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<document id="02190BB79643A1391FF6CD93BC937C68" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C429B15FF67F33B9CF0FAF0" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus citus Azevedo 2017, sp. nov." docType="treatment" docVersion="15" lastPageNumber="50" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="50" updateTime="1698455438235" updateUser="plazi">
<mods:mods id="5C33551FDAEEFD37191BD44CE5C84F5A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="474C4C28C391F6AC1D3864F2EF3577F6">
<mods:title id="2EA300C7D6980E41DB15F46DBF0CD871">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
</mods:titleInfo>
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<mods:namePart id="3D89E61386BE9BB959936A817A20BF9F">Azevedo, Celso O.</mods:namePart>
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<mods:title id="885D8569EE0C7532B422B993A41A531F">Zootaxa</mods:title>
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<mods:part id="DFEB88D2DC837E496779078CB5B0A338">
<mods:date id="82DA877B10BD631F3E151B78E5D32C99">2017</mods:date>
<mods:detail id="2303A9D2B71F27DACCDF63086A4DD47D" type="pubDate">
<mods:number id="8F3C834091F5FAE4F89A61F7DC1FA00C">2017-10-13</mods:number>
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<mods:detail id="08140B94EEFB824D3619B7BF1BB3DCE4" type="volume">
<mods:number id="FDA81C547EC3709DF6AECB0A100316BB">4335</mods:number>
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<mods:number id="7FA39CB74CA6B974C262CAB3EC5BD306">1</mods:number>
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<mods:classification id="495DFEA0584B81096C82BF29076266F0">journal article</mods:classification>
<mods:identifier id="7A3FAD37F3733ED5981D018FD4F87185" type="CLB-Dataset">31852</mods:identifier>
<mods:identifier id="A0062FC69D73CFF4DE4BB2F2ECEAD726" type="DOI">10.11646/zootaxa.4335.1.1</mods:identifier>
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<mods:identifier id="147CE989234E116AB4B509C45E8F38AC" type="ISSN">1175-5326</mods:identifier>
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<treatment id="3B3687C07C429B15FF67F33B9CF0FAF0" ID-DOI="http://doi.org/10.5281/zenodo.5020470" ID-GBIF-Taxon="135435260" ID-Zenodo-Dep="5020470" LSID="urn:lsid:plazi:treatment:3B3687C07C429B15FF67F33B9CF0FAF0" httpUri="http://treatment.plazi.org/id/3B3687C07C429B15FF67F33B9CF0FAF0" lastPageNumber="50" pageId="49" pageNumber="50">
<subSubSection id="FB85655D7C429B15FF67F33B9EA4FEE0" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="B32036D67C429B15FF67F33B9DADFEC1" blockId="49.[151,712,260,319]" box="[151,712,260,286]" pageId="49" pageNumber="50">
<heading id="E86881BA7C429B15FF67F33B9DADFEC1" bold="true" box="[151,712,260,286]" fontSize="11" level="1" pageId="49" pageNumber="50" reason="1">
<emphasis id="81EBEAC47C429B15FF67F33B9DADFEC1" bold="true" box="[151,712,260,286]" pageId="49" pageNumber="50">
<taxonomicName id="749F4D557C429B15FF67F33B9D38FEC1" authority="Brito &amp; Azevedo" authorityName="Azevedo" authorityYear="2017" box="[151,605,260,286]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="citus" status="sp. nov.">
<emphasis id="81EBEAC47C429B15FF67F33B9EE7FEC1" bold="true" box="[151,386,260,286]" italics="true" pageId="49" pageNumber="50">Dissomphalus citus</emphasis>
Brito &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="9AD857BF7C429B15FD9AF33B9DADFEC1" box="[618,712,260,286]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B32036D67C429B15FF67F3199EA4FEE0" blockId="49.[151,712,260,319]" box="[151,449,294,319]" pageId="49" pageNumber="50">
(
<figureCitation id="2BA42A537C429B15FF6FF3199F90FEE0" box="[159,245,294,319]" captionStart="FIGURES 13 – 24" captionStartId="11.[151,264,1615,1638]" captionTargetBox="[151,1436,306,1594]" captionTargetId="figure@11.[151,1436,303,1594]" captionTargetPageId="11" captionText="FIGURES 13 – 24. Dissomphalus spp., ♂, head, dorsal view. 13. D. racarus sp. nov. 14. D. teraus sp. nov. 15. D. urazius sp. nov. 16. D. afestus sp. nov. 17. D. arbeius sp. nov. 18. D. raziogus sp. nov. 19. D. areius sp. nov. 20. D. aretisus sp. nov. 21. D. citus sp. nov. 22. D. sartus sp. nov. 23. D. teobus sp. nov. 24. D. terbius sp. nov.. Scale bars = 20 µm, except figs 14, 20 = 10 µm and fig. 15 = 50 µm." httpUri="https://zenodo.org/record/1010018/files/figure.png" pageId="49" pageNumber="50">Figs 21</figureCitation>
,
<figureCitation id="2BA42A537C429B15FEF1F3199E45FEE0" box="[257,288,294,319]" captionStart="FIGURES 45 – 56" captionStartId="14.[151,264,1544,1567]" captionTargetBox="[151,1436,193,1524]" captionTargetId="figure@14.[151,1436,193,1524]" captionTargetPageId="14" captionText="FIGURES 45 – 56. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 45. D. teraus sp. nov. 46. D. urazius sp. nov. 47. D. afestus sp. nov. 48. D. arbeius sp. nov. 49. D. raziogus sp. nov. 50. D. areius sp. nov. 51. D. aretisus sp. nov. 52. D. citus sp. nov. 53. D. sartus sp. nov. 54. D. teobus sp. nov. 55. D. terbius sp. nov. 56. D. abarus sp. nov.. Scale bars = 20 µm except, figs 45 and 46 = 50 µm." httpUri="https://zenodo.org/record/1010024/files/figure.png" pageId="49" pageNumber="50">52</figureCitation>
,
<figureCitation id="2BA42A537C429B15FEDBF3189E2CFEE0" box="[299,329,295,319]" captionStart="FIGURES 76 – 87" captionStartId="17.[151,264,1595,1618]" captionTargetBox="[159,1404,219,1523]" captionTargetId="figure@17.[147,1432,209,1558]" captionTargetPageId="17" captionText="FIGURES 76 – 87. Dissomphalus spp., ♂, hypopygium, internal view. 76. D. teraus sp. nov. 77. D. urazius sp. nov. 78. D. afestus sp. nov. 79. D. arbeius sp. nov. 80. D. raziogus sp. nov. 81. D. areius sp. nov. 82. D. aretisus sp. nov. 83. D. citus sp. nov. 84. D. sartus sp. nov. 85. D. teobus sp. nov. 86. D. terbius sp. nov. 87. D. abarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010030/files/figure.png" pageId="49" pageNumber="50">83</figureCitation>
,
<figureCitation id="2BA42A537C429B15FEA5F3189EDCFEE0" box="[341,441,295,319]" captionStart="FIGURES 174 – 183" captionStartId="46.[151,264,1785,1808]" captionTargetBox="[186,1392,390,1733]" captionTargetId="figure@46.[151,1436,375,1764]" captionTargetPageId="46" captionText="FIGURES 174 – 183. Dissomphalus spp., ♂, genitalia, except fig. 177 paramere. 174. D. bilobatus. Lateral view. 175 – 177. D. rufipalpis 175. Dorsal view. 176. Ventral view. 177. Dorsal view. 178 – 179. D. areius sp. nov. 178. Dorsal view. 179. Ventral view. 180 – 181. D. aretisus sp. nov. 180. Dorsal view. 181. Ventral view. 182 – 183. D. citus sp. nov. 182. Dorsal view. 183. Ventral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010046/files/figure.png" pageId="49" pageNumber="50">182–183</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C429B15FF67F3519EABFC84" pageId="49" pageNumber="50" type="description">
<paragraph id="B32036D67C429B15FF67F3519D86FCCC" blockId="49.[151,1437,366,1327]" pageId="49" pageNumber="50">
<emphasis id="81EBEAC47C429B15FF67F3519E4CFE58" bold="true" box="[151,297,366,391]" pageId="49" pageNumber="50">Description.</emphasis>
Male. Body length
<quantity id="74679B337C429B15FDF7F3519D0BFE59" box="[519,622,366,391]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="49" pageNumber="50" unit="mm" value="1.55">1.55 mm</quantity>
. Color: body castaneous. Head (
<figureCitation id="2BA42A537C429B15FC27F3519B4EFE58" box="[983,1067,366,391]" captionStart="FIGURES 13 – 24" captionStartId="11.[151,264,1615,1638]" captionTargetBox="[151,1436,306,1594]" captionTargetId="figure@11.[151,1436,303,1594]" captionTargetPageId="11" captionText="FIGURES 13 – 24. Dissomphalus spp., ♂, head, dorsal view. 13. D. racarus sp. nov. 14. D. teraus sp. nov. 15. D. urazius sp. nov. 16. D. afestus sp. nov. 17. D. arbeius sp. nov. 18. D. raziogus sp. nov. 19. D. areius sp. nov. 20. D. aretisus sp. nov. 21. D. citus sp. nov. 22. D. sartus sp. nov. 23. D. teobus sp. nov. 24. D. terbius sp. nov.. Scale bars = 20 µm, except figs 14, 20 = 10 µm and fig. 15 = 50 µm." httpUri="https://zenodo.org/record/1010018/files/figure.png" pageId="49" pageNumber="50">Fig. 21</figureCitation>
). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina very high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures large. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin carinate. Metasoma (
<figureCitation id="2BA42A537C429B15FC30F3E59B73FE2C" box="[960,1046,474,499]" captionStart="FIGURES 45 – 56" captionStartId="14.[151,264,1544,1567]" captionTargetBox="[151,1436,193,1524]" captionTargetId="figure@14.[151,1436,193,1524]" captionTargetPageId="14" captionText="FIGURES 45 – 56. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 45. D. teraus sp. nov. 46. D. urazius sp. nov. 47. D. afestus sp. nov. 48. D. arbeius sp. nov. 49. D. raziogus sp. nov. 50. D. areius sp. nov. 51. D. aretisus sp. nov. 52. D. citus sp. nov. 53. D. sartus sp. nov. 54. D. teobus sp. nov. 55. D. terbius sp. nov. 56. D. abarus sp. nov.. Scale bars = 20 µm except, figs 45 and 46 = 50 µm." httpUri="https://zenodo.org/record/1010024/files/figure.png" pageId="49" pageNumber="50">Fig. 52</figureCitation>
). Tergal process with lateral, very shallow, subcircular and small depression, with very small, low and posterad tubercle placed on center of depression, pit very small, with short and dense tuft of setae. Posterior hypopygeal margin strongly concave (
<figureCitation id="2BA42A537C429B15FA9FF01D9FD3FD80" captionStart="FIGURES 76 – 87" captionStartId="17.[151,264,1595,1618]" captionTargetBox="[159,1404,219,1523]" captionTargetId="figure@17.[147,1432,209,1558]" captionTargetPageId="17" captionText="FIGURES 76 – 87. Dissomphalus spp., ♂, hypopygium, internal view. 76. D. teraus sp. nov. 77. D. urazius sp. nov. 78. D. afestus sp. nov. 79. D. arbeius sp. nov. 80. D. raziogus sp. nov. 81. D. areius sp. nov. 82. D. aretisus sp. nov. 83. D. citus sp. nov. 84. D. sartus sp. nov. 85. D. teobus sp. nov. 86. D. terbius sp. nov. 87. D. abarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010030/files/figure.png" pageId="49" pageNumber="50">Fig. 83</figureCitation>
). Genitalia (
<figureCitation id="2BA42A537C429B15FECEF0799EBBFD80" box="[318,478,582,607]" captionStart="FIGURES 174 – 183" captionStartId="46.[151,264,1785,1808]" captionTargetBox="[186,1392,390,1733]" captionTargetId="figure@46.[151,1436,375,1764]" captionTargetPageId="46" captionText="FIGURES 174 – 183. Dissomphalus spp., ♂, genitalia, except fig. 177 paramere. 174. D. bilobatus. Lateral view. 175 – 177. D. rufipalpis 175. Dorsal view. 176. Ventral view. 177. Dorsal view. 178 – 179. D. areius sp. nov. 178. Dorsal view. 179. Ventral view. 180 – 181. D. aretisus sp. nov. 180. Dorsal view. 181. Ventral view. 182 – 183. D. citus sp. nov. 182. Dorsal view. 183. Ventral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010046/files/figure.png" pageId="49" pageNumber="50">Figs 182–183</figureCitation>
): paramere with dorsal margin entirely wide; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex long with sinuous, slender ventral projection, curved outward; additional inner ramus absent; basal stub long and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, subhorizontal and narrow, with apical margin rounded and dorsad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.
</paragraph>
<paragraph id="B32036D67C429B15FF37F1219EABFC84" blockId="49.[151,1437,366,1327]" pageId="49" pageNumber="50">
<emphasis id="81EBEAC47C429B15FF37F1219E22FCE8" bold="true" box="[199,327,798,823]" pageId="49" pageNumber="50">Variations.</emphasis>
Head, mesosoma and metasoma light castaneous; anterior margin of pronotal disc coarse; tergal process without depression.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C429B15FF37F1599CD6FB88" pageId="49" pageNumber="50" type="discussion">
<paragraph id="B32036D67C429B15FF37F1599CD6FB88" blockId="49.[151,1437,366,1327]" pageId="49" pageNumber="50">
<emphasis id="81EBEAC47C429B15FF37F1599E5EFCA0" bold="true" box="[199,315,870,895]" pageId="49" pageNumber="50">Remarks.</emphasis>
This species is easily recognized by having the aedeagal ventral ramus with slender ventral projection and the carina of the median clypeal tooth very high in profile and the ventral margin of the paramere thick with thick setae. This species is similar to
<taxonomicName id="749F4D557C429B15FD33F1909C5BFC19" box="[707,830,943,966]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="81EBEAC47C429B15FD33F1909C5BFC19" box="[707,830,943,966]" italics="true" pageId="49" pageNumber="50">D. magnus</emphasis>
</taxonomicName>
by having the dorsal margin of the paramere almost straight with distinct thick setae apical and longer than the basiparamere. However, this species has the aedeagal ventral ramus abruptly narrow apicad with one ventral slender projection and the apical lobes of the aedeagal dorsal body subhorizontal whereas
<taxonomicName id="749F4D557C429B15FDC2F6239DCAFBEC" box="[562,687,1052,1075]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="81EBEAC47C429B15FDC2F6239DCAFBEC" box="[562,687,1052,1075]" italics="true" pageId="49" pageNumber="50">D. magnus</emphasis>
</taxonomicName>
has the aedeagal ventral ramus divided in two tubular, sinuous, projections and the apical lobes of the aedeagal dorsal body horizontal.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C429B15FF37F65D9C59FAD4" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="B32036D67C429B15FF37F65D9C59FAD4" blockId="49.[151,1437,366,1327]" pageId="49" pageNumber="50">
<materialsCitation id="03F73C8B7C429B15FF37F65D9AFCFB40" ID-GBIF-Occurrence="1671715708" collectingDate="1999-04-08" collectingDateMax="1999-04-11" collectingDateMin="1999-04-08" collectionCode="MIUP" collectorName="Malaise, R &amp; Cambra, A &amp; Santos" country="Panama" elevation="1080" latitude="8.771111" location="Toro" longLatPrecision="21" longitude="-82.190834" municipality="Material" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="PANAMA" typeStatus="holotype">
<emphasis id="81EBEAC47C429B15FF37F65D9ED3FBA4" bold="true" box="[199,438,1122,1147]" pageId="49" pageNumber="50">
<collectingMunicipality id="5344ACAC7C429B15FF37F65D9E4AFBA4" box="[199,303,1122,1147]" pageId="49" pageNumber="50">Material</collectingMunicipality>
examined.
</emphasis>
<typeStatus id="6C2488747C429B15FE32F65D9D48FBA4" box="[450,557,1122,1147]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
,
<specimenCount id="A599FD5F7C429B15FDCEF65C9D30FBA5" box="[574,597,1123,1146]" pageId="49" pageNumber="50" type="male">♂</specimenCount>
,
<collectingRegion id="715BF8347C429B15FD96F65C9D85FBA4" box="[614,736,1123,1147]" country="Panama" name="Panama" pageId="49" pageNumber="50">
<collectingCountry id="CB8876467C429B15FD96F65C9D85FBA4" box="[614,736,1123,1147]" name="Panama" pageId="49" pageNumber="50">PANAMA</collectingCountry>
</collectingRegion>
, [
<collectingRegion id="715BF8347C429B15FD0AF65D9C3CFBA4" box="[762,857,1122,1147]" country="Panama" name="Chiriqui" pageId="49" pageNumber="50">
<emphasis id="81EBEAC47C429B15FD0AF65D9C3CFBA4" box="[762,857,1122,1147]" italics="true" pageId="49" pageNumber="50">Chiriquí</emphasis>
</collectingRegion>
]:
<taxonomicName id="749F4D557C429B15FC86F65C9B3EFBA4" authority=", Div. Cont." authorityName="Div. Cont." box="[886,1115,1122,1147]" class="Gastropoda" family="Achatinidae" genus="Fortuna" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="50" phylum="Mollusca" rank="genus">Fortuna, Div. Cont.</taxonomicName>
Chiriqui-B[ocas del]
<location id="B640600D7C429B15FA90F65C9AFDFBA5" LSID="urn:lsid:plazi:treatment:3B3687C07C429B15FF67F33B9CF0FAF0:B640600D7C429B15FA90F65C9AFDFBA5" box="[1376,1432,1123,1146]" country="Panama" latitude="8.771111" longLatPrecision="21" longitude="-82.190834" municipality="Material" name="Toro" pageId="49" pageNumber="50" stateProvince="PANAMA">Toro</location>
,
<elevation id="38B2D1E57C429B15FF67F6B89E5BFB41" box="[151,318,1159,1183]" metricMagnitude="3" metricUnit="m" metricValue="1.08" metricValueMax="1.09" metricValueMin="1.0699999999999998" pageId="49" pageNumber="50" unit="m" value="1080.0" valueMax="1090.0" valueMin="1070.0">
<quantity id="74679B337C429B15FF67F6B89E5BFB41" box="[151,318,1159,1183]" metricMagnitude="3" metricUnit="m" metricValue="1.08" metricValueMax="1.09" metricValueMin="1.0699999999999998" pageId="49" pageNumber="50" unit="m" value="1080.0" valueMax="1090.0" valueMin="1070.0">1070–1090 m</quantity>
</elevation>
, [
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,
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],
<collectingDate id="D765E9FE7C429B15FD54F6B99C20FB40" box="[676,837,1158,1183]" pageId="49" pageNumber="50" value="1999-04-08" valueMax="1999-04-11" valueMin="1999-04-08">
<date id="C72110167C429B15FD54F6B99C20FB40" box="[676,837,1158,1183]" pageId="49" pageNumber="50" value="1999-04-08" valueMax="1999-04-11" valueMin="1999-04-08">8–11.abr.1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C429B15FCABF6B99C87FB41" box="[859,994,1158,1183]" pageId="49" pageNumber="50">Malaise, R</collectorName>
.
<collectorName id="1E6A53007C429B15FC07F6B99B1BFB40" box="[1015,1150,1158,1183]" pageId="49" pageNumber="50">Cambra, A</collectorName>
.
<collectorName id="1E6A53007C429B15FB64F6B89BBAFB40" box="[1172,1247,1159,1183]" pageId="49" pageNumber="50">Santos</collectorName>
[col.] (
<collectionCode id="D58EAE137C429B15FAB4F6B89AF5FB40" box="[1348,1424,1159,1183]" pageId="49" pageNumber="50">MIUP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="03F73C8B7C429B15FF67F6949D59FB1C" ID-GBIF-Occurrence="1671715720" box="[151,572,1194,1219]" collectingDate="1999-04-08" collectingDateMax="1999-04-11" collectingDateMin="1999-04-08" collectionCode="MIUP" collectorName="Malaise, R &amp; Cambra, A &amp; Santos" country="Panama" elevation="1080" latitude="8.771111" location="Toro" longLatPrecision="21" longitude="-82.190834" municipality="Material" pageId="49" pageNumber="50" specimenCount="5" specimenCount-male="5" stateProvince="PANAMA" typeStatus="paratype">
<typeStatus id="6C2488747C429B15FF67F6949E6CFB1C" box="[151,265,1195,1219]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="A599FD5F7C429B15FEE2F6959E52FB1C" box="[274,311,1194,1219]" pageId="49" pageNumber="50" type="male">5♂</specimenCount>
, same data as holotype
</materialsCitation>
; 2♂, same of holotype except
<date id="C72110167C429B15FC7DF6959B4AFB1C" box="[909,1071,1194,1219]" pageId="49" pageNumber="50" value="1999-04-09" valueMax="1999-04-11" valueMin="1999-04-09">9–11.abr.1999</date>
, Tr[ampas] Amar[illas] (MIUP);
<materialsCitation id="03F73C8B7C429B15FF67F6F09C5DFAD4" ID-GBIF-Occurrence="1671715721" collectingDate="1999-10-17" collectingDateMax="1999-10-24" collectingDateMin="1999-10-17" collectionCode="MIUP" collectorName="Tr &amp; Malaise, A &amp; Santos" country="Panama" latitude="9.406944" location="Teribe" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Bocas del Toro" typeStatus="paratype">
<collectingRegion id="715BF8347C429B15FF67F6F09E5BFB39" box="[151,318,1230,1255]" country="Panama" name="Bocas del Toro" pageId="49" pageNumber="50">
<emphasis id="81EBEAC47C429B15FF67F6F09E5BFB39" box="[151,318,1230,1255]" italics="true" pageId="49" pageNumber="50">Bocas del Toro</emphasis>
</collectingRegion>
:
<specimenCount id="A599FD5F7C429B15FEBCF6F09E14FB38" box="[332,369,1231,1255]" pageId="49" pageNumber="50" type="male">1♂</specimenCount>
,
<collectingMunicipality id="5344ACAC7C429B15FE8DF6F19EA9FB38" box="[381,460,1230,1255]" pageId="49" pageNumber="50">Wekso</collectingMunicipality>
,
<location id="B640600D7C429B15FE26F6F19D45FB38" LSID="urn:lsid:plazi:treatment:3B3687C07C429B15FF67F33B9CF0FAF0:B640600D7C429B15FE26F6F19D45FB38" box="[470,544,1230,1255]" country="Panama" latitude="9.406944" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" name="Teribe" pageId="49" pageNumber="50" stateProvince="Bocas del Toro">Teribe</location>
, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [
<geoCoordinate id="D6AB50117C429B15FB82F6F19B9DFB38" box="[1138,1272,1230,1255]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="15" value="9.406944">09º24'25&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C429B15FAF4F6F19AF4FB38" box="[1284,1425,1230,1255]" direction="west" orientation="longitude" pageId="49" pageNumber="50" precision="15" value="-82.93889">82º56'20&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C429B15FF67F6CC9E2FFAD4" box="[151,330,1267,1291]" pageId="49" pageNumber="50" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">
<date id="C72110167C429B15FF67F6CC9E2FFAD4" box="[151,330,1267,1291]" pageId="49" pageNumber="50" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">17-24 oct. 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C429B15FEA5F6CC9E14FAD5" box="[341,369,1267,1290]" pageId="49" pageNumber="50">Tr</collectorName>
[ampa]
<collectorName id="1E6A53007C429B15FE35F6CD9D26FAD4" box="[453,579,1266,1291]" pageId="49" pageNumber="50">Malaise, A</collectorName>
.
<collectorName id="1E6A53007C429B15FDBEF6CD9DFAFAD4" box="[590,671,1266,1291]" pageId="49" pageNumber="50">Santos</collectorName>
[col.] (
<collectionCode id="D58EAE137C429B15FD14F6CC9C55FAD4" box="[740,816,1267,1291]" pageId="49" pageNumber="50">MIUP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C429B15FF37F7299CF0FAF0" box="[199,917,1302,1327]" pageId="49" pageNumber="50" type="distribution">
<paragraph id="B32036D67C429B15FF37F7299CF0FAF0" blockId="49.[151,1437,366,1327]" box="[199,917,1302,1327]" pageId="49" pageNumber="50">
<emphasis id="81EBEAC47C429B15FF37F7299E3CFAF0" bold="true" box="[199,345,1302,1327]" pageId="49" pageNumber="50">Distribution</emphasis>
(
<figureCitation id="2BA42A537C429B15FE98F7299EAEFAF0" box="[360,459,1302,1327]" captionStart="FIGURES 224 – 233" captionStartId="66.[151,264,1814,1837]" captionTargetBox="[182,1404,199,1768]" captionTargetId="figure@66.[169,1413,193,1773]" captionTargetPageId="66" captionText="FIGURES 224 – 233. Known distribution of Dissomphalus in Panama. 224. Provinces and Counties of Panama. 1. Bocas del Toro. 2. Chiriquí. 3. Ngöbe-Buglé County. 4. Veraguas. 5. Herrera. 6. Los Santos. 7. Colón. 8. Coclé. 9. Panama. 10. Guna Yala County. 11. Darién. 12. Emberá Woonan County. 225. D. abarus sp. nov., D. afestus sp. nov. and D. altivolans. 226. D. angulatus sp. nov., D. ardonius sp. nov. and D. aretisus sp. nov.. 227. D. apertus, D. areius sp. nov. and D. arteobius sp. nov.. 228. D. arbeius sp. nov., D. asitius sp. nov. and D. brasiliensis. 229. D. arteus sp. nov., D. cervoides and D. culteratus. 230. D. asfartus sp. nov.. 231. D. azagus sp. nov., D. bilobatus and D. coronatus. 232. D. borus sp. nov., D. citus sp. nov. and D. curvifoveatus. 233. D. declinatus, D. dilatatus and D. ettus sp. nov.." httpUri="https://zenodo.org/record/1010056/files/figure.png" pageId="49" pageNumber="50">Fig. 232</figureCitation>
).
<collectingRegion id="715BF8347C429B15FE2EF7289D52FAF1" box="[478,567,1303,1326]" country="Panama" name="Panama" pageId="49" pageNumber="50">
<collectingCountry id="CB8876467C429B15FE2EF7289D52FAF1" box="[478,567,1303,1326]" name="Panama" pageId="49" pageNumber="50">Panama</collectingCountry>
</collectingRegion>
(
<collectingRegion id="715BF8347C429B15FDB6F7289D95FAF1" box="[582,752,1302,1327]" country="Panama" name="Bocas del Toro" pageId="49" pageNumber="50">Bocas del Toro</collectingRegion>
and
<collectingRegion id="715BF8347C429B15FCD6F7299CECFAF0" box="[806,905,1302,1327]" country="Panama" name="Chiriqui" pageId="49" pageNumber="50">Chiriquí</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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