="18" pageNumber="2195">Tenerife</location>
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||
(
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<collectionCode id="AEACAE584565793740F38320FDBDF9ED" box="[463,530,1633,1657]" pageId="18" pageNumber="2195">UTM</collectionCode>
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coordinates: 28
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||
<specimenCode id="981B9EE64565793743FA8320FC9BF9ED" box="[710,820,1633,1657]" collectionCode="RCS" country="United Kingdom" httpUri="http://grbio.org/cool/bha5-99ng" name="Royal College of Surgeons" pageId="18" pageNumber="2195">RCS7859</specimenCode>
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,
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<specimenCode id="981B9EE64565793742038320FCF5F9ED" box="[831,858,1633,1657]" collectionCode="RCS" country="United Kingdom" httpUri="http://grbio.org/cool/bha5-99ng" name="Royal College of Surgeons" pageId="18" pageNumber="2195">28</specimenCode>
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||
<specimenCode id="981B9EE64565793742668320FC69F9ED" box="[858,966,1633,1657]" collectionCode="RCS" country="United Kingdom" httpUri="http://grbio.org/cool/bha5-99ng" name="Royal College of Surgeons" pageId="18" pageNumber="2195">RCS8356</specimenCode>
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and 28
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<specimenCode id="981B9EE64565793745258320FB27F9ED" box="[1049,1160,1633,1657]" collectionCode="RCS" country="United Kingdom" httpUri="http://grbio.org/cool/bha5-99ng" name="Royal College of Surgeons" pageId="18" pageNumber="2195">RCS8761</specimenCode>
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||
), deposited in
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<collectionCode id="AEACAE5845657937405F83C0FE0DF90D" box="[355,418,1665,1689]" country="0" httpUri="http://grbio.org/cool/ib7f-dbfe" name="Universidad Nacional de Tucuman, Coleccion de Mamiferos Lillo (Argentina)" pageId="18" pageNumber="2195">CML</collectionCode>
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(one shell), TFMC (
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<specimenCode id="981B9EE64565793743B783C0FD59F90D" box="[651,758,1665,1689]" collectionCode="MT" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="18" pageNumber="2195">MT 0134</specimenCode>
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||
/2) and AIT
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<caption id="9CC266154564793641AC812AFEB7FAA5" ID-DOI="http://doi.org/10.5281/zenodo.5217109" ID-Zenodo-Dep="5217109" httpUri="https://zenodo.org/record/5217109/files/figure.png" pageId="19" pageNumber="2196" startId="19.[144,211,1131,1153]" targetBox="[144,1104,168,1101]" targetPageId="19">
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||
<paragraph id="C802369D4564793641AC812AFEB7FAA5" blockId="19.[144,1104,1131,1329]" pageId="19" pageNumber="2196">
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Figure 9.
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<taxonomicName id="0FBD4D1E4564793641C0812AFE07FB15" authority="Yanes & Martín & Moro & Alonso & Ibáñez, 2009" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[252,424,1131,1153]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="2196" phylum="Mollusca" rank="species" species="bajamarensis" status="sp. nov.">
|
||
<emphasis id="FAC9EA8F4564793641C0812AFE07FB15" box="[252,424,1131,1153]" italics="true" pageId="19" pageNumber="2196">N. bajamarensis</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="E1FA57F445647936408F812DFDA1FB15" box="[435,526,1132,1153]" pageId="19" pageNumber="2196" rank="species">sp. nov.</taxonomicNameLabel>
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||
, paratypes from Barranco de Vargas, Bajamar. (A) Genital system; (B) inner anatomy of distal epiphallus and proximal penis, showing the penial papilla. A
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||
<subScript id="543934D845647936406D81F1FEF5FB54" attach="left" box="[337,346,1200,1216]" fontSize="7" pageId="19" pageNumber="2196">1</subScript>
|
||
–A
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||
<subScript id="543934D845647936404681F1FE2CFB54" attach="left" box="[378,387,1200,1216]" fontSize="7" pageId="19" pageNumber="2196">5</subScript>
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||
, parts of the penial appendix; ag, albumen gland; bc, bursa copulatrix; bd, bursa copulatrix duct; d, diverticulum; dp, distal penis; e, epiphallus; ec, epiphallar caecum; f, flagellum; go, genital orifice; o, free oviduct; p pa, penial papilla; pp, proximal penis; pr, penis retractor; r, retractor muscles; t, body tegument; v, vagina; vd, vas deferens.
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</paragraph>
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</caption>
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<paragraph id="C802369D4564793641AC80CAFEA6FA37" blockId="19.[144,265,1419,1443]" box="[144,265,1419,1443]" pageId="19" pageNumber="2196">
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||
<emphasis id="FAC9EA8F4564793641AC80CAFEA6FA37" box="[144,265,1419,1443]" italics="true" pageId="19" pageNumber="2196">Etymology</emphasis>
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||
</paragraph>
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||
<paragraph id="C802369D4564793641AC80F2FCABFA5F" blockId="19.[144,772,1459,1484]" box="[144,772,1459,1484]" pageId="19" pageNumber="2196">
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The specific name derives from that of the
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<typeStatus id="1706883F45647936434E80F5FD0CFA58" box="[626,675,1460,1484]" pageId="19" pageNumber="2196">type</typeStatus>
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locality.
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</paragraph>
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<paragraph id="C802369D4564793641AC80BAFBE0F9AF" blockId="19.[144,470,1531,1554]" lastBlockId="19.[144,1104,1566,1691]" pageId="19" pageNumber="2196">
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||
<emphasis id="FAC9EA8F4564793641AC80BAFE79F987" box="[144,470,1531,1555]" italics="true" pageId="19" pageNumber="2196">Geographic range and habitat</emphasis>
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Ground dweller endemic to Tenerife, present in about
|
||
<quantity id="0F459B784564793642348362FCFFF9AF" box="[776,848,1571,1595]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="2196" unit="km" value="25.0">25 km</quantity>
|
||
2 area along the north
|
||
</paragraph>
|
||
<paragraph id="C802369D4564793641AC8302FECBF90F" blockId="19.[144,1104,1566,1691]" pageId="19" pageNumber="2196">
|
||
and south facing slopes of the Anaga massif (northeast of the island), associated with lowland vegetation and some subtropical scrub, ranging in altitude between 140 and
|
||
<quantity id="0F459B784564793641AC83C2FF7BF90F" box="[144,212,1667,1691]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="2196" unit="m" value="600.0">600 m</quantity>
|
||
(
|
||
<figureCitation id="50862A184564793641DE83C2FEF9F90F" box="[226,342,1667,1691]" captionStart="Figure 1" captionStartId="11.[144,211,1524,1546]" captionTargetBox="[144,1104,173,1488]" captionTargetId="figure-84@11.[144,1104,173,1488]" captionTargetPageId="11" captionText="Figure 1. (A) Geographical location of the Azores, Madeira and the Canaries; arrows indicate possible colonization patterns used by land snails; position of sea mounts comes from Van den Broeck et al. 2008; (B) geographical distribution of the Napaeus species; symbols represent 1×1 km Universal Transverse Mercator squares; arrow points to the Pleistocene deposit from Mancha de La Laja-Bajamar, where the oldest Napaeus species (N. lajaensis) has been collected (Castillo et al., 2006)." figureDoi="http://doi.org/10.5281/zenodo.5217089" httpUri="https://zenodo.org/record/5217089/files/figure.png" pageId="19" pageNumber="2196">Figure 1B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="9CC266154563793141E8812CFE7EFB6D" ID-DOI="http://doi.org/10.5281/zenodo.5217111" ID-Zenodo-Dep="5217111" httpUri="https://zenodo.org/record/5217111/files/figure.png" pageId="20" pageNumber="2197" startId="20.[212,279,1133,1155]" targetBox="[212,1172,168,1103]" targetPageId="20">
|
||
<paragraph id="C802369D4563793141E8812CFE7EFB6D" blockId="20.[212,1172,1133,1273]" pageId="20" pageNumber="2197">
|
||
Figure 10. Holotype shell ornamentation details of the penultimate whorl (A, B, D, E) and that of the body whorl columellar zone, near the peristome (C). (A)
|
||
<taxonomicName id="0FBD4D1E4563793142AF81CAFB9AFB35" authority="Yanes & Martín & Moro & Alonso & Ibáñez, 2009" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[915,1077,1163,1185]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="2197" phylum="Mollusca" rank="species" species="bajamarensis" status="sp. nov.">
|
||
<emphasis id="FAC9EA8F4563793142AF81CAFB9AFB35" box="[915,1077,1163,1185]" italics="true" pageId="20" pageNumber="2197">N. bajamarensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E1FA57F445637931450081CDFB3FFB35" box="[1084,1168,1164,1185]" pageId="20" pageNumber="2197" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (B–C)
|
||
<taxonomicName id="0FBD4D1E45637931402B81E9FE0AFB2A" authority="Yanes & Martín & Moro & Alonso & Ibáñez, 2009" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[279,421,1192,1214]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="2197" phylum="Mollusca" rank="species" species="tafadaensis" status="sp. nov.">
|
||
<emphasis id="FAC9EA8F45637931402B81E9FE0AFB2A" box="[279,421,1192,1214]" italics="true" pageId="20" pageNumber="2197">N. tafadaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E1FA57F445637931409081E8FE50FB2A" box="[428,511,1193,1214]" pageId="20" pageNumber="2197" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (D)
|
||
<taxonomicName id="0FBD4D1E45637931430E81E9FD76FB2A" authority="Yanes & Martín & Moro & Alonso & Ibáñez, 2009" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[562,729,1192,1214]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="2197" phylum="Mollusca" rank="species" species="ornamentatus" status="sp. nov.">
|
||
<emphasis id="FAC9EA8F45637931430E81E9FD76FB2A" box="[562,729,1192,1214]" italics="true" pageId="20" pageNumber="2197">N. ornamentatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E1FA57F44563793143E381E8FC9DFB2A" box="[735,818,1193,1214]" pageId="20" pageNumber="2197" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (E)
|
||
<taxonomicName id="0FBD4D1E45637931425E81E9FC73FB2A" authority="Yanes & Martín & Moro & Alonso & Ibáñez, 2009" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[866,988,1192,1214]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="2197" phylum="Mollusca" rank="species" species="teobaldoi" status="sp. nov.">
|
||
<emphasis id="FAC9EA8F45637931425E81E9FC73FB2A" box="[866,988,1192,1214]" italics="true" pageId="20" pageNumber="2197">N. teobaldoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E1FA57F44563793142DF81E8FB99FB2A" box="[995,1078,1193,1214]" pageId="20" pageNumber="2197" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (F) shell ornamentation detail of the penultimate whorl of
|
||
<taxonomicName id="0FBD4D1E4563793143E38187FCFAFB48" baseAuthorityName="Webb and Berthelot" baseAuthorityYear="1833" box="[735,853,1222,1244]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="2197" phylum="Mollusca" rank="species" species="badiosus">
|
||
<emphasis id="FAC9EA8F4563793143E38187FCFAFB48" box="[735,853,1222,1244]" italics="true" pageId="20" pageNumber="2197">N. badiosus</emphasis>
|
||
</taxonomicName>
|
||
, from Barranco de Valle Seco (Tenerife); pt, peristome.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C802369D4563793141E88001FEEEFACC" blockId="20.[212,321,1344,1368]" box="[212,321,1344,1368]" pageId="20" pageNumber="2197">
|
||
<emphasis id="FAC9EA8F4563793141E88001FEEEFACC" box="[212,321,1344,1368]" italics="true" pageId="20" pageNumber="2197">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="C802369D4563793141E88029FBFEFA54" blockId="20.[212,1171,1384,1472]" pageId="20" pageNumber="2197">Shell elongated oval, weakly ornamented, sometimes with an irregular growth mode, tan brown coloured, with first whorls slightly darker than the rest. Epiphallus with a well-developed epiphallar caecum. Bursa copulatrix duct without diverticulum.</paragraph>
|
||
<paragraph id="C802369D4563793141E880B1FEFBF99C" blockId="20.[212,340,1520,1544]" box="[212,340,1520,1544]" pageId="20" pageNumber="2197">
|
||
<emphasis id="FAC9EA8F4563793141E880B1FEFBF99C" box="[212,340,1520,1544]" italics="true" pageId="20" pageNumber="2197">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="C802369D4563793141E88358FD6EF9C4" blockId="20.[212,1172,1560,1680]" pageId="20" pageNumber="2197">Two morphotypes of the species displayed small but clear conchological differences while the genital system remained the same.</paragraph>
|
||
<paragraph id="C802369D4563793041C28319FDF6F9A8" blockId="20.[212,1172,1560,1680]" lastBlockId="21.[144,1104,1348,1692]" lastPageId="21" lastPageNumber="2198" pageId="20" pageNumber="2197">
|
||
Typical form (
|
||
<figureCitation id="50862A184563793140928319FD80F9E4" box="[430,559,1624,1648]" captionStart="Figure 3" captionStartId="13.[144,211,1424,1446]" captionTargetBox="[150,1096,172,1387]" captionTargetId="figure-92@13.[144,1104,167,1394]" captionTargetPageId="13" captionText="Figure 3. Napaeus shell drawings. (A) N. consecoanus; (B) N. bertheloti; (C) N. subsimplex; (D) N. savinosa (syntype, NHM 1895.2.2.223); (E) N. inflatiusculus; (F) N. severus; (G) N. baeticatus; (H) N. baeticatus morphotype subbeticatus; (J) N. baeticatus morphotype halmyris; (K) N. interpunctatus; (L) N. servus; (M) N. tabidus; (N) N. tafadaensis sp. nov. (holotype, TFMC MT 0398); (O) N. tafadaensis sp. nov. morphotype “irregular” (N. tafadaensis paratype, AIT); (P) N. badiosus." figureDoi="http://doi.org/10.5281/zenodo.5217095" httpUri="https://zenodo.org/record/5217095/files/figure.png" pageId="20" pageNumber="2197">Figure 3N</figureCitation>
|
||
): shell obese (SB/SH index), elongated oval, coneshaped above, 6¼–6½ convex whorls, deeply marked suture (
|
||
<figureCitation id="50862A184563793142BA8339FC53F904" box="[902,1020,1656,1680]" captionStart="Figure 7" captionStartId="17.[144,211,1168,1190]" captionTargetBox="[144,1104,168,1139]" captionTargetId="figure-197@17.[144,1104,168,1139]" captionTargetPageId="17" captionText="Figure 7. Holotype shells (A–D) and a specimen (E). (A) N. bajamarensis sp. nov.; (B) N. tafadaensis sp. nov.; (C) N. ornamentatus sp. nov.; (D) N. teobaldoi sp. nov.; (E) N. ornamentatus, paratype from Mirador de Taguluche (L. Moro photograph)." figureDoi="http://doi.org/10.5281/zenodo.5217105" httpUri="https://zenodo.org/record/5217105/files/figure.png" pageId="20" pageNumber="2197">Figure 7B</figureCitation>
|
||
). Body whorl large (BH/SH index), protoconch weakly striated with 1½–1¾ whorls. Aperture long, wide (AH/SH and AB/SB indices, respectively), elliptical section at palatal side, slightly angular at union of columellar and palatal edges. Upper palatal side forms angle slightly greater than 90º with parietal side, aperture juts out considerably from body whorl beginning. Whitish discontinuous peristome expanded as lip, more developed in lower part of palatal edge, reflected in columellar edge, where it partially covers umbilical slit. Older specimens with callosity between peristome edges, nodule at union of parietal–palatal area.
|
||
</paragraph>
|
||
<caption id="9CC266154562793041AC811CFE4DFA92" ID-DOI="http://doi.org/10.5281/zenodo.5217117" ID-Zenodo-Dep="5217117" httpUri="https://zenodo.org/record/5217117/files/figure.png" pageId="21" pageNumber="2198" startId="21.[144,211,1117,1139]" targetBox="[144,1104,168,1087]" targetPageId="21">
|
||
<paragraph id="C802369D4562793041AC811CFE4DFA92" blockId="21.[144,1104,1117,1286]" pageId="21" pageNumber="2198">
|
||
Figure 11. Genital system. (A)
|
||
<taxonomicName id="0FBD4D1E4562793040E5811FFDC6FBE0" authority="Yanes & Martín & Moro & Alonso & Ibáñez, 2009" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[473,617,1118,1140]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="2198" phylum="Mollusca" rank="species" species="tafadaensis" status="sp. nov.">
|
||
<emphasis id="FAC9EA8F4562793040E5811FFDC6FBE0" box="[473,617,1118,1140]" italics="true" pageId="21" pageNumber="2198">N. tafadaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E1FA57F445627930434E811FFD68FBE7" box="[626,711,1118,1139]" pageId="21" pageNumber="2198" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype from Montaña Tafada; (B)
|
||
<taxonomicName id="0FBD4D1E4562793041AC813AFE9AFB05" authority="Yanes & Martín & Moro & Alonso & Ibáñez, 2009" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[144,309,1147,1169]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="2198" phylum="Mollusca" rank="species" species="ornamentatus" status="sp. nov.">
|
||
<emphasis id="FAC9EA8F4562793041AC813AFE9AFB05" box="[144,309,1147,1169]" italics="true" pageId="21" pageNumber="2198">N. ornamentatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E1FA57F445627930407C813DFE39FB05" box="[320,406,1148,1169]" pageId="21" pageNumber="2198" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype from Mirador de Taguluche; (C)
|
||
<taxonomicName id="0FBD4D1E456279304251813AFC44FB05" authority="Yanes & Martín & Moro & Alonso & Ibáñez, 2009" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[877,1003,1147,1169]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="2198" phylum="Mollusca" rank="species" species="teobaldoi" status="sp. nov.">
|
||
<emphasis id="FAC9EA8F456279304251813AFC44FB05" box="[877,1003,1147,1169]" italics="true" pageId="21" pageNumber="2198">N. teobaldoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E1FA57F44562793042C9813DFBE3FB05" box="[1013,1100,1148,1169]" pageId="21" pageNumber="2198" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype from Barranco Hondo. A
|
||
<subScript id="543934D84562793040C581E3FDADFB26" attach="left" box="[505,514,1186,1202]" fontSize="7" pageId="21" pageNumber="2198">1</subScript>
|
||
, A
|
||
<subScript id="543934D845627930431E81E3FD84FB26" attach="left" box="[546,555,1186,1202]" fontSize="7" pageId="21" pageNumber="2198">5</subScript>
|
||
, parts of the penial appendix; ag, albumen gland; bc, bursa copulatrix; di, diaphragm; dp, distal penis; e, epiphallus; ec, epiphallar caecum; f, flagellum; go, genital orifice; o, free oviduct; par, penial appendix retractor; pp, proximal penis; v, vagina; vd, vas deferens.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C802369D4562793041868305FBE1F908" blockId="21.[144,1104,1348,1692]" pageId="21" pageNumber="2198">
|
||
Second morphotype, “irregular” (
|
||
<figureCitation id="50862A184562793043018305FD1CF9C8" box="[573,691,1604,1628]" captionStart="Figure 3" captionStartId="13.[144,211,1424,1446]" captionTargetBox="[150,1096,172,1387]" captionTargetId="figure-92@13.[144,1104,167,1394]" captionTargetPageId="13" captionText="Figure 3. Napaeus shell drawings. (A) N. consecoanus; (B) N. bertheloti; (C) N. subsimplex; (D) N. savinosa (syntype, NHM 1895.2.2.223); (E) N. inflatiusculus; (F) N. severus; (G) N. baeticatus; (H) N. baeticatus morphotype subbeticatus; (J) N. baeticatus morphotype halmyris; (K) N. interpunctatus; (L) N. servus; (M) N. tabidus; (N) N. tafadaensis sp. nov. (holotype, TFMC MT 0398); (O) N. tafadaensis sp. nov. morphotype “irregular” (N. tafadaensis paratype, AIT); (P) N. badiosus." figureDoi="http://doi.org/10.5281/zenodo.5217095" httpUri="https://zenodo.org/record/5217095/files/figure.png" pageId="21" pageNumber="2198">Figure 3O</figureCitation>
|
||
): shell with irregular growth mode, body whorl and palatal side of aperture flatter, shell smaller (with about six whorls only). Morphotype found in two lower localities (140 and
|
||
<quantity id="0F459B7845627930422D83C5FCFAF908" box="[785,853,1668,1692]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="21" pageNumber="2198" unit="m" value="350.0">350 m</quantity>
|
||
altitude, respectively).
|
||
</paragraph>
|
||
<caption id="9CC266154561793341E881DCFC73FABD" ID-DOI="http://doi.org/10.5281/zenodo.5217119" ID-Zenodo-Dep="5217119" httpUri="https://zenodo.org/record/5217119/files/figure.png" pageId="22" pageNumber="2199" startId="22.[212,279,1181,1203]" targetBox="[212,1172,168,1151]" targetPageId="22">
|
||
<paragraph id="C802369D4561793341E881DCFC73FABD" blockId="22.[212,1172,1181,1321]" pageId="22" pageNumber="2199">
|
||
Figure 12. Shell ornamentation details of the penultimate whorl (A–C, E, F) and that of the two last whorls (D). (A)
|
||
<taxonomicName id="0FBD4D1E4561793340F181FAFDE9FB45" authorityName="Henriquez" authorityYear="1993" box="[461,582,1211,1233]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="2199" phylum="Mollusca" rank="species" species="tenoensis">
|
||
<emphasis id="FAC9EA8F4561793340F181FAFDE9FB45" box="[461,582,1211,1233]" italics="true" pageId="22" pageNumber="2199">N. tenoensis</emphasis>
|
||
</taxonomicName>
|
||
holotype, from Tamargo (Tenerife); (B)
|
||
<taxonomicName id="0FBD4D1E4561793342D981FAFBF8FB45" authorityName="Alonso and Ibanez" authorityYear="1993" box="[997,1111,1211,1233]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="2199" phylum="Mollusca" rank="species" species="boucheti">
|
||
<emphasis id="FAC9EA8F4561793342D981FAFBF8FB45" box="[997,1111,1211,1233]" italics="true" pageId="22" pageNumber="2199">N. boucheti</emphasis>
|
||
</taxonomicName>
|
||
holotype, from Llano del Lance (La Palma); (C)
|
||
<taxonomicName id="0FBD4D1E4561793343A38199FCBBFB7A" baseAuthorityName="Webb and Berthelot" baseAuthorityYear="1833" box="[671,788,1240,1262]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="2199" phylum="Mollusca" rank="species" species="obesatus">
|
||
<emphasis id="FAC9EA8F4561793343A38199FCBBFB7A" box="[671,788,1240,1262]" italics="true" pageId="22" pageNumber="2199">N. obesatus</emphasis>
|
||
</taxonomicName>
|
||
, from La Milagrosa (Gran Canaria); (D)
|
||
<taxonomicName id="0FBD4D1E4561793341C381B7FECAFA98" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[255,357,1270,1292]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="2199" phylum="Mollusca" rank="species" species="tabidus">
|
||
<emphasis id="FAC9EA8F4561793341C381B7FECAFA98" box="[255,357,1270,1292]" italics="true" pageId="22" pageNumber="2199">N. tabidus</emphasis>
|
||
</taxonomicName>
|
||
, from La Laguna (Tenerife); (E)
|
||
<taxonomicName id="0FBD4D1E4561793343FF81B7FCCBFA98" baseAuthorityName="Mousson" baseAuthorityYear="1872" box="[707,868,1270,1292]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="2199" phylum="Mollusca" rank="species" species="maffioteanus">
|
||
<emphasis id="FAC9EA8F4561793343FF81B7FCCBFA98" box="[707,868,1270,1292]" italics="true" pageId="22" pageNumber="2199">N. maffioteanus</emphasis>
|
||
</taxonomicName>
|
||
, from Barranco de La Aldea (Gran Canaria); (F)
|
||
<taxonomicName id="0FBD4D1E4561793340988052FD9FFABD" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[420,560,1299,1321]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="2199" phylum="Mollusca" rank="species" species="propinquus">
|
||
<emphasis id="FAC9EA8F4561793340988052FD9FFABD" box="[420,560,1299,1321]" italics="true" pageId="22" pageNumber="2199">N. propinquus</emphasis>
|
||
</taxonomicName>
|
||
, from Barranco de Las Galgas (Tenerife).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C802369D4561793341C28020FBFFFA4D" blockId="22.[212,1172,1377,1689]" pageId="22" pageNumber="2199">
|
||
Shell colour tan brown, first whorls slightly darker than rest. Ornamentation characterized by weak, radial oblique ribs, more-or-less regular on almost all shell (
|
||
<figureCitation id="50862A184561793341E780E0FEF9FA2D" box="[219,342,1441,1465]" captionStart="Figure 7" captionStartId="17.[144,211,1168,1190]" captionTargetBox="[144,1104,168,1139]" captionTargetId="figure-197@17.[144,1104,168,1139]" captionTargetPageId="17" captionText="Figure 7. Holotype shells (A–D) and a specimen (E). (A) N. bajamarensis sp. nov.; (B) N. tafadaensis sp. nov.; (C) N. ornamentatus sp. nov.; (D) N. teobaldoi sp. nov.; (E) N. ornamentatus, paratype from Mirador de Taguluche (L. Moro photograph)." figureDoi="http://doi.org/10.5281/zenodo.5217105" httpUri="https://zenodo.org/record/5217105/files/figure.png" pageId="22" pageNumber="2199">Figures 7B</figureCitation>
|
||
,
|
||
<figureCitation id="50862A1845617933405D80E0FE3FFA2D" box="[353,400,1441,1465]" captionStart="Figure 10" captionStartId="20.[212,279,1133,1155]" captionTargetBox="[212,1172,168,1103]" captionTargetId="figure-150@20.[212,1172,167,1103]" captionTargetPageId="20" captionText="Figure 10. Holotype shell ornamentation details of the penultimate whorl (A, B, D, E) and that of the body whorl columellar zone, near the peristome (C). (A) N. bajamarensis sp. nov.; (B–C) N. tafadaensis sp. nov.; (D) N. ornamentatus sp. nov.; (E) N. teobaldoi sp. nov.; (F) shell ornamentation detail of the penultimate whorl of N. badiosus, from Barranco de Valle Seco (Tenerife); pt, peristome." figureDoi="http://doi.org/10.5281/zenodo.5217111" httpUri="https://zenodo.org/record/5217111/files/figure.png" pageId="22" pageNumber="2199">10B</figureCitation>
|
||
). Shell shiny despite ribs. At columellar zone of body whorl, ribs sinuous, irregularly undulating, several ribs interrupted forming granules (
|
||
<figureCitation id="50862A184561793342F98080FBEBFA4D" box="[965,1092,1473,1497]" captionStart="Figure 10" captionStartId="20.[212,279,1133,1155]" captionTargetBox="[212,1172,168,1103]" captionTargetId="figure-150@20.[212,1172,167,1103]" captionTargetPageId="20" captionText="Figure 10. Holotype shell ornamentation details of the penultimate whorl (A, B, D, E) and that of the body whorl columellar zone, near the peristome (C). (A) N. bajamarensis sp. nov.; (B–C) N. tafadaensis sp. nov.; (D) N. ornamentatus sp. nov.; (E) N. teobaldoi sp. nov.; (F) shell ornamentation detail of the penultimate whorl of N. badiosus, from Barranco de Valle Seco (Tenerife); pt, peristome." figureDoi="http://doi.org/10.5281/zenodo.5217111" httpUri="https://zenodo.org/record/5217111/files/figure.png" pageId="22" pageNumber="2199">Figure 10C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C802369D4561793241C280A0FC86FE2B" blockId="22.[212,1172,1377,1689]" lastBlockId="23.[144,1105,167,511]" lastPageId="23" lastPageNumber="2200" pageId="22" pageNumber="2199">
|
||
Genital system (
|
||
<figureCitation id="50862A1845617933408780A0FDECFA6D" box="[443,579,1505,1529]" captionStart="Figure 11" captionStartId="21.[144,211,1117,1139]" captionTargetBox="[144,1104,168,1087]" captionTargetId="figure-219@21.[144,1104,168,1087]" captionTargetPageId="21" captionText="Figure 11. Genital system. (A) N. tafadaensis sp. nov., paratype from Montaña Tafada; (B) N. ornamentatus sp. nov., paratype from Mirador de Taguluche; (C) N. teobaldoi sp. nov., paratype from Barranco Hondo. A1, A5, parts of the penial appendix; ag, albumen gland; bc, bursa copulatrix; di, diaphragm; dp, distal penis; e, epiphallus; ec, epiphallar caecum; f, flagellum; go, genital orifice; o, free oviduct; par, penial appendix retractor; pp, proximal penis; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5217117" httpUri="https://zenodo.org/record/5217117/files/figure.png" pageId="22" pageNumber="2199">Figure 11A</figureCitation>
|
||
;
|
||
<specimenCount id="DEBBFD1445617933436E80A0FD50FA6D" box="[594,767,1505,1529]" pageId="22" pageNumber="2199" type="generic">four specimens</specimenCount>
|
||
dissected of typical form, three of “irregular” morphotype). Atrium very short. Penis mainly tubular, similar in length to epiphallus, four times longer than A
|
||
<subScript id="543934D84561793343B2836CFD37F9AB" attach="left" box="[654,664,1581,1599]" fontSize="8" pageId="22" pageNumber="2199">1</subScript>
|
||
portion of penial appendix. Penis with three portions; distal portion located between atrium and insertion of penial retractor muscle; slender, tubular, intermediate portion slightly longer than each of the others; swollen, proximal portion with small penial papilla and inner thick, ring-shaped folds. Epiphallus tubular, opening distally on penis, with two regions defined by presence of well-developed caecum, for moulding and initial accommodation of spermatophoral spur-like hook. Proximal region widened, slender distally, tubular. Flagellum very short. Vas deferens opens laterally on proximal end of epiphallus. Penial appendix arising near distal end of penis, slightly longer than penis and epiphallus together, and shorter than that of other
|
||
<taxonomicName id="0FBD4D1E4560793240678406FDBFFECB" box="[347,528,327,351]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="FAC9EA8F4560793240678406FE18FECB" box="[347,439,327,351]" italics="true" pageId="23" pageNumber="2200">Napaeus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
Male genital system with two slender retractor muscles, penial and appendicular, inserting closely, but not fused, on diaphragm walls. Penial muscle inserts between distal and intermediate penis portions, appendicular muscle inserts laterally near proximal top of penial appendix part A
|
||
<subScript id="543934D845607932422484F3FC8DFE50" attach="left" box="[792,802,434,452]" fontSize="8" pageId="23" pageNumber="2200">1</subScript>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C802369D4560793241868489FCB3FE6B" blockId="23.[144,1105,167,511]" pageId="23" pageNumber="2200">Free oviduct longer than vagina. Bursa copulatrix duct without diverticulum. Bursa duct short, less than half of penial appendix length.</paragraph>
|
||
<paragraph id="C802369D4560793241AC876CFF5CFDD1" blockId="23.[144,243,557,581]" box="[144,243,557,581]" pageId="23" pageNumber="2200">
|
||
<emphasis id="FAC9EA8F4560793241AC876CFF5CFDD1" box="[144,243,557,581]" italics="true" pageId="23" pageNumber="2200">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="C802369D4560793241AC8714FC2DFCF9" blockId="23.[143,1105,597,877]" pageId="23" pageNumber="2200">
|
||
<emphasis id="FAC9EA8F4560793241AC8714FEF6FDF9" box="[144,345,597,621]" italics="true" pageId="23" pageNumber="2200">Shell morphology.</emphasis>
|
||
The shell of
|
||
<taxonomicName id="0FBD4D1E4560793240D68714FD2CFDF9" authority="Yanes & Martín & Moro & Alonso & Ibáñez, 2009" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[490,643,597,621]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="tafadaensis" status="sp. nov.">
|
||
<emphasis id="FAC9EA8F4560793240D68714FD2CFDF9" box="[490,643,597,621]" italics="true" pageId="23" pageNumber="2200">N. tafadaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E1FA57F44560793243B78714FD4CFDF8" box="[651,739,597,620]" pageId="23" pageNumber="2200" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="50862A184560793243CE8714FC25FDF9" box="[754,906,597,621]" captionStart="Figure 3" captionStartId="13.[144,211,1424,1446]" captionTargetBox="[150,1096,172,1387]" captionTargetId="figure-92@13.[144,1104,167,1394]" captionTargetPageId="13" captionText="Figure 3. Napaeus shell drawings. (A) N. consecoanus; (B) N. bertheloti; (C) N. subsimplex; (D) N. savinosa (syntype, NHM 1895.2.2.223); (E) N. inflatiusculus; (F) N. severus; (G) N. baeticatus; (H) N. baeticatus morphotype subbeticatus; (J) N. baeticatus morphotype halmyris; (K) N. interpunctatus; (L) N. servus; (M) N. tabidus; (N) N. tafadaensis sp. nov. (holotype, TFMC MT 0398); (O) N. tafadaensis sp. nov. morphotype “irregular” (N. tafadaensis paratype, AIT); (P) N. badiosus." figureDoi="http://doi.org/10.5281/zenodo.5217095" httpUri="https://zenodo.org/record/5217095/files/figure.png" pageId="23" pageNumber="2200">Figure 3N, O</figureCitation>
|
||
) is comparable in dimensions with those of
|
||
<taxonomicName id="0FBD4D1E4560793240988734FDBEFD19" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[420,529,629,653]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="tabidus">
|
||
<emphasis id="FAC9EA8F4560793240988734FDBEFD19" box="[420,529,629,653]" italics="true" pageId="23" pageNumber="2200">N. tabidus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="50862A184560793243238734FD3AFD19" box="[543,661,629,653]" captionStart="Figure 3" captionStartId="13.[144,211,1424,1446]" captionTargetBox="[150,1096,172,1387]" captionTargetId="figure-92@13.[144,1104,167,1394]" captionTargetPageId="13" captionText="Figure 3. Napaeus shell drawings. (A) N. consecoanus; (B) N. bertheloti; (C) N. subsimplex; (D) N. savinosa (syntype, NHM 1895.2.2.223); (E) N. inflatiusculus; (F) N. severus; (G) N. baeticatus; (H) N. baeticatus morphotype subbeticatus; (J) N. baeticatus morphotype halmyris; (K) N. interpunctatus; (L) N. servus; (M) N. tabidus; (N) N. tafadaensis sp. nov. (holotype, TFMC MT 0398); (O) N. tafadaensis sp. nov. morphotype “irregular” (N. tafadaensis paratype, AIT); (P) N. badiosus." figureDoi="http://doi.org/10.5281/zenodo.5217095" httpUri="https://zenodo.org/record/5217095/files/figure.png" pageId="23" pageNumber="2200">Figure 3M</figureCitation>
|
||
) and
|
||
<taxonomicName id="0FBD4D1E4560793243EF8734FCE0FD19" baseAuthorityName="Webb and Berthelot" baseAuthorityYear="1833" box="[723,847,629,653]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="badiosus">
|
||
<emphasis id="FAC9EA8F4560793243EF8734FCE0FD19" box="[723,847,629,653]" italics="true" pageId="23" pageNumber="2200">N. badiosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="50862A184560793242608734FC64FD19" box="[860,971,629,653]" captionStart="Figure 3" captionStartId="13.[144,211,1424,1446]" captionTargetBox="[150,1096,172,1387]" captionTargetId="figure-92@13.[144,1104,167,1394]" captionTargetPageId="13" captionText="Figure 3. Napaeus shell drawings. (A) N. consecoanus; (B) N. bertheloti; (C) N. subsimplex; (D) N. savinosa (syntype, NHM 1895.2.2.223); (E) N. inflatiusculus; (F) N. severus; (G) N. baeticatus; (H) N. baeticatus morphotype subbeticatus; (J) N. baeticatus morphotype halmyris; (K) N. interpunctatus; (L) N. servus; (M) N. tabidus; (N) N. tafadaensis sp. nov. (holotype, TFMC MT 0398); (O) N. tafadaensis sp. nov. morphotype “irregular” (N. tafadaensis paratype, AIT); (P) N. badiosus." figureDoi="http://doi.org/10.5281/zenodo.5217095" httpUri="https://zenodo.org/record/5217095/files/figure.png" pageId="23" pageNumber="2200">Figure 3P</figureCitation>
|
||
), but clearly more slender than that of
|
||
<taxonomicName id="0FBD4D1E45607932409687D4FD89FD39" baseAuthorityName="Webb and Berthelot" baseAuthorityYear="1833" box="[426,550,661,685]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="badiosus">
|
||
<emphasis id="FAC9EA8F45607932409687D4FD89FD39" box="[426,550,661,685]" italics="true" pageId="23" pageNumber="2200">N. badiosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="50862A1845607932430887D4FD0BFD39" box="[564,676,661,685]" captionStart="Figure 6" captionStartId="16.[212,279,1357,1379]" captionTargetBox="[214,1168,170,1326]" captionTargetId="figure-114@16.[212,1172,168,1327]" captionTargetPageId="16" captionText="Figure 6. Scatter plots of some shell measurements for the new species and the nearest species of their respective conchological group. AH, aperture height; AS, aperture surface (plane view); FH, first whorls height; FP, first whorls perimeter; FS, first whorls surface (plane view); SB, shell breadth; SH, shell height." figureDoi="http://doi.org/10.5281/zenodo.5217103" httpUri="https://zenodo.org/record/5217103/files/figure.png" pageId="23" pageNumber="2200">Figure 6E</figureCitation>
|
||
).
|
||
<taxonomicName id="0FBD4D1E45607932438B87D4FCE1FD39" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[695,846,661,685]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="tafadaensis">
|
||
<emphasis id="FAC9EA8F45607932438B87D4FCE1FD39" box="[695,846,661,685]" italics="true" pageId="23" pageNumber="2200">N. tafadaensis</emphasis>
|
||
</taxonomicName>
|
||
has similar ornamentation to
|
||
<taxonomicName id="0FBD4D1E4560793241E187F4FEF6FD59" baseAuthorityName="Webb and Berthelot" baseAuthorityYear="1833" box="[221,345,693,717]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="badiosus">
|
||
<emphasis id="FAC9EA8F4560793241E187F4FEF6FD59" box="[221,345,693,717]" italics="true" pageId="23" pageNumber="2200">N. badiosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="50862A1845607932405987F4FDE4FD59" box="[357,587,693,717]" captionStart="Figure 10" captionStartId="20.[212,279,1133,1155]" captionTargetBox="[212,1172,168,1103]" captionTargetId="figure-150@20.[212,1172,167,1103]" captionTargetPageId="20" captionText="Figure 10. Holotype shell ornamentation details of the penultimate whorl (A, B, D, E) and that of the body whorl columellar zone, near the peristome (C). (A) N. bajamarensis sp. nov.; (B–C) N. tafadaensis sp. nov.; (D) N. ornamentatus sp. nov.; (E) N. teobaldoi sp. nov.; (F) shell ornamentation detail of the penultimate whorl of N. badiosus, from Barranco de Valle Seco (Tenerife); pt, peristome." figureDoi="http://doi.org/10.5281/zenodo.5217111" httpUri="https://zenodo.org/record/5217111/files/figure.png" pageId="23" pageNumber="2200">Figures 10B and 10F</figureCitation>
|
||
, respectively) while the radial oblique ribs of the body whorl are discontinuous and granulated in
|
||
<taxonomicName id="0FBD4D1E4560793243A98794FCAEFD79" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[661,769,725,749]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="tabidus">
|
||
<emphasis id="FAC9EA8F4560793243A98794FCAEFD79" box="[661,769,725,749]" italics="true" pageId="23" pageNumber="2200">N. tabidus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="50862A184560793242318794FC23FD79" box="[781,908,725,749]" captionStart="Figure 12" captionStartId="22.[212,279,1181,1203]" captionTargetBox="[212,1172,168,1151]" captionTargetId="figure-208@22.[212,1172,167,1151]" captionTargetPageId="22" captionText="Figure 12. Shell ornamentation details of the penultimate whorl (A–C, E, F) and that of the two last whorls (D). (A) N. tenoensis holotype, from Tamargo (Tenerife); (B) N. boucheti holotype, from Llano del Lance (La Palma); (C) N. obesatus, from La Milagrosa (Gran Canaria); (D) N. tabidus, from La Laguna (Tenerife); (E) N. maffioteanus, from Barranco de La Aldea (Gran Canaria); (F) N. propinquus, from Barranco de Las Galgas (Tenerife)." figureDoi="http://doi.org/10.5281/zenodo.5217119" httpUri="https://zenodo.org/record/5217119/files/figure.png" pageId="23" pageNumber="2200">Figure 12D</figureCitation>
|
||
).
|
||
<taxonomicName id="0FBD4D1E4560793242A18794FBA6FD79" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[925,1033,725,749]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="tabidus">
|
||
<emphasis id="FAC9EA8F4560793242A18794FBA6FD79" box="[925,1033,725,749]" italics="true" pageId="23" pageNumber="2200">N. tabidus</emphasis>
|
||
</taxonomicName>
|
||
shows variability in shell dimensions but normally has a bigger aperture than
|
||
<taxonomicName id="0FBD4D1E45607932428A87B4FBFFFC99" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[950,1104,757,781]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="tafadaensis">
|
||
<emphasis id="FAC9EA8F45607932428A87B4FBFFFC99" box="[950,1104,757,781]" italics="true" pageId="23" pageNumber="2200">N. tafadaensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="50862A184560793241AA8654FEABFCB9" box="[150,260,789,813]" captionStart="Figure 6" captionStartId="16.[212,279,1357,1379]" captionTargetBox="[214,1168,170,1326]" captionTargetId="figure-114@16.[212,1172,168,1327]" captionTargetPageId="16" captionText="Figure 6. Scatter plots of some shell measurements for the new species and the nearest species of their respective conchological group. AH, aperture height; AS, aperture surface (plane view); FH, first whorls height; FP, first whorls perimeter; FS, first whorls surface (plane view); SB, shell breadth; SH, shell height." figureDoi="http://doi.org/10.5281/zenodo.5217103" httpUri="https://zenodo.org/record/5217103/files/figure.png" pageId="23" pageNumber="2200">Figure 6F</figureCitation>
|
||
). The other species of the
|
||
<taxonomicName id="0FBD4D1E4560793243228654FD05FCB9" baseAuthorityName="Webb and Berthelot" baseAuthorityYear="1833" box="[542,682,789,813]" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="baeticatus">
|
||
<emphasis id="FAC9EA8F4560793243228654FD05FCB9" box="[542,682,789,813]" italics="true" pageId="23" pageNumber="2200">N. baeticatus</emphasis>
|
||
</taxonomicName>
|
||
group are clearly bigger, while the species of the
|
||
<taxonomicName id="0FBD4D1E4560793240308674FE29FCD9" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[268,390,821,845]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="nanodes">
|
||
<emphasis id="FAC9EA8F4560793240308674FE29FCD9" box="[268,390,821,845]" italics="true" pageId="23" pageNumber="2200">N. nanodes</emphasis>
|
||
</taxonomicName>
|
||
group are smaller and have prominent radial oblique striation. Lastly, the species of the
|
||
<taxonomicName id="0FBD4D1E4560793240A08614FDBEFCF9" baseAuthorityName="Webb and Berthelot" baseAuthorityYear="1833" box="[412,529,853,877]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="variatus">
|
||
<emphasis id="FAC9EA8F4560793240A08614FDBEFCF9" box="[412,529,853,877]" italics="true" pageId="23" pageNumber="2200">N. variatus</emphasis>
|
||
</taxonomicName>
|
||
group are evidently more slender.
|
||
</paragraph>
|
||
<paragraph id="C802369D4560793241AC86D4FE39FB98" blockId="23.[144,1104,917,1037]" pageId="23" pageNumber="2200">
|
||
<emphasis id="FAC9EA8F4560793241AC86D4FE0EFC39" box="[144,417,917,941]" italics="true" pageId="23" pageNumber="2200">Genital system anatomy.</emphasis>
|
||
The epiphallar caecum is present in
|
||
<taxonomicName id="0FBD4D1E45607932425486D4FC4DFC39" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[872,994,917,941]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="tabidus">
|
||
<emphasis id="FAC9EA8F45607932425486D4FC4DFC39" box="[872,994,917,941]" italics="true" pageId="23" pageNumber="2200">N. tabidus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="AC2C4B6C4560793242C486D4FF5FFC59" author="Odhner NH" pageId="23" pageNumber="2200" pagination="1 - 116" refId="ref12541" refString="Odhner NH. 1932. Beitrage zur Malakozoologie der Kanarischen Inseln. Lamellibranchien, Cephalopoden, Gastropoden. Ark Zool. 23 A: 1 - 116." type="journal article" year="1932">Odhner 1932: 62</bibRefCitation>
|
||
, figure 21) and
|
||
<taxonomicName id="0FBD4D1E4560793240A186F4FD94FC59" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[413,571,949,973]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="tafadaensis">
|
||
<emphasis id="FAC9EA8F4560793240A186F4FD94FC59" box="[413,571,949,973]" italics="true" pageId="23" pageNumber="2200">N. tafadaensis</emphasis>
|
||
</taxonomicName>
|
||
, and lacking in
|
||
<taxonomicName id="0FBD4D1E4560793243D186F4FCC2FC59" baseAuthorityName="Webb and Berthelot" baseAuthorityYear="1833" box="[749,877,949,973]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="badiosus">
|
||
<emphasis id="FAC9EA8F4560793243D186F4FCC2FC59" box="[749,877,949,973]" italics="true" pageId="23" pageNumber="2200">N. badiosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="AC2C4B6C45607932424086F4FBA1FC59" author="Krause A" box="[892,1038,949,973]" pageId="23" pageNumber="2200" pagination="20 - 29" refId="ref12485" refString="Krause A. 1895. Landschnecken von Tenerifa. Nach Sammlungen von Dr. Aurel Krause. Nachrichtenbl dtsch malakozool Ges. 27: 20 - 29." type="journal article" year="1895">Krause 1895</bibRefCitation>
|
||
: pl. 1, figure 9). The bursa duct of
|
||
<taxonomicName id="0FBD4D1E4560793240ED8694FDDFFC79" authorityName="Yanes & Martín & Moro & Alonso & Ibáñez" authorityYear="2009" box="[465,624,981,1005]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="tafadaensis">
|
||
<emphasis id="FAC9EA8F4560793240ED8694FDDFFC79" box="[465,624,981,1005]" italics="true" pageId="23" pageNumber="2200">N. tafadaensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0FBD4D1E4560793243968694FC84FC79" baseAuthorityName="Webb and Berthelot" baseAuthorityYear="1833" box="[682,811,981,1005]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="badiosus">
|
||
<emphasis id="FAC9EA8F4560793243968694FC84FC79" box="[682,811,981,1005]" italics="true" pageId="23" pageNumber="2200">N. badiosus</emphasis>
|
||
</taxonomicName>
|
||
lacks a diverticulum; this is present in
|
||
<taxonomicName id="0FBD4D1E45607932402186B4FE3FFB99" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[285,400,1013,1037]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="tabidus">
|
||
<emphasis id="FAC9EA8F45607932402186B4FE3FFB99" box="[285,400,1013,1037]" italics="true" pageId="23" pageNumber="2200">N. tabidus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |