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<document id="A04885CC8E11CAAA0984ABCD2C98FA08" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C4D9B1BFF67F6C99C99FD12" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus ilarius Azevedo 2017, sp. nov." docType="treatment" docVersion="15" lastPageNumber="64" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="63" updateTime="1698455438235" updateUser="plazi">
<mods:mods id="1A893F7FC0639A71918D9B13E9712BD8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8FE2BD3B23E29290CBA3C4966A38C664">
<mods:title id="E6A9E831D287200447691D405D046FC7">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
</mods:titleInfo>
<mods:name id="D7EBA0290A8ED68C7D5EDAE0661DB2A7" type="personal">
<mods:role id="5392D43A92A5B87123CE55ECEA4D3593">
<mods:roleTerm id="D98B2133722750D2126CE75820CBF2F4">Author</mods:roleTerm>
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<mods:namePart id="455CCB44E51799F0572826A0CEA82AD9">Azevedo, Celso O.</mods:namePart>
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<mods:title id="0A47963763DC9C1256DB132C9B668F16">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CC63AA7BC415F5BD0CD28C62838455A8">
<mods:date id="7F120DFD15B9A424F79DA830993D387C">2017</mods:date>
<mods:detail id="624587A24603546ED0A79858BA4C16E2" type="pubDate">
<mods:number id="6D8B88A9FFC3C4B2923789E92BB09E04">2017-10-13</mods:number>
</mods:detail>
<mods:detail id="13CF0102FD9ECD3AD46580FB3A7E8CA8" type="volume">
<mods:number id="79B35CF4B5682A9CAACFA0FFA4BEADD7">4335</mods:number>
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<mods:detail id="6AC8097B91133000A04589A384FD8D4D" type="issue">
<mods:number id="EE007BD437FC5A350618C35E6D7D8DFD">1</mods:number>
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<mods:classification id="B9EC625B3EFC98F2624EFBF9C769DBA5">journal article</mods:classification>
<mods:identifier id="0C2A9FBD451A9956A00CB7AB30D64284" type="CLB-Dataset">31852</mods:identifier>
<mods:identifier id="13FB080FD1BBFDC23A51C3B65C1A8655" type="DOI">10.11646/zootaxa.4335.1.1</mods:identifier>
<mods:identifier id="813A19E2F7C82E459EDEB22597C528BD" type="GBIF-Dataset">f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf</mods:identifier>
<mods:identifier id="58B57511887DF6165C70638144D9A068" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5B625FB053522713A64C273C9D2F0E7D" type="Zenodo-Dep">1010014</mods:identifier>
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<treatment id="3B3687C07C4D9B1BFF67F6C99C99FD12" ID-DOI="http://doi.org/10.5281/zenodo.5020526" ID-GBIF-Taxon="135435282" ID-Zenodo-Dep="5020526" LSID="urn:lsid:plazi:treatment:3B3687C07C4D9B1BFF67F6C99C99FD12" httpUri="http://treatment.plazi.org/id/3B3687C07C4D9B1BFF67F6C99C99FD12" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
<subSubSection id="FB85655D7C4D9B1AFF67F6C99EA4FAED" pageId="62" pageNumber="63" type="nomenclature">
<paragraph id="B32036D67C4D9B1AFF67F6C99DB0FACE" blockId="62.[151,725,1270,1330]" box="[151,725,1270,1297]" pageId="62" pageNumber="63">
<heading id="E86881BA7C4D9B1AFF67F6C99DB0FACE" bold="true" box="[151,725,1270,1297]" fontSize="11" level="1" pageId="62" pageNumber="63" reason="1">
<emphasis id="81EBEAC47C4D9B1AFF67F6C99DB0FACE" bold="true" box="[151,725,1270,1297]" pageId="62" pageNumber="63">
<taxonomicName id="749F4D557C4D9B1AFF67F6C99D15FACE" authority="Brito &amp; Azevedo" authorityName="Azevedo" authorityYear="2017" box="[151,624,1270,1297]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ilarius" status="sp. nov.">
<emphasis id="81EBEAC47C4D9B1AFF67F6C99EF2FACF" bold="true" box="[151,407,1270,1296]" italics="true" pageId="62" pageNumber="63">Dissomphalus ilarius</emphasis>
Brito &amp; Azevedo
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C4D9B1AFD87F6C89DB0FACE" box="[631,725,1271,1297]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B32036D67C4D9B1AFF67F7269EA4FAED" blockId="62.[151,725,1270,1330]" box="[151,449,1305,1330]" pageId="62" pageNumber="63">
(
<figureCitation id="2BA42A537C4D9B1AFF6FF7269F90FAED" box="[159,245,1305,1330]" captionStart="FIGURES 25 – 32" captionStartId="12.[151,264,1546,1569]" captionTargetBox="[151,1436,555,1525]" captionTargetId="figure@12.[151,1436,555,1525]" captionTargetPageId="12" captionText="FIGURES 25 – 32. Dissomphalus spp., ♂, head in dorsal view. 25. D. abarus sp. nov. 26. D. arteobius sp. nov. 27. D. borus sp. nov. 28. D. geortus sp. nov. 29. D. ilarius sp. nov. 30. D. isortus sp. nov. 31. D. trisus sp. nov. 32. D. ubracus sp. nov.. Scale bars = 20 µm, except fig. 31 = 50 µm and fig. 32 = 100 µm." httpUri="https://zenodo.org/record/1010020/files/figure.png" pageId="62" pageNumber="63">Figs 29</figureCitation>
,
<figureCitation id="2BA42A537C4D9B1AFEF1F7269E45FAED" box="[257,288,1305,1330]" captionStart="FIGURES 57 – 63" captionStartId="15.[151,264,1212,1235]" captionTargetBox="[151,1436,197,1188]" captionTargetId="figure@15.[151,1436,193,1191]" captionTargetPageId="15" captionText="FIGURES 57 – 63. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 57. D. arteobius sp. nov. 58. D. borus sp. nov. 59. D. geortus sp. nov. 60. D. ilarius sp. nov. 61. D. isortus sp. nov. 62. D. trisus sp. nov. 63. D. ubracus sp. nov. Scale bars = 20 µm except, fig. 62 = 50 µm and fig. 63 = 200 µm." httpUri="https://zenodo.org/record/1010026/files/figure.png" pageId="62" pageNumber="63">60</figureCitation>
,
<figureCitation id="2BA42A537C4D9B1AFEDBF7259E2CFAED" box="[299,329,1306,1330]" captionStart="FIGURES 88 – 94" captionStartId="18.[151,264,1101,1124]" captionTargetBox="[190,1378,187,1060]" captionTargetId="figure@18.[179,1393,179,1082]" captionTargetPageId="18" captionText="FIGURES 88 – 94. Dissomphalus spp., ♂, hypopygium, internal view. 88. D. arteobius sp. nov. 89. D. borus sp. nov. 90. D. geortus sp. nov. 91. D. ilarius sp. nov. 92. D. isortus sp. nov. 93. D. trisus sp. nov. 94. D. ubracus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010032/files/figure.png" pageId="62" pageNumber="63">91</figureCitation>
,
<figureCitation id="2BA42A537C4D9B1AFEA5F7269EDCFAED" box="[341,441,1305,1330]" captionStart="FIGURES 211 – 222" captionStartId="61.[151,264,1820,1843]" captionTargetBox="[153,1424,424,1779]" captionTargetId="figure@61.[151,1436,411,1799]" captionTargetPageId="61" captionText="FIGURES 211 – 222. Dissomphalus spp., ♂, genitalia. 211 – 212. D. borus sp. nov. 211. Dorsal view. 212. Ventral view. 213 – 214. D. geortus sp. nov. 213. Dorsal view. 214. Ventral view. 215 – 216. D. ilarius sp. nov. 215. Dorsal view. 216. Ventral view. 217 – 218. D. isortus sp. nov. 217. Dorsal view. 218. Ventral view. 219 – 220. D. trisus sp. nov. 219. Dorsal view. 220. Ventral view. 221 – 222. D. ubracus sp. nov. 221. Dorsal view. 222. Ventral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010052/files/figure.png" pageId="62" pageNumber="63">215–216</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C4D9B1AFF67F75E9E70F892" pageId="62" pageNumber="63" type="description">
<paragraph id="B32036D67C4D9B1AFF67F75E9FAEF8D9" blockId="62.[151,1437,1377,2014]" pageId="62" pageNumber="63">
<emphasis id="81EBEAC47C4D9B1AFF67F75E9E4CFAA5" bold="true" box="[151,297,1377,1402]" pageId="62" pageNumber="63">Description.</emphasis>
Male. Body length
<quantity id="74679B337C4D9B1AFDE6F75D9DE4FAA6" box="[534,641,1378,1402]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="62" pageNumber="63" unit="mm" value="2.18">2.18 mm</quantity>
. Color: head and mesosoma castaneous; metasoma light castaneous. Head (
<figureCitation id="2BA42A537C4D9B1AFF11F7BA9E51FA41" box="[225,308,1413,1438]" captionStart="FIGURES 25 – 32" captionStartId="12.[151,264,1546,1569]" captionTargetBox="[151,1436,555,1525]" captionTargetId="figure@12.[151,1436,555,1525]" captionTargetPageId="12" captionText="FIGURES 25 – 32. Dissomphalus spp., ♂, head in dorsal view. 25. D. abarus sp. nov. 26. D. arteobius sp. nov. 27. D. borus sp. nov. 28. D. geortus sp. nov. 29. D. ilarius sp. nov. 30. D. isortus sp. nov. 31. D. trisus sp. nov. 32. D. ubracus sp. nov.. Scale bars = 20 µm, except fig. 31 = 50 µm and fig. 32 = 100 µm." httpUri="https://zenodo.org/record/1010020/files/figure.png" pageId="62" pageNumber="63">Fig. 29</figureCitation>
). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one angulate very short tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (
<figureCitation id="2BA42A537C4D9B1AFE7DF7CE9E84F9D5" box="[397,481,1521,1546]" captionStart="FIGURES 57 – 63" captionStartId="15.[151,264,1212,1235]" captionTargetBox="[151,1436,197,1188]" captionTargetId="figure@15.[151,1436,193,1191]" captionTargetPageId="15" captionText="FIGURES 57 – 63. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 57. D. arteobius sp. nov. 58. D. borus sp. nov. 59. D. geortus sp. nov. 60. D. ilarius sp. nov. 61. D. isortus sp. nov. 62. D. trisus sp. nov. 63. D. ubracus sp. nov. Scale bars = 20 µm except, fig. 62 = 50 µm and fig. 63 = 200 µm." httpUri="https://zenodo.org/record/1010026/files/figure.png" pageId="62" pageNumber="63">Fig. 60</figureCitation>
). Tergal process lateral, without depression, with small, very low and dorsad tubercle, pit absent, with short and sparse tuft of setae. Posterior hypopygeal margin straight or nearly so (
<figureCitation id="2BA42A537C4D9B1AFB3EF42A9A44F9F1" box="[1230,1313,1557,1582]" captionStart="FIGURES 88 – 94" captionStartId="18.[151,264,1101,1124]" captionTargetBox="[190,1378,187,1060]" captionTargetId="figure@18.[179,1393,179,1082]" captionTargetPageId="18" captionText="FIGURES 88 – 94. Dissomphalus spp., ♂, hypopygium, internal view. 88. D. arteobius sp. nov. 89. D. borus sp. nov. 90. D. geortus sp. nov. 91. D. ilarius sp. nov. 92. D. isortus sp. nov. 93. D. trisus sp. nov. 94. D. ubracus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010032/files/figure.png" pageId="62" pageNumber="63">Fig. 91</figureCitation>
). Genitalia (
<figureCitation id="2BA42A537C4D9B1AFF6FF4069E5AF98D" box="[159,319,1593,1618]" captionStart="FIGURES 211 – 222" captionStartId="61.[151,264,1820,1843]" captionTargetBox="[153,1424,424,1779]" captionTargetId="figure@61.[151,1436,411,1799]" captionTargetPageId="61" captionText="FIGURES 211 – 222. Dissomphalus spp., ♂, genitalia. 211 – 212. D. borus sp. nov. 211. Dorsal view. 212. Ventral view. 213 – 214. D. geortus sp. nov. 213. Dorsal view. 214. Ventral view. 215 – 216. D. ilarius sp. nov. 215. Dorsal view. 216. Ventral view. 217 – 218. D. isortus sp. nov. 217. Dorsal view. 218. Ventral view. 219 – 220. D. trisus sp. nov. 219. Dorsal view. 220. Ventral view. 221 – 222. D. ubracus sp. nov. 221. Dorsal view. 222. Ventral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010052/files/figure.png" pageId="62" pageNumber="63">Figs 215–216</figureCitation>
): paramere with dorsal margin wide basally; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex long, simple, parallel; additional inner ramus absent; basal stub very short and very narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, horizontal and wide, with apical margin abruptly acute and posterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.
</paragraph>
<paragraph id="B32036D67C4D9B1AFF37F52E9E70F892" blockId="62.[151,1437,1377,2014]" pageId="62" pageNumber="63">
<emphasis id="81EBEAC47C4D9B1AFF37F52E9E22F8F5" bold="true" box="[199,327,1809,1834]" pageId="62" pageNumber="63">Variations.</emphasis>
Head dark castaneous; median clypeal tooth inconspicuous; pit of tubercle of tergal process with short setae.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C4D9B1BFF37F5669EBBFEC3" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="discussion">
<paragraph id="B32036D67C4D9B1BFF37F5669EBBFEC3" blockId="62.[151,1437,1377,2014]" lastBlockId="63.[151,1437,151,717]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
<emphasis id="81EBEAC47C4D9B1AFF37F5669E5EF8AD" bold="true" box="[199,315,1881,1906]" pageId="62" pageNumber="63">Remarks.</emphasis>
This species is easily recognized by having the paramere very expanded basally and four narrow projections that connect the apical lobes of the aedeagal dorsal body with the median portion of the aedeagal dorsal body and the tergal process with one minute tubercle without pit with few setae. This species is similar to the
<taxonomicName id="749F4D557C4D9B1AFA71F59C9EB1F801" authority=", Azevedo" authorityName="Azevedo" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="microtuberculatus">
<emphasis id="81EBEAC47C4D9B1AFA71F59C9E07F801" italics="true" pageId="62" pageNumber="63">D. microtuberculatus</emphasis>
, Azevedo
</taxonomicName>
by having the tergal process with two minutes tubercles. However, this species has the basiparamere shorter than the paramere, the apical lobes of the aedeagal dorsal body without flagella and the apex of the aedeagal ventral ramus progressively narrowing apicad, whereas
<taxonomicName id="749F4D557C4C9B1BFC4CF2829BCCFF0A" box="[956,1193,188,213]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="microtuberculatus">
<emphasis id="81EBEAC47C4C9B1BFC4CF2829BCCFF0A" box="[956,1193,188,213]" italics="true" pageId="63" pageNumber="64">D. microtuberculatus</emphasis>
</taxonomicName>
has the basiparamere longer than the paramere, the apical lobes of the aedeagal dorsal body with flagella and the apex of the aedeagal ventral ramus abruptly acute.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C4C9B1BFF37F3189CCBFD77" pageId="63" pageNumber="64" type="materials_examined">
<paragraph id="B32036D67C4C9B1BFF37F3189CCBFD77" blockId="63.[151,1437,151,717]" pageId="63" pageNumber="64">
<materialsCitation id="03F73C8B7C4C9B1BFF37F3189DD3FE57" ID-GBIF-Occurrence="1671715827" collectingDate="1999-09-26" collectingDateMax="1999-09-30" collectingDateMin="1999-09-26" collectionCode="MIUP" collectorName="El Cope &amp; de Division &amp; Omar Torrijos H &amp; Tr &amp; Malaise, A &amp; Santos, P" country="Panama" county="Material" elevation="620" latitude="8.698889" location="El Cope" longLatPrecision="21" longitude="-80.654724" municipality="Cocle Pr" pageId="63" pageNumber="64" specimenCount="1" specimenCount-male="1" stateProvince="Cocle" typeStatus="holotype">
<emphasis id="81EBEAC47C4C9B1BFF37F3189ED1FE9F" bold="true" box="[199,436,295,320]" pageId="63" pageNumber="64">
<collectingCounty id="5A414E5A7C4C9B1BFF37F3189E4AFE9F" box="[199,303,295,320]" pageId="63" pageNumber="64">Material</collectingCounty>
examined.
</emphasis>
<typeStatus id="6C2488747C4C9B1BFE4EF3189D4CFE9F" box="[446,553,295,320]" pageId="63" pageNumber="64" type="holotype">Holotype</typeStatus>
,
<specimenCount id="A599FD5F7C4C9B1BFDC8F3179D2AFEE0" box="[568,591,296,319]" pageId="63" pageNumber="64" type="male">♂</specimenCount>
,
<collectingCountry id="CB8876467C4C9B1BFDAEF3179DBDFE9F" box="[606,728,296,320]" name="Panama" pageId="63" pageNumber="64">PANAMA</collectingCountry>
,
<emphasis id="81EBEAC47C4C9B1BFD17F3189CD2FEE0" box="[743,951,295,320]" italics="true" pageId="63" pageNumber="64">
<collectingMunicipality id="5344ACAC7C4C9B1BFD17F3189C2AFEE0" box="[743,847,295,320]" pageId="63" pageNumber="64">
<collectingRegion id="715BF8347C4C9B1BFD17F3189C4DFE9F" box="[743,808,295,320]" country="Panama" name="Cocle" pageId="63" pageNumber="64">Coclé</collectingRegion>
<collectingRegion id="715BF8347C4C9B1BFCC3F3179C2AFEE0" box="[819,847,296,319]" country="Brazil" name="Para" pageId="63" pageNumber="64">Pr</collectingRegion>
</collectingMunicipality>
[ovincia]
</emphasis>
:
<location id="B640600D7C4C9B1BFC3AF3189B4EFE9F" LSID="urn:lsid:plazi:treatment:3B3687C07C4D9B1BFF67F6C99C99FD12:B640600D7C4C9B1BFC3AF3189B4EFE9F" box="[970,1067,295,320]" country="Panama" county="Material" latitude="8.698889" longLatPrecision="21" longitude="-80.654724" municipality="Cocle Pr" name="El Cope" pageId="63" pageNumber="64" stateProvince="Cocle">
<collectorName id="1E6A53007C4C9B1BFC3AF3189B4EFE9F" box="[970,1067,295,320]" pageId="63" pageNumber="64">El Copé</collectorName>
</location>
,
<location id="B640600D7C4C9B1BFBCAF3179B06FEE0" LSID="urn:lsid:plazi:treatment:3B3687C07C4D9B1BFF67F6C99C99FD12:B640600D7C4C9B1BFBCAF3179B06FEE0" box="[1082,1123,296,319]" country="Panama" county="Material" latitude="8.698889" longLatPrecision="21" longitude="-80.654724" municipality="Cocle Pr" name="Par" pageId="63" pageNumber="64" stateProvince="Cocle">Par</location>
[que]
<location id="B640600D7C4C9B1BFB55F3189BB1FE9F" LSID="urn:lsid:plazi:treatment:3B3687C07C4D9B1BFF67F6C99C99FD12:B640600D7C4C9B1BFB55F3189BB1FE9F" box="[1189,1236,295,320]" country="Panama" county="Material" latitude="8.698889" longLatPrecision="21" longitude="-80.654724" municipality="Cocle Pr" name="Nac" pageId="63" pageNumber="64" stateProvince="Cocle">Nac</location>
[ional
<location id="B640600D7C4C9B1BFAEEF3189F9EFEBA" LSID="urn:lsid:plazi:treatment:3B3687C07C4D9B1BFF67F6C99C99FD12:B640600D7C4C9B1BFAEEF3189F9EFEBA" country="Panama" county="Material" latitude="8.698889" longLatPrecision="21" longitude="-80.654724" municipality="Cocle Pr" name="General de Division" pageId="63" pageNumber="64" stateProvince="Cocle">
General
<collectorName id="1E6A53007C4C9B1BFA72F3189F9EFEBA" pageId="63" pageNumber="64">de División</collectorName>
</location>
]
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[errera],
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<quantity id="74679B337C4C9B1BFDC9F3739DA7FEBB" box="[569,706,332,357]" metricMagnitude="2" metricUnit="m" metricValue="6.2" metricValueMax="6.7" metricValueMin="5.7" pageId="63" pageNumber="64" unit="m" value="620.0" valueMax="670.0" valueMin="570.0">570–670 m</quantity>
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.s.n.m., [
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],
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,
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[ampa]
<collectorName id="1E6A53007C4C9B1BFF67F3509E7DFE57" box="[151,280,367,392]" pageId="63" pageNumber="64">Malaise, A</collectorName>
.
<collectorName id="1E6A53007C4C9B1BFED6F34F9EF7FE58" box="[294,402,368,392]" pageId="63" pageNumber="64">Santos, P</collectorName>
. González [col.] (
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)
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.
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<typeStatus id="6C2488747C4C9B1BFD34F34F9C50FE57" box="[708,821,368,392]" pageId="63" pageNumber="64" type="paratype">Paratypes</typeStatus>
:
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<emphasis id="81EBEAC47C4C9B1BFCB3F3509CC7FE57" box="[835,930,367,392]" italics="true" pageId="63" pageNumber="64">Chiriqui</emphasis>
</collectingRegion>
:
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,
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Fortuna Dam,
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<quantity id="74679B337C4C9B1BFB49F34F9B94FE57" box="[1209,1265,368,392]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="63" pageNumber="64" unit="m" value="1200.0">1200</quantity>
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m, wet forest, [
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,
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],
<collectingDate id="D765E9FE7C4C9B1BFE14F3AB9DA9FE72" box="[484,716,404,429]" pageId="63" pageNumber="64" value="1982-06-14" valueMax="1982-07-16" valueMin="1982-06-14">
<date id="C72110167C4C9B1BFE14F3AB9DA9FE72" box="[484,716,404,429]" pageId="63" pageNumber="64" value="1982-06-14" valueMax="1982-07-16" valueMin="1982-06-14">14–VI–16.VII–1982</date>
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, F[light] I[nterception] T[rap],
<collectorName id="1E6A53007C4C9B1BFBBCF3AA9BC4FE72" box="[1100,1185,404,429]" pageId="63" pageNumber="64">B. Gill</collectorName>
[col.] (
<collectionCode id="D58EAE137C4C9B1BFAF2F3AB9A33FE73" box="[1282,1366,404,428]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15193" name="Royal Alberta Museum" pageId="63" pageNumber="64">PMAE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C4C9B1BFA82F3AB9DC1FE2B" ID-GBIF-Occurrence="1671715797" collectingDate="1999-04-08" collectingDateMax="1999-04-11" collectingDateMin="1999-04-08" collectionCode="MIUP" collectorName="Malaise, R &amp; Cambra, A &amp; Santos" country="Panama" elevation="1080" latitude="8.77125" location="Toro" longLatPrecision="1" longitude="-82.19092" pageId="63" pageNumber="64" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="paratype">
<specimenCount id="A599FD5F7C4C9B1BFA82F3AB9AF3FE73" box="[1394,1430,404,428]" pageId="63" pageNumber="64" type="male">1♂</specimenCount>
,
<taxonomicName id="749F4D557C4C9B1BFF67F3879E18FE0F" authority=", Div. Cont." authorityName="Div. Cont." box="[151,381,439,464]" class="Gastropoda" family="Achatinidae" genus="Fortuna" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="64" phylum="Mollusca" rank="genus">Fortuna, Div. Cont.</taxonomicName>
Chiriqui-B[ocas del]
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,
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<quantity id="74679B337C4C9B1BFD3EF3879C14FE10" box="[718,881,440,464]" metricMagnitude="3" metricUnit="m" metricValue="1.08" metricValueMax="1.09" metricValueMin="1.0699999999999998" pageId="63" pageNumber="64" unit="m" value="1080.0" valueMax="1090.0" valueMin="1070.0">1070–1090 m</quantity>
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, [
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,
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],
<collectingDate id="D765E9FE7C4C9B1BFB06F3889AFDFE0F" box="[1270,1432,439,464]" pageId="63" pageNumber="64" value="1999-04-08" valueMax="1999-04-11" valueMin="1999-04-08">
<date id="C72110167C4C9B1BFB06F3889AFDFE0F" box="[1270,1432,439,464]" pageId="63" pageNumber="64" value="1999-04-08" valueMax="1999-04-11" valueMin="1999-04-08">8–11.abr.1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C4C9B1BFF67F3E39E70FE2B" box="[151,277,476,501]" pageId="63" pageNumber="64">Malaise, R</collectorName>
.
<collectorName id="1E6A53007C4C9B1BFED2F3E39EC7FE2B" box="[290,418,476,501]" pageId="63" pageNumber="64">Cambra, A</collectorName>
.
<collectorName id="1E6A53007C4C9B1BFE5FF3E39E9FFE2B" box="[431,506,476,500]" pageId="63" pageNumber="64">Santos</collectorName>
[col.] (
<collectionCode id="D58EAE137C4C9B1BFDA0F3E39DF9FE2B" box="[592,668,476,500]" pageId="63" pageNumber="64">MIUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C4C9B1BFD41F3E29EDAFDE3" ID-GBIF-Occurrence="1671715832" collectingDate="1998-11-30" collectingDateMax="1998-12-04" collectingDateMin="1998-11-30" collectionCode="MIUP" collectorName="Malaise, R &amp; Cambra, S &amp; Bermudez" country="Iceland" elevation="90" latitude="7.4691668" location="Cerro La Equis" longLatPrecision="21" longitude="-81.75667" municipality="Coiba" pageId="63" pageNumber="64" specimenCount="18" specimenCount-male="18" stateProvince="Veraguas Pr" typeStatus="paratype">
<emphasis id="81EBEAC47C4C9B1BFD41F3E29CC3FE2B" box="[689,934,477,500]" italics="true" pageId="63" pageNumber="64">
<collectingRegion id="715BF8347C4C9B1BFD41F3E29C5BFE2B" box="[689,830,477,500]" isEnumeration="true" pageId="63" pageNumber="64">
<collectingRegion id="715BF8347C4C9B1BFD41F3E29C7CFE2B" box="[689,793,477,500]" country="Panama" name="Veraguas" pageId="63" pageNumber="64">Veraguas</collectingRegion>
<collectingRegion id="715BF8347C4C9B1BFCD2F3E29C5BFE2B" box="[802,830,477,500]" country="Brazil" name="Para" pageId="63" pageNumber="64">Pr</collectingRegion>
</collectingRegion>
[ovincia]
</emphasis>
:
<specimenCount id="A599FD5F7C4C9B1BFC47F3E39CBFFE2B" box="[951,986,476,500]" pageId="63" pageNumber="64" type="male">9♂</specimenCount>
P[arque] N[acional]
<collectingMunicipality id="5344ACAC7C4C9B1BFB3DF3E39A77FE2A" box="[1229,1298,476,501]" pageId="63" pageNumber="64">Coiba</collectingMunicipality>
,
<collectingCountry id="CB8876467C4C9B1BFAD0F3E39A2DFE2A" box="[1312,1352,476,501]" name="Iceland" pageId="63" pageNumber="64">Isla</collectingCountry>
Coiba,
<location id="B640600D7C4C9B1BFF67F03F9E21FDC7" LSID="urn:lsid:plazi:treatment:3B3687C07C4D9B1BFF67F6C99C99FD12:B640600D7C4C9B1BFF67F03F9E21FDC7" box="[151,324,511,536]" country="Iceland" latitude="7.4691668" longLatPrecision="21" longitude="-81.75667" municipality="Coiba" name="Cerro La Equis" pageId="63" pageNumber="64" stateProvince="Veraguas Pr">Cerro La Equis</location>
,
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<quantity id="74679B337C4C9B1BFEBFF03F9EEDFDC8" box="[335,392,512,536]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="63" pageNumber="64" unit="m" value="90.0">90 m</quantity>
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(
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) and
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<quantity id="74679B337C4C9B1BFE0AF03F9D5FFDC7" box="[506,570,512,536]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="63" pageNumber="64" unit="m" value="200.0">200m</quantity>
</elevation>
(
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) [
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,
<geoCoordinate id="D6AB50117C4C9B1BFCE5F3C09CC4FDC7" box="[789,929,511,536]" direction="west" orientation="longitude" pageId="63" pageNumber="64" precision="15" value="-81.75667">81º45'24&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C4C9B1BFC44F3C09BEBFDC7" box="[948,1166,511,536]" pageId="63" pageNumber="64" value="1998-11-30" valueMax="1998-12-04" valueMin="1998-11-30">
<date id="C72110167C4C9B1BFC44F3C09BEBFDC7" box="[948,1166,511,536]" pageId="63" pageNumber="64" value="1998-11-30" valueMax="1998-12-04" valueMin="1998-11-30">30–nov.–4dic.1998</date>
</collectingDate>
,
<collectorName id="1E6A53007C4C9B1BFB68F3C09A70FDC8" box="[1176,1301,511,536]" pageId="63" pageNumber="64">Malaise, R</collectorName>
.
<collectorName id="1E6A53007C4C9B1BFAEFF3C09AFDFDC7" box="[1311,1432,511,536]" pageId="63" pageNumber="64">Cambra, S</collectorName>
.
<collectorName id="1E6A53007C4C9B1BFF67F01B9E6FFDE2" box="[151,266,548,573]" pageId="63" pageNumber="64">Bermudez</collectorName>
[col.] (
<collectionCode id="D58EAE137C4C9B1BFE94F01B9ED3FDE3" box="[356,438,548,572]" pageId="63" pageNumber="64">MIUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C4C9B1BFE33F01B9C05FDBF" ID-GBIF-Occurrence="1671715806" collectingDate="1999-11-13" collectingDateMax="1999-11-17" collectingDateMin="1999-11-13" collectionCode="MIUP" collectorName="Tr &amp; Malaise, A &amp; Santos, P &amp; Gonzalez" country="Iceland" latitude="8.518194" location="Altos De Piedra" longLatPrecision="1" longitude="-81.108444" pageId="63" pageNumber="64" specimenCount="1" specimenCount-male="1" stateProvince="Pr" typeStatus="paratype">
<specimenCount id="A599FD5F7C4C9B1BFE33F01B9E84FDE3" box="[451,481,548,572]" pageId="63" pageNumber="64" type="male">1♂</specimenCount>
,
<collectingRegion id="715BF8347C4C9B1BFE00F01B9D6EFDE2" box="[496,523,548,573]" country="Brazil" name="Para" pageId="63" pageNumber="64">Pr</collectingRegion>
[ovincia]
<collectingRegion id="715BF8347C4C9B1BFD89F01B9D84FDE3" box="[633,737,548,572]" country="Argentina" name="Santa Fe" pageId="63" pageNumber="64">Santa Fé</collectingRegion>
,
<location id="B640600D7C4C9B1BFD00F01B9CCBFDE2" LSID="urn:lsid:plazi:treatment:3B3687C07C4D9B1BFF67F6C99C99FD12:B640600D7C4C9B1BFD00F01B9CCBFDE2" box="[752,942,548,573]" country="Iceland" latitude="8.518194" longLatPrecision="1" longitude="-81.108444" name="Altos De Piedra" pageId="63" pageNumber="64" stateProvince="Pr">Altos De Piedra</location>
, [
<geoCoordinate id="D6AB50117C4C9B1BFC33F01B9B37FDE2" box="[963,1106,548,573]" direction="north" orientation="latitude" pageId="63" pageNumber="64" precision="1" value="8.518194">08°31'5.5&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C4C9B1BFB92F01B9A63FDE2" box="[1122,1286,548,573]" direction="west" orientation="longitude" pageId="63" pageNumber="64" precision="1" value="-81.108444">81°06'30.4&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C4C9B1BFAECF01B9FB7FDBF" pageId="63" pageNumber="64" value="1999-11-13" valueMax="1999-11-17" valueMin="1999-11-13">
<date id="C72110167C4C9B1BFAECF01B9FB7FDBF" pageId="63" pageNumber="64" value="1999-11-13" valueMax="1999-11-17" valueMin="1999-11-13">13–17 nov. 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C4C9B1BFF2EF0779F9FFD80" box="[222,250,584,607]" pageId="63" pageNumber="64">Tr</collectorName>
[ampa]
<collectorName id="1E6A53007C4C9B1BFEBEF0789EA8FDBF" box="[334,461,583,608]" pageId="63" pageNumber="64">Malaise, A</collectorName>
.
<collectorName id="1E6A53007C4C9B1BFE29F0779D26FD80" box="[473,579,584,608]" pageId="63" pageNumber="64">Santos, P</collectorName>
.
<collectorName id="1E6A53007C4C9B1BFDBEF0789DDDFDBF" box="[590,696,583,608]" pageId="63" pageNumber="64">González</collectorName>
[col.] (
<collectionCode id="D58EAE137C4C9B1BFCFCF0779C3DFDBF" box="[780,856,584,608]" pageId="63" pageNumber="64">MIUP</collectionCode>
)
</materialsCitation>
;
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<specimenCount id="A599FD5F7C4C9B1BFC9CF0779CF4FDBF" box="[876,913,584,608]" pageId="63" pageNumber="64" type="male">1♂</specimenCount>
, [
<collectingCounty id="5A414E5A7C4C9B1BFC55F0789CB7FDBF" box="[933,978,583,608]" pageId="63" pageNumber="64">Rio</collectingCounty>
]
<collectingMunicipality id="5344ACAC7C4C9B1BFC11F0789B00FDBF" box="[993,1125,583,608]" pageId="63" pageNumber="64">Calovébora</collectingMunicipality>
, [
<location id="B640600D7C4C9B1BFB88F0789BC0FDBF" LSID="urn:lsid:plazi:treatment:3B3687C07C4D9B1BFF67F6C99C99FD12:B640600D7C4C9B1BFB88F0789BC0FDBF" box="[1144,1189,583,608]" country="Iceland" county="Rio" latitude="8.7" longLatPrecision="1300" longitude="-81.21667" municipality="Calovebora" name="Rio" pageId="63" pageNumber="64" stateProvince="Pr">Rio</location>
]
<location id="B640600D7C4C9B1BFB44F0789A40FDBF" LSID="urn:lsid:plazi:treatment:3B3687C07C4D9B1BFF67F6C99C99FD12:B640600D7C4C9B1BFB44F0789A40FDBF" box="[1204,1317,583,608]" country="Iceland" county="Rio" latitude="8.7" longLatPrecision="1300" longitude="-81.21667" municipality="Calovebora" name="Guazarito" pageId="63" pageNumber="64" stateProvince="Pr">Guazarito</location>
, [
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,
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],
<collectingDate id="D765E9FE7C4C9B1BFEE2F0539EB6FD5B" box="[274,467,620,645]" pageId="63" pageNumber="64" value="2000-02-15" valueMax="2000-02-18" valueMin="2000-02-15">
<date id="C72110167C4C9B1BFEE2F0539EB6FD5B" box="[274,467,620,645]" pageId="63" pageNumber="64" value="2000-02-15" valueMax="2000-02-18" valueMin="2000-02-15">15–18 feb. 2000</date>
</collectingDate>
,
<collectorName id="1E6A53007C4C9B1BFE12F0539DD5FD5A" box="[482,688,620,645]" pageId="63" pageNumber="64">Col. S. Bermudez</collectorName>
(
<collectionCode id="D58EAE137C4C9B1BFD33F0539C6AFD5B" box="[707,783,620,644]" pageId="63" pageNumber="64">MIUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C4C9B1BFCD6F0539B3FFD5A" ID-GBIF-Occurrence="1671715840" box="[806,1114,620,645]" collectingDate="1999-09-26" collectingDateMax="1999-09-30" collectingDateMin="1999-09-26" collectionCode="MIUP" collectorName="El Cope &amp; de Division &amp; Omar Torrijos H &amp; Tr &amp; Malaise, A &amp; Santos, P" country="Panama" county="Material" elevation="620" latitude="8.698889" location="El Cope" longLatPrecision="21" longitude="-80.654724" municipality="Cocle Pr" pageId="63" pageNumber="64" specimenCount="1" specimenCount-male="1" stateProvince="Cocle" typeStatus="paratype">
<specimenCount id="A599FD5F7C4C9B1BFCD6F0539C2CFD5B" box="[806,841,620,644]" pageId="63" pageNumber="64" type="male">1♂</specimenCount>
same data as holotype
</materialsCitation>
;
<materialsCitation id="03F73C8B7C4C9B1BFB98F0529CCBFD77" ID-GBIF-Occurrence="1671715817" collectingDate="1996-06-03" collectingDateMax="1996-06-07" collectingDateMin="1996-06-03" collectionCode="CNCI" collectorName="J. Ashe &amp; R. Brooks" country="Panama" elevation="600" latitude="7.85" location="Cana" longLatPrecision="1302" longitude="-87.61667" pageId="63" pageNumber="64" specimenCount="2" specimenCount-male="2" stateProvince="Darien" typeStatus="paratype">
<collectingRegion id="715BF8347C4C9B1BFB98F0529BD2FD5B" box="[1128,1207,621,644]" country="Panama" name="Darien" pageId="63" pageNumber="64">
<emphasis id="81EBEAC47C4C9B1BFB98F0529BD2FD5B" box="[1128,1207,621,644]" italics="true" pageId="63" pageNumber="64">Darién</emphasis>
</collectingRegion>
:
<specimenCount id="A599FD5F7C4C9B1BFB3AF0539B95FD5B" box="[1226,1264,620,644]" pageId="63" pageNumber="64" type="male">2♂</specimenCount>
,
<location id="B640600D7C4C9B1BFB0FF0539A5EFD5B" LSID="urn:lsid:plazi:treatment:3B3687C07C4D9B1BFF67F6C99C99FD12:B640600D7C4C9B1BFB0FF0539A5EFD5B" box="[1279,1339,620,644]" country="Panama" latitude="7.85" longLatPrecision="1302" longitude="-87.61667" name="Cana" pageId="63" pageNumber="64" stateProvince="Darien">Cana</location>
,
<elevation id="38B2D1E57C4C9B1BFABAF0539AF2FD5B" box="[1354,1431,620,645]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="63" pageNumber="64" unit="m" value="600.0">
<quantity id="74679B337C4C9B1BFABAF0539AF2FD5B" box="[1354,1431,620,645]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="63" pageNumber="64" unit="m" value="600.0">600 m</quantity>
</elevation>
, [
<geoCoordinate id="D6AB50117C4C9B1BFF6EF0B09F9AFD77" box="[158,255,655,680]" direction="north" orientation="latitude" pageId="63" pageNumber="64" precision="925" value="7.85">07°51'N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C4C9B1BFEFBF0AF9E17FD77" box="[267,370,656,680]" direction="west" orientation="longitude" pageId="63" pageNumber="64" precision="925" value="-87.61667">87°37'W</geoCoordinate>
],
<collectingDate id="D765E9FE7C4C9B1BFE74F0B09D70FD77" box="[388,533,655,680]" pageId="63" pageNumber="64" value="1996-06-03" valueMax="1996-06-07" valueMin="1996-06-03">
<date id="C72110167C4C9B1BFE74F0B09D70FD77" box="[388,533,655,680]" pageId="63" pageNumber="64" value="1996-06-03" valueMax="1996-06-07" valueMin="1996-06-03">3–7.VI.1996</date>
</collectingDate>
,
<collectorName id="1E6A53007C4C9B1BFDD0F0AF9D16FD77" box="[544,627,655,680]" pageId="63" pageNumber="64">J. Ashe</collectorName>
&amp;
<collectorName id="1E6A53007C4C9B1BFD66F0AF9C6DFD77" box="[662,776,655,680]" pageId="63" pageNumber="64">R. Brooks</collectorName>
[col.] (
<collectionCode id="D58EAE137C4C9B1BFCAAF0AF9CC6FD77" box="[858,931,656,680]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="63" pageNumber="64">CNCI</collectionCode>
);
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C4C9B1BFF37F08B9C99FD12" box="[199,1020,692,717]" pageId="63" pageNumber="64" type="distribution">
<paragraph id="B32036D67C4C9B1BFF37F08B9C99FD12" blockId="63.[151,1437,151,717]" box="[199,1020,692,717]" pageId="63" pageNumber="64">
<emphasis id="81EBEAC47C4C9B1BFF37F08B9E3CFD12" bold="true" box="[199,345,692,717]" pageId="63" pageNumber="64">Distribution</emphasis>
(
<figureCitation id="2BA42A537C4C9B1BFE98F08B9EAEFD12" box="[360,459,692,717]" captionStart="FIGURES 234 – 243" captionStartId="67.[151,264,1856,1879]" captionTargetBox="[174,1439,203,1822]" captionTargetId="figure@67.[162,1447,193,1827]" captionTargetPageId="67" captionText="FIGURES 234 – 243. Known distribution of Dissomphalus in Panama. 234. D. galeatus, D. geniculatus and D. geortus sp. nov.. 235. D. gilvipes, D. ibocius sp. nov. and D. ibrastus sp. nov.. 236. D. guttus and D. ilarius sp. nov.. 237. D. infissus, D. intradentatus and D. iozus sp. nov.. 238. D. isortus sp. nov., D. isteus sp. nov. and D. latus. 239. D. persontaus, D. pilus and D. pronus. 240. D. plaumanni and D. politus. 241. D. piscicercus, D. ramosus and D. raziogus sp. nov.. 242. D. punctatus, D. strepsus and D. subdeformis. 243. D. racarus sp. nov., D. strabus and D. tisinus sp. nov.." httpUri="https://zenodo.org/record/1010060/files/figure.png" pageId="63" pageNumber="64">Fig. 236</figureCitation>
).
<collectingRegion id="715BF8347C4C9B1BFE2EF08A9D52FD13" box="[478,567,693,716]" country="Panama" name="Panama" pageId="63" pageNumber="64">
<collectingCountry id="CB8876467C4C9B1BFE2EF08A9D52FD13" box="[478,567,693,716]" name="Panama" pageId="63" pageNumber="64">Panama</collectingCountry>
</collectingRegion>
(
<collectingRegion id="715BF8347C4C9B1BFDB5F08B9DD5FD13" box="[581,688,692,716]" country="Panama" name="Veraguas" pageId="63" pageNumber="64">Veraguas</collectingRegion>
,
<collectingRegion id="715BF8347C4C9B1BFD4CF08B9C6EFD12" box="[700,779,692,717]" country="Panama" name="Darien" pageId="63" pageNumber="64">Darién</collectingRegion>
,
<collectingRegion id="715BF8347C4C9B1BFCE6F08B9C32FD12" box="[790,855,692,717]" country="Panama" name="Cocle" pageId="63" pageNumber="64">Coclé</collectingRegion>
and
<collectingRegion id="715BF8347C4C9B1BFC7EF08B9C94FD12" box="[910,1009,692,717]" country="Panama" name="Chiriqui" pageId="63" pageNumber="64">Chiriquí</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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