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<document id="083554438A055F60D12DCA603E646EE3" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C539B05FF67F4A29CE1FC82" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus racarus Azevedo 2017, sp. nov." docType="treatment" docVersion="15" lastPageNumber="34" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="33" updateTime="1698455438235" updateUser="plazi">
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<mods:title id="E636928B57B8CF02DF3E616D0852657D">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
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<mods:namePart id="84B6C6401481974415052AAAC32952D6">Azevedo, Celso O.</mods:namePart>
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<mods:date id="4F70C869CA7FD3736CC5E0CA35CBCD30">2017</mods:date>
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<mods:number id="78D01A4919E5F7418BC8F92E05C9D2DB">2017-10-13</mods:number>
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<mods:identifier id="B43FC118D98EC2C440DAAF6D8081A396" type="DOI">10.11646/zootaxa.4335.1.1</mods:identifier>
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<treatment id="3B3687C07C539B05FF67F4A29CE1FC82" ID-DOI="http://doi.org/10.5281/zenodo.5020382" ID-GBIF-Taxon="135435291" ID-Zenodo-Dep="5020382" LSID="urn:lsid:plazi:treatment:3B3687C07C539B05FF67F4A29CE1FC82" httpUri="http://treatment.plazi.org/id/3B3687C07C539B05FF67F4A29CE1FC82" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<subSubSection id="FB85655D7C539B04FF67F4A29EA4F907" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="B32036D67C539B04FF67F4A29D8EF968" blockId="32.[151,747,1693,1752]" box="[151,747,1693,1719]" pageId="32" pageNumber="33">
<heading id="E86881BA7C539B04FF67F4A29D8EF968" bold="true" box="[151,747,1693,1719]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="81EBEAC47C539B04FF67F4A29D8EF968" bold="true" box="[151,747,1693,1719]" pageId="32" pageNumber="33">
<taxonomicName id="749F4D557C539B04FF67F4A29DE5F968" authority="Brito &amp; Azevedo" authorityName="Azevedo" authorityYear="2017" box="[151,640,1693,1719]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="racarus" status="sp. nov.">
<emphasis id="81EBEAC47C539B04FF67F4A29EC3F968" bold="true" box="[151,422,1693,1719]" italics="true" pageId="32" pageNumber="33">Dissomphalus racarus</emphasis>
Brito &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="9AD857BF7C539B04FD7EF4A29D8EF968" box="[654,747,1693,1719]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B32036D67C539B04FF67F4809EA4F907" blockId="32.[151,747,1693,1752]" box="[151,449,1727,1752]" pageId="32" pageNumber="33">
(
<figureCitation id="2BA42A537C539B04FF6FF4809F90F907" box="[159,245,1727,1752]" captionStart="FIGURES 13 – 24" captionStartId="11.[151,264,1615,1638]" captionTargetBox="[151,1436,306,1594]" captionTargetId="figure@11.[151,1436,303,1594]" captionTargetPageId="11" captionText="FIGURES 13 – 24. Dissomphalus spp., ♂, head, dorsal view. 13. D. racarus sp. nov. 14. D. teraus sp. nov. 15. D. urazius sp. nov. 16. D. afestus sp. nov. 17. D. arbeius sp. nov. 18. D. raziogus sp. nov. 19. D. areius sp. nov. 20. D. aretisus sp. nov. 21. D. citus sp. nov. 22. D. sartus sp. nov. 23. D. teobus sp. nov. 24. D. terbius sp. nov.. Scale bars = 20 µm, except figs 14, 20 = 10 µm and fig. 15 = 50 µm." httpUri="https://zenodo.org/record/1010018/files/figure.png" pageId="32" pageNumber="33">Figs 13</figureCitation>
,
<figureCitation id="2BA42A537C539B04FEF1F4FF9E45F907" box="[257,288,1728,1752]" captionStart="FIGURES 33 – 44" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[151,1436,553,1886]" captionTargetId="figure@13.[151,1436,553,1886]" captionTargetPageId="13" captionText="FIGURES 33 – 44. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 33. D. isteus sp. nov. 34. D. azagus sp. nov. 35. D. asfartus sp. nov. 36. D. iozus sp. nov. 37. D. asitius sp. nov. 38. D. ibocius sp. nov. 39. D. tisinus sp. nov. 40. D. ardonius sp. nov. 41. D. arteus sp. nov. 42. D. ettus sp. nov. 43. D. ibrastus sp. nov. 44. D. racarus sp. nov.. Scale bars = 20 µm, except fig. 35 = 200 µm." httpUri="https://zenodo.org/record/1010022/files/figure.png" pageId="32" pageNumber="33">44</figureCitation>
,
<figureCitation id="2BA42A537C539B04FEDBF4809E2FF907" box="[299,330,1727,1752]" captionStart="FIGURES 64 – 75" captionStartId="16.[151,264,1938,1961]" captionTargetBox="[175,1408,606,1889]" captionTargetId="figure@16.[151,1436,569,1917]" captionTargetPageId="16" captionText="FIGURES 64 – 75. Dissomphalus spp., ♂, hypopygium, internal view. 64. D. isteus sp. nov. 65. D. azagus sp. nov. 66. D. asfartus sp. nov. 67. D. iozus sp. nov. 68. D. asitius sp. nov. 69. D. ibocius sp. nov. 70. D. tisinus sp. nov. 71. D. ardonius sp. nov. 72. D. arteus sp. nov. 73. D. ettus sp. nov. 74. D. ibrastus sp. nov. 75. D. racarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010028/files/figure.png" pageId="32" pageNumber="33">75</figureCitation>
,
<figureCitation id="2BA42A537C539B04FEA5F4FF9EDCF907" box="[341,441,1728,1752]" captionStart="FIGURES 136 – 147" captionStartId="31.[151,264,1782,1805]" captionTargetBox="[157,1424,384,1751]" captionTargetId="figure@31.[151,1436,373,1762]" captionTargetPageId="31" captionText="FIGURES 136 – 147. Dissomphalus spp., ♂, genitalia, except fig. 147 apical lobes of aedeagal dorsal body. 136 – 137. D. arteus sp. nov. 136. Dorsal view. 137. Ventral view. 138 – 139. D. ettus sp. nov. 138. Dorsal view. 139. Ventral view. 140 – 141. D. ibrastus sp. nov. 140. Dorsal view. 141. Ventral view. 142 – 144. D. racarus sp. nov. 142. Dorsal view. 143. Ventral view. 143. Lateral view. 145 – 147. D. teraus sp. nov. 145. Dorsal view. 146. Ventral view. 147. Apical lobes of aedeagal dorsal body, ventral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010040/files/figure.png" pageId="32" pageNumber="33">142–144</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C539B05FF67F53A9F91FEBA" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="B32036D67C539B05FF67F53A9BB4FEC2" blockId="32.[151,1437,1797,2038]" lastBlockId="33.[151,1437,151,861]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="81EBEAC47C539B04FF67F53A9E4CF8C1" bold="true" box="[151,297,1797,1822]" pageId="32" pageNumber="33">Description.</emphasis>
Male. Body length
<quantity id="74679B337C539B04FDF7F5399D0BF8C2" box="[519,622,1798,1822]" metricMagnitude="-3" metricUnit="m" metricValue="4.78" pageId="32" pageNumber="33" unit="mm" value="4.78">4.78 mm</quantity>
. Color: body castaneous. Head (
<figureCitation id="2BA42A537C539B04FC27F53A9B4EF8C1" box="[983,1067,1797,1822]" captionStart="FIGURES 13 – 24" captionStartId="11.[151,264,1615,1638]" captionTargetBox="[151,1436,306,1594]" captionTargetId="figure@11.[151,1436,303,1594]" captionTargetPageId="11" captionText="FIGURES 13 – 24. Dissomphalus spp., ♂, head, dorsal view. 13. D. racarus sp. nov. 14. D. teraus sp. nov. 15. D. urazius sp. nov. 16. D. afestus sp. nov. 17. D. arbeius sp. nov. 18. D. raziogus sp. nov. 19. D. areius sp. nov. 20. D. aretisus sp. nov. 21. D. citus sp. nov. 22. D. sartus sp. nov. 23. D. teobus sp. nov. 24. D. terbius sp. nov.. Scale bars = 20 µm, except figs 14, 20 = 10 µm and fig. 15 = 50 µm." httpUri="https://zenodo.org/record/1010018/files/figure.png" pageId="32" pageNumber="33">Fig. 13</figureCitation>
). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin coarse. Metasoma (
<figureCitation id="2BA42A537C539B04FC54F54E9C9CF855" box="[932,1017,1905,1930]" captionStart="FIGURES 33 – 44" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[151,1436,553,1886]" captionTargetId="figure@13.[151,1436,553,1886]" captionTargetPageId="13" captionText="FIGURES 33 – 44. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 33. D. isteus sp. nov. 34. D. azagus sp. nov. 35. D. asfartus sp. nov. 36. D. iozus sp. nov. 37. D. asitius sp. nov. 38. D. ibocius sp. nov. 39. D. tisinus sp. nov. 40. D. ardonius sp. nov. 41. D. arteus sp. nov. 42. D. ettus sp. nov. 43. D. ibrastus sp. nov. 44. D. racarus sp. nov.. Scale bars = 20 µm, except fig. 35 = 200 µm." httpUri="https://zenodo.org/record/1010022/files/figure.png" pageId="32" pageNumber="33">Fig. 44</figureCitation>
). Tergal process lateral with shallow, subcircular and large depression, with small, low and dorsad tubercle placed on anterior area of depression, pit large, with short and sparse tuft of setae. Posterior hypopygeal margin sinuous (
<figureCitation id="2BA42A537C539B04FBE3F5869B02F80D" box="[1043,1127,1977,2002]" captionStart="FIGURES 64 – 75" captionStartId="16.[151,264,1938,1961]" captionTargetBox="[175,1408,606,1889]" captionTargetId="figure@16.[151,1436,569,1917]" captionTargetPageId="16" captionText="FIGURES 64 – 75. Dissomphalus spp., ♂, hypopygium, internal view. 64. D. isteus sp. nov. 65. D. azagus sp. nov. 66. D. asfartus sp. nov. 67. D. iozus sp. nov. 68. D. asitius sp. nov. 69. D. ibocius sp. nov. 70. D. tisinus sp. nov. 71. D. ardonius sp. nov. 72. D. arteus sp. nov. 73. D. ettus sp. nov. 74. D. ibrastus sp. nov. 75. D. racarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010028/files/figure.png" pageId="32" pageNumber="33">Fig. 75</figureCitation>
). Genitalia (
<figureCitation id="2BA42A537C539B04FB00F5869AF4F80D" box="[1264,1425,1977,2002]" captionStart="FIGURES 136 – 147" captionStartId="31.[151,264,1782,1805]" captionTargetBox="[157,1424,384,1751]" captionTargetId="figure@31.[151,1436,373,1762]" captionTargetPageId="31" captionText="FIGURES 136 – 147. Dissomphalus spp., ♂, genitalia, except fig. 147 apical lobes of aedeagal dorsal body. 136 – 137. D. arteus sp. nov. 136. Dorsal view. 137. Ventral view. 138 – 139. D. ettus sp. nov. 138. Dorsal view. 139. Ventral view. 140 – 141. D. ibrastus sp. nov. 140. Dorsal view. 141. Ventral view. 142 – 144. D. racarus sp. nov. 142. Dorsal view. 143. Ventral view. 143. Lateral view. 145 – 147. D. teraus sp. nov. 145. Dorsal view. 146. Ventral view. 147. Apical lobes of aedeagal dorsal body, ventral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010040/files/figure.png" pageId="32" pageNumber="33">Figs 142–144</figureCitation>
): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, entirely wide with small projection apicad; cross section laminar; surface vertical; inner margin sinuous, outer margin sinuous; apex short, simple, parallel; additional inner ramus long; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe, long, vertical and very narrow, with apical margin crenulate and posterad; inner pair, stout, membranous and setose. Apodeme not extending beyond genital ring.
</paragraph>
<paragraph id="B32036D67C529B05FF37F3189F91FEBA" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="34">
<emphasis id="81EBEAC47C529B05FF37F3189E22FE9F" bold="true" box="[199,327,295,320]" pageId="33" pageNumber="34">Variations.</emphasis>
Body length 3.31 (min.) mm and
<quantity id="74679B337C529B05FD14F3189C29FEE0" box="[740,844,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="6.06" pageId="33" pageNumber="34" unit="mm" value="6.06">6.06 mm</quantity>
(max.). Depression of tergal process deep or very shallow.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C529B05FF37F3509BCEFD5A" pageId="33" pageNumber="34" type="discussion">
<paragraph id="B32036D67C529B05FF37F3509BCEFD5A" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="34">
<emphasis id="81EBEAC47C529B05FF37F3509E5EFE57" bold="true" box="[199,315,367,392]" pageId="33" pageNumber="34">Remarks.</emphasis>
This species is easily recognized by having the ventral margin of the apical lobes of the aedeagal dorsal body deeply excavated, the apex of the basal with apex projected forward, the aedeagal ventral ramus vertical and the apex with a small projection dorsad. This species is similar to
<taxonomicName id="749F4D557C529B05FBD5F3869BEAFE0F" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[1061,1167,441,464]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="arteus" status="sp. nov.">
<emphasis id="81EBEAC47C529B05FBD5F3869BEAFE0F" box="[1061,1167,441,464]" italics="true" pageId="33" pageNumber="34">D. arteus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C529B05FB69F3889B90FE0F" box="[1177,1269,439,464]" pageId="33" pageNumber="34" rank="species">
<emphasis id="81EBEAC47C529B05FB69F3889B90FE0F" bold="true" box="[1177,1269,439,464]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
by having the dorsal margin of the paramere sinuous, the apex of the basal bar projected forward and the inner margin of the aedeagal ventral ramus crenulate. However, this species has the apex of the basal bar crenulate, the ventral margin of the aedeagal dorsal body deeply excavated and the apical margin of the aedeagal dorsal body with small dorsad projection, whereas
<taxonomicName id="749F4D557C529B05FE8AF0769E86FDBF" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[378,483,585,608]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="arteus" status="sp. nov.">
<emphasis id="81EBEAC47C529B05FE8AF0769E86FDBF" box="[378,483,585,608]" italics="true" pageId="33" pageNumber="34">D. arteus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C529B05FE1BF0789D20FDBF" box="[491,581,583,608]" pageId="33" pageNumber="34" rank="species">
<emphasis id="81EBEAC47C529B05FE1BF0789D20FDBF" bold="true" box="[491,581,583,608]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
has the apex of the basal bar smooth, the ventral margin of the apical lobes of the aedeagal dorsal body not excavated and the aedeagal ventral ramus without projection.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C529B05FF37F0B09B6DFCE7" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="B32036D67C529B05FF37F0B09B6DFCE7" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="34">
<materialsCitation id="03F73C8B7C529B05FF37F0B09E52FD2F" ID-GBIF-Occurrence="1671715834" collectingDate="2000-01-17" collectingDateMax="2016-11" collectingDateMin="2000-01-17" collectionCode="MIUP" country="Panama" county="Material" elevation="80" latitude="8.0" location="Rancho Frio" longLatPrecision="1302" longitude="-77.75" municipality="Darien Pr" pageId="33" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="PANAMA" typeStatus="holotype">
<emphasis id="81EBEAC47C529B05FF37F0B09EDFFD77" bold="true" box="[199,442,655,680]" pageId="33" pageNumber="34">
<collectingCounty id="5A414E5A7C529B05FF37F0B09E4AFD77" box="[199,303,655,680]" pageId="33" pageNumber="34">Material</collectingCounty>
examined.
</emphasis>
<typeStatus id="6C2488747C529B05FE3AF0B09D50FD77" box="[458,565,655,680]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
,
<specimenCount id="A599FD5F7C529B05FDBAF0AF9D04FD78" box="[586,609,656,679]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
,
<collectingRegion id="715BF8347C529B05FD87F0AF9D94FD77" box="[631,753,656,680]" country="Panama" name="Panama" pageId="33" pageNumber="34">
<collectingCountry id="CB8876467C529B05FD87F0AF9D94FD77" box="[631,753,656,680]" name="Panama" pageId="33" pageNumber="34">PANAMA</collectingCountry>
</collectingRegion>
,
<emphasis id="81EBEAC47C529B05FCF5F0AF9C8CFD78" box="[773,1001,656,679]" italics="true" pageId="33" pageNumber="34">
<collectingMunicipality id="5344ACAC7C529B05FCF5F0AF9CE4FD78" box="[773,897,656,679]" pageId="33" pageNumber="34">
<collectingRegion id="715BF8347C529B05FCF5F0AF9C31FD78" box="[773,852,656,679]" country="Panama" name="Darien" pageId="33" pageNumber="34">Darién</collectingRegion>
<collectingRegion id="715BF8347C529B05FC95F0AF9CE4FD78" box="[869,897,656,679]" country="Brazil" name="Para" pageId="33" pageNumber="34">Pr</collectingRegion>
</collectingMunicipality>
[ovincia]
</emphasis>
: P[arque] N[acional]
<collectingRegion id="715BF8347C529B05FB0BF0B09A2FFD77" box="[1275,1354,655,680]" country="Panama" name="Darien" pageId="33" pageNumber="34">Darién</collectingRegion>
, Pirre, Est[ación]
<location id="B640600D7C529B05FEE3F08B9EC6FD12" LSID="urn:lsid:plazi:treatment:3B3687C07C539B05FF67F4A29CE1FC82:B640600D7C529B05FEE3F08B9EC6FD12" box="[275,419,692,717]" country="Panama" county="Material" latitude="8.0" longLatPrecision="1302" longitude="-77.75" municipality="Darien Pr" name="Rancho Frio" pageId="33" pageNumber="34" stateProvince="PANAMA">Rancho Frio</location>
,
<elevation id="38B2D1E57C529B05FE40F08B9E88FD13" box="[432,493,692,716]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="33" pageNumber="34" unit="m" value="80.0">
<quantity id="74679B337C529B05FE40F08B9E88FD13" box="[432,493,692,716]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="33" pageNumber="34" unit="m" value="80.0">80 m</quantity>
</elevation>
, [
<geoCoordinate id="D6AB50117C529B05FDF2F08B9D06FD13" box="[514,611,692,716]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="8.0">08°00'N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C529B05FD80F08B9DB2FD12" box="[624,727,692,717]" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="-77.75">77°45'W</geoCoordinate>
],
<collectingDate id="D765E9FE7C529B05FD1CF08B9C5EFD13" box="[748,827,692,717]" pageId="33" pageNumber="34" value="2016-11">
<date id="C72110167C529B05FD1CF08B9C5EFD13" box="[748,827,692,717]" pageId="33" pageNumber="34" value="2016-11">16 nov</date>
</collectingDate>
[20]
<collectingDate id="D765E9FE7C529B05FC80F08B9C8CFD13" box="[880,1001,692,716]" pageId="33" pageNumber="34" value="2000-01-17">
<date id="C72110167C529B05FC80F08B9C8CFD13" box="[880,1001,692,716]" pageId="33" pageNumber="34" value="2000-01-17">00–17 ene</date>
</collectingDate>
2001, Malaise, R. Cambra, A. Santos [col.] (
<collectionCode id="D58EAE137C529B05FF13F0E79E4AFD2F" box="[227,303,728,752]" pageId="33" pageNumber="34">MIUP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="03F73C8B7C529B05FEB3F0E79B39FCCB" ID-GBIF-Occurrence="1671715866" collectingDate="1999-10-17" collectingDateMax="1999-10-24" collectingDateMin="1999-10-17" collectionCode="MIUP" collectorName="Tr &amp; Malaise, A &amp; Santos" country="Panama" latitude="9.406944" location="Teribe" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" pageId="33" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Bocas del Toro" typeStatus="paratype">
<typeStatus id="6C2488747C529B05FEB3F0E79ED1FD2F" box="[323,436,728,752]" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
:
<collectingRegion id="715BF8347C529B05FE4FF0E69D0CFD2F" box="[447,617,727,752]" country="Panama" name="Bocas del Toro" pageId="33" pageNumber="34">
<emphasis id="81EBEAC47C529B05FE4FF0E69D0CFD2F" box="[447,617,727,752]" italics="true" pageId="33" pageNumber="34">Bocas del Toro</emphasis>
</collectingRegion>
:
<specimenCount id="A599FD5F7C529B05FD89F0E79DF8FD2F" box="[633,669,728,752]" pageId="33" pageNumber="34" type="male">1♂</specimenCount>
,
<collectingMunicipality id="5344ACAC7C529B05FD5BF0E89D9FFD2F" box="[683,762,727,752]" pageId="33" pageNumber="34">Wekso</collectingMunicipality>
,
<location id="B640600D7C529B05FCF6F0E89C35FD2F" LSID="urn:lsid:plazi:treatment:3B3687C07C539B05FF67F4A29CE1FC82:B640600D7C529B05FCF6F0E89C35FD2F" box="[774,848,727,752]" country="Panama" latitude="9.406944" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" name="Teribe" pageId="33" pageNumber="34" stateProvince="Bocas del Toro">Teribe</location>
, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [
<geoCoordinate id="D6AB50117C529B05FF6EF0C39E46FCCA" box="[158,291,764,789]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="15" value="9.406944">09º24'25&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C529B05FEC1F0C39EDBFCCA" box="[305,446,764,789]" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="15" value="-82.93889">82º56'20&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C529B05FE21F0C39DEEFCCB" box="[465,651,764,788]" pageId="33" pageNumber="34" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">
<date id="C72110167C529B05FE21F0C39DEEFCCB" box="[465,651,764,788]" pageId="33" pageNumber="34" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">17–24 oct. 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C529B05FD67F0C29DD1FCCB" box="[663,692,765,788]" pageId="33" pageNumber="34">Tr</collectorName>
[ap]
<collectorName id="1E6A53007C529B05FD16F0C39C00FCCB" box="[742,869,764,789]" pageId="33" pageNumber="34">Malaise, A</collectorName>
.
<collectorName id="1E6A53007C529B05FC81F0C39CA7FCCA" box="[881,962,764,789]" pageId="33" pageNumber="34">Santos</collectorName>
[col.] (
<collectionCode id="D58EAE137C529B05FBF8F0C39B31FCCB" box="[1032,1108,764,788]" pageId="33" pageNumber="34">MIUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C529B05FB98F0C39AF2FCCA" ID-GBIF-Occurrence="1671715848" box="[1128,1431,764,789]" collectingDate="2000-01-17" collectingDateMax="2016-11" collectingDateMin="2000-01-17" collectionCode="MIUP" country="Panama" county="Material" elevation="80" latitude="8.0" location="Rancho Frio" longLatPrecision="1302" longitude="-77.75" municipality="Darien Pr" pageId="33" pageNumber="34" specimenCount="4" specimenCount-male="4" stateProvince="PANAMA" typeStatus="paratype">
<specimenCount id="A599FD5F7C529B05FB98F0C39BEBFCCB" box="[1128,1166,764,788]" pageId="33" pageNumber="34" type="male">4♂</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="03F73C8B7C529B05FF67F11F9B6DFCE7" ID-GBIF-Occurrence="1671715829" box="[151,1032,799,824]" collectingDate="1924-03-26" collectingDateMax="2016-11" collectingDateMin="2000-01-17" collectionCode="MIUP" collectorName="A. Santos" country="Panama" county="Material" elevation="450" latitude="8.0" location="Rancho Frio" longLatPrecision="1302" longitude="-77.75" municipality="Darien Pr" pageId="33" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="PANAMA" typeStatus="paratype">
<specimenCount id="A599FD5F7C529B05FF67F11F9FDEFCE7" box="[151,187,800,824]" pageId="33" pageNumber="34" type="male">1♂</specimenCount>
, same data as holotype except
<elevation id="38B2D1E57C529B05FDE6F1209D3DFCE7" box="[534,600,799,824]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="33" pageNumber="34" unit="m" value="450.0">
<quantity id="74679B337C529B05FDE6F1209D3DFCE7" box="[534,600,799,824]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="33" pageNumber="34" unit="m" value="450.0">450m</quantity>
</elevation>
,
<collectingDate id="D765E9FE7C529B05FD93F1209D9CFCE8" box="[611,761,799,824]" pageId="33" pageNumber="34" value="1924-03-26">
<date id="C72110167C529B05FD93F1209D9CFCE8" box="[611,761,799,824]" pageId="33" pageNumber="34" value="1924-03-26">24–26 marzo</date>
</collectingDate>
, col.
<collectorName id="1E6A53007C529B05FCC4F11F9CC7FCE7" box="[820,930,800,824]" pageId="33" pageNumber="34">A. Santos</collectorName>
(
<collectionCode id="D58EAE137C529B05FC40F11F9C99FCE7" box="[944,1020,800,824]" pageId="33" pageNumber="34">MIUP</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C529B05FF37F17B9CE1FC82" box="[199,900,836,861]" pageId="33" pageNumber="34" type="distribution">
<paragraph id="B32036D67C529B05FF37F17B9CE1FC82" blockId="33.[151,1437,151,861]" box="[199,900,836,861]" pageId="33" pageNumber="34">
<emphasis id="81EBEAC47C529B05FF37F17B9E3CFC82" bold="true" box="[199,345,836,861]" pageId="33" pageNumber="34">Distribution</emphasis>
(
<figureCitation id="2BA42A537C529B05FE98F17B9EAEFC83" box="[360,459,836,861]" captionStart="FIGURES 234 – 243" captionStartId="67.[151,264,1856,1879]" captionTargetBox="[174,1439,203,1822]" captionTargetId="figure@67.[162,1447,193,1827]" captionTargetPageId="67" captionText="FIGURES 234 – 243. Known distribution of Dissomphalus in Panama. 234. D. galeatus, D. geniculatus and D. geortus sp. nov.. 235. D. gilvipes, D. ibocius sp. nov. and D. ibrastus sp. nov.. 236. D. guttus and D. ilarius sp. nov.. 237. D. infissus, D. intradentatus and D. iozus sp. nov.. 238. D. isortus sp. nov., D. isteus sp. nov. and D. latus. 239. D. persontaus, D. pilus and D. pronus. 240. D. plaumanni and D. politus. 241. D. piscicercus, D. ramosus and D. raziogus sp. nov.. 242. D. punctatus, D. strepsus and D. subdeformis. 243. D. racarus sp. nov., D. strabus and D. tisinus sp. nov.." httpUri="https://zenodo.org/record/1010060/files/figure.png" pageId="33" pageNumber="34">Fig. 243</figureCitation>
).
<collectingRegion id="715BF8347C529B05FE2EF17A9D52FC83" box="[478,567,837,860]" country="Panama" name="Panama" pageId="33" pageNumber="34">
<collectingCountry id="CB8876467C529B05FE2EF17A9D52FC83" box="[478,567,837,860]" name="Panama" pageId="33" pageNumber="34">Panama</collectingCountry>
</collectingRegion>
(
<collectingRegion id="715BF8347C529B05FDB6F17B9D95FC83" box="[582,752,836,861]" country="Panama" name="Bocas del Toro" pageId="33" pageNumber="34">Bocas del Toro</collectingRegion>
and
<collectingRegion id="715BF8347C529B05FCD6F17B9C12FC82" box="[806,887,836,861]" country="Panama" name="Darien" pageId="33" pageNumber="34">Darién</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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